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<document id="DA891460FB5C67D954A269EEF42C0F4E" ID-DOI="10.11865/zs.2021102" ID-ISSN="2095-6827" ID-Zenodo-Dep="5366340" ID-ZooBank="7AA28522-9CBA-4A0E-A97B-B6A40C6DBB7B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630256903413" checkinUser="diego" docAuthor="Wang, Luyu, Lu, Tian, Cai, Ducheng, Barrion, Alberto Tomas, Heong, Kong-Luen, Li, Shuqiang &amp; Zhang, Zhisheng" docDate="2021" docId="0391BA55FF8AFF923DE4F8C82D01DD15" docLanguage="en" docName="Review of the wolf spiders from Hainan Island, China (Araneae_ Lycosidae).pdf" docOrigin="Zoological Systematics 46 (1)" docStyle="DocumentStyle:07E7829F32901B127B59E960199959BD.1:ZoolSyst.2014-.journal_article" docStyleId="07E7829F32901B127B59E960199959BD" docStyleName="ZoolSyst.2014-.journal_article" docStyleVersion="1" docTitle="Pardosa pusiola" docType="treatment" docVersion="3" lastPageNumber="50" masterDocId="FFA8C22DFFABFFB03D6CFFF32E53DC79" masterDocTitle="Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae)" masterLastPageNumber="74" masterPageNumber="16" pageNumber="49" updateTime="1699199438376" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DCD0F0F283F114155D7FC1D505019AD7">Review of the wolf spiders from Hainan Island, China (Araneae: Lycosidae)</mods:title>
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<mods:namePart id="8E9DA17908F1C2DCBF9012BAADBCAF93">Wang, Luyu</mods:namePart>
<mods:affiliation id="B02D2FDA6BB727CBFFF242C59FFACEB7">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China</mods:affiliation>
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<mods:namePart id="1F437EC6C415FBCBFD62C0E83036DB82">Lu, Tian</mods:namePart>
<mods:affiliation id="27A35049474C3CE79091CC0095C3C66C">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China</mods:affiliation>
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<mods:namePart id="65853E0784407892EC2F4A6AE43DA4A3">Cai, Ducheng</mods:namePart>
<mods:affiliation id="AD7E947FFD475E379A330AE7B028D160">Institute of Tropical Agriculture and Forestry, Hainan University, Haikou, Hainan 570228, China</mods:affiliation>
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<mods:namePart id="6F22182618256C7B3DE101C0294E9D6A">Barrion, Alberto Tomas</mods:namePart>
<mods:affiliation id="F604B2D3923961D246A1FFF07A601F51">Crop and Environmental Science Division, International Rice Research Institute (IRRI), Los Baños 4030, Laguna, Philippines</mods:affiliation>
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<mods:namePart id="11E2D65882114F67140DF3D5A8F3B8EC">Heong, Kong-Luen</mods:namePart>
<mods:affiliation id="7E16670477BC4C1E05E89D450BCF747D">Institute of Biological Sciences and Museum of Natural History, University of the Philippines Los Baños, 4031, Laguna, Philippines</mods:affiliation>
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<mods:namePart id="D817D89405A097F16CC7A4943A6BDD23">Li, Shuqiang</mods:namePart>
<mods:affiliation id="4E66105C0B21DB84F71E2A46DB0682C4">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China Corresponding author, E-mail: zhangzs 327 @ qq. com</mods:affiliation>
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<mods:namePart id="2759B15DB6FCEE11F90D196857039E74">Zhang, Zhisheng</mods:namePart>
<mods:affiliation id="D080A6E4613C993940ECD0D9CCAE6FC2">Key Laboratory of Eco-environments in Three Gorges Reservoir Region (Ministry of Education), School of Life Sciences, Southwest University, Chongqing 400715, China</mods:affiliation>
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<mods:title id="EFBB8E26922267961C2A538ECBBC3165">Zoological Systematics</mods:title>
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<mods:date id="486EAAEE4A878C4E0E4661F651FAA022">2021</mods:date>
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<paragraph id="8B870B43FF8AFF913DE4F8C82CB1DB2A" blockId="33.[136,1330,1851,1940]" box="[136,738,1851,1875]" pageId="33" pageNumber="49">
<heading id="D0CFBC2FFF8AFF913DE4F8C82CB1DB2A" box="[136,738,1851,1875]" fontSize="10" level="2" pageId="33" pageNumber="49" reason="2">
<taxonomicName id="4C3870C0FF8AFF913DE4F8C82FBADB2A" ID-CoL="4DJZP" authority="(Thorell, 1891)" baseAuthorityName="Thorell" baseAuthorityYear="1891" box="[136,489,1851,1875]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="49" phylum="Arthropoda" rank="species" species="pusiola">
<emphasis id="B94CD751FF8AFF913DE4F8C82FBADB2A" bold="true" box="[136,489,1851,1875]" pageId="33" pageNumber="49">
<emphasis id="B94CD751FF8AFF913DE4F8C82F6ADB2A" bold="true" box="[136,313,1851,1875]" italics="true" pageId="33" pageNumber="49">Pardosa pusiola</emphasis>
(
<bibRefCitation id="EFA976B2FF8AFF913C2BF8C82FB2DB2A" author="Thorell" box="[327,481,1851,1875]" pageId="33" pageNumber="49" pagination="1 - 149" refId="ref18238" refString="Thorell, T. 1891. Spindlar fran Nikobarerna och andra delar af sodra Asien. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 24 (2): 1 - 149." type="journal article" year="1891">Thorell, 1891</bibRefCitation>
)
</emphasis>
</taxonomicName>
(
<figureCitation id="130317C6FF8AFF913F67F8C82CD8DB2A" box="[523,651,1851,1875]" captionStart="Figure 45" captionStartId="35.[155,218,1995,2017]" captionText="Figure 45. Pardosa pusiola (Thorell, 1891), male and female from Hainan. A. Left male palp, ventral view; B. Same, retrolateral view; C. Left male palp, bulbus, ventral view; D. Same, retrolateral view; E. Epigyne, ventral view; F. Vulva, dorsal view. Scale bars = 0.2mm." pageId="33" pageNumber="49">Figs 45AF</figureCitation>
,
<figureCitation id="130317C6FF8AFF913FFAF8C82C88DB2A" box="[662,731,1851,1875]" captionStart="Figure 46" captionStartId="36.[155,218,1974,1996]" captionTargetBox="[189,1398,499,1933]" captionTargetId="figure-156@36.[189,1398,499,1933]" captionTargetPageId="36" captionText="Figure 46. Pardosa pusiola (Thorell, 1891), male and female from Hainan. A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Left male palp, bulbus, ventral view; D. Same, retrolateral view; E. Left male palp, ventral view; F. Same, retrolateral view; G. Epigyne, ventral view; H. Same, cleaned the dorsal tissue; I. Vulva, dorsal view. Scale bars:A=1.0mm; B=2.0mm; CI=0.2mm." figureDoi="http://doi.org/10.5281/zenodo.5366380" httpUri="https://zenodo.org/record/5366380/files/figure.png" pageId="33" pageNumber="49">46AI</figureCitation>
)
</heading>
</paragraph>
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<subSubSection id="C32258C8FF8AFF913DE4F8AD2A43DBEA" pageId="33" pageNumber="49" type="reference_group">
<paragraph id="8B870B43FF8AFF913DE4F8AD2B61DB0D" blockId="33.[136,1330,1851,1940]" box="[136,1330,1885,1908]" pageId="33" pageNumber="49">
<treatmentCitationGroup id="AB282C6DFF8AFF913DE4F8AD2B61DB0D" box="[136,1330,1885,1908]" pageId="33" pageNumber="49">
<treatmentCitation id="0A992D52FF8AFF913DE4F8AD2F96DB0A" author="Thorell" box="[136,453,1885,1908]" page="65" pageId="33" pageNumber="49" year="1891">
<taxonomicName id="4C3870C0FF8AFF913DE4F8AD2F96DB0A" authority="Thorell, 1891: 65" authorityName="Thorell" authorityPageNumber="65" authorityYear="1891" box="[136,453,1885,1908]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="49" phylum="Arthropoda" rank="species" species="pusiola">
<emphasis id="B94CD751FF8AFF913DE4F8AD2F4ADB0A" box="[136,281,1886,1907]" italics="true" pageId="33" pageNumber="49">Lycosa pusiola</emphasis>
<bibRefCitation id="EFA976B2FF8AFF913C72F8AD2F96DB0A" author="Thorell" box="[286,453,1885,1908]" pageId="33" pageNumber="49" pagination="1 - 149" refId="ref18238" refString="Thorell, T. 1891. Spindlar fran Nikobarerna och andra delar af sodra Asien. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 24 (2): 1 - 149." type="journal article" year="1891">Thorell, 1891: 65</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<specimenCount id="9D3EC0CAFF8AFF913CB8F8AD2FB9DB0D" box="[468,490,1886,1908]" pageId="33" pageNumber="49" type="male"></specimenCount>
<specimenCount id="9D3EC0CAFF8AFF913C85F8AD2FABDB0D" box="[489,504,1886,1908]" pageId="33" pageNumber="49" type="female"></specimenCount>
). For full list of publications and synonyms concerning this species, see
<bibRefCitation id="EFA976B2FF8AFF9139DDF8AD2B7DDB0D" author="WSC &amp; World Spider Catalog" box="[1201,1326,1886,1908]" pageId="33" pageNumber="49" refId="ref18550" refString="WSC. 2020. World Spider Catalog. Version 21.5. Natural History Museum Bern. Available from: http: // wsc. nmbe. ch (accessed on 9 November 2020). doi: 10.24436 / 2." type="book" year="2020">WSC (2020)</bibRefCitation>
.
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</paragraph>
<paragraph id="8B870B43FF8AFF913DE4F88D2A43DBEA" blockId="33.[136,1330,1851,1940]" box="[136,1040,1917,1940]" pageId="33" pageNumber="49">
<taxonomicName id="4C3870C0FF8AFF913DE4F88D2CA4DBEA" ID-CoL="4DJP9" authority="Barrion, Barrion-Dupo &amp; Heong, 2012: 15" authorityName="Barrion, Barrion-Dupo &amp; Heong" authorityPageNumber="15" authorityYear="2012" box="[136,759,1917,1940]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="49" phylum="Arthropoda" rank="species" species="daxiansongi">
<emphasis id="B94CD751FF8AFF913DE4F88D2F07DBEA" box="[136,340,1918,1939]" italics="true" pageId="33" pageNumber="49">Pardosa daxiansongi</emphasis>
Barrion, Barrion-Dupo &amp; Heong, 2012: 15
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, figs 16AD (
<specimenCount id="9D3EC0CAFF8AFF913EEEF88D2DC1DBED" box="[898,914,1918,1940]" pageId="33" pageNumber="49" type="female"></specimenCount>
).
<emphasis id="B94CD751FF8AFF913ED4F88D2A43DBEA" bold="true" box="[952,1040,1918,1939]" pageId="33" pageNumber="49">syn. nov.</emphasis>
</paragraph>
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<paragraph id="8B870B43FF8AFF913DD1F8402BF9DB94" blockId="33.[136,1452,1970,2064]" pageId="33" pageNumber="49">
Description. See Wang &amp; Zhang (2014) for both sexes. Habitus, male palp and epigyne as in
<figureCitation id="130317C6FF8AFF9139F3F8402B4CDBB3" box="[1183,1311,1970,1995]" captionStart="Figure 45" captionStartId="35.[155,218,1995,2017]" captionText="Figure 45. Pardosa pusiola (Thorell, 1891), male and female from Hainan. A. Left male palp, ventral view; B. Same, retrolateral view; C. Left male palp, bulbus, ventral view; D. Same, retrolateral view; E. Epigyne, ventral view; F. Vulva, dorsal view. Scale bars = 0.2mm." pageId="33" pageNumber="49">Figs 45AF</figureCitation>
,
<figureCitation id="130317C6FF8AFF913845F8402B22DBB2" box="[1321,1393,1971,1995]" captionStart="Figure 46" captionStartId="36.[155,218,1974,1996]" captionTargetBox="[189,1398,499,1933]" captionTargetId="figure-156@36.[189,1398,499,1933]" captionTargetPageId="36" captionText="Figure 46. Pardosa pusiola (Thorell, 1891), male and female from Hainan. A. Male habitus, dorsal view; B. Female habitus, dorsal view; C. Left male palp, bulbus, ventral view; D. Same, retrolateral view; E. Left male palp, ventral view; F. Same, retrolateral view; G. Epigyne, ventral view; H. Same, cleaned the dorsal tissue; I. Vulva, dorsal view. Scale bars:A=1.0mm; B=2.0mm; CI=0.2mm." figureDoi="http://doi.org/10.5281/zenodo.5366380" httpUri="https://zenodo.org/record/5366380/files/figure.png" pageId="33" pageNumber="49">46AI</figureCitation>
. Material examined.
<specimenCount id="9D3EC0CAFF8AFF913CFBF8262FEBDB94" box="[407,440,2005,2029]" pageId="33" pageNumber="49" type="female">1♀</specimenCount>
(
<typeStatus id="5483B5E1FF8AFF913CA4F8262C74DB94" box="[456,551,2005,2029]" pageId="33" pageNumber="49" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C3870C0FF8AFF913F23F8252CBEDB94" authorityName="Barrion, Barrion-Dupo &amp; Heong" authorityYear="2012" box="[591,749,2005,2029]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="33" pageNumber="49" phylum="Arthropoda" rank="species" species="daxiansongi">
<emphasis id="B94CD751FF8AFF913F23F8252CBEDB94" box="[591,749,2005,2029]" italics="true" pageId="33" pageNumber="49">P. daxiansongi</emphasis>
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), Dapo, Dapo Village,
<date id="FF862D83FF8AFF913E80F8262AFBDB94" box="[1004,1192,2005,2029]" pageId="33" pageNumber="49" value="2011-03-25" valueMax="2011-04-09" valueMin="2011-03-25">25.III9.IV. 2011</date>
, A.T. Barrion
<emphasis id="B94CD751FF8AFF913829F8252B2FDB94" box="[1349,1404,2005,2029]" italics="true" pageId="33" pageNumber="49">et al.</emphasis>
leg.
</paragraph>
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(HANU);
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,
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Danzhou City, near Tropical Plants Garden,
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, elev.
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&amp;
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leg. (
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<paragraph id="8B870B43FF89FF923DD1FEFC2AFFDD5E" blockId="34.[136,1452,236,364]" box="[189,1196,271,296]" pageId="34" pageNumber="50">
Distribution. South
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,
<collectingCountry id="F32F4BD3FF89FF923CB2FEFC2C0EDD5E" box="[478,605,271,295]" name="Bangladesh" pageId="34" pageNumber="50">Bangladesh</collectingCountry>
,
<collectingCountry id="F32F4BD3FF89FF923F04FEFC2CEBDD5E" box="[616,696,271,295]" name="Bhutan" pageId="34" pageNumber="50">Bhutan</collectingCountry>
,
<collectingCountry id="F32F4BD3FF89FF923FAEFEFC2CA8DD5E" box="[706,763,271,295]" name="India" pageId="34" pageNumber="50">India</collectingCountry>
,
<collectingCountry id="F32F4BD3FF89FF923E6AFEFC2D3DDD5E" box="[774,878,271,295]" name="Indonesia" pageId="34" pageNumber="50">Indonesia</collectingCountry>
,
<collectingCountry id="F32F4BD3FF89FF923E17FEE32DB7DD51" box="[891,996,272,296]" name="Myanmar" pageId="34" pageNumber="50">Myanmar</collectingCountry>
,
<collectingCountry id="F32F4BD3FF89FF923E83FEFC2A62DD5E" box="[1007,1073,271,295]" name="Nepal" pageId="34" pageNumber="50">Nepal</collectingCountry>
,
<collectingCountry id="F32F4BD3FF89FF923950FEFC2AFBDD5E" box="[1084,1192,271,295]" name="Sri Lanka" pageId="34" pageNumber="50">Sri Lanka</collectingCountry>
.
</paragraph>
<paragraph id="8B870B43FF89FF923DD1FEC12D01DD15" blockId="34.[136,1452,236,364]" pageId="34" pageNumber="50">
Remarks. We examined the
<typeStatus id="5483B5E1FF89FF923C86FEC12C14DD33" box="[490,583,306,330]" pageId="34" pageNumber="50" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C3870C0FF89FF923F00FEC02CB7DD33" box="[620,740,306,330]" class="Arachnida" family="Lycosidae" genus="Pardosa" kingdom="Animalia" order="Araneae" pageId="34" pageNumber="50" phylum="Arthropoda" rank="species" species="daxiangi">
<emphasis id="B94CD751FF89FF923F00FEC02CB7DD33" box="[620,740,306,330]" italics="true" pageId="34" pageNumber="50">P. daxiangi</emphasis>
</taxonomicName>
as well as several specimens from other provinces in South
<collectingCountry id="F32F4BD3FF89FF923800FEC12BFFDD33" box="[1388,1452,306,330]" name="China" pageId="34" pageNumber="50">China</collectingCountry>
and Southeast Asia, confirming this is a widely distributed species.
</paragraph>
</subSubSection>
</treatment>
</document>