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<document id="056197B05520A7293B5BB8C8EAA97E5B" ID-CLB-Dataset="29741" ID-DOI="10.11646/zootaxa.4443.1.1" ID-GBIF-Dataset="fb772898-fe24-438e-9a4a-1a98c5b393a8" ID-ISSN="1175-5326" ID-Zenodo-Dep="1305628" ID-ZooBank="urn:lsid:zoobank.org:pub:8CF7D069-783C-4D20-8A10-6987529AB4BE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530775745819" checkinUser="plazi" docAuthor="Chris J. Hodgson &amp; Douglas J. Williams" docDate="2018" docId="03A687D07B140368EEA8F92FFE0DF9AD" docLanguage="en" docName="zootaxa.4443.1.1.pdf.imf" docOrigin="Zootaxa 4443 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Paralecanium maculatum Takahashi 1950" docType="treatment" docVersion="7" lastPageNumber="111" masterDocId="FF9FFFA87B780306EE3FFFACFFC6FFFD" masterDocTitle="Revision of the soft scale genus Paralecanium (Hemiptera: Coccomorpha: Coccidae) with the introduction of three new genera and twenty new species" masterLastPageNumber="162" masterPageNumber="1" pageNumber="109" updateTime="1698552844880" updateUser="ExternalLinkService">
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<mods:title id="D9A2BD18D6BCBA749A2070A22FB09C32">Revision of the soft scale genus Paralecanium (Hemiptera: Coccomorpha: Coccidae) with the introduction of three new genera and twenty new species</mods:title>
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<mods:namePart id="10551C2C85C156387203DAA0EBCCB1D8">Chris J. Hodgson</mods:namePart>
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<mods:namePart id="ECA29F5E262D7685261FA215E952837A">Douglas J. Williams</mods:namePart>
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<mods:typeOfResource id="06DC1C823F1D55956451345984F28523">text</mods:typeOfResource>
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<mods:title id="4EDFEB14806E25E897E090601A531FDB">Zootaxa</mods:title>
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<mods:part id="B9FE64A7F29F679D48A02E1C198F1302">
<mods:date id="80D404803B604A0AAB5DC136001F9FC3">2018</mods:date>
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<mods:number id="29EF499FCE0B3B688E937471F93E1BE9">2018-07-05</mods:number>
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<mods:number id="8C428CF40B72CE9D341634BEF25045EE">4443</mods:number>
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<mods:number id="19A6F2618F1FDB760A6F7A0CFC04F0C8">1</mods:number>
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<mods:classification id="763489ECA4D359DDE49D4A0E9C2BCAEA">journal article</mods:classification>
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<treatment id="03A687D07B140368EEA8F92FFE0DF9AD" ID-DOI="http://doi.org/10.5281/zenodo.5981528" ID-GBIF-Taxon="146071299" ID-Zenodo-Dep="5981528" LSID="urn:lsid:plazi:treatment:03A687D07B140368EEA8F92FFE0DF9AD" httpUri="http://treatment.plazi.org/id/03A687D07B140368EEA8F92FFE0DF9AD" lastPageId="110" lastPageNumber="111" pageId="108" pageNumber="109">
<subSubSection id="C315654D7B14036AEEA8F92FFF3FF943" pageId="108" pageNumber="109" type="nomenclature">
<paragraph id="8BB036C67B14036AEEA8F92FFD24F960" blockId="108.[151,738,1667,1726]" box="[151,738,1667,1693]" pageId="108" pageNumber="109">
<heading id="D0F881AA7B14036AEEA8F92FFD24F960" bold="true" box="[151,738,1667,1693]" fontSize="11" level="1" pageId="108" pageNumber="109" reason="1">
<emphasis id="B97BEAD47B14036AEEA8F92FFD24F960" bold="true" box="[151,738,1667,1693]" pageId="108" pageNumber="109">
<taxonomicName id="4C0F4D457B14036AEEA8F92FFD9CF960" ID-CoL="4CY9D" authority="Takahashi" authorityName="Takahashi" authorityYear="1950" box="[151,602,1667,1693]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="maculatum">
<emphasis id="B97BEAD47B14036AEEA8F92FFE14F960" bold="true" box="[151,466,1667,1693]" italics="true" pageId="108" pageNumber="109">Paralecanium maculatum</emphasis>
Takahashi
</taxonomicName>
rev. comb.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB036C67B14036AEEA8F909FF3FF943" blockId="108.[151,738,1667,1726]" box="[151,249,1701,1726]" pageId="108" pageNumber="109">
(
<figureCitation id="13342A437B14036AEEA0F909FF37F943" box="[159,241,1701,1726]" captionStart="FIGURE 45" captionStartId="109.[151,250,1953,1975]" captionTargetBox="[226,1352,229,1912]" captionTargetId="figure@109.[207,1380,193,1932]" captionTargetPageId="109" captionText="FIGURE 45. Paralecanium maculatum Takahashi, adult female." httpUri="https://zenodo.org/record/1305718/files/figure.png" pageId="108" pageNumber="109">Fig. 45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C315654D7B14036AEEA8F941FCF3F8BC" pageId="108" pageNumber="109" type="reference_group">
<paragraph id="8BB036C67B14036AEEA8F941FD72F8DF" blockId="108.[151,1436,1773,1857]" pageId="108" pageNumber="109">
<materialsCitation id="3B673C9B7B14036AEEA8F941FD76F8DF" ID-GBIF-Occurrence="1872522441" collectionCode="EIHU" country="Indonesia" location="Riau Islands" pageId="108" pageNumber="109" specimenCount="3" stateProvince="Riau Islands" typeStatus="syntype">
<taxonomicName id="4C0F4D457B14036AEEA8F941FD99F8FE" ID-CoL="4CY9D" authority="Takahashi, 1950 a" authorityName="Takahashi" authorityYear="1950" box="[151,607,1773,1795]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="maculatum">
<emphasis id="B97BEAD47B14036AEEA8F941FE59F8FE" box="[151,415,1773,1795]" italics="true" pageId="108" pageNumber="109">Paralecanium maculatum</emphasis>
<bibRefCitation id="EF9E4B377B14036AEF96F941FD99F8FE" box="[425,607,1773,1795]" pageId="108" pageNumber="109" refId="ref104609" refString="Takahashi, R. (1950 a) Some species of Coccidae from the Riouw Islands - Part I. (Homoptera). Insecta Matsumurana, 17, 65 - 72." type="journal article">Takahashi, 1950a</bibRefCitation>
</taxonomicName>
, 71.
<typeStatus id="54B488647B14036AECAAF942FD01F8FE" box="[661,711,1774,1795]" pageId="108" pageNumber="109">Type</typeStatus>
data:
<collectingCountry id="F31876567B14036AED35F941FC5EF8FE" box="[778,920,1773,1795]" name="Indonesia" pageId="108" pageNumber="109">INDONESIA</collectingCountry>
:
<collectingRegion id="49CBF8247B14036AED9BF941FBE1F8FE" box="[932,1063,1773,1795]" country="Indonesia" name="Kepulauan Riau" pageId="108" pageNumber="109">Riau Islands</collectingRegion>
, host-plant not indicated.
<typeStatus id="54B488647B14036AEB07F941FA51F8FE" box="[1336,1431,1773,1795]" pageId="108" pageNumber="109" type="syntype">Syntypes</typeStatus>
, female.
<typeStatus id="54B488647B14036AEF27F8A1FE8CF8DF" box="[280,330,1805,1826]" pageId="108" pageNumber="109">Type</typeStatus>
depository:
<collectionCode id="ED1EAE037B14036AEFF6F8A1FDCEF8DF" box="[457,520,1805,1826]" country="Japan" httpUri="http://grbio.org/cool/5rw3-8mvc" name="Entomological Institute, Hokkaido University" pageId="108" pageNumber="109">EIHU</collectionCode>
, Sapporo, Japan
</materialsCitation>
.
</paragraph>
<paragraph id="8BB036C67B14036AEEA8F887FCF3F8BC" blockId="108.[151,1436,1773,1857]" box="[151,821,1835,1857]" pageId="108" pageNumber="109">
<taxonomicName id="4C0F4D457B14036AEEA8F887FE59F8BC" ID-CoL="6VQ5V" authorityName="Ali" authorityYear="1971" box="[151,415,1835,1857]" class="Insecta" family="Coccidae" genus="Platylecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="maculatum">
<emphasis id="B97BEAD47B14036AEEA8F887FE59F8BC" box="[151,415,1835,1857]" italics="true" pageId="108" pageNumber="109">Platylecanium maculatum</emphasis>
</taxonomicName>
;
<bibRefCitation id="EF9E4B377B14036AEF92F887FDC9F8BC" box="[429,527,1835,1857]" pageId="108" pageNumber="109" refId="ref102242" refString="Ali, S. M. (1971) A catalogue of the Oriental Coccoidea (Part V) (Insecta: Homoptera: Coccoidea) (with an index). Indian Museum Bulletin, 6, 7 - 82." type="journal article">Ali, 1971</bibRefCitation>
, 41. Change of combination.
</paragraph>
</subSubSection>
<subSubSection id="C315654D7B140368EEA8F8C1FD6EFF29" lastPageId="110" lastPageNumber="111" pageId="108" pageNumber="109" type="materials_examined">
<paragraph id="8BB036C67B14036AEEA8F8C1FC45F857" blockId="108.[151,1436,1901,1998]" pageId="108" pageNumber="109">
<emphasis id="B97BEAD47B14036AEEA8F8C1FD79F87B" bold="true" box="[151,703,1901,1926]" pageId="108" pageNumber="109">
Material examined.
<typeStatus id="54B488647B14036AEFB2F8C1FE3CF87B" box="[397,506,1901,1926]" pageId="108" pageNumber="109" type="holotype">Holotype</typeStatus>
f:
<collectingCountry id="F31876567B14036AEC1FF8C1FD79F87B" box="[544,703,1901,1926]" name="Indonesia" pageId="108" pageNumber="109">INDONESIA</collectingCountry>
</emphasis>
, Riouw Is., Rampang (almost certainly
<emphasis id="B97BEAD47B14036AEAB0F8C1FB2EF87B" bold="true" box="[1167,1256,1901,1926]" pageId="108" pageNumber="109">
<collectingRegion id="49CBF8247B14036AEAB0F8C1FB0FF87B" box="[1167,1225,1901,1926]" country="Indonesia" name="Riau" pageId="108" pageNumber="109">Riau</collectingRegion>
Is
</emphasis>
, Kempang), no host,
<date id="FFB110067B14036AEEEBF83EFE84F857" box="[212,322,1937,1962]" pageId="108" pageNumber="109" value="1946-01">Jan. 1946</date>
, R. Takahashi (
<collectionCode id="ED1EAE037B14036AEFD2F83EFDF2F857" box="[493,564,1938,1962]" country="Japan" httpUri="http://grbio.org/cool/5rw3-8mvc" name="Entomological Institute, Hokkaido University" pageId="108" pageNumber="109">EIHU</collectionCode>
, Sapporo,
<collectingCountry id="F31876567B14036AEC94F83EFD37F857" box="[683,753,1938,1962]" name="Japan" pageId="108" pageNumber="109">Japan</collectingCountry>
): 1/1adf (fg).
</paragraph>
<paragraph id="8BB036C67B14036AEEF8F81AFC53F833" blockId="108.[151,1436,1901,1998]" box="[199,917,1973,1998]" pageId="108" pageNumber="109">Note. The description is based on the single available specimen.</paragraph>
<caption id="DF70664E7B15036BEEA8F80DFCECF84A" box="[151,810,1953,1975]" httpUri="https://zenodo.org/record/1305718/files/figure.png" pageId="109" pageNumber="110" startId="109.[151,250,1953,1975]" targetBox="[226,1352,229,1912]" targetPageId="109">
<paragraph id="8BB036C67B15036BEEA8F80DFCECF84A" blockId="109.[151,810,1953,1975]" box="[151,810,1953,1975]" pageId="109" pageNumber="110">
<emphasis id="B97BEAD47B15036BEEA8F80DFEE6F84A" bold="true" box="[151,288,1953,1975]" pageId="109" pageNumber="110">FIGURE 45.</emphasis>
<taxonomicName id="4C0F4D457B15036BEF1AF80DFD5DF84A" authority="Takahashi" authorityName="Takahashi" authorityYear="1950" box="[293,667,1953,1975]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="109" pageNumber="110" phylum="Arthropoda" rank="species" species="maculatum">
<emphasis id="B97BEAD47B15036BEF1AF80DFDEAF84A" box="[293,556,1953,1975]" italics="true" pageId="109" pageNumber="110">Paralecanium maculatum</emphasis>
Takahashi
</taxonomicName>
, adult female.
</paragraph>
</caption>
<paragraph id="8BB036C67B160368EEF8FF3BFD6EFF29" blockId="110.[151,1437,151,1616]" pageId="110" pageNumber="111">
<emphasis id="B97BEAD47B160368EEF8FF3BFDEAFF4D" bold="true" box="[199,556,151,176]" pageId="110" pageNumber="111">Slide-mounted adult female.</emphasis>
Body oval, more pointed anteriorly than posteriorly; venter somewhat asymmetrical. Length
<quantity id="4CF79B237B160368EFADFF10FE3EFF29" box="[402,504,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" pageId="110" pageNumber="111" unit="mm" value="2.05">2.05 mm</quantity>
, width
<quantity id="4CF79B237B160368EC75FF10FD65FF29" box="[586,675,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="110" pageNumber="111" unit="mm" value="1.4">1.4 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C315654D7B160368EEF8FF4DFC56FACD" pageId="110" pageNumber="111" type="description">
<paragraph id="8BB036C67B160368EEF8FF4DFEF7FD55" blockId="110.[151,1437,151,1616]" pageId="110" pageNumber="111">
<emphasis id="B97BEAD47B160368EEF8FF4DFEEFFF05" box="[199,297,225,248]" italics="true" pageId="110" pageNumber="111">Dorsum.</emphasis>
Derm fairly uniformly sclerotised, with moderately large areaolations throughout, otherwise smooth without minute pale spots. Marginal radial lines obscure but positions indicated by distribution of pores and setae, with 11 lines anteriorly between anterior stigmatic clefts, a line from each cleft, each side with 3 lines between clefts and 11 on abdomen. Abdominal clear areas not apparent. Dorsal setae each rather short, stout, parallel-sided and blunt, 6.57.0 µm long; frequent submarginally within radial lines but absent more medially. Dorsal pores of 3 sizes present: (i) large dorsal pores, each 4.55.5 µm wide without obvious micropores, present more or less throughout apart from in a fairly wide marginal band, with about 3748 pores; (ii) smaller simple pores, each about 2.0 µm wide, sparsely present throughout, and (iii) a minute pore, appearing as a clear spot about 1 µm wide; all three pore
<typeStatus id="54B488647B160368EF28FDACFE94FDE5" box="[279,338,512,536]" pageId="110" pageNumber="111">types</typeStatus>
perhaps in a sparse polygonal pattern. Anal plates with anterior border subequal to posterior border, slightly longer than wide, each with 1 seta in middle of each plate and?1 short setae along outer margin near apex, plus 1 small pore medially; length of plates about 100 µm, combined width about 75 µm. Anogenital fold with 1 pair of small setae along anterior margin, each lateral margin with 1 seta at anterior end and another at posterior end.
</paragraph>
<paragraph id="8BB036C67B160368EEF8FD19FADBFC7D" blockId="110.[151,1437,151,1616]" pageId="110" pageNumber="111">
<emphasis id="B97BEAD47B160368EEF8FD19FEE4FD31" box="[199,290,693,716]" italics="true" pageId="110" pageNumber="111">Margin.</emphasis>
Marginal ornamentation represented by a series of 57 small, radial sclerotised blocks between marginal setae. Marginal setae fan-shaped, mostly clearly wider than long; width of each 2535 µm, length 2025 µm; with about 55 setae anteriorly between anterior stigmatic clefts, each side with 1721 between stigmatic clefts, and 4750 along abdominal margin. Stigmatic clefts rather shallow, each with a narrow opening, apparently without a sclerotised inner margin and with a single stigmatic spine; each spine 1820 µm long, parallel sided with a blunt apex. Eyespots each more or less round; width of socket 2528 µm; diameter of lens 1617 µm.
</paragraph>
<paragraph id="8BB036C67B160368EEF8FC21FC56FACD" blockId="110.[151,1437,151,1616]" pageId="110" pageNumber="111">
<emphasis id="B97BEAD47B160368EEF8FC21FED6FC59" box="[199,272,909,932]" italics="true" pageId="110" pageNumber="111">Venter.</emphasis>
With a broad dark marginal border, about 105165 µm wide, with dark radial fingers. Multilocular discpores hard to discern but present in bands on either side of genital opening and on preceding 2 segments, each side with possibly 7 on abdominal segment VII, 15 on segment VI and 1 or 2 on segment V. Spiracular disc-pores present in a narrow band between margin and each spiracle, with
<quantity id="4CF79B237B160368EDA2FC54FC1FFBED" box="[925,985,1015,1040]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="110" pageNumber="111" unit="in" value="18.0">18 in</quantity>
each anterior band and
<quantity id="4CF79B237B160368EAC7FC5BFA98FBED" box="[1272,1374,1015,1040]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" metricValueMax="6.35" metricValueMin="4.826" pageId="110" pageNumber="111" unit="in" value="22.0" valueMax="25.0" valueMin="19.0">1925 in</quantity>
each posterior band. Ventral microducts minute, present at least near mouthparts but probably present throughout. Ventral setae: pairs of longer setae medially in abdominal segments VVII, otherwise not visible. Antennae well developed, each with 6 segments; total length of each about 270 µm; apical segment 6065 µm long; apical seta about 18 µm long. Clypeolabral shield small, about 85 µm long. Spiracles small; width of each peritreme 1316 µm. Legs well developed; dimensions of hind leg (µm): coxa 7580, trochanter + femur 110115, tibia + tarsus 108112 (some with a hint of segmentation); claw 1012. Tarsal digitules slightly longer than claw digitules, with capitate apices; claw digitules both broad but one possibly slightly narrower than other; claw without a denticle; setal distribution: coxa 5; trochanter 1; femur 2; tibia 1 and tarsus 3.
</paragraph>
</subSubSection>
<subSubSection id="C315654D7B160368EEF8FA90FE0DF9AD" pageId="110" pageNumber="111" type="discussion">
<paragraph id="8BB036C67B160368EEF8FA90FCD6F9D0" blockId="110.[151,1437,151,1616]" pageId="110" pageNumber="111">
<emphasis id="B97BEAD47B160368EEF8FA90FE89FAA8" bold="true" box="[199,335,1340,1365]" pageId="110" pageNumber="111">Comments.</emphasis>
Of the
<taxonomicName id="4C0F4D457B160368EF93FA90FD8CFAA8" box="[428,586,1340,1365]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="B97BEAD47B160368EF93FA90FD8CFAA8" box="[428,586,1340,1365]" italics="true" pageId="110" pageNumber="111">Paralecanium</emphasis>
</taxonomicName>
species which have only a single stigmatic spine in each spiracular cleft, adult female
<taxonomicName id="4C0F4D457B160368EF0DFACDFE09FA85" authorityName="Takahashi" authorityYear="1950" box="[306,463,1375,1400]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="maculatum">
<emphasis id="B97BEAD47B160368EF0DFACDFE09FA85" box="[306,463,1375,1400]" italics="true" pageId="110" pageNumber="111">P. maculatum</emphasis>
</taxonomicName>
are most similar to those of
<taxonomicName id="4C0F4D457B160368ED11FACDFC77FA85" box="[814,945,1377,1400]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="acinaces">
<emphasis id="B97BEAD47B160368ED11FACDFC77FA85" box="[814,945,1377,1400]" italics="true" pageId="110" pageNumber="111">P. acinaces</emphasis>
</taxonomicName>
, described as new above, but differ most obviously in the shape and size of the dorsal setae, those on
<taxonomicName id="4C0F4D457B160368ED7CFA29FC1BFA60" authorityName="Takahashi" authorityYear="1950" box="[835,989,1412,1437]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="maculatum">
<emphasis id="B97BEAD47B160368ED7CFA29FC1BFA60" box="[835,989,1412,1437]" italics="true" pageId="110" pageNumber="111">P. maculatum</emphasis>
</taxonomicName>
being rather short, stout and very blunt whereas those on
<taxonomicName id="4C0F4D457B160368EF5EFA05FE26FA3D" box="[353,480,1449,1472]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="acinaces">
<emphasis id="B97BEAD47B160368EF5EFA05FE26FA3D" box="[353,480,1449,1472]" italics="true" pageId="110" pageNumber="111">P. acinaces</emphasis>
</taxonomicName>
are at least twice as long, with a sharp apex, and often appear blade-like. These two species also differ in (character-states for
<taxonomicName id="4C0F4D457B160368EC58FA61FD22FA19" box="[615,740,1485,1508]" class="Insecta" family="Coccidae" genus="Paralecanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="110" pageNumber="111" phylum="Arthropoda" rank="species" species="acinaces">
<emphasis id="B97BEAD47B160368EC58FA61FD22FA19" box="[615,740,1485,1508]" italics="true" pageId="110" pageNumber="111">P. acinaces</emphasis>
</taxonomicName>
in brackets): (i) the large dorsal pores numbering about 40 per side (fewer than 25); (ii) the dorsal setae are not thought to extend into the submedial area (present in the submedial area), and (iii) the form of their marginal ornamentation.
</paragraph>
<paragraph id="8BB036C67B160368EEF8F99BFE0DF9AD" blockId="110.[151,1437,151,1616]" box="[199,459,1591,1616]" pageId="110" pageNumber="111">
<emphasis id="B97BEAD47B160368EEF8F99BFE88F9AD" bold="true" box="[199,334,1591,1616]" pageId="110" pageNumber="111">Host-plant.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>