168 lines
20 KiB
XML
168 lines
20 KiB
XML
<document id="D852F73567EB05540E8C56F129540B25" ID-CLB-Dataset="28092" ID-DOI="10.11646/zootaxa.4504.1.2" ID-GBIF-Dataset="cffb097a-824a-4a4d-9e24-1962e7e14021" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606146" ID-ZooBank="477150EE-9E89-4F03-879D-38D418C2FFEB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553577904264" checkinUser="plazi" docAuthor="Mey, Wolfram & Ospina-Torres, Rodulfo" docDate="2018" docId="03B587A0FA7B3628AFD388BACE11B485" docLanguage="en" docName="zootaxa.4504.1.2.pdf" docOrigin="Zootaxa 4504 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Marilia bogotaensis Mey & Ospina-Torres 2018, spec. nov." docType="treatment" docVersion="8" lastPageNumber="36" masterDocId="FF8CFFD8FA763625AF448B6BCD47B315" masterDocTitle="Contribution to the Trichoptera fauna of the river La Vieja, Bogotá, Colombia (Insecta: Trichoptera)" masterLastPageNumber="40" masterPageNumber="23" pageNumber="36" updateTime="1698712709265" updateUser="ExternalLinkService">
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<mods:mods id="5D86B8F792209B12089B880C845DDD87" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8307AA734C4A4395F0A05F379C878EFF">
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<mods:title id="91F3354A19CD9E0A38CB747A3862E52D">Contribution to the Trichoptera fauna of the river La Vieja, Bogotá, Colombia (Insecta: Trichoptera)</mods:title>
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<mods:name id="F548EE6E141C60168D366B408831EAAA" type="personal">
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<mods:roleTerm id="A6CE173866B101E334F48B5E926C6F66">Author</mods:roleTerm>
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<mods:namePart id="279CCD2F5F062BF93DD9488E7AF31D78">Mey, Wolfram</mods:namePart>
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<mods:name id="55B8651614209A92CD9CC25C63DCF1BA" type="personal">
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<mods:roleTerm id="E99700807C6CA3A95C3A6A3B0EFA880A">Author</mods:roleTerm>
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<mods:namePart id="8A3757B7421F46DC689459F899A2AC7C">Ospina-Torres, Rodulfo</mods:namePart>
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<mods:typeOfResource id="66E42F803E344E11B305849724385771">text</mods:typeOfResource>
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<mods:title id="6553B6B6D05F3B080A56F1CF77490809">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="50590E503712A222F8F478F39F3FB85F">
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<mods:date id="39C45CED140A507DF9566479BB4D8CC7">2018</mods:date>
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<mods:detail id="EFB835F5B6E31DAD030EE690C8808CF7" type="pubDate">
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<mods:number id="C2F21A20C4B53109DBD3354353C75AA4">2018-10-23</mods:number>
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<mods:detail id="09186D5BC4206B4B30ED902114DF2E52" type="volume">
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<mods:number id="6252A40FA4B38697CCBC7A78F2BC4775">4504</mods:number>
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</mods:detail>
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<mods:detail id="139C8DC6CA3FA4E7A657106AC78AD510" type="issue">
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<mods:number id="CA7185AFF9B7510F80A0235C23982C19">1</mods:number>
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<mods:classification id="3C74CF83609A876397C19BBFBC6FBB0B">journal article</mods:classification>
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<mods:identifier id="DBB45BBF44410F0E18DF1B361B7578B3" type="CLB-Dataset">28092</mods:identifier>
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<mods:identifier id="C274963E80A718DAAF938D1C8072D6AE" type="DOI">10.11646/zootaxa.4504.1.2</mods:identifier>
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<treatment id="03B587A0FA7B3628AFD388BACE11B485" ID-DOI="http://doi.org/10.5281/zenodo.5960836" ID-GBIF-Taxon="154494775" ID-Zenodo-Dep="5960836" LSID="urn:lsid:plazi:treatment:03B587A0FA7B3628AFD388BACE11B485" httpUri="http://treatment.plazi.org/id/03B587A0FA7B3628AFD388BACE11B485" lastPageNumber="36" pageId="13" pageNumber="36">
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<subSubSection id="C306653DFA7B3628AFD388BACF42B0FE" box="[151,517,977,1003]" pageId="13" pageNumber="36" type="nomenclature">
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<paragraph id="8BA336B6FA7B3628AFD388BACF42B0FE" blockId="13.[151,517,977,1037]" box="[151,517,977,1003]" pageId="13" pageNumber="36">
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<heading id="D0EB81DAFA7B3628AFD388BACF42B0FE" bold="true" box="[151,517,977,1003]" fontSize="11" level="1" pageId="13" pageNumber="36" reason="1">
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<emphasis id="B968EAA4FA7B3628AFD388BACF42B0FE" bold="true" box="[151,517,977,1003]" pageId="13" pageNumber="36">
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<taxonomicName id="4C1C4D35FA7B3628AFD388BACCC1B0FE" authority="Mey & Ospina-Torres, 2018" authorityName="Mey & Ospina-Torres" authorityYear="2018" box="[151,390,977,1003]" class="Insecta" family="Odontoceridae" genus="Marilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="bogotaensis" status="sp. nov.">
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<emphasis id="B968EAA4FA7B3628AFD388BACCC1B0FE" bold="true" box="[151,390,977,1003]" italics="true" pageId="13" pageNumber="36">Marilia bogotaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25B57DFFA7B3628AECA88BACF42B0FE" box="[398,517,977,1003]" pageId="13" pageNumber="36" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C306653DFA7B3628AFD3889FCEEEB76D" pageId="13" pageNumber="36" type="description">
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<paragraph id="8BA336B6FA7B3628AFD3889FCCCFB718" blockId="13.[151,517,977,1037]" box="[151,392,1012,1037]" pageId="13" pageNumber="36">
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(
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<figureCitation id="13272A33FA7B3628AFDB889FCDB5B719" box="[159,242,1012,1037]" captionStart="FIGURES 10–13" captionStartId="11.[151,264,1883,1905]" captionTargetBox="[201,1392,205,1769]" captionTargetId="figure@11.[189,1404,193,1871]" captionTargetPageId="11" captionText="FIGURES 10–13. Trichoptera spp. 10A–10C, Contulma spec., female genitalia, scale = 0.3 mm: 10A, left lateral; 10B, dorsal; 10C, ventral. 11A–11D, Phylloicus monserratensis spec. nov., male genitalia, scale = 0.5 mm: 11A, left lateral; 11B, dorsal; 11C, ventral; 11D, phallic apparatus, ventral. 12A–12D, Marilia bogotaensis spec. nov., male genitalia, scale: 0.3 mm: 12A, left lateral; 12B, ventral; 12C, dorsal; 12D, phallic apparatus, left lateral. 13A–13B, Oecetis chapinera spec. nov., scale = 0.25 mm: 13A, left lateral, with phallic apparatus in situ; 13B, ventral." httpUri="https://zenodo.org/record/2606156/files/figure.png" pageId="13" pageNumber="36">Figs 12</figureCitation>
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A–12D, 15F)
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</paragraph>
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<paragraph id="8BA336B6FA7B3628AFD38F57C865B741" blockId="13.[151,1437,1084,1936]" box="[151,1314,1084,1109]" pageId="13" pageNumber="36">
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<materialsCitation id="3B743CEBFA7B3628AFD38F57C865B741" ID-GBIF-Occurrence="2028137678" box="[151,1314,1084,1109]" collectingDate="2017-02-06" collectionCode="ICN" collectorName="W. Mey" country="Colombia" location="Quebrada La Vieja" municipality="Chapinero" pageId="13" pageNumber="36" specimenCount="1" specimenCount-male="1" stateProvince="Bogota" typeStatus="holotype">
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<typeStatus id="54A78814FA7B3628AFD38F57CDB8B740" box="[151,255,1084,1109]" pageId="13" pageNumber="36" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D1AFD3FFA7B3628AE428F56CC5AB741" box="[262,285,1085,1108]" pageId="13" pageNumber="36" type="male">♂</specimenCount>
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,
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<collectingCountry id="F30B7626FA7B3628AE6D8F57CCDDB740" box="[297,410,1084,1109]" name="Colombia" pageId="13" pageNumber="36">Colombia</collectingCountry>
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,
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<collectingRegion id="49D8F854FA7B3628AEE28F57CCBDB741" box="[422,506,1084,1108]" country="Colombia" name="Bogota" pageId="13" pageNumber="36">Bogotá</collectingRegion>
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,
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<collectingMunicipality id="6BC7ACCCFA7B3628AD408F57CF3BB740" box="[516,636,1084,1109]" pageId="13" pageNumber="36">Chapinero</collectingMunicipality>
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,
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<location id="8EC3606DFA7B3628ADC38F57CE19B740" LSID="urn:lsid:plazi:treatment:03B587A0FA7B3628AFD388BACE11B485:8EC3606DFA7B3628ADC38F57CE19B740" box="[647,862,1084,1109]" country="Colombia" municipality="Chapinero" name="Quebrada La Vieja" pageId="13" pageNumber="36" stateProvince="Bogota">Quebrada La Vieja</location>
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,
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<date id="FFA21076FA7B3628AC2D8F57CE89B740" box="[873,974,1084,1109]" pageId="13" pageNumber="36" value="2017-02-06">
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<collectingDate id="EFE6E99EFA7B3628AC2D8F57CE89B740" box="[873,974,1084,1109]" pageId="13" pageNumber="36" value="2017-02-06">6.ii.2017</collectingDate>
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</date>
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,
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<collectingMethod id="525D4EA1FA7B3628AC9E8F57C97AB741" box="[986,1085,1084,1109]" pageId="13" pageNumber="36">hand net</collectingMethod>
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, leg.
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<collectorName id="26E95360FA7B3628AB3C8F56C995B740" box="[1144,1234,1085,1109]" pageId="13" pageNumber="36">W. Mey</collectorName>
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(
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<collectionCode id="ED0DAE73FA7B3628ABA48F57C851B741" box="[1248,1302,1084,1108]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="13" pageNumber="36">ICN</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BA336B6FA7B3628AF838F0BCEEEB76D" blockId="13.[151,1437,1084,1936]" box="[199,937,1119,1144]" pageId="13" pageNumber="36">
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<materialsCitation id="3B743CEBFA7B3628AF838F0BCEE2B76D" ID-GBIF-Occurrence="2028137687" box="[199,933,1119,1144]" collectingDate="2017-02-06" collectionCode="MfN" collectorName="W. Mey" pageId="13" pageNumber="36" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
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<typeStatus id="54A78814FA7B3628AF838F0BCC7EB76D" box="[199,313,1120,1144]" pageId="13" pageNumber="36" type="paratype">Paratypes</typeStatus>
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: same data,
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<specimenCount id="9D1AFD3FFA7B3628AE848F0BCCA5B76D" box="[448,482,1120,1144]" pageId="13" pageNumber="36" type="male">2♂</specimenCount>
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[pinned],
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<date id="FFA21076FA7B3628AD138F34CFFBB76D" box="[599,700,1119,1144]" pageId="13" pageNumber="36" value="2017-02-06">
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<collectingDate id="EFE6E99EFA7B3628AD138F34CFFBB76D" box="[599,700,1119,1144]" pageId="13" pageNumber="36" value="2017-02-06">6.ii.2017</collectingDate>
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</date>
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, leg.
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<collectorName id="26E95360FA7B3628ADBC8F0BCE16B76D" box="[760,849,1120,1144]" pageId="13" pageNumber="36">W. Mey</collectorName>
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(
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<collectionCode id="ED0DAE73FA7B3628AC248F34CEDAB76D" box="[864,925,1119,1144]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum für Naturkunde" pageId="13" pageNumber="36" type="Museum">MfN</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C306653DFA7B3628AF838FEFCE74B789" box="[199,819,1156,1181]" pageId="13" pageNumber="36" type="etymology">
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<paragraph id="8BA336B6FA7B3628AF838FEFCE74B789" blockId="13.[151,1437,1084,1936]" box="[199,819,1156,1181]" pageId="13" pageNumber="36">
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<emphasis id="B968EAA4FA7B3628AF838FEFCC09B789" bold="true" box="[199,334,1156,1180]" pageId="13" pageNumber="36">Etymology.</emphasis>
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The specific name is derived from
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<collectingRegion id="49D8F854FA7B3628AD9F8FEFCE77B789" box="[731,816,1156,1180]" country="Colombia" name="Bogota" pageId="13" pageNumber="36">Bogotá</collectingRegion>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C306653DFA7B3628AF838FCCCCD7B580" pageId="13" pageNumber="36" type="description">
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<paragraph id="8BA336B6FA7B3628AF838FCCC9ABB68D" blockId="13.[151,1437,1084,1936]" pageId="13" pageNumber="36">
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<emphasis id="B968EAA4FA7B3628AF838FCCCC1EB7D5" bold="true" box="[199,345,1191,1216]" pageId="13" pageNumber="36">Description.</emphasis>
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Male (
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<figureCitation id="13272A33FA7B3628AEEE8FCCCF49B7D5" box="[426,526,1191,1216]" captionStart="FIGURES 15" captionStartId="14.[151,264,1539,1561]" captionTargetBox="[185,1404,198,1508]" captionTargetId="figure@14.[175,1411,193,1517]" captionTargetPageId="14" captionText="FIGURES 15A–15F. Trichoptera spp., photos of adults, dorsal view. 15A, Atopsyche viejae spec. nov., male paratype; 15B, Atopsyche flavoantennata spec. nov., female paratype; 15C, Smicridea colombosaucia spec. nov., male paratype; 15D, Polycentropus joergenseni Ulmer 1909, male; 15E, Phylloicus monserratensis spec. nov., male paratype; 15F, Marilia bogotaensis spec. nov., male paratype." httpUri="https://zenodo.org/record/2606160/files/figure.png" pageId="13" pageNumber="36">Fig. 15F</figureCitation>
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). Length of each forewing
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<quantity id="4CE49B53FA7B3628AC078FC3CEC0B7AA" box="[835,903,1192,1216]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="13" pageNumber="36" unit="mm" value="9.0">9 mm</quantity>
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, wing span
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<quantity id="4CE49B53FA7B3628AB548FC3C9CAB7AA" box="[1040,1165,1192,1216]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" pageId="13" pageNumber="36" unit="mm" value="19.5" valueMax="20.0" valueMin="19.0">19–20 mm</quantity>
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. Head brown, frons and vertex densely covered by grey-white hairs; maxillary and labial palpi dark brown, scapes bulbous, with grey-white hairs, antennae nearly twice as long as forewings, dark brown, white-ringed at articulations of flagellomeres, eyes small, widely separated middorsally; forewings with brown veins and grey-white hairs on wing membrane, large brown spot on bifurcation of cubitus and crossvein
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<emphasis id="B968EAA4FA7B3628ADBA8E52CE77B645" box="[766,816,1337,1360]" italics="true" pageId="13" pageNumber="36">cu-a</emphasis>
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; hindwings white, Cu1a+1b seemingly running onto M3+4, Cu2 and A1 with short, basal stalk, jugum and jugal margins of hindwings with long, thin hairs. Forelegs brown, middle and hind legs yellow-brown, tarsal spines black, spurs 2.4.4. Abdomen grey-brown.
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</paragraph>
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<paragraph id="8BA336B6FA7B3628AF838ECFCC6FB565" blockId="13.[151,1437,1084,1936]" pageId="13" pageNumber="36">
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Male genitalia (
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<figureCitation id="13272A33FA7B3628AE3E8ECFCC96B6A9" box="[378,465,1444,1469]" captionStart="FIGURES 10–13" captionStartId="11.[151,264,1883,1905]" captionTargetBox="[201,1392,205,1769]" captionTargetId="figure@11.[189,1404,193,1871]" captionTargetPageId="11" captionText="FIGURES 10–13. Trichoptera spp. 10A–10C, Contulma spec., female genitalia, scale = 0.3 mm: 10A, left lateral; 10B, dorsal; 10C, ventral. 11A–11D, Phylloicus monserratensis spec. nov., male genitalia, scale = 0.5 mm: 11A, left lateral; 11B, dorsal; 11C, ventral; 11D, phallic apparatus, ventral. 12A–12D, Marilia bogotaensis spec. nov., male genitalia, scale: 0.3 mm: 12A, left lateral; 12B, ventral; 12C, dorsal; 12D, phallic apparatus, left lateral. 13A–13B, Oecetis chapinera spec. nov., scale = 0.25 mm: 13A, left lateral, with phallic apparatus in situ; 13B, ventral." httpUri="https://zenodo.org/record/2606156/files/figure.png" pageId="13" pageNumber="36">Figs 12</figureCitation>
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A–12D): Segment IX in lateral view vertical anteriorly, separated on lateral side into tergum and sternum by broad, pleural sclerite; tergum X straight and membranous dorsally, apicolateral sides sclerotized and folded medioventrad, lateral hooks absent; preanal appendages triangular, somewhat twisted in basal half, nearly as long as tergum X; inferior appendages elongate, each with apical segment small; phallic apparatus with short and slightly curved phallotheca, phallotremal sclerite spatulate in lateral view, U-shaped in ventral view.
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</paragraph>
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<paragraph id="8BA336B6FA7B3628AF838D17CCD7B580" blockId="13.[151,1437,1084,1936]" box="[199,400,1660,1685]" pageId="13" pageNumber="36">Female: unknown</paragraph>
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</subSubSection>
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<subSubSection id="C306653DFA7B3628AF838DF4CE11B485" pageId="13" pageNumber="36" type="diagnosis">
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<paragraph id="8BA336B6FA7B3628AF838DF4CE11B485" blockId="13.[151,1437,1084,1936]" pageId="13" pageNumber="36">
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<emphasis id="B968EAA4FA7B3628AF838DF4CC06B5AD" bold="true" box="[199,321,1695,1720]" pageId="13" pageNumber="36">Diagnosis.</emphasis>
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The new species shares with
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<taxonomicName id="4C1C4D35FA7B3628ADD58DCACED7B5AD" authority="Flint 1983" authorityName="Flint" authorityYear="1983" box="[657,912,1695,1720]" class="Insecta" family="Odontoceridae" genus="Marilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="lateralis">
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<emphasis id="B968EAA4FA7B3628ADD58DCACE53B5AD" box="[657,788,1696,1720]" italics="true" pageId="13" pageNumber="36">M. lateralis</emphasis>
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<bibRefCitation id="EF8D4B47FA7B3628AC588DF4CED7B5AD" author="Flint, O. S. Jr." box="[796,912,1695,1720]" pageId="13" pageNumber="36" pagination="1 - 100" refId="ref7447" refString="Flint, O. S. Jr. (1983) Studies of Neotropical caddisflies, XXXIII: New species from austral South America (Trichoptera). Smithsonian Contributions to Zoology, 377, 1 - 100. https: // doi. org / 10.5479 / si. 00810282.377" type="journal article" year="1983">Flint 1983</bibRefCitation>
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</taxonomicName>
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the sclerotized, apicolateral sides of tergum X and the straight inferior appendages. However, preanal appendages and phallic apparatus are of different morphology. Also, the small eyes of the male exclude the species from a closer relationship with
|
||
<taxonomicName id="4C1C4D35FA7B3628AA4B8D82C8D2B415" authorityName="Flint" authorityYear="1983" box="[1295,1429,1768,1792]" class="Insecta" family="Odontoceridae" genus="Marilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="lateralis">
|
||
<emphasis id="B968EAA4FA7B3628AA4B8D82C8D2B415" box="[1295,1429,1768,1792]" italics="true" pageId="13" pageNumber="36">M. lateralis</emphasis>
|
||
</taxonomicName>
|
||
, where eyes meet mid-dorsally. Similar genitalia and small eyes are present in
|
||
<taxonomicName id="4C1C4D35FA7B3628AB5D8C66C85EB431" authority="Flint 1991" authorityName="Flint" authorityYear="1991" box="[1049,1305,1804,1829]" class="Insecta" family="Odontoceridae" genus="Marilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="microps">
|
||
<emphasis id="B968EAA4FA7B3628AB5D8C66C9DDB431" box="[1049,1178,1805,1828]" italics="true" pageId="13" pageNumber="36">M. microps</emphasis>
|
||
Flint 1991
|
||
</taxonomicName>
|
||
which was described from
|
||
<collectingRegion id="49D8F854FA7B3628AE0F8C44CCD3B45D" box="[331,404,1839,1864]" country="Colombia" name="Choco" pageId="13" pageNumber="36">Choco</collectingRegion>
|
||
and
|
||
<collectingRegion id="49D8F854FA7B3628AE8A8C44CF06B45D" box="[462,577,1839,1864]" country="Colombia" name="Antioquia" pageId="13" pageNumber="36">Antioquia</collectingRegion>
|
||
, but it does not exhibit triangular preanal appendages, a character which is present in another Colombian species,
|
||
<taxonomicName id="4C1C4D35FA7B3628AD248C3ECEC7B479" authority="Banks 1913" authorityName="Banks" authorityYear="1913" box="[608,896,1876,1901]" class="Insecta" family="Odontoceridae" genus="Marilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="13" pageNumber="36" phylum="Arthropoda" rank="species" species="modesta">
|
||
<emphasis id="B968EAA4FA7B3628AD248C3ECFAFB479" box="[608,744,1876,1900]" italics="true" pageId="13" pageNumber="36">M. modesta</emphasis>
|
||
<bibRefCitation id="EF8D4B47FA7B3628ADB08C3FCEC7B479" author="Banks, N." box="[756,896,1876,1901]" pageId="13" pageNumber="36" pagination="201 - 242" refId="ref7177" refString="Banks, N. (1913) Synopses and descriptions of exotic Neuroptera. Transactions of the American Entomological Society, 39, 201 - 242, pls. 23 - 26." type="journal article" year="1913">Banks 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
. The phallic apparatus of the latter species is strongly bent basally and much longer than in the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |