201 lines
21 KiB
XML
201 lines
21 KiB
XML
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<mods:title id="80942127726541DBF85913087AE2618B">Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane</mods:title>
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<mods:name id="8B2D06AFF66835C434CD419D4C9E3EDB" type="personal">
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<mods:namePart id="54E962849F5F0643E6556D8AEAC6A705">Ketwetsuriya, Chatchalerm</mods:namePart>
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<mods:namePart id="1D6BA88844B38A6142291ACB8F0FA6B6">Karapunar, Baran</mods:namePart>
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<mods:namePart id="A88F08913C290A3DB9F01BB16CEECFD8">Charoentitirat, Thasinee</mods:namePart>
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<mods:namePart id="4291E80619D08B0BE59343C4DB58C8F4">Nützel, Al- Exander</mods:namePart>
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<mods:title id="8557BEFC970303A2EB4D43F163A46D74">Zootaxa</mods:title>
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<mods:part id="560C1EA272B838AD3E49C5008144B74B">
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<mods:date id="4077F0AB4BF333A35EADE762A4D3CF65">2020</mods:date>
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<mods:number id="6561D3D7025CA5E931393E49F722FC34">2020-04-17</mods:number>
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<mods:number id="820D83CCF2B5E903DC12E1F09DD93095">4766</mods:number>
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<mods:identifier id="7A3898505C9EC7468D9DC561ADA22AA1" type="DOI">10.11646/zootaxa.4766.1.1</mods:identifier>
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<treatment id="03B587AB4F2C1565FF5179A3FC35FB42" ID-DOI="http://doi.org/10.5281/zenodo.3803899" ID-GBIF-Taxon="163737130" ID-Zenodo-Dep="3803899" LSID="urn:lsid:plazi:treatment:03B587AB4F2C1565FF5179A3FC35FB42" httpUri="http://treatment.plazi.org/id/03B587AB4F2C1565FF5179A3FC35FB42" lastPageNumber="28" pageId="27" pageNumber="28">
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<subSubSection id="C30665364F2C1565FF5179A3FE9EFE72" box="[151,341,477,503]" pageId="27" pageNumber="28" type="nomenclature">
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<paragraph id="8BA336BD4F2C1565FF5179A3FE9EFE72" blockId="27.[151,341,477,539]" box="[151,341,477,503]" pageId="27" pageNumber="28">
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<heading id="D0EB81D14F2C1565FF5179A3FE9EFE72" bold="true" box="[151,341,477,503]" fontSize="10" level="2" pageId="27" pageNumber="28" reason="2">
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<emphasis id="B968EAAF4F2C1565FF5179A3FE9EFE72" bold="true" box="[151,341,477,503]" pageId="27" pageNumber="28">
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<taxonomicName id="4C1C4D3E4F2C1565FF5179A3FEDFFE72" authorityName="McCoy" authorityYear="1844" box="[151,276,477,503]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F2C1565FF5179A3FEDFFE72" bold="true" box="[151,276,477,503]" italics="true" pageId="27" pageNumber="28">Naticopsis</emphasis>
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</taxonomicName>
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sp. 2
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C30665364F2C1565FF517A7FFEE0FD9E" box="[151,299,513,539]" pageId="27" pageNumber="28" type="description">
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<paragraph id="8BA336BD4F2C1565FF517A7FFEE0FD9E" blockId="27.[151,341,477,539]" box="[151,299,513,539]" pageId="27" pageNumber="28">
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(
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<figureCitation id="13272A384F2C1565FF667A7FFF3BFD9E" box="[160,240,513,539]" captionStart="FIGURE 20" captionStartId="28.[151,250,1717,1742]" captionTargetBox="[179,1409,309,1693]" captionTargetId="figure@28.[179,1409,309,1693]" captionTargetPageId="28" captionText="FIGURE 20. A–C, Naticopsis sp. 1, ESKU-19-LP 47. D–F, Naticopsis sp. 2, ESKU-19-LP 46. G–I, Naticopsis? sp. 3, ESKU- 19-LP 32. J, Naticopsis? sp. 4, ESKU-19-LP 52. K–L, Naticopsis cf. heshanensis Pan & Erwin, 2002, ESKU-19-LP 44. Scale bars represent: 2 mm (A–J); 5 mm (K–L)." figureDoi="http://doi.org/10.5281/zenodo.3763947" httpUri="https://zenodo.org/record/3763947/files/figure.png" pageId="27" pageNumber="28">Fig. 20</figureCitation>
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D–F)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F2C1565FF517A36FE41FD2E" pageId="27" pageNumber="28" type="materials_examined">
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<paragraph id="8BA336BD4F2C1565FF517A36FDB6FDE6" blockId="27.[151,1437,584,1223]" box="[151,637,584,611]" pageId="27" pageNumber="28">
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<emphasis id="B968EAAF4F2C1565FF517A36FECDFDE6" bold="true" box="[151,262,584,611]" pageId="27" pageNumber="28">Material.</emphasis>
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One specimen: ESKU-19-
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<collectionCode id="ED0DAE784F2C1565FDF77A37FD98FDE6" box="[561,595,585,611]" country="Russia" name="Laboratory of Palaeontology" pageId="27" pageNumber="28">LP</collectionCode>
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46.
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</paragraph>
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<paragraph id="8BA336BD4F2C1565FF017A13FE41FD2E" blockId="27.[151,1437,584,1223]" pageId="27" pageNumber="28">
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Dimensions (mm): ESKU-19-
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<collectionCode id="ED0DAE784F2C1565FDDD7A13FDF6FD02" box="[539,573,621,647]" country="Russia" name="Laboratory of Palaeontology" pageId="27" pageNumber="28">LP</collectionCode>
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46: height = 9.3; width = 10.3; apical angle = 112
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<superScript id="7C699BF54F2C1565FB417A12FB47FDFF" attach="none" box="[1159,1164,620,634]" fontSize="6" pageId="27" pageNumber="28">º</superScript>
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; apertural height = 7.5; apertural width = 7.7.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F2C1565FF017ACAFF22FCDA" pageId="27" pageNumber="28" type="description">
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<paragraph id="8BA336BD4F2C1565FF017ACAFF22FCDA" blockId="27.[151,1437,584,1223]" pageId="27" pageNumber="28">
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<emphasis id="B968EAAF4F2C1565FF017ACAFE92FD4A" bold="true" box="[199,345,692,719]" pageId="27" pageNumber="28">Description.</emphasis>
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Moderately low-spired shell, naticiform, consisting of three rapidly expanding whorls; whorls smooth, strongly convex, rounded, embracing at periphery; spire low but distinctly elevated; body-whorl inflated, very large and much higher than spire with height about 90% of total height; periphery at about mid-whorl; base convex; suture shallow but impressed; anomphalous; aperture evenly rounded in cross section, lip thin, inner lip arched.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F2C1565FF017B17FC35FB42" pageId="27" pageNumber="28" type="discussion">
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<paragraph id="8BA336BD4F2C1565FF017B17FC35FB42" blockId="27.[151,1437,584,1223]" pageId="27" pageNumber="28">
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<emphasis id="B968EAAF4F2C1565FF017B17FEF0FC06" bold="true" box="[199,315,873,899]" pageId="27" pageNumber="28">Remarks.</emphasis>
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<taxonomicName id="4C1C4D3E4F2C1565FE847B14FE73FC06" authorityName="McCoy" authorityYear="1844" box="[322,440,874,899]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F2C1565FE847B14FE73FC06" box="[322,440,874,899]" italics="true" pageId="27" pageNumber="28">Naticopsis</emphasis>
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</taxonomicName>
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sp. 2 differs from
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<taxonomicName id="4C1C4D3E4F2C1565FD4F7B14FD34FC06" authorityName="McCoy" authorityYear="1844" box="[649,767,874,899]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F2C1565FD4F7B14FD34FC06" box="[649,767,874,899]" italics="true" pageId="27" pageNumber="28">Naticopsis</emphasis>
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</taxonomicName>
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sp.
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<quantity id="4CE49B584F2C1565FCEB7B17FC93FC06" box="[813,856,873,899]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="27" pageNumber="28" unit="in" value="1.0">1 in</quantity>
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having more flattened upper whorl surface, lacking callus, having a higher whorl expansion rate and a rounded aperture. It resembles
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<taxonomicName id="4C1C4D3E4F2C1565FBF27BF0FE64FC4E" authority="Worthen in Meek & Worthen (1873)" authorityName="Worthen in Meek & Worthen" authorityYear="1873" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="subovata">
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<emphasis id="B968EAAF4F2C1565FBF27BF0FADCFC22" box="[1076,1303,909,935]" italics="true" pageId="27" pageNumber="28">Naticopsis subovata</emphasis>
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Worthen
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<emphasis id="B968EAAF4F2C1565FA407BF0FA57FC22" box="[1414,1436,910,935]" italics="true" pageId="27" pageNumber="28">in</emphasis>
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<bibRefCitation id="EF8D4B4C4F2C1565FF517BCFFE64FC4E" author="Meek & Worthen" box="[151,431,945,971]" firstAuthor="Meek" pageId="27" pageNumber="28" pagination="321 - 619" refId="ref26472" refString="Meek, F. B. & Worthen, A. H. (1873) Descriptions of invertebrates from the Carboniferous System. Illinois Geological Survey, 5 (2), 321 - 619." type="journal article" year="1873">Meek & Worthen (1873)</bibRefCitation>
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</taxonomicName>
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as illustrated by
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<bibRefCitation id="EF8D4B4C4F2C1565FDB27BCFFCD2FC4E" author="Knight" box="[628,793,945,971]" firstAuthor="Knight" pageId="27" pageNumber="28" pagination="359 - 392" refId="ref25163" refString="Knight, J. B. (1933 b) The gastropods of the St. Louis, Missouri, Pennsylvanian outlier: VI. The Neritidae. Journal of Paleontology, 7 (4), 359 - 392." type="journal article" year="1933" yearSuffix="b">
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Knight (1933
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<emphasis id="B968EAAF4F2C1565FCCD7BCFFCD2FC4E" box="[779,793,945,971]" italics="true" pageId="27" pageNumber="28">b</emphasis>
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</bibRefCitation>
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, p. 379, pl. 43, fig. 2a–j) in size, shell shape and pleural angle.
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<taxonomicName id="4C1C4D3E4F2C1565FF227BA8FDDCFC6A" authority="Knight, 1933 b" authorityName="Knight" authorityYear="1933" box="[228,535,981,1007]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="wortheni">
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<emphasis id="B968EAAF4F2C1565FF227BA8FF3CFC6A" box="[228,247,982,1007]" italics="true" pageId="27" pageNumber="28">N</emphasis>
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.
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<emphasis id="B968EAAF4F2C1565FEC07BABFEA1FC6A" box="[262,362,981,1007]" italics="true" pageId="27" pageNumber="28">wortheni</emphasis>
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<bibRefCitation id="EF8D4B4C4F2C1565FEB57BABFDDCFC6A" author="Knight" box="[371,535,981,1007]" firstAuthor="Knight" pageId="27" pageNumber="28" pagination="359 - 392" refId="ref25163" refString="Knight, J. B. (1933 b) The gastropods of the St. Louis, Missouri, Pennsylvanian outlier: VI. The Neritidae. Journal of Paleontology, 7 (4), 359 - 392." type="journal article" year="1933" yearSuffix="b">
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Knight, 1933
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<emphasis id="B968EAAF4F2C1565FDCF7BABFDDCFC6A" box="[521,535,981,1007]" italics="true" pageId="27" pageNumber="28">b</emphasis>
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</bibRefCitation>
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</taxonomicName>
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(p. 377, pl. 43, fig. 3a–k) is also similar.
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<bibRefCitation id="EF8D4B4C4F2C1565FC3E7BABFB6CFC6A" author="Knight" box="[1016,1191,981,1007]" firstAuthor="Knight" pageId="27" pageNumber="28" pagination="359 - 392" refId="ref25163" refString="Knight, J. B. (1933 b) The gastropods of the St. Louis, Missouri, Pennsylvanian outlier: VI. The Neritidae. Journal of Paleontology, 7 (4), 359 - 392." type="journal article" year="1933" yearSuffix="b">
|
||
Knight (1933
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<emphasis id="B968EAAF4F2C1565FB567BABFB55FC6A" box="[1168,1182,981,1007]" italics="true" pageId="27" pageNumber="28">b</emphasis>
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||
)
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</bibRefCitation>
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mentioned that these two species,
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<taxonomicName id="4C1C4D3E4F2C1565FEEF7B84FE7BFB96" authorityName="Worthen in Meek & Worthen" authorityYear="1873" box="[297,432,1017,1043]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="subovata">
|
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<emphasis id="B968EAAF4F2C1565FEEF7B84FEF7FB96" box="[297,316,1018,1043]" italics="true" pageId="27" pageNumber="28">N</emphasis>
|
||
.
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<emphasis id="B968EAAF4F2C1565FE8D7B87FE7BFB96" box="[331,432,1017,1043]" italics="true" pageId="27" pageNumber="28">subovata</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C1C4D3E4F2C1565FE2E7B84FDBBFB96" authorityName="Knight" authorityYear="1933" box="[488,624,1017,1043]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="wortheni">
|
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<emphasis id="B968EAAF4F2C1565FE2E7B84FE30FB96" box="[488,507,1018,1043]" italics="true" pageId="27" pageNumber="28">N</emphasis>
|
||
.
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<emphasis id="B968EAAF4F2C1565FDCC7B87FDBBFB96" box="[522,624,1017,1043]" italics="true" pageId="27" pageNumber="28">wortheni</emphasis>
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</taxonomicName>
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<emphasis id="B968EAAF4F2C1565FDB67B87FDBFFB96" box="[624,628,1017,1043]" italics="true" pageId="27" pageNumber="28">,</emphasis>
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are very close to each other but
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<taxonomicName id="4C1C4D3E4F2C1565FC2E7B84FBA5FB96" authorityName="Worthen in Meek & Worthen" authorityYear="1873" box="[1000,1134,1017,1043]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="subovata">
|
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<emphasis id="B968EAAF4F2C1565FC2E7B84FC30FB96" box="[1000,1019,1018,1043]" italics="true" pageId="27" pageNumber="28">N</emphasis>
|
||
.
|
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<emphasis id="B968EAAF4F2C1565FBCF7B87FBA5FB96" box="[1033,1134,1017,1043]" italics="true" pageId="27" pageNumber="28">subovata</emphasis>
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</taxonomicName>
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can be differentiated from
|
||
<taxonomicName id="4C1C4D3E4F2C1565FF517C60FED7FBB2" authorityName="Knight" authorityYear="1933" box="[151,284,1053,1079]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="wortheni">
|
||
<emphasis id="B968EAAF4F2C1565FF517C60FF61FBB2" box="[151,170,1054,1079]" italics="true" pageId="27" pageNumber="28">N</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F2C1565FF7E7C63FED7FBB2" box="[184,284,1053,1079]" italics="true" pageId="27" pageNumber="28">wortheni</emphasis>
|
||
</taxonomicName>
|
||
in having a swelling just above the mid-whorl height, and by the shape of the columellar lip, which is evenly crescent-shaped in
|
||
<taxonomicName id="4C1C4D3E4F2C1565FE057C3CFD81FBDE" authorityName="Worthen in Meek & Worthen" authorityYear="1873" box="[451,586,1089,1115]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="subovata">
|
||
<emphasis id="B968EAAF4F2C1565FE057C3CFE1DFBDE" box="[451,470,1090,1115]" italics="true" pageId="27" pageNumber="28">N</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F2C1565FE237C3FFD81FBDE" box="[485,586,1089,1115]" italics="true" pageId="27" pageNumber="28">subovata</emphasis>
|
||
</taxonomicName>
|
||
while it is slightly angled in
|
||
<taxonomicName id="4C1C4D3E4F2C1565FC5E7C3CFBD5FBDE" authorityName="Knight" authorityYear="1933" box="[920,1054,1089,1115]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="wortheni">
|
||
<emphasis id="B968EAAF4F2C1565FC5E7C3CFC60FBDE" box="[920,939,1090,1115]" italics="true" pageId="27" pageNumber="28">N</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F2C1565FC7C7C3FFBD5FBDE" box="[954,1054,1089,1115]" italics="true" pageId="27" pageNumber="28">wortheni</emphasis>
|
||
</taxonomicName>
|
||
. Although the parietal area of the present specimen is obscure it seems closer to
|
||
<taxonomicName id="4C1C4D3E4F2C1565FD617C18FCE5FBFA" authorityName="Worthen in Meek & Worthen" authorityYear="1873" box="[679,814,1125,1151]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="subovata">
|
||
<emphasis id="B968EAAF4F2C1565FD617C18FD71FBFA" box="[679,698,1126,1151]" italics="true" pageId="27" pageNumber="28">N</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F2C1565FD0F7C1BFCE5FBFA" box="[713,814,1125,1151]" italics="true" pageId="27" pageNumber="28">subovata</emphasis>
|
||
</taxonomicName>
|
||
. The present specimen is also similar to the specimens illustrated and described as
|
||
<taxonomicName id="4C1C4D3E4F2C1565FE0C7CF4FD23FB26" authority="Wanner, 1922" authorityName="Wanner" authorityYear="1922" box="[458,744,1161,1187]" class="Gastropoda" family="Anomphalidae" genus="Naticopsis" higherTaxonomySource="CoL" kingdom="Animalia" order="Trochida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="praealta">
|
||
<emphasis id="B968EAAF4F2C1565FE0C7CF4FE16FB26" box="[458,477,1162,1187]" italics="true" pageId="27" pageNumber="28">N</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F2C1565FE2F7CF7FD83FB26" box="[489,584,1161,1187]" italics="true" pageId="27" pageNumber="28">praealta</emphasis>
|
||
<bibRefCitation id="EF8D4B4C4F2C1565FD887CF7FD23FB26" author="Wanner" box="[590,744,1161,1187]" firstAuthor="Wanner" pageId="27" pageNumber="28" pagination="1 - 82" refId="ref28413" refString="Wanner, C. (1922) Die Gastropoden und Lamellibranchiaten der Dyas von Timor. Palaontologie von Timor, 11, 1 - 82." type="journal article" year="1922">Wanner, 1922</bibRefCitation>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF8D4B4C4F2C1565FCC97CF7FC68FB26" author="Batten" box="[783,931,1161,1187]" firstAuthor="Batten" pageId="27" pageNumber="28" pagination="1 - 26" refId="ref23086" refString="Batten, R. L. (1979) Permian gastropods from Perak, Malaysia. Part 2. The trochids, patellids, and neritids. American Museum Novitates, 2685, 1 - 26." type="journal article" year="1979">Batten (1979</bibRefCitation>
|
||
; p. 14, fig. 16a–b) from the Permian of
|
||
<collectingRegion id="49D8F85F4F2C1565FA907CF7FA52FB26" box="[1366,1433,1161,1187]" country="Malaysia" name="Perak" pageId="27" pageNumber="28">Perak</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30B762D4F2C1565FF517CD3FF35FB42" box="[151,254,1197,1223]" name="Malaysia" pageId="27" pageNumber="28">Malaysia</collectingCountry>
|
||
but the latter specimens are more slender and relatively high-spired.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |