325 lines
40 KiB
XML
325 lines
40 KiB
XML
<document id="D00610EC21A3C974F8292817D3C6E763" ID-CLB-Dataset="22833" ID-DOI="10.11646/zootaxa.4766.1.1" ID-GBIF-Dataset="ed4d1bf6-284a-4898-952a-58943d5a3026" ID-ISSN="1175-5334" ID-Zenodo-Dep="3763907" ID-ZooBank="urn:lsid:zoobank.org:pub:B1B5DA41-5035-4783-8D47-28857B6305AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1587670592452" checkinUser="carolina" docAuthor="Ketwetsuriya, Chatchalerm, Karapunar, Baran, Charoentitirat, Thasinee & Nützel, Al- Exander" docDate="2020" docId="03B587AB4F27156CFF517E0AFC43FA63" docLanguage="en" docName="zootaxa.4766.1.1.pdf.imf" docOrigin="Zootaxa 4766 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Worthenia humiligrada Ketwetsuriya & Karapunar & Charoentitirat & Nützel 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="19" masterDocId="FF8CFFD34F37157EFFC6787EFFCBFF85" masterDocTitle="Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane" masterLastPageNumber="47" masterPageNumber="1" pageNumber="17" updateTime="1698836971451" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="4CDD0E3987A0CD99DC8106EA7D5C374F">
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<mods:title id="52DF8718ED2BC7BCD9BA75D36558E091">Middle Permian (Roadian) gastropods from the Khao Khad Formation, Central Thailand: Implications for palaeogeography of the Indochina Terrane</mods:title>
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<mods:roleTerm id="55C9E261ACF9CB32506EF936EA0E2235">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C2A8BE1DAACD491DD418F9FEA38F8141">Ketwetsuriya, Chatchalerm</mods:namePart>
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<mods:namePart id="C767BEB726298B3C8F68FB1D28339F95">Karapunar, Baran</mods:namePart>
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<mods:namePart id="1B3A7FD0AC58E082C659B79F7C16D2B5">Charoentitirat, Thasinee</mods:namePart>
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<mods:roleTerm id="BB4C3BE7120249549C8CA65B32E81891">Author</mods:roleTerm>
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<mods:namePart id="DB09F96945487E5684A31A0851C190BE">Nützel, Al- Exander</mods:namePart>
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<mods:typeOfResource id="5C91A7B98DA701EE9E559CCE68195CDD">text</mods:typeOfResource>
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<mods:title id="5CD644EE22A1F2C7C9054C65B9011C8A">Zootaxa</mods:title>
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<mods:part id="C4A61F0DFFBB35071DBF28A56AC459A3">
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<mods:date id="3A6293182D415A8AD29BB99B5CD7C472">2020</mods:date>
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<mods:number id="3B234E87CBB11327AC3BECB6DF707C6D">2020-04-17</mods:number>
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<mods:number id="D5C99C304F92340E4365C43C9B004E68">4766</mods:number>
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<mods:identifier id="3FE693B2B5DFE92FFA47EFFFF27D60A5" type="DOI">10.11646/zootaxa.4766.1.1</mods:identifier>
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<treatment id="03B587AB4F27156CFF517E0AFC43FA63" ID-DOI="http://doi.org/10.5281/zenodo.3803875" ID-GBIF-Taxon="163737212" ID-Zenodo-Dep="3803875" LSID="urn:lsid:plazi:treatment:03B587AB4F27156CFF517E0AFC43FA63" httpUri="http://treatment.plazi.org/id/03B587AB4F27156CFF517E0AFC43FA63" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
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<subSubSection id="C30665364F27156EFF517E0AFDD3F90B" box="[151,536,1652,1679]" pageId="16" pageNumber="17" type="nomenclature">
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<paragraph id="8BA336BD4F27156EFF517E0AFDD3F90B" blockId="16.[151,536,1652,1714]" box="[151,536,1652,1679]" pageId="16" pageNumber="17">
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<heading id="D0EB81D14F27156EFF517E0AFDD3F90B" bold="true" box="[151,536,1652,1679]" fontSize="10" level="2" pageId="16" pageNumber="17" reason="2">
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<emphasis id="B968EAAF4F27156EFF517E0AFDD3F90B" bold="true" box="[151,536,1652,1679]" pageId="16" pageNumber="17">
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<taxonomicName id="4C1C4D3E4F27156EFF517E0AFE78F90A" authority="Ketwetsuriya & Karapunar & Charoentitirat & Nützel, 2020" authorityName="Ketwetsuriya & Karapunar & Charoentitirat & Nützel" authorityYear="2020" box="[151,435,1652,1679]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="humiligrada" status="sp. nov.">
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<emphasis id="B968EAAF4F27156EFF517E0AFE78F90A" bold="true" box="[151,435,1652,1679]" italics="true" pageId="16" pageNumber="17">Worthenia humiligrada</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57D44F27156EFE7C7E0AFDD3F90B" box="[442,536,1652,1678]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F27156EFF517EE6FEEEF937" box="[151,293,1688,1714]" pageId="16" pageNumber="17" type="description">
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<paragraph id="8BA336BD4F27156EFF517EE6FEEEF937" blockId="16.[151,536,1652,1714]" box="[151,293,1688,1714]" pageId="16" pageNumber="17">
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(
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<figureCitation id="13272A384F27156EFF667EE6FF25F937" box="[160,238,1688,1714]" captionStart="FIGURE 14" captionStartId="17.[151,250,1945,1970]" captionTargetBox="[155,1432,181,1920]" captionTargetId="figure@17.[155,1432,181,1920]" captionTargetPageId="17" captionText="FIGURE 14. A–I, Worthenia humiligrada sp. nov. A–D, ESKU-19-LP 22. E–F, ESKU-19-LP 33. G–H, ESKU-19-LP 23. I, ESKU-19-LP 34. J–N, Worthenia cf. pagoda Mansuy, 1912, J, ESKU-19-LP 17. K–N, ESKU-19-LP 66. O–S, Worthenia sp., ESKU-19-LP 88. Scale bars represent: 2 mm (A, B, G, H, I, K, L, O); 1 mm (C, D, J, M, N, P–S); 5 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.3763935" httpUri="https://zenodo.org/record/3763935/files/figure.png" pageId="16" pageNumber="17">Fig. 14</figureCitation>
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A–I)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F27156EFF517EA5FF21F89C" pageId="16" pageNumber="17" type="etymology">
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<paragraph id="8BA336BD4F27156EFF517EA5FF21F89C" blockId="16.[151,1436,1755,2033]" pageId="16" pageNumber="17">
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<emphasis id="B968EAAF4F27156EFF517EA5FED5F970" bold="true" box="[151,286,1755,1781]" pageId="16" pageNumber="17">Etymology.</emphasis>
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From Latin humilis, meaning low, and Latin gradus, meaning step, referring to the low-stepped whorl profile.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30665364F27156EFF017F5DFDABF874" pageId="16" pageNumber="17" type="materials_examined">
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<paragraph id="8BA336BD4F27156EFF017F5DFDCFF8B8" blockId="16.[151,1436,1755,2033]" box="[199,516,1827,1853]" pageId="16" pageNumber="17">
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<materialsCitation id="3B743CE04F27156EFF017F5DFDCFF8B8" ID-GBIF-Occurrence="2608527135" box="[199,516,1827,1853]" collectionCode="LP" pageId="16" pageNumber="17" specimenCode="ESKU-19-LP 22" specimenCount="1" typeStatus="holotype">
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<emphasis id="B968EAAF4F27156EFF017F5DFEF0F8B8" bold="true" box="[199,315,1827,1853]" pageId="16" pageNumber="17">
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<typeStatus id="54A7881F4F27156EFF017F5DFEFDF8B8" box="[199,310,1827,1853]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="DBBA9EC64F27156EFE847F5DFE34F8B8" box="[322,511,1827,1853]" pageId="16" pageNumber="17">ESKU-19-LP 22</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BA336BD4F27156EFF017F39FD64F8E4" blockId="16.[151,1436,1755,2033]" box="[199,687,1862,1889]" pageId="16" pageNumber="17">
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<emphasis id="B968EAAF4F27156EFF017F39FE8DF8E4" bold="true" box="[199,326,1863,1889]" pageId="16" pageNumber="17">
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<typeStatus id="54A7881F4F27156EFF017F39FE8AF8E4" box="[199,321,1863,1889]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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ESKU-19-
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<collectionCode id="ED0DAE784F27156EFE047F39FE2FF8E4" box="[450,484,1863,1889]" country="Russia" name="Laboratory of Palaeontology" pageId="16" pageNumber="17">LP</collectionCode>
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23, 33, 34, 35, 74
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</paragraph>
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<paragraph id="8BA336BD4F27156EFF017F15FDABF874" blockId="16.[151,1436,1755,2033]" pageId="16" pageNumber="17">
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Dimensions (mm): ESKU-19-
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<collectionCode id="ED0DAE784F27156EFDDE7F15FDF1F800" box="[536,570,1899,1925]" country="Russia" name="Laboratory of Palaeontology" pageId="16" pageNumber="17">LP</collectionCode>
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22: height = 6.8; width = 7.0; apical angle = 98º. ESKU-19-
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<collectionCode id="ED0DAE784F27156EFB2F7F15FAC0F800" box="[1257,1291,1899,1925]" country="Russia" name="Laboratory of Palaeontology" pageId="16" pageNumber="17">LP</collectionCode>
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23: height = 10.5; width = 12.7. ESKU-19-
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<collectionCode id="ED0DAE784F27156EFE237FF1FDCCF82C" box="[485,519,1935,1961]" country="Russia" name="Laboratory of Palaeontology" pageId="16" pageNumber="17">LP</collectionCode>
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33: height =
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<emphasis id="B968EAAF4F27156EFD5F7FEEFD6EF82C" box="[665,677,1936,1961]" italics="true" pageId="16" pageNumber="17">c</emphasis>
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. 16.0; width = 13.8; apical angle = 92º. ESKU-19-
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<collectionCode id="ED0DAE784F27156EFB137FF1FB3CF82C" box="[1237,1271,1935,1961]" country="Russia" name="Laboratory of Palaeontology" pageId="16" pageNumber="17">LP</collectionCode>
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34: height =
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<emphasis id="B968EAAF4F27156EFA4F7FEEFA5EF82C" box="[1417,1429,1936,1961]" italics="true" pageId="16" pageNumber="17">c</emphasis>
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. 9.0; width = 11.3; apical angle = 110º. ESKU-19-
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<collectionCode id="ED0DAE784F27156EFD7A7FCDFD15F848" box="[700,734,1971,1997]" country="Russia" name="Laboratory of Palaeontology" pageId="16" pageNumber="17">LP</collectionCode>
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35: height =
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<emphasis id="B968EAAF4F27156EFCB47FCAFCB5F848" box="[882,894,1972,1997]" italics="true" pageId="16" pageNumber="17">c</emphasis>
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. 10.9; width = 13.6; apical angle = 110º. ESKU- 19-
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<collectionCode id="ED0DAE784F27156EFF7F7FA9FF17F874" box="[185,220,2007,2033]" country="Russia" name="Laboratory of Palaeontology" pageId="16" pageNumber="17">LP</collectionCode>
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74: height =
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<emphasis id="B968EAAF4F27156EFEB47FA6FEB5F874" box="[370,382,2008,2033]" italics="true" pageId="16" pageNumber="17">c</emphasis>
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. 1.7; width =
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<emphasis id="B968EAAF4F27156EFDDD7FA6FDECF874" box="[539,551,2008,2033]" italics="true" pageId="16" pageNumber="17">c</emphasis>
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. 3.1.
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</paragraph>
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</subSubSection>
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<caption id="DF6366354F26156FFF517FE7FACBF87F" ID-DOI="http://doi.org/10.5281/zenodo.3763935" ID-Zenodo-Dep="3763935" httpUri="https://zenodo.org/record/3763935/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1945,1970]" targetBox="[155,1432,181,1920]" targetPageId="17">
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<paragraph id="8BA336BD4F26156FFF517FE7FACBF87F" blockId="17.[151,1437,1945,2042]" pageId="17" pageNumber="18">
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<emphasis id="B968EAAF4F26156FFF517FE7FEE9F837" bold="true" box="[151,290,1945,1970]" pageId="17" pageNumber="18">FIGURE 14.</emphasis>
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A–I,
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<taxonomicName id="4C1C4D3E4F26156FFEA67FE4FD85F837" authority="Ketwetsuriya & Karapunar & Charoentitirat & Nützel, 2020" authorityName="Ketwetsuriya & Karapunar & Charoentitirat & Nützel" authorityYear="2020" box="[352,590,1946,1970]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="humiligrada" status="sp. nov.">
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<emphasis id="B968EAAF4F26156FFEA67FE4FD85F837" box="[352,590,1946,1970]" italics="true" pageId="17" pageNumber="18">Worthenia humiligrada</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57D44F26156FFD907FE4FD6CF834" box="[598,679,1946,1969]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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A–D, ESKU-19-LP 22. E–F, ESKU-19-LP 33. G–H, ESKU-19-LP 23. I, ESKU-19-LP 34. J–N,
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<taxonomicName id="4C1C4D3E4F26156FFE407FC0FD36F853" authority="Mansuy, 1912" authorityName="Mansuy" authorityYear="1912" box="[390,765,1981,2006]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="pagoda">
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<emphasis id="B968EAAF4F26156FFE407FC0FE24F853" box="[390,495,1982,2006]" italics="true" pageId="17" pageNumber="18">Worthenia</emphasis>
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cf.
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<emphasis id="B968EAAF4F26156FFDD17FC0FDA8F853" box="[535,611,1982,2006]" italics="true" pageId="17" pageNumber="18">pagoda</emphasis>
|
||
<bibRefCitation id="EF8D4B4C4F26156FFDAD7FC0FD36F853" author="Mansuy" box="[619,765,1982,2006]" firstAuthor="Mansuy" pageId="17" pageNumber="18" pagination="1 - 146" refId="ref25972" refString="Mansuy, H. (1912) Paleontologie. In: Mansuy, H. & Deprat, J. (Eds.), Etude geologique du Yun-Nan oriental. Actes de la Societe Geologique de l'Indo-Chine, 1 (2), pp. 1 - 146." type="journal article" year="1912">Mansuy, 1912</bibRefCitation>
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</taxonomicName>
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, J, ESKU-19-LP 17. K–N, ESKU-19-LP 66. O–S,
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<taxonomicName id="4C1C4D3E4F26156FFACF7FC0FAB9F853" authorityName="de Koninck" authorityYear="1883" box="[1289,1394,1982,2006]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
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<emphasis id="B968EAAF4F26156FFACF7FC0FAB9F853" box="[1289,1394,1982,2006]" italics="true" pageId="17" pageNumber="18">Worthenia</emphasis>
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</taxonomicName>
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sp., ESKU-19-LP 88. Scale bars represent: 2 mm (A, B, G, H, I, K, L, O); 1 mm (C, D, J, M, N, P–S); 5 mm (E, F).
|
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</paragraph>
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||
</caption>
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<subSubSection id="C30665364F25156CFF0178E7FE74FF7F" pageId="18" pageNumber="19" type="distribution">
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<paragraph id="8BA336BD4F25156CFF0178E7FE74FF7F" blockId="18.[151,1437,152,1511]" pageId="18" pageNumber="19">
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<emphasis id="B968EAAF4F25156CFF0178E7FD6AFF36" bold="true" box="[199,673,152,179]" pageId="18" pageNumber="19">Type locality and stratigraphical range.</emphasis>
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Erawan Hill, Chong Sarika sub-district, Phatthana Nikhom district, located about
|
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<quantity id="4CE49B584F25156CFEF178C3FEB4FF53" box="[311,383,188,215]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="18" pageNumber="19" unit="km" value="13.0">13 km</quantity>
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east of
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<collectingRegion id="49D8F85F4F25156CFE1C78C2FD69FF53" box="[474,674,188,214]" country="Thailand" name="Lop Buri" pageId="18" pageNumber="19">Lopburi Province</collectingRegion>
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||
, Central
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<collectingCountry id="F30B762D4F25156CFCCF78C2FCA6FF53" box="[777,877,188,214]" name="Thailand" pageId="18" pageNumber="19">Thailand</collectingCountry>
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(
|
||
<figureCitation id="13272A384F25156CFCBB78C2FC0BFF52" box="[893,960,188,215]" captionStart="FIGURE" captionStartId="1.[151,168,1975,2001]" captionTargetBox="[243,1345,1164,1941]" captionTargetId="figure@1.[236,1350,1160,1948]" captionTargetPageId="1" captionText="FIGURE 1. Geological map of the study location at Erawan Hill, Chong Sarika sub-district, Phatthana Nikhom district, located about 13 km east of Lopburi Province, Central Thailand." figureDoi="http://doi.org/10.5281/zenodo.3763909" httpUri="https://zenodo.org/record/3763909/files/figure.png" pageId="18" pageNumber="19">Fig. 1</figureCitation>
|
||
), Khao Khad Formation,
|
||
<collectingRegion id="49D8F85F4F25156CFB2178C2FA8DFF53" box="[1255,1350,188,214]" country="Thailand" name="Sara Buri" pageId="18" pageNumber="19">Saraburi</collectingRegion>
|
||
Group, Middle Permian, Roadian.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30665364F25156CFF01797AFAD4FD77" pageId="18" pageNumber="19" type="description">
|
||
<paragraph id="8BA336BD4F25156CFF01797AFAD4FD77" blockId="18.[151,1437,152,1511]" pageId="18" pageNumber="19">
|
||
<emphasis id="B968EAAF4F25156CFF01797AFE92FE9A" bold="true" box="[199,345,260,287]" pageId="18" pageNumber="19">Description.</emphasis>
|
||
Shell low-spired for genus, gradate, trochiform; first whorl planispiral; early teleoconch whorls (about first two or three whorls) low-spired, smooth, convex, rounded; second and third whorls with fine spiral and axial ribs; suture distinct and impressed; later whorls sharply angulated at about mid-whorl face, the middle angulation separates subsutural ramp from subvertical lower whorl face; subsutural ramp slightly convex near adapical suture to concave on the remaining part; ramp inclining at an angle of 35–50°; subsutural ramp ornamented with up to eight spiral ribs and equally spaced prosocyrt fine collabral ribs or strengthened growth lines; spiral ribs and collabral ribs form nodes at intersections, forming fine reticulate pattern; ornamentation more prominent near adapical suture; lower whorl face subvertical, concave, ornamented with up to six strong spiral cords and numerous faint prosocyrt collabral threads or growth lines; mid angulation forms crest-like periphery of whorl where selenizone situated; selenizone sharply angular covered by v-shaped lunulae; lunulae form nodes when intersecting with angulation; selenizone bordered with fine spiral ribs, upper rib represents lowest spiral rib of subsutural ramp, while lower rib represents uppermost spiral rib of the subvertical lower whorl face; intersections of lunulae and angulation nodular; basal edge sharply angulated; base convex, ornamented by several evenly spaced spiral ribs and very weak opisthocyrt collabral threads; suture shallow, indistinct; narrowly phaneromphalous; aperture unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30665364F25156CFF017A83FC43FA63" pageId="18" pageNumber="19" type="discussion">
|
||
<paragraph id="8BA336BD4F25156CFF017A83FC76FC22" blockId="18.[151,1437,152,1511]" pageId="18" pageNumber="19">
|
||
<emphasis id="B968EAAF4F25156CFF017A83FEF0FC92" bold="true" box="[199,315,765,791]" pageId="18" pageNumber="19">Remarks.</emphasis>
|
||
The convex early teleoconch whorls, the position and the ornamentation pattern of the selenizone suggest that this species belongs to the genus
|
||
<taxonomicName id="4C1C4D3E4F25156CFD527B5EFC13FCBE" authority="de Koninck, 1883" authorityName="de Koninck" authorityYear="1883" box="[660,984,800,827]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAAF4F25156CFD527B5EFCC3FCBF" box="[660,776,800,826]" italics="true" pageId="18" pageNumber="19">Worthenia</emphasis>
|
||
<bibRefCitation id="EF8D4B4C4F25156CFCC87B5EFC13FCBE" author="Koninck, L. G. de" box="[782,984,800,827]" firstAuthor="de Koninck" pageId="18" pageNumber="19" pagination="1 - 240" refId="ref25592" refString="Koninck, L. G. de (1883) Faune du calcaire carbonifere de la Belgique. 4. Gasteropodes (suite en fin). Annales du Musee Royale d'Histoire Naturelle Belgique, 8, 1 - 240." type="journal article" year="1883">de Koninck, 1883</bibRefCitation>
|
||
</taxonomicName>
|
||
. The distinguishing characters of
|
||
<taxonomicName id="4C1C4D3E4F25156CFA897B5EFE9CFCDB" authority="Ketwetsuriya & Karapunar & Charoentitirat & Nützel, 2020" authorityName="Ketwetsuriya & Karapunar & Charoentitirat & Nützel" authorityYear="2020" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="humiligrada" status="sp. nov.">
|
||
<emphasis id="B968EAAF4F25156CFA897B5EFE9CFCDB" italics="true" pageId="18" pageNumber="19">Worthenia humiligrada</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B57D44F25156CFE987B3BFE7FFCDA" box="[350,436,837,863]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are the relatively low spire, wide umbilicus, weak subsutural convexity of the ramp, sharp angulation at the basal edge and nodular selenizone. The combination of these characters distinguishes
|
||
<taxonomicName id="4C1C4D3E4F25156CFAE37B17FF11FC23" authority="Ketwetsuriya & Karapunar & Charoentitirat & Nützel, 2020" authorityName="Ketwetsuriya & Karapunar & Charoentitirat & Nützel" authorityYear="2020" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="humiligrada" status="sp. nov.">
|
||
<emphasis id="B968EAAF4F25156CFAE37B17FAF7FC07" box="[1317,1340,873,898]" italics="true" pageId="18" pageNumber="19">W</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F25156CFA8D7B16FF11FC23" italics="true" pageId="18" pageNumber="19">humiligrada</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B57D44F25156CFF277BF3FEF3FC22" box="[225,312,909,935]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
from the other species belonging to the genus
|
||
<taxonomicName id="4C1C4D3E4F25156CFC847BF2FC7DFC23" authorityName="de Koninck" authorityYear="1883" box="[834,950,908,934]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAAF4F25156CFC847BF2FC7DFC23" box="[834,950,908,934]" italics="true" pageId="18" pageNumber="19">Worthenia</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA336BD4F25156CFF017BCEFC43FA63" blockId="18.[151,1437,152,1511]" pageId="18" pageNumber="19">
|
||
There are some similar Permian species in Southeast Asia which have been assigned to
|
||
<taxonomicName id="4C1C4D3E4F25156CFB5E7BCEFAC7FC4F" authorityName="de Koninck" authorityYear="1883" box="[1176,1292,944,970]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B968EAAF4F25156CFB5E7BCEFAC7FC4F" box="[1176,1292,944,970]" italics="true" pageId="18" pageNumber="19">Worthenia</emphasis>
|
||
</taxonomicName>
|
||
, for instance
|
||
<taxonomicName id="4C1C4D3E4F25156CFF517BABFDDEFC6A" authority="(Mansuy, 1912)" baseAuthorityName="Mansuy" baseAuthorityYear="1912" box="[151,533,980,1007]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="multicarinata">
|
||
<emphasis id="B968EAAF4F25156CFF517BABFF65FC6B" box="[151,174,981,1006]" italics="true" pageId="18" pageNumber="19">W</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F25156CFF797BAAFE9CFC6B" box="[191,343,980,1006]" italics="true" pageId="18" pageNumber="19">multicarinata</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8D4B4C4F25156CFEAF7BABFDC6FC6A" author="Mansuy" box="[361,525,981,1007]" firstAuthor="Mansuy" pageId="18" pageNumber="19" pagination="1 - 146" refId="ref25972" refString="Mansuy, H. (1912) Paleontologie. In: Mansuy, H. & Deprat, J. (Eds.), Etude geologique du Yun-Nan oriental. Actes de la Societe Geologique de l'Indo-Chine, 1 (2), pp. 1 - 146." type="journal article" year="1912">Mansuy, 1912</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from the Carboniferous of
|
||
<collectingRegion id="49D8F85F4F25156CFC937BABFC65FC6A" box="[853,942,981,1007]" country="China" name="Yunnan" pageId="18" pageNumber="19">Yunnan</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30B762D4F25156CFC7D7BAAFC35FC6B" box="[955,1022,980,1006]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF8D4B4C4F25156CFBC97BABFB65FC6A" author="Mansuy" box="[1039,1198,981,1007]" firstAuthor="Mansuy" pageId="18" pageNumber="19" pagination="1 - 146" refId="ref25972" refString="Mansuy, H. (1912) Paleontologie. In: Mansuy, H. & Deprat, J. (Eds.), Etude geologique du Yun-Nan oriental. Actes de la Societe Geologique de l'Indo-Chine, 1 (2), pp. 1 - 146." type="journal article" year="1912">Mansuy 1912</bibRefCitation>
|
||
) and the Permian of
|
||
<collectingCountry id="F30B762D4F25156CFF517B86FEC1FB97" box="[151,266,1016,1042]" name="Cambodia" pageId="18" pageNumber="19">Cambodia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF8D4B4C4F25156CFEDD7B86FE64FB96" author="Delpey" box="[283,431,1016,1043]" firstAuthor="Delpey" pageId="18" pageNumber="19" pagination="345 - 369" refId="ref23861" refString="Delpey, G. (1941) Les Gasteropodes permiens du Cambodge. Journal de Conchyologie, 84, 255 - 278, 345 - 369." type="journal article" year="1941">Delpey 1941</bibRefCitation>
|
||
) and
|
||
<collectingCountry id="F30B762D4F25156CFE347B86FD92FB97" box="[498,601,1016,1042]" name="Malaysia" pageId="18" pageNumber="19">Malaysia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF8D4B4C4F25156CFDAF7B87FD33FB96" author="Batten" box="[617,760,1017,1043]" firstAuthor="Batten" pageId="18" pageNumber="19" pagination="1 - 44" refId="ref23042" refString="Batten, R. L. (1972) Permian gastropods and chitons from Perak, Malaysia. Part 1. Chitons, bellerophontids, euomphalids and pleurotomarians. Bulletin of the American Museum of Natural History, 147, 1 - 44." type="journal article" year="1972">Batten 1972</bibRefCitation>
|
||
, p. 32, figs 36–38) is very similar in shape but it has more prominent subsutural nodes on the upper whorl face, a distinctly impressed suture and its basal edge is rounded angular and ornamented with nodes. The
|
||
<typeStatus id="54A7881F4F25156CFDA17C3FFD5CFBDE" box="[615,663,1089,1115]" pageId="18" pageNumber="19">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="4C1C4D3E4F25156CFCFF7C3FFB23FBDE" authority="(Stuckenberg, 1905)" baseAuthorityName="Stuckenberg" baseAuthorityYear="1905" box="[825,1256,1088,1115]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="schirjaevensis">
|
||
<emphasis id="B968EAAF4F25156CFCFF7C3FFC9BFBDF" box="[825,848,1089,1114]" italics="true" pageId="18" pageNumber="19">W</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F25156CFC987C3EFC36FBDF" box="[862,1021,1088,1114]" italics="true" pageId="18" pageNumber="19">schirjaevensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8D4B4C4F25156CFBCA7C3EFB2BFBDE" author="Stuckenberg" box="[1036,1248,1088,1115]" firstAuthor="Stuckenberg" pageId="18" pageNumber="19" pagination="1 - 115" refId="ref27737" refString="Stuckenberg, A. (1905) Die Fauna der obercarbonischen Suite des Wolgadurchbruches bei Samara. Geologie Kommitte Trudy Memoir, 23, 1 - 115." type="journal article" year="1905">Stuckenberg, 1905</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from the Upper Carboniferous of
|
||
<collectingRegion id="49D8F85F4F25156CFE987C1BFE7EFBFA" box="[350,437,1125,1151]" country="Russia" name="Samara" pageId="18" pageNumber="19">Samara</collectingRegion>
|
||
,
|
||
<collectingCountry id="F30B762D4F25156CFE077C1AFDC7FBFB" box="[449,524,1124,1150]" name="Russia" pageId="18" pageNumber="19">Russia</collectingCountry>
|
||
and the specimens assigned to
|
||
<taxonomicName id="4C1C4D3E4F25156CFCB67C1BFBFEFBFB" baseAuthorityName="Stuckenberg" baseAuthorityYear="1905" box="[880,1077,1124,1151]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="schirjaevensis">
|
||
<emphasis id="B968EAAF4F25156CFCB67C1BFC4CFBFB" box="[880,903,1125,1150]" italics="true" pageId="18" pageNumber="19">W</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F25156CFC507C1AFBFEFBFB" box="[918,1077,1124,1150]" italics="true" pageId="18" pageNumber="19">schirjaevensis</emphasis>
|
||
</taxonomicName>
|
||
from the Permian of
|
||
<collectingCountry id="F30B762D4F25156CFAEF7C1AFA57FBFB" box="[1321,1436,1124,1150]" name="Cambodia" pageId="18" pageNumber="19">Cambodia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF8D4B4C4F25156CFF597CF6FEF8FB26" author="Delpey" box="[159,307,1160,1187]" firstAuthor="Delpey" pageId="18" pageNumber="19" pagination="345 - 369" refId="ref23861" refString="Delpey, G. (1941) Les Gasteropodes permiens du Cambodge. Journal de Conchyologie, 84, 255 - 278, 345 - 369." type="journal article" year="1941">Delpey 1941</bibRefCitation>
|
||
, fig. 27) has a very similar whorl profile and ornamentation but they differ from
|
||
<taxonomicName id="4C1C4D3E4F25156CFB027CF7FABFFB27" authority="Ketwetsuriya & Karapunar & Charoentitirat & Nützel, 2020" authorityName="Ketwetsuriya & Karapunar & Charoentitirat & Nützel" authorityYear="2020" box="[1220,1396,1160,1187]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="humiligrada" status="sp. nov.">
|
||
<emphasis id="B968EAAF4F25156CFB027CF7FB10FB27" box="[1220,1243,1161,1186]" italics="true" pageId="18" pageNumber="19">W</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F25156CFB2C7CF6FABFFB27" box="[1258,1396,1160,1186]" italics="true" pageId="18" pageNumber="19">humiligrada</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B57D44F25156CFABA7CF7FF0CFB42" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by having a higher spire. According to
|
||
<bibRefCitation id="EF8D4B4C4F25156CFD5E7CD3FCE5FB42" author="Batten" box="[664,814,1197,1223]" firstAuthor="Batten" pageId="18" pageNumber="19" pagination="1 - 44" refId="ref23042" refString="Batten, R. L. (1972) Permian gastropods and chitons from Perak, Malaysia. Part 1. Chitons, bellerophontids, euomphalids and pleurotomarians. Bulletin of the American Museum of Natural History, 147, 1 - 44." type="journal article" year="1972">Batten (1972</bibRefCitation>
|
||
, fig. 38),
|
||
<taxonomicName id="4C1C4D3E4F25156CFC647CD3FA92FB42" authority="(Stuckenberg, 1905)" baseAuthorityName="Stuckenberg" baseAuthorityYear="1905" box="[930,1369,1196,1223]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="schirjaevensis">
|
||
<emphasis id="B968EAAF4F25156CFC647CD3FC72FB43" box="[930,953,1197,1222]" italics="true" pageId="18" pageNumber="19">W</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F25156CFC0C7CD2FBA2FB43" box="[970,1129,1196,1222]" italics="true" pageId="18" pageNumber="19">schirjaevensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8D4B4C4F25156CFBBC7CD2FA9AFB42" author="Stuckenberg" box="[1146,1361,1196,1223]" firstAuthor="Stuckenberg" pageId="18" pageNumber="19" pagination="1 - 115" refId="ref27737" refString="Stuckenberg, A. (1905) Die Fauna der obercarbonischen Suite des Wolgadurchbruches bei Samara. Geologie Kommitte Trudy Memoir, 23, 1 - 115." type="journal article" year="1905">Stuckenberg, 1905</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
lacks nodes on the selenizone.
|
||
<taxonomicName id="4C1C4D3E4F25156CFE757CAFFD35FB6F" authority="Winters, 1963" authorityName="Winters" authorityYear="1963" box="[435,766,1232,1259]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="arizonensis">
|
||
<emphasis id="B968EAAF4F25156CFE757CAFFE01FB6F" box="[435,458,1233,1258]" italics="true" pageId="18" pageNumber="19">W</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F25156CFE1F7CAFFD92FB6F" box="[473,601,1233,1258]" italics="true" pageId="18" pageNumber="19">arizonensis</emphasis>
|
||
<bibRefCitation id="EF8D4B4C4F25156CFDA67CAEFD35FB6F" author="Winters" box="[608,766,1232,1258]" firstAuthor="Winters" pageId="18" pageNumber="19" pagination="1 - 99" refId="ref28656" refString="Winters, S. S. (1963) Supai Formation (Permian) of eastern Arizona. Geological Society of America Memoir, 89, 1 - 99. https: // doi. org / 10.1130 / MEM 89 - p 1" type="journal article" year="1963">Winters, 1963</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Permian of
|
||
<collectingRegion id="49D8F85F4F25156CFC377CAEFB80FB6F" box="[1009,1099,1232,1258]" country="United States of America" name="Arizona" pageId="18" pageNumber="19">Arizona</collectingRegion>
|
||
is another species with planispiral early whorl but differs from
|
||
<taxonomicName id="4C1C4D3E4F25156CFDD37C8BFD0FFA8B" authority="Ketwetsuriya & Karapunar & Charoentitirat & Nützel, 2020" authorityName="Ketwetsuriya & Karapunar & Charoentitirat & Nützel" authorityYear="2020" box="[533,708,1268,1295]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="humiligrada" status="sp. nov.">
|
||
<emphasis id="B968EAAF4F25156CFDD37C8BFDE7FA8B" box="[533,556,1269,1294]" italics="true" pageId="18" pageNumber="19">W</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F25156CFDFC7C8AFD0FFA8B" box="[570,708,1268,1294]" italics="true" pageId="18" pageNumber="19">humiligrada</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25B57D44F25156CFD0C7C8BFCEBFA8A" box="[714,800,1269,1295]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by having a narrower upper whorl face, a very prominent subsutural convexity, by lacking the sharp angulation at the basal edge and by having a very narrow or no umbilicus.
|
||
<taxonomicName id="4C1C4D3E4F25156CFF517D43FE70FAD2" authority="Batten, 1989" authorityName="Batten" authorityYear="1989" box="[151,443,1340,1367]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="crenulata">
|
||
<emphasis id="B968EAAF4F25156CFF517D43FF65FAD3" box="[151,174,1341,1366]" italics="true" pageId="18" pageNumber="19">W</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F25156CFF7A7D42FEEDFAD3" box="[188,294,1340,1366]" italics="true" pageId="18" pageNumber="19">crenulata</emphasis>
|
||
<bibRefCitation id="EF8D4B4C4F25156CFEEA7D43FE70FAD2" author="Batten" box="[300,443,1341,1367]" firstAuthor="Batten" pageId="18" pageNumber="19" pagination="1 - 64" refId="ref23162" refString="Batten, R. L. (1989) Permian Gastropoda of the Southwestern United States. 7. Pleurotomariacea: Eotomariidae, Lophospiriidae, Gosseletinidae. American Museum Novitates, 2958, 1 - 64." type="journal article" year="1989">Batten, 1989</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Permian of southwestern
|
||
<collectingCountry id="F30B762D4F25156CFC877D42FC11FAD2" box="[833,986,1340,1367]" name="United States of America" pageId="18" pageNumber="19">United States</collectingCountry>
|
||
, with steeper upper whorl face and more prominent nodes on its selenizone. The specimens assigned to
|
||
<taxonomicName id="4C1C4D3E4F25156CFC937D1FFB59FAFE" authority="Chronic, 1952" authorityName="Chronic" authorityYear="1952" box="[853,1170,1376,1403]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="corrugata">
|
||
<emphasis id="B968EAAF4F25156CFC937D1FFCA7FAFF" box="[853,876,1377,1402]" italics="true" pageId="18" pageNumber="19">W</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F25156CFCBD7D1FFC20FAFF" box="[891,1003,1377,1402]" italics="true" pageId="18" pageNumber="19">corrugata</emphasis>
|
||
<bibRefCitation id="EF8D4B4C4F25156CFC347D1EFB59FAFE" author="Chronic" box="[1010,1170,1376,1403]" firstAuthor="Chronic" pageId="18" pageNumber="19" pagination="95 - 166" refId="ref23437" refString="Chronic, H. (1952) Molluscan fauna from the Permian Kaibab Formation, Walnut Canyon, Arizona. Geological Society America Bulletin, 63 (2), 95 - 166. https: // doi. org / 10.1130 / 0016 - 7606 (1952) 63 [95: MFFTPK] 2.0. CO; 2" type="journal article" year="1952">Chronic, 1952</bibRefCitation>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF8D4B4C4F25156CFB7B7D1FFA9CFAFE" author="Batten" box="[1213,1367,1377,1403]" firstAuthor="Batten" pageId="18" pageNumber="19" pagination="1 - 64" refId="ref23162" refString="Batten, R. L. (1989) Permian Gastropoda of the Southwestern United States. 7. Pleurotomariacea: Eotomariidae, Lophospiriidae, Gosseletinidae. American Museum Novitates, 2958, 1 - 64." type="journal article" year="1989">Batten (1989)</bibRefCitation>
|
||
differ from
|
||
<taxonomicName id="4C1C4D3E4F25156CFF157DFBFEB4FA1B" authorityName="Ketwetsuriya & Karapunar & Charoentitirat & Nützel" authorityYear="2020" box="[211,383,1412,1438]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="humiligrada">
|
||
<emphasis id="B968EAAF4F25156CFF157DFBFEB4FA1B" box="[211,383,1412,1438]" italics="true" pageId="18" pageNumber="19">W. humiligrada</emphasis>
|
||
</taxonomicName>
|
||
by having a stronger spiral ornament, a sharper collabral ornament and by the presence of axial folds on its upper whorl face. The specimens assigned to
|
||
<taxonomicName id="4C1C4D3E4F25156CFCD37DD7FC61FA47" authorityName="Chronic" authorityYear="1952" box="[789,938,1449,1475]" class="Gastropoda" family="Gosseletinidae" genus="Worthenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleurotomariida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="corrugata">
|
||
<emphasis id="B968EAAF4F25156CFCD37DD7FCE7FA47" box="[789,812,1449,1474]" italics="true" pageId="18" pageNumber="19">W</emphasis>
|
||
.
|
||
<emphasis id="B968EAAF4F25156CFCFC7DD7FC61FA47" box="[826,938,1449,1474]" italics="true" pageId="18" pageNumber="19">corrugata</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EF8D4B4C4F25156CFC157DD6FB19FA46" author="Kulas & Batten" box="[979,1234,1448,1475]" firstAuthor="Kulas" pageId="18" pageNumber="19" pagination="33 - 58" refId="ref25797" refString="Kulas, H. A. & Batten, R. L. (1997) Silicified gastropods from the Permian Phosphoria rock complex of Wyoming. Contributions to Geology, University of Wyoming, 31 (2), 33 - 58." type="journal article" year="1997">Kulas & Batten (1997)</bibRefCitation>
|
||
from the Permian of
|
||
<collectingRegion id="49D8F85F4F25156CFF737DB2FEE8FA63" box="[181,291,1484,1510]" country="United States of America" name="Wyoming" pageId="18" pageNumber="19">Wyoming</collectingRegion>
|
||
differ by having a narrow umbilicus and faint lunulae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |