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<document id="CB87DFABE3E9ACFEC1D4F92E339DE679" ID-DOI="10.11646/zootaxa.4996.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5074595" ID-ZooBank="A1982CE0-AD2F-496B-80AB-FB3C4FA69F7A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625556599242" checkinUser="plazi" docAuthor="Monniot, Francoise" docDate="2021" docId="03A6FD6CFF9DE83255E3310FFDBCFA92" docLanguage="en" docName="zootaxa.4996.3.2.pdf" docOrigin="Zootaxa 4996 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterygascidia kanakae Monniot 2021, sp. nov." docType="treatment" docVersion="4" lastPageNumber="452" masterDocId="FF9F8514FF9AE83B5574370CFFBCFFCC" masterDocTitle="New records of deep-sea ascidians (Tunicata, Ascidiacea) from the New Caledonia region" masterLastPageNumber="468" masterPageNumber="443" pageNumber="450" updateTime="1698992134275" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B1B44F4D73E5987C19F5D2AA8012FABA">New records of deep-sea ascidians (Tunicata, Ascidiacea) from the New Caledonia region</mods:title>
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<mods:date id="B9D86D9534B5E57DE8EAC9020ECB027D">2021</mods:date>
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<heading id="D0F8FB16FF9DE83C55E3310FFDB1F9D1" bold="true" box="[151,525,1539,1566]" fontSize="11" level="1" pageId="7" pageNumber="450" reason="1">
<emphasis id="B97B9068FF9DE83C55E3310FFDB1F9D1" bold="true" box="[151,525,1539,1566]" pageId="7" pageNumber="450">
<taxonomicName id="4C0F37F9FF9DE83C55E3310FFE14F9D2" authority="Monniot, 2021" authorityName="Monniot" authorityYear="2021" box="[151,424,1539,1566]" class="Ascidiacea" family="Agneziidae" genus="Pterygascidia" kingdom="Animalia" order="Enterogona" pageId="7" pageNumber="450" phylum="Chordata" rank="species" species="kanakae" status="sp. nov.">
<emphasis id="B97B9068FF9DE83C55E3310FFE14F9D2" bold="true" box="[151,424,1539,1566]" italics="true" pageId="7" pageNumber="450">Pterygascidia kanakae</emphasis>
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<taxonomicNameLabel id="A2482D13FF9DE83C54DB310FFDB1F9D1" box="[431,525,1539,1565]" pageId="7" pageNumber="450" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3151FF1FF9DE83255E3312BFDBCFA92" lastPageId="9" lastPageNumber="452" pageId="7" pageNumber="450" type="description">
<paragraph id="8BB04C7AFF9DE83C55E3312BFEA0F98D" blockId="7.[151,525,1539,1601]" box="[151,284,1575,1601]" pageId="7" pageNumber="450">
<figureCitation id="133450FFFF9DE83C55E3312BFEBDF98D" box="[151,257,1575,1601]" captionStart="FIGURE 8" captionStartId="8.[151,250,2008,2032]" captionTargetBox="[177,1409,669,1985]" captionTargetId="figure-308@8.[177,1411,669,1985]" captionTargetPageId="8" captionText="FIGURE 8. Pterygascidia kanakae sp. nov. A, specimen; B oral siphon; scale bars = 1cm. C, part of the branchial tissue." figureDoi="http://doi.org/10.5281/zenodo.5074613" httpUri="https://zenodo.org/record/5074613/files/figure.png" pageId="7" pageNumber="450">Figures 8</figureCitation>
,
<figureCitation id="133450FFFF9DE83C547A312BFEA0F98D" box="[270,284,1575,1601]" captionStart="FIGURE 9" captionStartId="9.[151,250,1134,1158]" captionTargetBox="[151,1435,196,1102]" captionTargetId="figure-15@9.[151,1435,196,1102]" captionTargetPageId="9" captionText="FIGURE 9. Pterygascidia kanakae sp. nov. A, anterior part of the body; B, anus and part of dorsal lamina; C, gut and gonads; D, testis lobes and sperm duct. Scale bars A and C = 1cm." figureDoi="http://doi.org/10.5281/zenodo.5074615" httpUri="https://zenodo.org/record/5074615/files/figure.png" pageId="7" pageNumber="450">9</figureCitation>
</paragraph>
<paragraph id="8BB04C7AFF9DE83C55E33163FB19F945" blockId="7.[151,1189,1647,1674]" box="[151,1189,1647,1674]" pageId="7" pageNumber="450">
<materialsCitation id="3B674627FF9DE83C55E33163FB1CF945" box="[151,1184,1647,1674]" collectionCode="MNHN" pageId="7" pageNumber="450" specimenCode="P1" specimenCount="1" typeStatus="holotype">
Station: Kanadeep 12-742-03. One specimen (
<typeStatus id="54B4F2D8FF9DE83C57E93163FCBBF945" box="[669,775,1647,1673]" pageId="7" pageNumber="450" type="holotype">Holotype</typeStatus>
<collectionCode id="ED1ED4BFFF9DE83C567A317CFCD8F946" box="[782,868,1648,1674]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="450">MNHN</collectionCode>
<specimenCode id="DBA9E401FF9DE83C561E3163FC38F945" box="[874,900,1647,1673]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="450">P1</specimenCode>
.PTE 3 slide A1. 1903)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB04C7AFF9DE83355E331B4FB8AFD49" blockId="7.[151,1437,1719,2034]" lastBlockId="8.[151,1437,150,645]" lastPageId="8" lastPageNumber="451" pageId="7" pageNumber="450">
Very soft and transparent
<quantity id="4CF7E19FFF9DE83C54CF31BBFDAAF91E" box="[443,534,1719,1746]" metricMagnitude="-1" metricUnit="m" metricValue="2.35" pageId="7" pageNumber="450" unit="cm" value="23.5">23.5 cm</quantity>
long (
<figureCitation id="133450FFFF9DE83C571431BBFD05F91D" box="[608,697,1719,1745]" captionStart="FIGURE 8" captionStartId="8.[151,250,2008,2032]" captionTargetBox="[177,1409,669,1985]" captionTargetId="figure-308@8.[177,1411,669,1985]" captionTargetPageId="8" captionText="FIGURE 8. Pterygascidia kanakae sp. nov. A, specimen; B oral siphon; scale bars = 1cm. C, part of the branchial tissue." figureDoi="http://doi.org/10.5281/zenodo.5074613" httpUri="https://zenodo.org/record/5074613/files/figure.png" pageId="7" pageNumber="450">Fig. 8A</figureCitation>
) the body has the general shape of the genus with the thorax and abdomen included in a single segment followed without a neat constriction by a wide and long peduncle.The anterior part of the body was a little damaged when collected, but most of the internal organs are preserved. The oral aperture opens at the base of the siphon developed into a hood which contains muscular fibres (
<figureCitation id="133450FFFF9DE83C51FD302FFB5FF8F1" box="[1161,1251,1827,1853]" captionStart="FIGURE 8" captionStartId="8.[151,250,2008,2032]" captionTargetBox="[177,1409,669,1985]" captionTargetId="figure-308@8.[177,1411,669,1985]" captionTargetPageId="8" captionText="FIGURE 8. Pterygascidia kanakae sp. nov. A, specimen; B oral siphon; scale bars = 1cm. C, part of the branchial tissue." figureDoi="http://doi.org/10.5281/zenodo.5074613" httpUri="https://zenodo.org/record/5074613/files/figure.png" pageId="7" pageNumber="450">Fig. 8B</figureCitation>
,
<figureCitation id="133450FFFF9DE83C5186302FFAABF8F1" box="[1266,1303,1827,1853]" captionStart="FIGURE 9" captionStartId="9.[151,250,1134,1158]" captionTargetBox="[151,1435,196,1102]" captionTargetId="figure-15@9.[151,1435,196,1102]" captionTargetPageId="9" captionText="FIGURE 9. Pterygascidia kanakae sp. nov. A, anterior part of the body; B, anus and part of dorsal lamina; C, gut and gonads; D, testis lobes and sperm duct. Scale bars A and C = 1cm." figureDoi="http://doi.org/10.5281/zenodo.5074615" httpUri="https://zenodo.org/record/5074615/files/figure.png" pageId="7" pageNumber="450">9A</figureCitation>
). The atrial aperture has eight round lobes above a few circular muscle fibres. No wing-shaped extensions of the tunic were seen associated with the atrial siphon, but the tunic is partly torn at this level.An average of 60 oral tentacles is distributed in two orders of size. The musculature is restricted to the thorax. The muscles are in transverse spaced fibres running from the endostyle to the middle of each body side and shorter transverse fibres are along the dorsal side. The dorsal tubercle opens in a slit. The dorsal lamina comprises pointed languets. The thin branchial tissue (
<figureCitation id="133450FFFF9DE83C519430DBFA8AF83D" box="[1248,1334,2007,2033]" captionStart="FIGURE 8" captionStartId="8.[151,250,2008,2032]" captionTargetBox="[177,1409,669,1985]" captionTargetId="figure-308@8.[177,1411,669,1985]" captionTargetPageId="8" captionText="FIGURE 8. Pterygascidia kanakae sp. nov. A, specimen; B oral siphon; scale bars = 1cm. C, part of the branchial tissue." figureDoi="http://doi.org/10.5281/zenodo.5074613" httpUri="https://zenodo.org/record/5074613/files/figure.png" pageId="7" pageNumber="450">Fig. 8C</figureCitation>
) extends to the posterior end of the body; about 50 longitudinal vessels were counted on the right side with a single stigma per mesh (
<figureCitation id="133450FFFF92E833547937B7FED9FF19" box="[269,357,187,213]" captionStart="FIGURE 8" captionStartId="8.[151,250,2008,2032]" captionTargetBox="[177,1409,669,1985]" captionTargetId="figure-308@8.[177,1411,669,1985]" captionTargetPageId="8" captionText="FIGURE 8. Pterygascidia kanakae sp. nov. A, specimen; B oral siphon; scale bars = 1cm. C, part of the branchial tissue." figureDoi="http://doi.org/10.5281/zenodo.5074613" httpUri="https://zenodo.org/record/5074613/files/figure.png" pageId="8" pageNumber="451">Fig. 8C</figureCitation>
). The digestive loop is long and straight; it begins with a short oesophagus in continuity with an olive-shaped horizontal stomach sided by two round glandular masses (
<figureCitation id="133450FFFF92E83356C637D3FBB6FF35" box="[946,1034,223,249]" captionStart="FIGURE 9" captionStartId="9.[151,250,1134,1158]" captionTargetBox="[151,1435,196,1102]" captionTargetId="figure-15@9.[151,1435,196,1102]" captionTargetPageId="9" captionText="FIGURE 9. Pterygascidia kanakae sp. nov. A, anterior part of the body; B, anus and part of dorsal lamina; C, gut and gonads; D, testis lobes and sperm duct. Scale bars A and C = 1cm." figureDoi="http://doi.org/10.5281/zenodo.5074615" httpUri="https://zenodo.org/record/5074615/files/figure.png" pageId="8" pageNumber="451">Fig. 9C</figureCitation>
). The state of the stomach wall does not allow seeing if it has ridges. The intestine ends in a smooth-edged anus (
<figureCitation id="133450FFFF92E8335684360FFBFAFED1" box="[1008,1094,259,285]" captionStart="FIGURE 9" captionStartId="9.[151,250,1134,1158]" captionTargetBox="[151,1435,196,1102]" captionTargetId="figure-15@9.[151,1435,196,1102]" captionTargetPageId="9" captionText="FIGURE 9. Pterygascidia kanakae sp. nov. A, anterior part of the body; B, anus and part of dorsal lamina; C, gut and gonads; D, testis lobes and sperm duct. Scale bars A and C = 1cm." figureDoi="http://doi.org/10.5281/zenodo.5074615" httpUri="https://zenodo.org/record/5074615/files/figure.png" pageId="8" pageNumber="451">Fig. 9B</figureCitation>
) The gonad is totally different from those described for
<taxonomicName id="4C0F37F9FF92E83354DA362BFD62FE8D" authority="(Van Name, 1918)" baseAuthorityName="Van Name" baseAuthorityYear="1918" box="[430,734,295,321]" class="Ascidiacea" family="Agneziidae" genus="Pterygascidia" kingdom="Animalia" order="Enterogona" pageId="8" pageNumber="451" phylum="Chordata" rank="species" species="longa">
<emphasis id="B97B9068FF92E83354DA362BFDB4FE8D" box="[430,520,295,321]" italics="true" pageId="8" pageNumber="451">P. longa</emphasis>
(
<bibRefCitation id="EF9E318BFF92E8335762362BFD6AFE8D" author="Van Name, W. G." box="[534,726,295,321]" pageId="8" pageNumber="451" pagination="49 - 174" refId="ref10436" refString="Van Name, W. G. (1918) Ascidians from the Philippines and adjacent waters. Bulletin of the United States National Museum, 100 (1), 49 - 174." type="journal article" year="1918">Van Name, 1918</bibRefCitation>
)
</taxonomicName>
. In the present specimen the gonad is hermaphrodite in a single organ, parallel to the intestine but not adherent to it (
<figureCitation id="133450FFFF92E833579A3647FCF4FEA9" box="[750,840,331,357]" captionStart="FIGURE 9" captionStartId="9.[151,250,1134,1158]" captionTargetBox="[151,1435,196,1102]" captionTargetId="figure-15@9.[151,1435,196,1102]" captionTargetPageId="9" captionText="FIGURE 9. Pterygascidia kanakae sp. nov. A, anterior part of the body; B, anus and part of dorsal lamina; C, gut and gonads; D, testis lobes and sperm duct. Scale bars A and C = 1cm." figureDoi="http://doi.org/10.5281/zenodo.5074615" httpUri="https://zenodo.org/record/5074615/files/figure.png" pageId="8" pageNumber="451">Fig. 9C</figureCitation>
). It begins in the bottom of the gut loop and extends parallel to the intestine up to the base of the atrial siphon. Several groups of testis vesicles(
<figureCitation id="133450FFFF92E83351093663FB6DFE45" box="[1149,1233,367,393]" captionStart="FIGURE 9" captionStartId="9.[151,250,1134,1158]" captionTargetBox="[151,1435,196,1102]" captionTargetId="figure-15@9.[151,1435,196,1102]" captionTargetPageId="9" captionText="FIGURE 9. Pterygascidia kanakae sp. nov. A, anterior part of the body; B, anus and part of dorsal lamina; C, gut and gonads; D, testis lobes and sperm duct. Scale bars A and C = 1cm." figureDoi="http://doi.org/10.5281/zenodo.5074615" httpUri="https://zenodo.org/record/5074615/files/figure.png" pageId="8" pageNumber="451">Fig. 9D</figureCitation>
) are applied along the axis of the long tubular ovary. The sperm duct issuing from each group of testis lobes connects to a common sperm duct running along the length of the ovary ending in four genital papillae. In
<taxonomicName id="4C0F37F9FF92E833513036BBFAF8FE1D" authority="Monniot, 2021" authorityName="Monniot" authorityYear="2021" box="[1092,1348,439,465]" class="Ascidiacea" family="Agneziidae" genus="Pterygascidia" kingdom="Animalia" order="Enterogona" pageId="8" pageNumber="451" phylum="Chordata" rank="species" species="kanakae" status="sp. nov.">
<emphasis id="B97B9068FF92E833513036BBFAF8FE1D" box="[1092,1348,439,465]" italics="true" pageId="8" pageNumber="451">Pterygascidia kanakae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2482D13FF92E833503F36BBFA20FE1C" box="[1355,1436,439,465]" pageId="8" pageNumber="451" rank="species">sp.nov.</taxonomicNameLabel>
male and female elements are associated in a single gland and represent a different organisation from the status described by several authors for
<taxonomicName id="4C0F37F9FF92E833548936F3FD62FDD5" baseAuthorityName="Van Name" baseAuthorityYear="1918" box="[509,734,511,537]" class="Ascidiacea" family="Agneziidae" genus="Pterygascidia" kingdom="Animalia" order="Enterogona" pageId="8" pageNumber="451" phylum="Chordata" rank="species" species="longa">
<emphasis id="B97B9068FF92E833548936F3FD62FDD5" box="[509,734,511,537]" italics="true" pageId="8" pageNumber="451">Pterygascidia longa</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9E318BFF92E833579F36F3FC27FDD5" author="Van Name, W. G." box="[747,923,511,537]" pageId="8" pageNumber="451" pagination="49 - 174" refId="ref10436" refString="Van Name, W. G. (1918) Ascidians from the Philippines and adjacent waters. Bulletin of the United States National Museum, 100 (1), 49 - 174." type="journal article" year="1918">Van name 1918</bibRefCitation>
;
<bibRefCitation id="EF9E318BFF92E83356D236F3FB90FDD5" author="Millar, R. H." box="[934,1068,511,537]" pageId="8" pageNumber="451" pagination="623 - 628" refId="ref9621" refString="Millar, R. H. (1963) The structure and relationships of the ascidian Ciallusia longa Van Name. Proceedings of the Zoological Society London, 141 (3), 623 - 628. https: // doi. org / 10.1111 / j. 1469 - 7998.1963. tb 06103. x" type="journal article" year="1963">Millar 1963</bibRefCitation>
;
<bibRefCitation id="EF9E318BFF92E833514236F3FB62FDD5" author="Tokioka, T." box="[1078,1246,511,537]" pageId="8" pageNumber="451" pagination="119 - 125" refId="ref10368" refString="Tokioka, T. (1971 a) The ascidian genera Pterygascidia Sluiter, 1904 and Ciallusia Van Name 1918. Zoologische Mededelingen, Leiden, 45 (10), 119 - 125." type="journal article" year="1971">Tokioka 1971a</bibRefCitation>
,
<bibRefCitation id="EF9E318BFF92E833519C36F3FA8DFDD5" author="Tokioka, T." box="[1256,1329,511,537]" pageId="8" pageNumber="451" pagination="191 - 198" refId="ref10403" refString="Tokioka, T. (1971 b) Redescription of Pterygascidia mirabilis Sluiter 1904 (Ascidiacea, Cionidae) based on the type specimen. Beaufortia, 18, 191 - 198." type="journal article" year="1971">1971b</bibRefCitation>
;
<bibRefCitation id="EF9E318BFF92E833504836F3FEEEFDF1" author="Monniot, F. &amp; Monniot, C." pageId="8" pageNumber="451" pagination="681 - 749" refId="ref10080" refString="Monniot, F. &amp; Monniot, C. (2003) Ascidies de la pente externe et bathyales de l'ouest Pacifique. Zoosystema, 25 (4), 681 - 749." type="journal article" year="2003">Monniot &amp; Monniot 2003</bibRefCitation>
<figureCitation id="133450FFFF92E833542C352FFE0FFDF1" box="[344,435,547,573]" captionStart="FIGURE 3" captionStartId="3.[151,250,1400,1424]" captionTargetBox="[151,1437,525,1377]" captionTargetId="figure-102@3.[151,1437,525,1377]" captionTargetPageId="3" captionText="FIGURE 3. Pharyngodictyon sp., scale bar = 1cm." figureDoi="http://doi.org/10.5281/zenodo.5074601" httpUri="https://zenodo.org/record/5074601/files/figure.png" pageId="8" pageNumber="451">Figs 3A</figureCitation>
,
<figureCitation id="133450FFFF92E83354CA352FFE5EFDF1" box="[446,482,547,573]" captionStart="FIGURE" captionStartId="4.[151,251,1225,1249]" captionTargetBox="[151,1435,406,1202]" captionTargetId="figure-66@4.[151,1435,406,1202]" captionTargetPageId="4" captionText="FIGURE. 4. Protoholozoa incrustans sp. nov. colony, scale bar = 1cm." figureDoi="http://doi.org/10.5281/zenodo.5074603" httpUri="https://zenodo.org/record/5074603/files/figure.png" pageId="8" pageNumber="451">4B</figureCitation>
;
<bibRefCitation id="EF9E318BFF92E8335499352FFDDEFDF1" author="Kott, P." box="[493,610,547,573]" pageId="8" pageNumber="451" pagination="47 - 52" refId="ref9572" refString="Kott, P. (2008) Cialusiidae (Ascidiacea, Tunicata), a monotypic family from deeper waters of the tropical Indo-West Pacific. Zoootaxa, 1742 (1), 47 - 52. https: // doi. org / 10.11646 / zootaxa. 1742.1.3" type="journal article" year="2008">Kott 2008</bibRefCitation>
). In
<taxonomicName id="4C0F37F9FF92E83357E5352FFD50FDF1" baseAuthorityName="Van Name" baseAuthorityYear="1918" box="[657,748,547,573]" class="Ascidiacea" family="Agneziidae" genus="Pterygascidia" kingdom="Animalia" order="Enterogona" pageId="8" pageNumber="451" phylum="Chordata" rank="species" species="longa">
<emphasis id="B97B9068FF92E83357E5352FFD50FDF1" box="[657,748,547,573]" italics="true" pageId="8" pageNumber="451">P. longa</emphasis>
</taxonomicName>
the testis vesicles are spread along a large part of the intestine wall, the common sperm duct emerges in the bottom of the gut loop to join the base of the tubular ovary and remains applied to the ovary passing the female papilla and ending in a single male papilla.
</paragraph>
<caption id="DF701CF2FF92E83355E330D4FAD8F83C" ID-DOI="http://doi.org/10.5281/zenodo.5074613" ID-Zenodo-Dep="5074613" box="[151,1380,2008,2033]" httpUri="https://zenodo.org/record/5074613/files/figure.png" pageId="8" pageNumber="451" startId="8.[151,250,2008,2032]" targetBox="[177,1409,669,1985]" targetPageId="8">
<paragraph id="8BB04C7AFF92E83355E330D4FAD8F83C" blockId="8.[151,1380,2008,2033]" box="[151,1380,2008,2033]" pageId="8" pageNumber="451">
<emphasis id="B97B9068FF92E83355E330D4FEAFF83C" bold="true" box="[151,275,2008,2032]" pageId="8" pageNumber="451">FIGURE 8.</emphasis>
<taxonomicName id="4C0F37F9FF92E833546D30D4FDBCF83C" authority="Monniot, 2021" authorityName="Monniot" authorityYear="2021" box="[281,512,2008,2032]" class="Ascidiacea" family="Agneziidae" genus="Pterygascidia" kingdom="Animalia" order="Enterogona" pageId="8" pageNumber="451" phylum="Chordata" rank="species" species="kanakae" status="sp. nov.">
<emphasis id="B97B9068FF92E833546D30D4FDBCF83C" box="[281,512,2008,2032]" italics="true" pageId="8" pageNumber="451">Pterygascidia kanakae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2482D13FF92E833577330D5FDEAF83C" box="[519,598,2009,2032]" pageId="8" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
A, specimen; B oral siphon; scale bars = 1cm. C, part of the branchial tissue.
</paragraph>
</caption>
<caption id="DF701CF2FF93E83255E33362FD5DFB66" ID-DOI="http://doi.org/10.5281/zenodo.5074615" ID-Zenodo-Dep="5074615" httpUri="https://zenodo.org/record/5074615/files/figure.png" pageId="9" pageNumber="452" startId="9.[151,250,1134,1158]" targetBox="[151,1435,196,1102]" targetPageId="9">
<paragraph id="8BB04C7AFF93E83255E33362FD5DFB66" blockId="9.[151,1436,1134,1194]" pageId="9" pageNumber="452">
<emphasis id="B97B9068FF93E83255E33362FEAFFB4A" bold="true" box="[151,275,1134,1158]" pageId="9" pageNumber="452">FIGURE 9.</emphasis>
<taxonomicName id="4C0F37F9FF93E832546E3362FDBDFB4A" authority="Monniot, 2021" authorityName="Monniot" authorityYear="2021" box="[282,513,1134,1158]" class="Ascidiacea" family="Agneziidae" genus="Pterygascidia" kingdom="Animalia" order="Enterogona" pageId="9" pageNumber="452" phylum="Chordata" rank="species" species="kanakae" status="sp. nov.">
<emphasis id="B97B9068FF93E832546E3362FDBDFB4A" box="[282,513,1134,1158]" italics="true" pageId="9" pageNumber="452">Pterygascidia kanakae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2482D13FF93E832577C3362FDEBFB49" box="[520,599,1134,1157]" pageId="9" pageNumber="452" rank="species">sp. nov.</taxonomicNameLabel>
A, anterior part of the body; B, anus and part of dorsal lamina; C, gut and gonads; D, testis lobes and sperm duct. Scale bars A and C = 1cm.
</paragraph>
</caption>
<paragraph id="8BB04C7AFF93E83255B333D5FDBCFA92" blockId="9.[151,1437,1240,1375]" pageId="9" pageNumber="452">
<bibRefCitation id="EF9E318BFF93E83255B333D5FEF6FB3E" author="Kott, P." box="[199,330,1240,1267]" pageId="9" pageNumber="452" pagination="47 - 52" refId="ref9572" refString="Kott, P. (2008) Cialusiidae (Ascidiacea, Tunicata), a monotypic family from deeper waters of the tropical Indo-West Pacific. Zoootaxa, 1742 (1), 47 - 52. https: // doi. org / 10.11646 / zootaxa. 1742.1.3" type="journal article" year="2008">Kott (2008)</bibRefCitation>
established the synonymy of the genera
<taxonomicName id="4C0F37F9FF93E832567F33D4FB87FB3E" authority="Sluiter, 1904" authorityName="Sluiter" authorityYear="1904" box="[779,1083,1240,1266]" class="Ascidiacea" family="Agneziidae" genus="Pterygascidia" kingdom="Animalia" order="Enterogona" pageId="9" pageNumber="452" phylum="Chordata" rank="genus">
<emphasis id="B97B9068FF93E832567F33D4FC1BFB3E" box="[779,935,1240,1266]" italics="true" pageId="9" pageNumber="452">Pterygascidia</emphasis>
<bibRefCitation id="EF9E318BFF93E83256D933D4FB87FB3E" author="Sluiter, C. P." box="[941,1083,1240,1266]" pageId="9" pageNumber="452" pagination="1 - 139" refId="ref10297" refString="Sluiter, C. P. (1904) Die Tunicaten der Siboga-Expedition I. Die socialen und holosomen Ascidien. Siboga Expedition, 56 A, 1 - 139." type="journal article" year="1904">Sluiter, 1904</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C0F37F9FF93E832510433D4FA24FB3E" authority="Van Name, 1918" authorityName="Van Name" authorityYear="1918" box="[1136,1432,1240,1267]" class="Ascidiacea" family="Agneziidae" genus="Ciallusia" kingdom="Animalia" order="Enterogona" pageId="9" pageNumber="452" phylum="Chordata" rank="genus">
<emphasis id="B97B9068FF93E832510433D4FB6BFB3E" box="[1136,1239,1240,1266]" italics="true" pageId="9" pageNumber="452">Ciallusia</emphasis>
<bibRefCitation id="EF9E318BFF93E83251A833D5FA24FB3E" author="Van Name, W. G." box="[1244,1432,1240,1267]" pageId="9" pageNumber="452" pagination="49 - 174" refId="ref10436" refString="Van Name, W. G. (1918) Ascidians from the Philippines and adjacent waters. Bulletin of the United States National Museum, 100 (1), 49 - 174." type="journal article" year="1918">Van Name, 1918</bibRefCitation>
</taxonomicName>
, a status generally accepted. Concerning the species, a doubt about the synonymy of
<taxonomicName id="4C0F37F9FF93E832514933F1FAEEFADA" authority="Sluiter, 1904" authorityName="Sluiter" authorityYear="1904" box="[1085,1362,1276,1302]" class="Ascidiacea" family="Agneziidae" genus="Pterygascidia" kingdom="Animalia" order="Enterogona" pageId="9" pageNumber="452" phylum="Chordata" rank="species" species="mirabilis">
<emphasis id="B97B9068FF93E832514933F1FB01FADA" box="[1085,1213,1276,1302]" italics="true" pageId="9" pageNumber="452">P. mirabilis</emphasis>
<bibRefCitation id="EF9E318BFF93E83251B033F0FAEEFADA" author="Sluiter, C. P." box="[1220,1362,1276,1302]" pageId="9" pageNumber="452" pagination="1 - 139" refId="ref10297" refString="Sluiter, C. P. (1904) Die Tunicaten der Siboga-Expedition I. Die socialen und holosomen Ascidien. Siboga Expedition, 56 A, 1 - 139." type="journal article" year="1904">Sluiter, 1904</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C0F37F9FF93E83250FC33F1FF6BFAF6" baseAuthorityName="Van Name" baseAuthorityYear="1918" class="Ascidiacea" family="Agneziidae" genus="Pterygascidia" kingdom="Animalia" order="Enterogona" pageId="9" pageNumber="452" phylum="Chordata" rank="species" species="longa">
<emphasis id="B97B9068FF93E83250FC33F1FF6BFAF6" italics="true" pageId="9" pageNumber="452">P. longa</emphasis>
</taxonomicName>
remains, especially that of
<bibRefCitation id="EF9E318BFF93E832577E322CFD79FAF7" author="Tokioka, T." box="[522,709,1312,1339]" pageId="9" pageNumber="452" pagination="191 - 198" refId="ref10403" refString="Tokioka, T. (1971 b) Redescription of Pterygascidia mirabilis Sluiter 1904 (Ascidiacea, Cionidae) based on the type specimen. Beaufortia, 18, 191 - 198." type="journal article" year="1971">Tokioka (1971b)</bibRefCitation>
who re-examined Sluiters material, the descriptions of the tunic extensions are slightly different.
</paragraph>
</subSubSection>
</treatment>
</document>