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<mods:title id="4B4918A1E28FC2AF33CCF8720E37539E">Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane</mods:title>
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<mods:namePart id="2A63A81DBA7AA8768775171B420A39B0">Ketwetsuriya, Chatchalerm</mods:namePart>
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<mods:namePart id="0DC630FF50818D7A1600C07AE554AB87">Karapunar, Baran</mods:namePart>
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<mods:namePart id="5D3DDBBD8291528AF1666DB0EADAF7C3">Charoentitirat, Thasinee</mods:namePart>
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<mods:namePart id="46F149BDE6CD5D7A5D17C059BDCE6D0C">Nützel, Al- Exander</mods:namePart>
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<treatment id="03B587AB4F3E1577FF5178E7FDF3FD2E" ID-DOI="http://doi.org/10.5281/zenodo.3803955" ID-GBIF-Taxon="163737107" ID-Zenodo-Dep="3803955" LSID="urn:lsid:plazi:treatment:03B587AB4F3E1577FF5178E7FDF3FD2E" httpUri="http://treatment.plazi.org/id/03B587AB4F3E1577FF5178E7FDF3FD2E" lastPageNumber="10" pageId="9" pageNumber="10">
<subSubSection id="C30665364F3E1577FF5178E7FE93FF31" box="[151,344,153,180]" pageId="9" pageNumber="10" type="nomenclature">
<paragraph id="8BA336BD4F3E1577FF5178E7FE93FF31" blockId="9.[151,344,153,216]" box="[151,344,153,180]" pageId="9" pageNumber="10">
<heading id="D0EB81D14F3E1577FF5178E7FE93FF31" bold="true" box="[151,344,153,180]" fontSize="10" level="2" pageId="9" pageNumber="10" reason="2">
<emphasis id="B968EAAF4F3E1577FF5178E7FE93FF31" bold="true" box="[151,344,153,180]" pageId="9" pageNumber="10">
<taxonomicName id="4C1C4D3E4F3E1577FF5178E7FEE6FF31" ID-CoL="7NZD9" authorityName="de Montfort" authorityYear="1808" box="[151,301,153,180]" class="Gastropoda" family="Bellerophontidae" genus="Bellerophon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Bellerophontida" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">
<emphasis id="B968EAAF4F3E1577FF5178E7FEE6FF31" bold="true" box="[151,301,153,180]" italics="true" pageId="9" pageNumber="10">Bellerophon</emphasis>
</taxonomicName>
sp.
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30665364F3E1577FF5178C3FED2FF5D" box="[151,281,189,216]" pageId="9" pageNumber="10" type="description">
<paragraph id="8BA336BD4F3E1577FF5178C3FED2FF5D" blockId="9.[151,344,153,216]" box="[151,281,189,216]" pageId="9" pageNumber="10">
(
<figureCitation id="13272A384F3E1577FF6678C3FF2AFF5D" box="[160,225,189,216]" captionStart="FIGURE 6" captionStartId="8.[151,250,1895,1920]" captionTargetBox="[151,1436,1136,1870]" captionTargetId="figure@8.[151,1436,1136,1870]" captionTargetPageId="8" captionText="FIGURE 6. AG, Bellerophon erawanensis sp. nov. AE, ESKU-19-LP 55, holotype; E, shell showing V-shaped slit at crest. F, ESKU-19-LP 54, paratype; shell showing growth lines and position of selenizone. G, ESKU-19-LP 56, paratype. HJ, Bellerophon sp. HI, ESKU-19-LP 58. J, ESKU-19-LP 59, shell fragment showing growth lines and position of selenizone. All scale bars represent 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3763919" httpUri="https://zenodo.org/record/3763919/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
HJ)
</paragraph>
</subSubSection>
<subSubSection id="C30665364F3E1577FF51797BFB6BFEC6" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8BA336BD4F3E1577FF51797BFE04FEA5" blockId="9.[151,1437,261,683]" box="[151,463,261,288]" pageId="9" pageNumber="10">
<emphasis id="B968EAAF4F3E1577FF51797BFECDFEA5" bold="true" box="[151,262,261,288]" pageId="9" pageNumber="10">Material.</emphasis>
ESKU-19-
<collectionCode id="ED0DAE784F3E1577FE45797BFE6EFE9A" box="[387,421,261,287]" country="Russia" name="Laboratory of Palaeontology" pageId="9" pageNumber="10">LP</collectionCode>
58.
</paragraph>
<paragraph id="8BA336BD4F3E1577FF017957FB6BFEC6" blockId="9.[151,1437,261,683]" box="[199,1184,297,324]" pageId="9" pageNumber="10">
Dimensions (mm): ESKU-19-
<collectionCode id="ED0DAE784F3E1577FDD07957FDF3FEC6" box="[534,568,297,323]" country="Russia" name="Laboratory of Palaeontology" pageId="9" pageNumber="10">LP</collectionCode>
58: height = 13.3; width (half) = 6.6; thickness = 12.1.
</paragraph>
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<subSubSection id="C30665364F3E1577FF017930FDF3FD2E" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8BA336BD4F3E1577FF017930FDF3FD2E" blockId="9.[151,1437,261,683]" pageId="9" pageNumber="10">
<emphasis id="B968EAAF4F3E1577FF017930FEF0FEED" bold="true" box="[199,315,334,360]" pageId="9" pageNumber="10">Remarks.</emphasis>
A single anomphalous bellerophontiform shell and a fragment show strengthened, thread-like growth lines and a distinct selenizone with slightly straight short lunulae, which are the general characteristics of the genus
<taxonomicName id="4C1C4D3E4F3E1577FF5179EBFEEBFE2A" authorityName="de Montfort" authorityYear="1808" box="[151,288,405,431]" class="Gastropoda" family="Bellerophontidae" genus="Bellerophon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Bellerophontida" pageId="9" pageNumber="10" phylum="Mollusca" rank="genus">
<emphasis id="B968EAAF4F3E1577FF5179EBFEEBFE2A" box="[151,288,405,431]" italics="true" pageId="9" pageNumber="10">Bellerophon</emphasis>
</taxonomicName>
. The involute shell is slightly elongated and inflated and the whorl profile is moderately well rounded. The present specimens resemble
<taxonomicName id="4C1C4D3E4F3E1577FDCF79C7FC29FE56" authority="de Koninck, 1863" authorityName="de Koninck" authorityYear="1863" box="[521,994,441,467]" class="Gastropoda" family="Bellerophontidae" genus="Bellerophon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Bellerophontida" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="jonesianus">
<emphasis id="B968EAAF4F3E1577FDCF79C7FCDAFE56" box="[521,785,441,467]" italics="true" pageId="9" pageNumber="10">Bellerophon jonesianus</emphasis>
<bibRefCitation id="EF8D4B4C4F3E1577FCDE79C7FC29FE56" author="Koninck, L. G. de" box="[792,994,441,467]" firstAuthor="de Koninck" pageId="9" pageNumber="10" refId="ref25516" refString="Koninck, L. G. de (1863) Meimoire sur les Fossiles Paleiozoiques Recueillis dans l'Inde par M. le Docteur Fleming. H. Dessaain, LieIge, 44 pp." type="book" year="1863">de Koninck, 1863</bibRefCitation>
</taxonomicName>
. The specimens assigned as
<taxonomicName id="4C1C4D3E4F3E1577FAE679C4FF14FE72" authorityName="de Koninck" authorityYear="1863" class="Gastropoda" family="Bellerophontidae" genus="Bellerophon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Bellerophontida" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="jonesianus">
<emphasis id="B968EAAF4F3E1577FAE679C4FAFAFE56" box="[1312,1329,442,467]" italics="true" pageId="9" pageNumber="10">B</emphasis>
. cf.
<emphasis id="B968EAAF4F3E1577FAA579C4FF14FE72" italics="true" pageId="9" pageNumber="10">jonesianus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF8D4B4C4F3E1577FECD79A0FE77FE72" author="Mansuy" box="[267,444,477,504]" firstAuthor="Mansuy" pageId="9" pageNumber="10" pagination="1 - 133" refId="ref26022" refString="Mansuy, H. (1913 a) Faunes des calcaires a Productus de l'Indochine. Memoires du Service Geologique de l'Indochine, 2 (4), 1 - 133." type="journal article" year="1913" yearSuffix="a">
Mansuy (1913
<emphasis id="B968EAAF4F3E1577FE6879A0FE77FE72" box="[430,444,478,503]" italics="true" pageId="9" pageNumber="10">a</emphasis>
</bibRefCitation>
, p. 103, pl. 11, figs 12) from the Permian of
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is the most similar form. The specimens assigned to
<taxonomicName id="4C1C4D3E4F3E1577FEA77A7CFE32FD9E" authorityName="de Koninck" authorityYear="1863" box="[353,505,514,540]" class="Gastropoda" family="Bellerophontidae" genus="Bellerophon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Bellerophontida" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="jonesianus">
<emphasis id="B968EAAF4F3E1577FEA77A7CFEB9FD9E" box="[353,370,514,539]" italics="true" pageId="9" pageNumber="10">B</emphasis>
.
<emphasis id="B968EAAF4F3E1577FE477A7CFE32FD9E" box="[385,505,514,539]" italics="true" pageId="9" pageNumber="10">jonesianus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF8D4B4C4F3E1577FDE37A7FFD09FD9E" author="Delpey" box="[549,706,513,539]" firstAuthor="Delpey" pageId="9" pageNumber="10" pagination="345 - 369" refId="ref23861" refString="Delpey, G. (1941) Les Gasteropodes permiens du Cambodge. Journal de Conchyologie, 84, 255 - 278, 345 - 369." type="journal article" year="1941">Delpey (1941</bibRefCitation>
, p. 346, figs 2122) from the Permian of
<collectingCountry id="F30B762D4F3E1577FB5B7A7FFAD8FD9E" box="[1181,1299,513,539]" name="Cambodia" pageId="9" pageNumber="10">Cambodia</collectingCountry>
, by
<bibRefCitation id="EF8D4B4C4F3E1577FA857A7CFF10FDBA" author="Waagen" firstAuthor="Waagen" pageId="9" pageNumber="10" pagination="1 - 183" refId="ref28294" refString="Waagen, W. (1880) Salt-Range Fossils. I. Productus Limestone fossils. 2. Pisces-Cephalopoda: supplement Gasteropoda. Memoirs of the Geological Survey of India (Palaeontologia Indica), 13, 1 - 183." type="journal article" year="1880">Waagen (1880</bibRefCitation>
, p. 135, pl. 8, figs 12) from the Permian of
<collectingCountry id="F30B762D4F3E1577FD107A5BFCFEFDBA" box="[726,821,549,575]" name="Pakistan" pageId="9" pageNumber="10">Pakistan</collectingCountry>
and by Nützel &amp;
<bibRefCitation id="EF8D4B4C4F3E1577FC397A5BFB0BFDBA" author="Nakazawa" box="[1023,1216,549,575]" firstAuthor="Nakazawa" pageId="9" pageNumber="10" pagination="103 - 169" refId="ref27024" refString="Nu ¨ tzel, A. &amp; Nakazawa, K. (2012) Permian (Capitanian) gastropods from the Akasaka Limestone (Gifu Prefecture, Japan). Journal of Systematic Palaeontology, 10, 103 - 169. https: // doi. org / 10.1080 / 14772019.2010.549659" type="journal article" year="2012">Nakazawa (2012</bibRefCitation>
, p. 109, fig. 8AH) from the Permian of
<collectingCountry id="F30B762D4F3E1577FE407A34FE0DFDE1" box="[390,454,586,612]" name="Japan" pageId="9" pageNumber="10">Japan</collectingCountry>
also resemble the present specimens. The present specimens resemble
<taxonomicName id="4C1C4D3E4F3E1577FB367A34FA57FDE6" authority="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel, 2020" authorityName="Ketwetsuriya &amp; Karapunar &amp; Charoentitirat &amp; Nützel" authorityYear="2020" box="[1264,1436,586,612]" class="Gastropoda" family="Bellerophontidae" genus="Bellerophon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Bellerophontida" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="erawanensis" status="sp. nov.">
<emphasis id="B968EAAF4F3E1577FB367A34FACAFDE6" box="[1264,1281,586,611]" italics="true" pageId="9" pageNumber="10">B</emphasis>
.
<emphasis id="B968EAAF4F3E1577FAD67A34FA57FDE6" box="[1296,1436,586,611]" italics="true" pageId="9" pageNumber="10">erawanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25B57D44F3E1577FF517A10FF24FD0D" box="[151,239,622,648]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
they have a more elongated shape, more distinct selenizone, stronger growth lines and wider interspaces between the thread-like growth lines.
</paragraph>
</subSubSection>
</treatment>
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