treatments-xml/data/03/91/6A/03916A48A244FFA7FF3CF94BFBF83F97.xml
2024-06-21 12:22:17 +02:00

541 lines
107 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="6CE34910C9AD9DF9A30794EFD3C99057" ID-CLB-Dataset="57521" ID-DOI="10.11646/zootaxa.5284.3.6" ID-GBIF-Dataset="568f2118-0a41-4aa0-8bb6-cb1b84862a29" ID-ISSN="1175-5326" ID-Zenodo-Dep="7929709" ID-ZooBank="E742B6DC-A5CB-4CB9-8DA2-D464FE3EC249" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683891156715" checkinUser="plazi" docAuthor="Felder, Darryl L. &amp; Dworschak, Peter C." docDate="2023" docId="03916A48A244FFA7FF3CF94BFBF83F97" docLanguage="en" docName="zootaxa.5284.3.6.pdf" docOrigin="Zootaxa 5284 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5284.3.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paragourretia sandrae Felder &amp; Dworschak 2023, n. sp." docType="treatment" docVersion="8" lastPageNumber="560" masterDocId="FFA81230A245FFA0FFABFF8CFFA93802" masterDocTitle="A new mud shrimp of the genus Paragourretia Sakai, 2004, from the Gulf of Mexico (Crustacea: Decapoda: Ctenochelidae)" masterLastPageNumber="563" masterPageNumber="553" pageNumber="554" updateTime="1699515614976" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="CE0064C392EA3F96A110247D8622CD00" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A12EAD0F8465BA13F03DCBFE9C41AF0F">
<mods:title id="919FA0D6105C4555060B150D90B757AE">A new mud shrimp of the genus Paragourretia Sakai, 2004, from the Gulf of Mexico (Crustacea: Decapoda: Ctenochelidae)</mods:title>
</mods:titleInfo>
<mods:name id="409E7897A4C6F401DD7B842E5505A324" type="personal">
<mods:role id="41635C2A61735F2A3BE1A72EF2A0ECE2">
<mods:roleTerm id="795E73D08D78F4F1B99164F64295B3EC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6FFB6795392121DA696B5A8F372F37EB">Felder, Darryl L.</mods:namePart>
<mods:affiliation id="E92473FF0E836292AC9F840688E40D33">Department of Biology, University of Louisiana, Lafayette, Louisiana 70504 - 2451, U. S. A. &amp; Department of Invertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560, U. S. A Dritte Zoologische Abteilung, Naturhistorisches Museum, Burgring 7, A 1010 Wien, Austria.</mods:affiliation>
</mods:name>
<mods:name id="8C134480C2F496F74A4DCD2BCBE46542" type="personal">
<mods:role id="4617B533D40C62A074DBEB6470173090">
<mods:roleTerm id="8F6B76C93AED024D013C0F16F15E4DA1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B65450B788F5C0DFB66390558A62B0BD">Dworschak, Peter C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="DB096708712121CA45ABB7D71054CEC9">text</mods:typeOfResource>
<mods:relatedItem id="55217A6119BAEA262EFB2A693A5A9D0F" type="host">
<mods:titleInfo id="D19B9221B21B6CD18DD34A577B9026FF">
<mods:title id="96B0E1A8B09BE1448EAA1790C2DA1234">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="00F40328CD0218071628B467E7EF6840">
<mods:date id="7B98D39843FE92EFF63FBA4C495DA519">2023</mods:date>
<mods:detail id="EBB154A03084E293E729F6D52363581F" type="pubDate">
<mods:number id="C4EAFA99D01F2D985CB9FEF073EA9D0C">2023-05-12</mods:number>
</mods:detail>
<mods:detail id="C400B6055785F1CF3AA1431BD3FD5161" type="volume">
<mods:number id="99E9D9049503623A5C8FF4F78B7C320A">5284</mods:number>
</mods:detail>
<mods:detail id="A88DB2B4FE914073E13C1B9B47F8FC4B" type="issue">
<mods:number id="50695B8FF1A7B31A29DFA96D4BEC4447">3</mods:number>
</mods:detail>
<mods:extent id="3492B71CDB13662C898D43779FC6F29D" unit="page">
<mods:start id="48D52093B57C357DB284AF6FCA6841EB">553</mods:start>
<mods:end id="740B914ECD4DBCAE09708AF6DB26A44B">563</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="0ECC511D01246B55290D03BA1B622FFD">
<mods:url id="8D0E69DAC61B27390AD7C581923D11E4">http://dx.doi.org/10.11646/zootaxa.5284.3.6</mods:url>
</mods:location>
<mods:classification id="2F7428D80F7119D67B953851A1120B85">journal article</mods:classification>
<mods:identifier id="AF47F201D974018ABCF3763163AD31AD" type="CLB-Dataset">57521</mods:identifier>
<mods:identifier id="766489C87FAB42117783153A2B6BF7E7" type="DOI">10.11646/zootaxa.5284.3.6</mods:identifier>
<mods:identifier id="1A98C43B36ADFB4D6AC8FFF660B91473" type="GBIF-Dataset">568f2118-0a41-4aa0-8bb6-cb1b84862a29</mods:identifier>
<mods:identifier id="823638DEFA16A0709B301B4752A3EF66" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4DDE3416FF89E4FA181F664C00661F0F" type="Zenodo-Dep">7929709</mods:identifier>
<mods:identifier id="E1BFA05A47C15838D66995FEB534EAE9" type="ZooBank">E742B6DC-A5CB-4CB9-8DA2-D464FE3EC249</mods:identifier>
</mods:mods>
<treatment id="03916A48A244FFA7FF3CF94BFBF83F97" ID-DOI="http://doi.org/10.5281/zenodo.7929445" ID-GBIF-Taxon="209622730" ID-Zenodo-Dep="7929445" LSID="urn:lsid:plazi:treatment:03916A48A244FFA7FF3CF94BFBF83F97" httpUri="http://treatment.plazi.org/id/03916A48A244FFA7FF3CF94BFBF83F97" lastPageId="7" lastPageNumber="560" pageId="1" pageNumber="554">
<subSubSection id="C32288D5A244FFA1FF3CF94BFB0C3F07" pageId="1" pageNumber="554" type="nomenclature">
<paragraph id="8B87DB5EA244FFA1FF3CF94BFE5D3EE3" blockId="1.[151,1189,1734,1833]" box="[151,500,1734,1761]" pageId="1" pageNumber="554">
<heading id="D0CF6C32A244FFA1FF3CF94BFE5D3EE3" bold="true" box="[151,500,1734,1761]" fontSize="11" level="1" pageId="1" pageNumber="554" reason="1">
<emphasis id="B94C074CA244FFA1FF3CF94BFE5D3EE3" bold="true" box="[151,500,1734,1761]" pageId="1" pageNumber="554">
<taxonomicName id="4C38A0DDA244FFA1FF3CF94BFE033EE3" ID-CoL="61ddf58d-a9fc-4bda-857e-68bb603f7fb6" authority="Felder &amp; Dworschak, 2023" authorityName="Felder &amp; Dworschak" authorityYear="2023" box="[151,426,1734,1761]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="554" phylum="Arthropoda" rank="species" species="sandrae" status="sp. nov.">
<emphasis id="B94C074CA244FFA1FF3CF94BFE033EE3" bold="true" box="[151,426,1734,1761]" italics="true" pageId="1" pageNumber="554">Paragourretia sandrae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27FBA37A244FFA1FE1AF94BFE5D3EE3" box="[433,500,1735,1761]" pageId="1" pageNumber="554" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B87DB5EA244FFA1FF3CF966FB0C3F07" blockId="1.[151,1189,1734,1833]" box="[151,1189,1770,1797]" pageId="1" pageNumber="554">
https://zoobank.org/
<uri id="FFA9D75CA244FFA1FED5F966FB0C3F07" box="[382,1189,1770,1797]" pageId="1" pageNumber="554">
urn:lsid:zoobank.org:act:
<uuid id="FF9EE18BA244FFA1FD24F966FB0C3F07" box="[655,1189,1770,1797]" pageId="1" pageNumber="554">4E6884D8-4B64-4A1F-B00E-529BE672BD98</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C32288D5A244FFA1FF3CF883FEF53F2B" box="[151,348,1807,1833]" pageId="1" pageNumber="554" type="description">
<paragraph id="8B87DB5EA244FFA1FF3CF883FEF53F2B" blockId="1.[151,1189,1734,1833]" box="[151,348,1807,1833]" pageId="1" pageNumber="554">
(
<figureCitation id="1303C7DBA244FFA1FF34F883FF4F3F2B" box="[159,230,1807,1833]" captionStart="FIGURE 1" captionStartId="4.[151,250,1756,1780]" captionTargetBox="[160,1424,182,1723]" captionTargetId="figure-29@4.[151,1436,181,1732]" captionTargetPageId="4" captionText="FIGURE 1. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, carapace front, eyestalks, antennular and antennal peduncles, dorsal surface; B, same, right lateral surface; C, right mandible, external surface; D, same, internal surface; E, right maxillule, external surface; F, posterior carapace and pleomeres 16, dorsal surface; G, sixth pleomere, telson, and uropods, right lateral surface; H, same, dorsal surface. Scale bars = 1 mm (AE, G, H); 3 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7929711" httpUri="https://zenodo.org/record/7929711/files/figure.png" pageId="1" pageNumber="554">Figs 1</figureCitation>
,
<figureCitation id="1303C7DBA244FFA1FF5FF883FEAD3F2B" box="[244,260,1807,1833]" captionStart="FIGURE 2" captionStartId="5.[151,250,1627,1651]" captionTargetBox="[152,1418,183,1602]" captionTargetId="figure-17@5.[151,1436,181,1604]" captionTargetPageId="5" captionText="FIGURE 2. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="1" pageNumber="554">2</figureCitation>
,
<figureCitation id="1303C7DBA244FFA1FEBBF883FEFD3F2B" box="[272,340,1807,1833]" captionStart="FIGURE 3" captionStartId="6.[151,250,1148,1172]" captionTargetBox="[151,1436,181,1124]" captionTargetId="figure-29@6.[151,1436,181,1124]" captionTargetPageId="6" captionText="FIGURE 3. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304): A, right lateral surface; B, dorsal surface. Paragourretia biffari (Blanco Rambla &amp; Liñero Arana, 1994), male, cl 10.6 mm, northwestern Gulf of Mexico, USNM 1540482 (ULLZ 4673): C, dorsal surface. P. biffari, female cl 6.0 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541735 (ULLZ 7305): D, dorsal surface. P. biffari, female holotype, cl 17.3 mm, Venezuela, USNM 259410: E, major (left) cheliped, internal surface; F, minor (right) cheliped, internal surface; G, posterior pleon and uropods, right ventrolateral, arrow indicating angular submarginal keel." figureDoi="http://doi.org/10.5281/zenodo.7929719" httpUri="https://zenodo.org/record/7929719/files/figure.png" pageId="1" pageNumber="554">3AC</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32288D5A244FFA1FF3CF8DBFE663FDF" pageId="1" pageNumber="554" type="materials_examined">
<paragraph id="8B87DB5EA244FFA1FF3CF8DBFE663FDF" blockId="1.[151,1437,1878,2013]" pageId="1" pageNumber="554">
<emphasis id="B94C074CA244FFA1FF3CF8DBFEE83F72" bold="true" box="[151,321,1878,1905]" pageId="1" pageNumber="554">Type material</emphasis>
.
<materialsCitation id="3B50D103A244FFA1FEF8F8DBFE663FDF" ID-GBIF-Occurrence="4106751301" collectingDate="2005-06-17" collectionCode="R, V" collectorName="D. L. Felder &amp; Robles, H &amp; Bracken-Grissom, S &amp; Fredericq, E &amp; Pelican" elevation="53" latitude="22.134" location="Banks" longLatPrecision="12" longitude="-91.3945" municipality="southwestern Gulf" pageId="1" pageNumber="554" specimenCode="USNM 1541885, ULLZ 7304" specimenCount="1" stateProvince="Mexico" typeStatus="holotype">
<typeStatus id="548365FCA244FFA1FEF8F8DBFE123F73" box="[339,443,1879,1905]" pageId="1" pageNumber="554" type="holotype">Holotype</typeStatus>
(southwestern Gulf of
<collectingRegion id="49FC15BCA244FFA1FD67F8DBFC8E3F73" box="[716,807,1879,1905]" country="Mexico" name="Mexico" pageId="1" pageNumber="554">Mexico</collectingRegion>
): male, cl 4.4 mm; dredge station NSF-II-096, muddy eroded calcareous rubble,
<location id="8EE78D85A244FFA1FE15F8F7FD2C3F97" LSID="urn:lsid:plazi:treatment:03916A48A244FFA7FF3CF94BFBF83F97:8EE78D85A244FFA1FE15F8F7FD2C3F97" box="[446,645,1915,1941]" latitude="22.134" longLatPrecision="12" longitude="-91.3945" municipality="southwestern Gulf" name="Banks" pageId="1" pageNumber="554" stateProvince="Mexico">Campeche Banks</location>
,
<geoCoordinate id="EE0CBD99A244FFA1FD3BF8F7FC8C3F97" box="[656,805,1915,1941]" degrees="22" direction="north" minutes="08.04" orientation="latitude" pageId="1" pageNumber="554" precision="9" value="22.134">22° 08.04´N</geoCoordinate>
,
<geoCoordinate id="EE0CBD99A244FFA1FC9AF8F7FC633F97" box="[817,970,1915,1941]" degrees="91" direction="west" minutes="23.67" orientation="longitude" pageId="1" pageNumber="554" precision="9" value="-91.3945">91° 23.67´W</geoCoordinate>
, 53 m,
<collectingDate id="EFC20476A244FFA1FBB7F8F7FB1C3F97" box="[1052,1205,1914,1941]" pageId="1" pageNumber="554" value="2005-06-17">
<date id="FF86FD9EA244FFA1FBB7F8F7FB1C3F97" box="[1052,1205,1914,1941]" pageId="1" pageNumber="554" value="2005-06-17">17 June 2005</date>
</collectingDate>
; coll
<collectorName id="26CDBE88A244FFA1FB58F8F7FADE3F97" box="[1267,1399,1915,1941]" pageId="1" pageNumber="554">D.L. Felder</collectorName>
, R.
<collectorName id="26CDBE88A244FFA1FF3CF813FEA03FBB" box="[151,265,1951,1977]" pageId="1" pageNumber="554">Robles, H</collectorName>
.
<collectorName id="26CDBE88A244FFA1FEB8F813FE5E3FBB" box="[275,503,1951,1977]" pageId="1" pageNumber="554">Bracken-Grissom, S</collectorName>
.
<collectorName id="26CDBE88A244FFA1FDA9F813FD273FBB" box="[514,654,1951,1977]" pageId="1" pageNumber="554">Fredericq, E</collectorName>
. Garcia and colleagues aboard
<emphasis id="B94C074CA244FFA1FC4EF813FBC33FBB" box="[997,1130,1951,1977]" italics="true" pageId="1" pageNumber="554">
R/V
<collectorName id="26CDBE88A244FFA1FBBFF813FBC33FBB" box="[1044,1130,1951,1977]" pageId="1" pageNumber="554">Pelican</collectorName>
</emphasis>
;
<specimenCode id="DB9E7325A244FFA1FBDCF813FA993FBB" box="[1143,1328,1950,1977]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="554" type="Museum">USNM 1541885</specimenCode>
(=
<specimenCode id="DB9E7325A244FFA1FAFEF813FF7B3FDF" collectionCode="ULLZ" country="USA" httpUri="http://grbio.org/cool/ggzt-j23h" name="University of Louisiana at Layafette, Zoological Collection" pageId="1" pageNumber="554">ULLZ 7304</specimenCode>
, photograph voucher).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32288D5A247FFA2FF6CFF1AFB7039D3" pageId="2" pageNumber="555" type="diagnosis">
<paragraph id="8B87DB5EA247FFA2FF6CFF1AFB7039D3" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="555">
<emphasis id="B94C074CA247FFA2FF6CFF1AFE9338B2" bold="true" box="[199,314,150,176]" pageId="2" pageNumber="555">Diagnosis</emphasis>
. Carapace with narrow triangular spiniform rostrum reaching beyond midlength of eyestalks. Eyestalk with subterminal cornea in distal third, diameter spanning over half of eyestalk width. Major chela with merus inferior keel bearing proximal hooked spine and adjacent ancillary spine, otherwise microdenticulate. First pleonal tergite crossed dorsally by smooth transverse furrow in anterior half, posterior sclerite narrowing anteriorly to acute middorsal terminus, sixth with neither angular nor arched projection in anterior third of lateral margin. Telson length slightly exceeding width, lateral margins converging posterior to weak lateral lobes, posterior margin truncate, lacking median spine. Uropodal endopod elongate ovoid with low longitudinal ridge; exopod anterodorsal plate obscure to obsolescent, lateral margin lacking incision. Diagnostic sequence data for the COI mitochondrial gene is provided under GenBank www.ncbi.nlm.nih.gov/genbank) accession number OQ413315.
</paragraph>
</subSubSection>
<subSubSection id="C32288D5A247FFA7FF6CFE56FA29398B" lastPageId="7" lastPageNumber="560" pageId="2" pageNumber="555" type="description">
<paragraph id="8B87DB5EA247FFA2FF6CFE56FC393AAB" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="555">
<emphasis id="B94C074CA247FFA2FF6CFE56FEF839F6" bold="true" box="[199,337,474,500]" pageId="2" pageNumber="555">Description</emphasis>
. Carapace front (
<figureCitation id="1303C7DBA247FFA2FDB2FE57FD3239F7" box="[537,667,475,501]" captionStart="FIGURE 1" captionStartId="4.[151,250,1756,1780]" captionTargetBox="[160,1424,182,1723]" captionTargetId="figure-29@4.[151,1436,181,1732]" captionTargetPageId="4" captionText="FIGURE 1. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, carapace front, eyestalks, antennular and antennal peduncles, dorsal surface; B, same, right lateral surface; C, right mandible, external surface; D, same, internal surface; E, right maxillule, external surface; F, posterior carapace and pleomeres 16, dorsal surface; G, sixth pleomere, telson, and uropods, right lateral surface; H, same, dorsal surface. Scale bars = 1 mm (AE, G, H); 3 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7929711" httpUri="https://zenodo.org/record/7929711/files/figure.png" pageId="2" pageNumber="555">Figs 1A, B</figureCitation>
;
<figureCitation id="1303C7DBA247FFA2FD02FE57FD4739F7" box="[681,750,475,501]" captionStart="FIGURE 3" captionStartId="6.[151,250,1148,1172]" captionTargetBox="[151,1436,181,1124]" captionTargetId="figure-29@6.[151,1436,181,1124]" captionTargetPageId="6" captionText="FIGURE 3. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304): A, right lateral surface; B, dorsal surface. Paragourretia biffari (Blanco Rambla &amp; Liñero Arana, 1994), male, cl 10.6 mm, northwestern Gulf of Mexico, USNM 1540482 (ULLZ 4673): C, dorsal surface. P. biffari, female cl 6.0 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541735 (ULLZ 7305): D, dorsal surface. P. biffari, female holotype, cl 17.3 mm, Venezuela, USNM 259410: E, major (left) cheliped, internal surface; F, minor (right) cheliped, internal surface; G, posterior pleon and uropods, right ventrolateral, arrow indicating angular submarginal keel." figureDoi="http://doi.org/10.5281/zenodo.7929719" httpUri="https://zenodo.org/record/7929719/files/figure.png" pageId="2" pageNumber="555">3A, B</figureCitation>
) with acute, narrowly triangular rostrum, deflected between eyestalks in lateral view, terminally spiniform, reaching beyond mid-length of eyestalks, rostral base flanked laterally by low, obtuse shoulders forming outer angles of orbits, orbital margin laterally lined by several small setal pits; postantennal carapace margin transverse, near vertical, strongly inset from front, slightly arched transverse row of four small setal pits posterior to margin; dorsal oval not defined; linea thalassinica extending full length of carapace; hepatic boss weakly developed; cardiac prominence poorly defined.
</paragraph>
<paragraph id="8B87DB5EA247FFA2FF6CFD3FFBED3B5F" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="555">
Eyestalks (
<figureCitation id="1303C7DBA247FFA2FEE8FD3FFE143ACF" box="[323,445,691,717]" captionStart="FIGURE 1" captionStartId="4.[151,250,1756,1780]" captionTargetBox="[160,1424,182,1723]" captionTargetId="figure-29@4.[151,1436,181,1732]" captionTargetPageId="4" captionText="FIGURE 1. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, carapace front, eyestalks, antennular and antennal peduncles, dorsal surface; B, same, right lateral surface; C, right mandible, external surface; D, same, internal surface; E, right maxillule, external surface; F, posterior carapace and pleomeres 16, dorsal surface; G, sixth pleomere, telson, and uropods, right lateral surface; H, same, dorsal surface. Scale bars = 1 mm (AE, G, H); 3 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7929711" httpUri="https://zenodo.org/record/7929711/files/figure.png" pageId="2" pageNumber="555">Fig. 1A, B</figureCitation>
) elongate, narrowing distally to subtriangular tips not reaching to second (penultimate) article of antennular peduncle, carried slightly deflected, swollen proximally in lateral view, distal part dorsoventrally flattened, lateral margin developed into narrow crest laterally; subterminal cornea defined as rounded dorsal swelling in distal third of eyestalk, diameter spanning over half of eyestalk width, poorly faceted, pigmentation (
<figureCitation id="1303C7DBA247FFA2FAB7FC93FA3A3B3B" box="[1308,1427,799,825]" captionStart="FIGURE 3" captionStartId="6.[151,250,1148,1172]" captionTargetBox="[151,1436,181,1124]" captionTargetId="figure-29@6.[151,1436,181,1124]" captionTargetPageId="6" captionText="FIGURE 3. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304): A, right lateral surface; B, dorsal surface. Paragourretia biffari (Blanco Rambla &amp; Liñero Arana, 1994), male, cl 10.6 mm, northwestern Gulf of Mexico, USNM 1540482 (ULLZ 4673): C, dorsal surface. P. biffari, female cl 6.0 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541735 (ULLZ 7305): D, dorsal surface. P. biffari, female holotype, cl 17.3 mm, Venezuela, USNM 259410: E, major (left) cheliped, internal surface; F, minor (right) cheliped, internal surface; G, posterior pleon and uropods, right ventrolateral, arrow indicating angular submarginal keel." figureDoi="http://doi.org/10.5281/zenodo.7929719" httpUri="https://zenodo.org/record/7929719/files/figure.png" pageId="2" pageNumber="555">Fig. 3A, B</figureCitation>
) filling cornea in life, dispersed into multiple coalesced dark spots with preservation.
</paragraph>
<paragraph id="8B87DB5EA247FFA2FF6CFCEBFB6E3C5B" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="555">
Antennular peduncle (
<figureCitation id="1303C7DBA247FFA2FE66FCEBFDF83B83" box="[461,593,871,897]" captionStart="FIGURE 1" captionStartId="4.[151,250,1756,1780]" captionTargetBox="[160,1424,182,1723]" captionTargetId="figure-29@4.[151,1436,181,1732]" captionTargetPageId="4" captionText="FIGURE 1. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, carapace front, eyestalks, antennular and antennal peduncles, dorsal surface; B, same, right lateral surface; C, right mandible, external surface; D, same, internal surface; E, right maxillule, external surface; F, posterior carapace and pleomeres 16, dorsal surface; G, sixth pleomere, telson, and uropods, right lateral surface; H, same, dorsal surface. Scale bars = 1 mm (AE, G, H); 3 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7929711" httpUri="https://zenodo.org/record/7929711/files/figure.png" pageId="2" pageNumber="555">Fig. 1A, B</figureCitation>
) much shorter and distinctly heavier than antennal peduncle, reaching approximately to distal end of fourth article of antennal peduncle; first (basal) article swollen, more than twice length of second, third article approximately 1 and 1.5 times length of second; second and third articles bearing few minute marginal setae. Antennular flagella missing. Antennal peduncle (fifth articles and flagella missing) distinctly overreaching antennular peduncle; first article with short produced ventrolateral process bearing excretory pore; length of second article about twice width, distal articulation to third article overreached dorsally by short bladelike scaphocerite; fourth article slightly exceeding combined lengths of first two, all sparsely setose.
</paragraph>
<paragraph id="8B87DB5EA247FFA2FF6CFBEFFE273DE7" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="555">
Mandible (
<figureCitation id="1303C7DBA247FFA2FEE8FBEFFE143C7F" box="[323,445,1123,1149]" captionStart="FIGURE 1" captionStartId="4.[151,250,1756,1780]" captionTargetBox="[160,1424,182,1723]" captionTargetId="figure-29@4.[151,1436,181,1732]" captionTargetPageId="4" captionText="FIGURE 1. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, carapace front, eyestalks, antennular and antennal peduncles, dorsal surface; B, same, right lateral surface; C, right mandible, external surface; D, same, internal surface; E, right maxillule, external surface; F, posterior carapace and pleomeres 16, dorsal surface; G, sixth pleomere, telson, and uropods, right lateral surface; H, same, dorsal surface. Scale bars = 1 mm (AE, G, H); 3 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7929711" httpUri="https://zenodo.org/record/7929711/files/figure.png" pageId="2" pageNumber="555">Fig. 1C, D</figureCitation>
) with palp of three articles, proximal articles short, little if any longer than broad, third article elongate, heavy, arched, distal setae elongate, densely plumose; gnathal lobe subquadrate, distolateral shoulder angular, incisor process with 4 well-defined subacute triangular teeth on cutting margin, tips corneous, molar process on internal surface forming internal lip of concavity accommodating flexed palp, lip originating proximal and internal to incisor teeth. Maxillule (
<figureCitation id="1303C7DBA247FFA2FD81FB7FFDD73D0F" box="[554,638,1267,1293]" captionStart="FIGURE 1" captionStartId="4.[151,250,1756,1780]" captionTargetBox="[160,1424,182,1723]" captionTargetId="figure-29@4.[151,1436,181,1732]" captionTargetPageId="4" captionText="FIGURE 1. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, carapace front, eyestalks, antennular and antennal peduncles, dorsal surface; B, same, right lateral surface; C, right mandible, external surface; D, same, internal surface; E, right maxillule, external surface; F, posterior carapace and pleomeres 16, dorsal surface; G, sixth pleomere, telson, and uropods, right lateral surface; H, same, dorsal surface. Scale bars = 1 mm (AE, G, H); 3 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7929711" httpUri="https://zenodo.org/record/7929711/files/figure.png" pageId="2" pageNumber="555">Fig. 1E</figureCitation>
) endopodal palp narrow, biarticulate, distal article deflected, over half length of proximal, bearing long terminal seta; proximal endite forming broad subangular lobe, uniformly fringed by dense row of setae along most of mesial margin, setae heavier, some spiniform, at distal prominence; distal endite elongate, terminally broadened, setation of mesial margin including closely set overlain rows of strongly spiniform setae. Maxilla (
<figureCitation id="1303C7DBA247FFA2FEE2FA0FFE0D3D9F" box="[329,420,1411,1437]" captionStart="FIGURE 2" captionStartId="5.[151,250,1627,1651]" captionTargetBox="[152,1418,183,1602]" captionTargetId="figure-17@5.[151,1436,181,1604]" captionTargetPageId="5" captionText="FIGURE 2. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="2" pageNumber="555">Fig. 2A</figureCitation>
) margins setose, endopod constricted distally to form narrow terminus; first and second endites each longitudinally subdivided, mesially directed distal margins fringed by long stiff setae; exopod forming broad scaphognathite.
</paragraph>
<paragraph id="8B87DB5EA247FFA2FF6CFA63FEDA3EBF" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="555">
First maxilliped (
<figureCitation id="1303C7DBA247FFA2FE25FA63FE4E3E0B" box="[398,487,1519,1545]" captionStart="FIGURE 2" captionStartId="5.[151,250,1627,1651]" captionTargetBox="[152,1418,183,1602]" captionTargetId="figure-17@5.[151,1436,181,1604]" captionTargetPageId="5" captionText="FIGURE 2. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="2" pageNumber="555">Fig. 2B</figureCitation>
) margins setose, endopod small, narrow, less than half length of exopod, partially concealed between base of distal endite and exopod; proximal endite a subangular lobe, field of dense stiff setae terminally; distal endite subrectangular, margins and most of external surface densely setose, strongest setae mesially directed, originating distal and along mesial margin; exopod elongate, arcuate, margins setose, strongest setae terminal and along convex lateral margin; epipod broad, not reaching beyond exopod, anterior and posterior lobes subtriangular.
</paragraph>
<paragraph id="8B87DB5EA247FFA2FF6CF94BFD0C3F97" blockId="2.[151,1437,150,2013]" pageId="2" pageNumber="555">
Second maxilliped (
<figureCitation id="1303C7DBA247FFA2FE0DF94BFE553EE3" box="[422,508,1735,1761]" captionStart="FIGURE 2" captionStartId="5.[151,250,1627,1651]" captionTargetBox="[152,1418,183,1602]" captionTargetId="figure-17@5.[151,1436,181,1604]" captionTargetPageId="5" captionText="FIGURE 2. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="2" pageNumber="555">Fig. 2C</figureCitation>
) much smaller than third, both rami setose; endopod merus subrectangular, slightly broadened proximally, weakly arched, length about 3 times width, length about twice combined length of propodus and dactylus; propodus robustly subcylindrical, most of surface setose, length about 1 ½ times breadth; dactylus short, broader than long, rounded terminus bearing long stiff setae; exopod narrow, straplike, carried closely against internal surface of endopod, not reaching end of endopod merus, terminally rounded with stiff marginal setae; short digitiform epipod evident near base of exopod.
</paragraph>
<paragraph id="8B87DB5EA247FFA3FF6CF813FBF1398B" blockId="2.[151,1437,150,2013]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="556" pageId="2" pageNumber="555">
Third maxilliped (
<figureCitation id="1303C7DBA247FFA2FE38F813FDA03FBB" box="[403,521,1951,1977]" captionStart="FIGURE 2" captionStartId="5.[151,250,1627,1651]" captionTargetBox="[152,1418,183,1602]" captionTargetId="figure-17@5.[151,1436,181,1604]" captionTargetPageId="5" captionText="FIGURE 2. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="2" pageNumber="555">Fig. 2D, E</figureCitation>
) with narrow biarticulate exopod, length approximately equal to ischium, bearing long stiff terminal seta; basis bearing spine on inner surface just proximal to articulation with ischium; endopod fringed by long setae, densest on mesial margins of ischium, propodus, and dactylus, ischium subrectangular, length more than twice breadth, internal surface with slightly arched longitudinal row of 13 spiniform teeth forming strong crista dentata, distalmost tooth forming spine, several additional small ancillary teeth trailing row proximally; merus subquadrate, length approximately twice width, length over three-fourths that of ischium, flexor margin with few long setae and strong distal spine; carpus almost as broad as propodus, both much longer than broad, flexor margins of both distinctly convex; dactylus subcylindrical, digitiform, length exceeding two times breadth, with dense subterminal to terminal field of long setae originating primarily from internal surface.
</paragraph>
<paragraph id="8B87DB5EA246FFA3FF6CFE1FFD073B3B" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="556">
First pereopods (
<figureCitation id="1303C7DBA246FFA3FE2DFE1FFDAB39AF" box="[390,514,403,429]" captionStart="FIGURE 2" captionStartId="5.[151,250,1627,1651]" captionTargetBox="[152,1418,183,1602]" captionTargetId="figure-17@5.[151,1436,181,1604]" captionTargetPageId="5" captionText="FIGURE 2. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="3" pageNumber="556">Figs 2F, G</figureCitation>
;
<figureCitation id="1303C7DBA246FFA3FDBBFE1FFDFD39AF" box="[528,596,403,429]" captionStart="FIGURE 3" captionStartId="6.[151,250,1148,1172]" captionTargetBox="[151,1436,181,1124]" captionTargetId="figure-29@6.[151,1436,181,1124]" captionTargetPageId="6" captionText="FIGURE 3. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304): A, right lateral surface; B, dorsal surface. Paragourretia biffari (Blanco Rambla &amp; Liñero Arana, 1994), male, cl 10.6 mm, northwestern Gulf of Mexico, USNM 1540482 (ULLZ 4673): C, dorsal surface. P. biffari, female cl 6.0 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541735 (ULLZ 7305): D, dorsal surface. P. biffari, female holotype, cl 17.3 mm, Venezuela, USNM 259410: E, major (left) cheliped, internal surface; F, minor (right) cheliped, internal surface; G, posterior pleon and uropods, right ventrolateral, arrow indicating angular submarginal keel." figureDoi="http://doi.org/10.5281/zenodo.7929719" httpUri="https://zenodo.org/record/7929719/files/figure.png" pageId="3" pageNumber="556">3A, B</figureCitation>
) strongly heterochelous; major (right) cheliped (
<figureCitation id="1303C7DBA246FFA3FBD4FE1FFB7D39AF" box="[1151,1236,403,429]" captionStart="FIGURE 2" captionStartId="5.[151,250,1627,1651]" captionTargetBox="[152,1418,183,1602]" captionTargetId="figure-17@5.[151,1436,181,1604]" captionTargetPageId="5" captionText="FIGURE 2. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="3" pageNumber="556">Fig. 2F</figureCitation>
) ischium slender, superior margin sinuous, inferior marginal carina armed by row of small denticles, merus superior margin smoothly convex, inferior (flexor) margin forming keel with triangular proximal spine, smaller more distal sharp spine, denticulate margin followed by smooth margin over distal half; carpus short, subtriangular, much broader than long, superior and inferior margins smoothly keeled, superior terminating distally at low rounded corner, inferior terminating distally in obtusely angular rounded corner; propodus heavy, more inflated than carpus, upper margin of palm 3 times length of carpus, length of fixed finger about 2/3 to 3/4 length of palm, palm superior margin keeled proximally, propodus inferior marginal keel weakly serrate, extending onto fixed finger, fixed finger weakly bowed ventrally, prehensile edge with few proximal microdenticles, otherwise unarmed, weak submarginal longitudinal furrow to internal side of tip not notably upturned; dactyl superior margin smooth arched over most of length, prehensile edge with minute notch in proximal 1/4, broadly triangular tooth centered proximal to mid-length, remainder mostly smooth, tip weakly deflected.
</paragraph>
<paragraph id="8B87DB5EA246FFA3FF6CFCCFFE593C13" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="556">
Minor cheliped (
<figureCitation id="1303C7DBA246FFA3FE28FCCFFE723B5F" box="[387,475,835,861]" captionStart="FIGURE 2" captionStartId="5.[151,250,1627,1651]" captionTargetBox="[152,1418,183,1602]" captionTargetId="figure-17@5.[151,1436,181,1604]" captionTargetPageId="5" captionText="FIGURE 2. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="3" pageNumber="556">Fig. 2G</figureCitation>
) ischium narrowly elongate, superior margin sinuous, inferior margin weakly serrate; merus subovoid, superior margin strongly convex, inferior margin with smooth keel with prominent proximal spine (possibly doublet, as damaged); carpus superior margin nearly straight, inferior margin arcuate, article length subequal to breadth; propodus palm longer than fixed finger, palm length over twice breadth, 2.5 times as long as carpus; fixed finger and dactylus with unarmed prehensile edges, gape minimal, weakly hooked tips acute, crossed distally when closely opposed.
</paragraph>
<paragraph id="8B87DB5EA246FFA3FF6CFB97FDAF3CA3" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="556">
Second pereopod (
<figureCitation id="1303C7DBA246FFA3FE30FB97FE5D3C37" box="[411,500,1051,1077]" captionStart="FIGURE 2" captionStartId="5.[151,250,1627,1651]" captionTargetBox="[152,1418,183,1602]" captionTargetId="figure-17@5.[151,1436,181,1604]" captionTargetPageId="5" captionText="FIGURE 2. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="3" pageNumber="556">Fig. 2H</figureCitation>
) chelate, superior margin of merus terminating distally in setal tuft including long stiff seta, flexor margin of merus, distal flexor margin of carpus, inferior margin of propodus lined by long regularly spaced setae, those of fixed finger becoming distally short, stiff, more hooked; outer surface of dactylus and fixed finger bearing tufts of stiff setae.
</paragraph>
<paragraph id="8B87DB5EA246FFA3FF6CFB27FEE03D7B" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="556">
Third pereopod (
<figureCitation id="1303C7DBA246FFA3FE2AFB27FE653CC7" box="[385,460,1195,1221]" captionStart="FIGURE 2" captionStartId="5.[151,250,1627,1651]" captionTargetBox="[152,1418,183,1602]" captionTargetId="figure-17@5.[151,1436,181,1604]" captionTargetPageId="5" captionText="FIGURE 2. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="3" pageNumber="556">Fig. 2I</figureCitation>
) merus length near four times width; carpus elongate, length over twice breadth, inferior margin distally with dense field of long stiff setae; propodus with superior margin ending distally in dense tuft of long setae, inferior margin weakly subdivided into eight low lobes, each surmounted by tuft of long stiff setae, proximal lobe slightly produced to form rounded heel, most of outer surface covered by tufts of setae; dactylus lanciform, slightly hooked, tip slightly twisted from arc of flexure, outer and inner surface with dense distal tuft of short stiff setae.
</paragraph>
<paragraph id="8B87DB5EA246FFA3FF6CFA0FFAD63E0B" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="556">
Fourth pereopod not subchelate, coxa mobile, (
<figureCitation id="1303C7DBA246FFA3FD45FA0FFCEB3D9F" box="[750,834,1411,1437]" captionStart="FIGURE 2" captionStartId="5.[151,250,1627,1651]" captionTargetBox="[152,1418,183,1602]" captionTargetId="figure-17@5.[151,1436,181,1604]" captionTargetPageId="5" captionText="FIGURE 2. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="3" pageNumber="556">Fig. 2J</figureCitation>
), propodus length approximately three times width, inferior (flexor) margin lined by closely spaced long setae, inferodistal corner with dense field of stiff setae, superodistal corner including long stiff seta greatly exceeding full length of dactylus; dactylus approximately half length of propodus, weakly hooked, tapering distally to corneous tip, flexor margin will dense row of stiff setae.
</paragraph>
<paragraph id="8B87DB5EA246FFA3FF6CF99FFB6B3EBF" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="556">
Fifth pereopod (
<figureCitation id="1303C7DBA246FFA3FE21F99FFE403E2F" box="[394,489,1555,1581]" captionStart="FIGURE 2" captionStartId="5.[151,250,1627,1651]" captionTargetBox="[152,1418,183,1602]" captionTargetId="figure-17@5.[151,1436,181,1604]" captionTargetPageId="5" captionText="FIGURE 2. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="3" pageNumber="556">Fig. 2K</figureCitation>
) obscurely subchelate terminally amid dense setation, propodus length slightly exceeding three times breadth, with dense inner and outer fields of long setae proximal to and overlying articulation of dactylus, another along distal half of inferior margin proximal to fixed finger, fixed finger slightly hooked, narrowly acute, terminating in corneous spine; dactylus elongate, approximately half length of propodus, distally twisted from arc of flexure, ending distally in upturned tip, appearing sinuous in lateral outline.
</paragraph>
<paragraph id="8B87DB5EA246FFA3FF6CF94BFA8D3EE3" blockId="3.[151,1437,151,2013]" box="[199,1316,1735,1761]" pageId="3" pageNumber="556">Gills limited to paired arthrobranchs on third maxilliped and each of first through fourth pereopods.</paragraph>
<paragraph id="8B87DB5EA246FFA3FF6CF967FDB53FDF" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="556">
Pleonal tergites glossy smooth, enamel-like dorsally (
<figureCitation id="1303C7DBA246FFA3FC85F967FC2C3F07" box="[814,901,1771,1797]" captionStart="FIGURE 1" captionStartId="4.[151,250,1756,1780]" captionTargetBox="[160,1424,182,1723]" captionTargetId="figure-29@4.[151,1436,181,1732]" captionTargetPageId="4" captionText="FIGURE 1. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, carapace front, eyestalks, antennular and antennal peduncles, dorsal surface; B, same, right lateral surface; C, right mandible, external surface; D, same, internal surface; E, right maxillule, external surface; F, posterior carapace and pleomeres 16, dorsal surface; G, sixth pleomere, telson, and uropods, right lateral surface; H, same, dorsal surface. Scale bars = 1 mm (AE, G, H); 3 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7929711" httpUri="https://zenodo.org/record/7929711/files/figure.png" pageId="3" pageNumber="556">Fig. 1F</figureCitation>
;
<figureCitation id="1303C7DBA246FFA3FC3FF967FC753F07" box="[916,988,1771,1797]" captionStart="FIGURE 3" captionStartId="6.[151,250,1148,1172]" captionTargetBox="[151,1436,181,1124]" captionTargetId="figure-29@6.[151,1436,181,1124]" captionTargetPageId="6" captionText="FIGURE 3. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304): A, right lateral surface; B, dorsal surface. Paragourretia biffari (Blanco Rambla &amp; Liñero Arana, 1994), male, cl 10.6 mm, northwestern Gulf of Mexico, USNM 1540482 (ULLZ 4673): C, dorsal surface. P. biffari, female cl 6.0 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541735 (ULLZ 7305): D, dorsal surface. P. biffari, female holotype, cl 17.3 mm, Venezuela, USNM 259410: E, major (left) cheliped, internal surface; F, minor (right) cheliped, internal surface; G, posterior pleon and uropods, right ventrolateral, arrow indicating angular submarginal keel." figureDoi="http://doi.org/10.5281/zenodo.7929719" httpUri="https://zenodo.org/record/7929719/files/figure.png" pageId="3" pageNumber="556">3A, B</figureCitation>
), few setae and obscure tracks of setal pits laterally. First pleonal tergite crossed by smoothly depressed transverse furrow in anterior half, posterior 2/3 covered by triangular dorsal sclerite with acute anterior, middorsal terminus. Second tergite slightly longer than first, almost twice length of third. Third to fifth tergites slightly decreasing in length to posterior. Sixth tergite approximately 2/3 length of second, dorsally with posterolateral track of fine short setae extending anteriorly as row of setal pits converging anteriorly, ventrolaterally with neither angular nor arched projection or keel in anterior half of marginal to submarginal region.
</paragraph>
<caption id="DF478BD6A241FFA4FF3CF950FDA73F86" ID-DOI="http://doi.org/10.5281/zenodo.7929711" ID-Zenodo-Dep="7929711" httpUri="https://zenodo.org/record/7929711/files/figure.png" pageId="4" pageNumber="557" startId="4.[151,250,1756,1780]" targetBox="[160,1424,182,1723]" targetPageId="4" targetType="figure">
<paragraph id="8B87DB5EA241FFA4FF3CF950FDA73F86" blockId="4.[151,1437,1756,1924]" pageId="4" pageNumber="557">
<emphasis id="B94C074CA241FFA4FF3CF950FEA43EF6" bold="true" box="[151,269,1756,1780]" pageId="4" pageNumber="557">FIGURE 1</emphasis>
.
<taxonomicName id="4C38A0DDA241FFA4FEB1F951FDAA3EF6" authority="Felder &amp; Dworschak, 2023" authorityName="Felder &amp; Dworschak" authorityYear="2023" box="[282,515,1756,1780]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="557" phylum="Arthropoda" rank="species" species="sandrae" status="sp. nov.">
<emphasis id="B94C074CA241FFA4FEB1F951FDAA3EF6" box="[282,515,1756,1780]" italics="true" pageId="4" pageNumber="557">Paragourretia sandrae</emphasis>
</taxonomicName>
<emphasis id="B94C074CA241FFA4FDA2F950FDEA3EF6" bold="true" box="[521,579,1756,1780]" pageId="4" pageNumber="557">
<taxonomicNameLabel id="A27FBA37A241FFA4FDA2F950FDEA3EF6" box="[521,579,1756,1780]" pageId="4" pageNumber="557" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, carapace front, eyestalks, antennular and antennal peduncles, dorsal surface; B, same, right lateral surface; C, right mandible, external surface; D, same, internal surface; E, right maxillule, external surface; F, posterior carapace and pleomeres 16, dorsal surface; G, sixth pleomere, telson, and uropods, right lateral surface; H, same, dorsal surface. Scale bars = 1 mm (AE, G, H); 3 mm (F).
</paragraph>
</caption>
<caption id="DF478BD6A240FFA5FF3CF9D7FC513F49" ID-DOI="http://doi.org/10.5281/zenodo.7929717" ID-Zenodo-Dep="7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="5" pageNumber="558" startId="5.[151,250,1627,1651]" targetBox="[152,1418,183,1602]" targetPageId="5" targetType="figure">
<paragraph id="8B87DB5EA240FFA5FF3CF9D7FC513F49" blockId="5.[151,1437,1627,1868]" pageId="5" pageNumber="558">
<emphasis id="B94C074CA240FFA5FF3CF9D7FEA43E71" bold="true" box="[151,269,1627,1651]" pageId="5" pageNumber="558">FIGURE 2</emphasis>
.
<taxonomicName id="4C38A0DDA240FFA5FEB1F9D0FDAA3E76" authority="Felder &amp; Dworschak, 2023" authorityName="Felder &amp; Dworschak" authorityYear="2023" box="[282,515,1628,1652]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="558" phylum="Arthropoda" rank="species" species="sandrae" status="sp. nov.">
<emphasis id="B94C074CA240FFA5FEB1F9D0FDAA3E76" box="[282,515,1628,1652]" italics="true" pageId="5" pageNumber="558">Paragourretia sandrae</emphasis>
</taxonomicName>
<emphasis id="B94C074CA240FFA5FDA2F9D0FDEA3E76" bold="true" box="[521,579,1628,1652]" pageId="5" pageNumber="558">
<taxonomicNameLabel id="A27FBA37A240FFA5FDA2F9D0FDEA3E76" box="[521,579,1628,1652]" pageId="5" pageNumber="558" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="DF478BD6A243FFA6FF3CFBF0FB053D4A" ID-DOI="http://doi.org/10.5281/zenodo.7929719" ID-Zenodo-Dep="7929719" httpUri="https://zenodo.org/record/7929719/files/figure.png" pageId="6" pageNumber="559" startId="6.[151,250,1148,1172]" targetBox="[151,1436,181,1124]" targetPageId="6" targetType="figure">
<paragraph id="8B87DB5EA243FFA6FF3CFBF0FB053D4A" blockId="6.[151,1436,1148,1352]" pageId="6" pageNumber="559">
<emphasis id="B94C074CA243FFA6FF3CFBF0FEA43C96" bold="true" box="[151,269,1148,1172]" pageId="6" pageNumber="559">FIGURE 3</emphasis>
.
<taxonomicName id="4C38A0DDA243FFA6FEB1FBF1FDAA3C96" authority="Felder &amp; Dworschak, 2023" authorityName="Felder &amp; Dworschak" authorityYear="2023" box="[282,515,1148,1172]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="559" phylum="Arthropoda" rank="species" species="sandrae" status="sp. nov.">
<emphasis id="B94C074CA243FFA6FEB1FBF1FDAA3C96" box="[282,515,1148,1172]" italics="true" pageId="6" pageNumber="559">Paragourretia sandrae</emphasis>
</taxonomicName>
<emphasis id="B94C074CA243FFA6FDA2FBF0FDEA3C96" bold="true" box="[521,579,1148,1172]" pageId="6" pageNumber="559">
<taxonomicNameLabel id="A27FBA37A243FFA6FDA2FBF0FDEA3C96" box="[521,579,1148,1172]" pageId="6" pageNumber="559" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304): A, right lateral surface; B, dorsal surface.
<taxonomicName id="4C38A0DDA243FFA6FCCBFB2DFF7D3CDE" authority="(Blanco Rambla &amp; Linero Arana, 1994)" baseAuthorityName="Blanco Rambla &amp; Linero Arana" baseAuthorityYear="1994" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="559" phylum="Arthropoda" rank="species" species="biffari">
<emphasis id="B94C074CA243FFA6FCCBFB2DFB903CBA" box="[864,1081,1184,1208]" italics="true" pageId="6" pageNumber="559">Paragourretia biffari</emphasis>
(
<bibRefCitation id="EFA9A6AFA243FFA6FBE2FB2CFF643CDE" author="Blanco Rambla, J. P. &amp; Linero Arana, I." pageId="6" pageNumber="559" pagination="16 - 29" refId="ref5558" refString="Blanco Rambla, J. P. &amp; Linero Arana, I. (1994) New records and new species of ghost shrimps (Crustacea: Thalassinidea) from Venezuela. Bulletin of Marine Science, 55, 16 - 29." type="journal article" year="1994">Blanco Rambla &amp; Liñero Arana, 1994</bibRefCitation>
)
</taxonomicName>
, male, cl 10.6 mm, northwestern Gulf of Mexico, USNM 1540482 (ULLZ 4673): C, dorsal surface.
<taxonomicName id="4C38A0DDA243FFA6FB73FB49FA983CDE" baseAuthorityName="Blanco Rambla &amp; Linero Arana" baseAuthorityYear="1994" box="[1240,1329,1220,1244]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="559" phylum="Arthropoda" rank="species" species="biffari">
<emphasis id="B94C074CA243FFA6FB73FB49FA983CDE" box="[1240,1329,1220,1244]" italics="true" pageId="6" pageNumber="559">P. biffari</emphasis>
</taxonomicName>
, female cl 6.0 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541735 (ULLZ 7305): D, dorsal surface.
<taxonomicName id="4C38A0DDA243FFA6FB5AFB65FAE03D02" baseAuthorityName="Blanco Rambla &amp; Linero Arana" baseAuthorityYear="1994" box="[1265,1353,1256,1280]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="559" phylum="Arthropoda" rank="species" species="biffari">
<emphasis id="B94C074CA243FFA6FB5AFB65FAE03D02" box="[1265,1353,1256,1280]" italics="true" pageId="6" pageNumber="559">P. biffari</emphasis>
</taxonomicName>
, female holotype, cl 17.3 mm, Venezuela, USNM 259410: E, major (left) cheliped, internal surface; F, minor (right) cheliped, internal surface; G, posterior pleon and uropods, right ventrolateral, arrow indicating angular submarginal keel.
</paragraph>
</caption>
<paragraph id="8B87DB5EA243FFA6FF6CFAFAFD5D3EB3" blockId="6.[151,1437,1398,2037]" pageId="6" pageNumber="559">
First pleopod uniramous (
<figureCitation id="1303C7DBA243FFA6FE49FAFAFD9F3D92" box="[482,566,1398,1424]" captionStart="FIGURE 2" captionStartId="5.[151,250,1627,1651]" captionTargetBox="[152,1418,183,1602]" captionTargetId="figure-17@5.[151,1436,181,1604]" captionTargetPageId="5" captionText="FIGURE 2. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="6" pageNumber="559">Fig. 2L</figureCitation>
), less than half length of third pleopod, biarticulate, distal article slightly longer than proximal, broadened distal end of article terminating in strong hooked spine over-reaching shorter straight distally directed spine. Second pleopod (
<figureCitation id="1303C7DBA243FFA6FDC9FA32FD163DDA" box="[610,703,1470,1496]" captionStart="FIGURE 2" captionStartId="5.[151,250,1627,1651]" captionTargetBox="[152,1418,183,1602]" captionTargetId="figure-17@5.[151,1436,181,1604]" captionTargetPageId="5" captionText="FIGURE 2. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="6" pageNumber="559">Fig. 2M</figureCitation>
) biramous, approximately twice length of first; exopod narrowly ovoid, not reaching end of endopod when flexed against it, margins bearing long setae; endopod broader than exopod, lateral and distal margins with long setae, mesial margin with long narrow appendix interna bearing minute terminal hooks and shorter robust appendix masculina bearing long distal setae. Third (
<figureCitation id="1303C7DBA243FFA6FBDCF9A6FB793E46" box="[1143,1232,1578,1604]" captionStart="FIGURE 2" captionStartId="5.[151,250,1627,1651]" captionTargetBox="[152,1418,183,1602]" captionTargetId="figure-17@5.[151,1436,181,1604]" captionTargetPageId="5" captionText="FIGURE 2. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, right maxilla, external surface, setae not shown; B, right first maxilliped, external surface, setae not shown; C, right second maxilliped, internal surface; D, right third maxilliped, external surface; E, same, internal surface, setae not shown; F, major (right) cheliped, external surface; G, minor (left) cheliped, external surface; H, left second pereopod, external surface; I, left third pereopod, external surface; J, right fourth pereopod, terminal articles, external surface; K, right fifth pereopod, terminal articles, external surface; L, left first pleopod, lateral surface; M, right second pleopod, posterior surface; N, endopod of right third pleopod, anterior surface, setae not shown. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7929717" httpUri="https://zenodo.org/record/7929717/files/figure.png" pageId="6" pageNumber="559">Fig. 2N</figureCitation>
) to fifth pleopods larger than second, forming biramous, posteriorly cupped fans, endopod of each an elongate ovoid with elongate digitiform appendix interna projecting distinctly from mesial margin, opposed surfaces on appendix internae of two sides each with small field of microscopic hook setae.
</paragraph>
<paragraph id="8B87DB5EA243FFA6FF6CF936FF583F67" blockId="6.[151,1437,1398,2037]" pageId="6" pageNumber="559">
Telson (
<figureCitation id="1303C7DBA243FFA6FE88F936FE003ED7" box="[291,425,1722,1749]" captionStart="FIGURE 1" captionStartId="4.[151,250,1756,1780]" captionTargetBox="[160,1424,182,1723]" captionTargetId="figure-29@4.[151,1436,181,1732]" captionTargetPageId="4" captionText="FIGURE 1. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, carapace front, eyestalks, antennular and antennal peduncles, dorsal surface; B, same, right lateral surface; C, right mandible, external surface; D, same, internal surface; E, right maxillule, external surface; F, posterior carapace and pleomeres 16, dorsal surface; G, sixth pleomere, telson, and uropods, right lateral surface; H, same, dorsal surface. Scale bars = 1 mm (AE, G, H); 3 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7929711" httpUri="https://zenodo.org/record/7929711/files/figure.png" pageId="6" pageNumber="559">Figs 1G, H</figureCitation>
;
<figureCitation id="1303C7DBA243FFA6FE12F936FE563ED6" box="[441,511,1722,1748]" captionStart="FIGURE 3" captionStartId="6.[151,250,1148,1172]" captionTargetBox="[151,1436,181,1124]" captionTargetId="figure-29@6.[151,1436,181,1124]" captionTargetPageId="6" captionText="FIGURE 3. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304): A, right lateral surface; B, dorsal surface. Paragourretia biffari (Blanco Rambla &amp; Liñero Arana, 1994), male, cl 10.6 mm, northwestern Gulf of Mexico, USNM 1540482 (ULLZ 4673): C, dorsal surface. P. biffari, female cl 6.0 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541735 (ULLZ 7305): D, dorsal surface. P. biffari, female holotype, cl 17.3 mm, Venezuela, USNM 259410: E, major (left) cheliped, internal surface; F, minor (right) cheliped, internal surface; G, posterior pleon and uropods, right ventrolateral, arrow indicating angular submarginal keel." figureDoi="http://doi.org/10.5281/zenodo.7929719" httpUri="https://zenodo.org/record/7929719/files/figure.png" pageId="6" pageNumber="559">3A, B</figureCitation>
) elongate subhexagonal, narrowing posteriorly, length slightly exceeding width, broadest where lateral margins protrude as weak lobes in anterior 1/3, margins beyond converging posteriorly, dorsally with each posterolateral region bearing 2 submarginal tufts of long setae, posterior margin truncate, lacking median spine, posterolateral corners each with short track of long setae, shorter setae along margin between corners.
</paragraph>
<paragraph id="8B87DB5EA243FFA7FF6CF8E2FCE238FB" blockId="6.[151,1437,1398,2037]" lastBlockId="7.[151,1437,151,1941]" lastPageId="7" lastPageNumber="560" pageId="6" pageNumber="559">
Uropods (
<figureCitation id="1303C7DBA243FFA6FE92F8E2FE173F8B" box="[313,446,1902,1929]" captionStart="FIGURE 1" captionStartId="4.[151,250,1756,1780]" captionTargetBox="[160,1424,182,1723]" captionTargetId="figure-29@4.[151,1436,181,1732]" captionTargetPageId="4" captionText="FIGURE 1. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, carapace front, eyestalks, antennular and antennal peduncles, dorsal surface; B, same, right lateral surface; C, right mandible, external surface; D, same, internal surface; E, right maxillule, external surface; F, posterior carapace and pleomeres 16, dorsal surface; G, sixth pleomere, telson, and uropods, right lateral surface; H, same, dorsal surface. Scale bars = 1 mm (AE, G, H); 3 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7929711" httpUri="https://zenodo.org/record/7929711/files/figure.png" pageId="6" pageNumber="559">Figs 1G, H</figureCitation>
;
<figureCitation id="1303C7DBA243FFA6FE67F8E2FDB83F8A" box="[460,529,1902,1928]" captionStart="FIGURE 3" captionStartId="6.[151,250,1148,1172]" captionTargetBox="[151,1436,181,1124]" captionTargetId="figure-29@6.[151,1436,181,1124]" captionTargetPageId="6" captionText="FIGURE 3. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304): A, right lateral surface; B, dorsal surface. Paragourretia biffari (Blanco Rambla &amp; Liñero Arana, 1994), male, cl 10.6 mm, northwestern Gulf of Mexico, USNM 1540482 (ULLZ 4673): C, dorsal surface. P. biffari, female cl 6.0 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541735 (ULLZ 7305): D, dorsal surface. P. biffari, female holotype, cl 17.3 mm, Venezuela, USNM 259410: E, major (left) cheliped, internal surface; F, minor (right) cheliped, internal surface; G, posterior pleon and uropods, right ventrolateral, arrow indicating angular submarginal keel." figureDoi="http://doi.org/10.5281/zenodo.7929719" httpUri="https://zenodo.org/record/7929719/files/figure.png" pageId="6" pageNumber="559">3A, B</figureCitation>
) with endopod broad, ovoid, about 1.5 times longer than broad, dorsal surface with weak longitudinal ridge, sparse submarginal setation distally, and single tuft of several stiffer longer setae posterolaterally, posterior margin with continuous fringe of long setae; exopod broadly ovoid, length approximately 1.3 times breadth, anterodorsal plate obscure to obsolescent, pair of weak, arcuate longitudinal ridges diverging distally from common basal origin, anterolateral of two ridges producing slight vertical offset where intersecting lateral margin (obscure in lateral view), margin lacking distinct notch or incision, anterolateral margin weakly serrate, posterior margin with continuous fringe of long setae.
</paragraph>
<paragraph id="8B87DB5EA242FFA7FF6CFE8EFA2D3943" blockId="7.[151,1437,151,1941]" pageId="7" pageNumber="560">
<emphasis id="B94C074CA242FFA7FF6CFE8EFEB9391E" bold="true" box="[199,272,258,284]" pageId="7" pageNumber="560">Color.</emphasis>
Primarily translucent pale yellowish integument (
<figureCitation id="1303C7DBA242FFA7FC9AFE8FFC0F391F" box="[817,934,259,285]" captionStart="FIGURE 3" captionStartId="6.[151,250,1148,1172]" captionTargetBox="[151,1436,181,1124]" captionTargetId="figure-29@6.[151,1436,181,1124]" captionTargetPageId="6" captionText="FIGURE 3. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304): A, right lateral surface; B, dorsal surface. Paragourretia biffari (Blanco Rambla &amp; Liñero Arana, 1994), male, cl 10.6 mm, northwestern Gulf of Mexico, USNM 1540482 (ULLZ 4673): C, dorsal surface. P. biffari, female cl 6.0 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541735 (ULLZ 7305): D, dorsal surface. P. biffari, female holotype, cl 17.3 mm, Venezuela, USNM 259410: E, major (left) cheliped, internal surface; F, minor (right) cheliped, internal surface; G, posterior pleon and uropods, right ventrolateral, arrow indicating angular submarginal keel." figureDoi="http://doi.org/10.5281/zenodo.7929719" httpUri="https://zenodo.org/record/7929719/files/figure.png" pageId="7" pageNumber="560">Fig. 3A, B</figureCitation>
), more opaque portions of chelipeds and other pereopods whitish, especially at thickened articulations, traces of pinkish violet proximal to fingers of chelipeds.
</paragraph>
<paragraph id="8B87DB5EA242FFA7FF6CFEC6FA29398B" blockId="7.[151,1437,151,1941]" pageId="7" pageNumber="560">
<emphasis id="B94C074CA242FFA7FF6CFEC6FE5F3966" bold="true" box="[199,502,330,357]" pageId="7" pageNumber="560">Habitat and Distribution.</emphasis>
Known from only 53 m depth on the Campeche Banks of the western
<collectingRegion id="49FC15BCA242FFA7FB54FEC7FAF03967" box="[1279,1369,331,357]" country="Mexico" name="Yucatan" pageId="7" pageNumber="560">Yucatan</collectingRegion>
shelf,
<location id="8EE78D85A242FFA7FF3CFEE3FECC398B" LSID="urn:lsid:plazi:treatment:03916A48A244FFA7FF3CF94BFBF83F97:8EE78D85A242FFA7FF3CFEE3FECC398B" box="[151,357,367,393]" municipality="Banks" name="southwestern Gulf" pageId="7" pageNumber="560" stateProvince="Campeche">southwestern Gulf</location>
of
<collectingRegion id="49FC15BCA242FFA7FE20FEE3FE4A398B" box="[395,483,367,393]" country="Mexico" name="Mexico" pageId="7" pageNumber="560">Mexico</collectingRegion>
, in muddy calcareous rubble comprised in part of dead coral and molluscan shells.
</paragraph>
</subSubSection>
<subSubSection id="C32288D5A242FFA7FF6CFE1FFCE239D3" pageId="7" pageNumber="560" type="etymology">
<paragraph id="8B87DB5EA242FFA7FF6CFE1FFCE239D3" blockId="7.[151,1437,151,1941]" pageId="7" pageNumber="560">
<emphasis id="B94C074CA242FFA7FF6CFE1FFEE739AF" bold="true" box="[199,334,403,429]" pageId="7" pageNumber="560">Etymology.</emphasis>
The species name “sandrae” honors Sandra Cheryl Collier, for her generous contributions to Smithsonian Institution research efforts in Florida and
<collectingCountry id="F32F9BCEA242FFA7FD57FE3BFCEE39D3" box="[764,839,439,465]" name="Belize" pageId="7" pageNumber="560">Belize</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32288D5A242FFA7FF6CFE57FBF83F97" pageId="7" pageNumber="560" type="discussion">
<paragraph id="8B87DB5EA242FFA7FF6CFE57FC9E3B83" blockId="7.[151,1437,151,1941]" pageId="7" pageNumber="560">
<emphasis id="B94C074CA242FFA7FF6CFE57FE9239F7" bold="true" box="[199,315,475,501]" pageId="7" pageNumber="560">Remarks.</emphasis>
In accord with recent revisions (
<bibRefCitation id="EFA9A6AFA242FFA7FD04FE57FCDE39F7" author="Poore, G. C. B. &amp; Dworschak, P. C. &amp; Robles, R. &amp; Mantelatto, F. &amp; Felder, D. L." box="[687,887,475,501]" pageId="7" pageNumber="560" pagination="73 - 146" refId="ref6503" refString="Poore, G. C. B., Dworschak, P. C., Robles, R., Mantelatto, F. &amp; Felder, D. L. (2019) A new classification of families and genera of Callianassoidea (Crustacea: Decapoda: Axiidea) derived from a molecular phylogeny with morphological support. Memoirs of Museum Victoria, 78, 73 - 146. https: // doi. org / 10.24199 / j. mmv. 2019.78.05" type="journal article" year="2019">
Poore
<emphasis id="B94C074CA242FFA7FD51FE57FC9A39F7" box="[762,819,475,501]" italics="true" pageId="7" pageNumber="560">et al.</emphasis>
2019
</bibRefCitation>
),
<taxonomicName id="4C38A0DDA242FFA7FC20FE57FB8539F6" authorityName="Sakai" authorityYear="2004" box="[907,1068,475,500]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B94C074CA242FFA7FC20FE57FB8539F6" box="[907,1068,475,500]" italics="true" pageId="7" pageNumber="560">Paragourretia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C38A0DDA242FFA7FBCFFE57FF663A1B" authority="de Saint Laurent, 1973" authorityName="de Saint Laurent" authorityYear="1973" class="Malacostraca" family="Ctenochelidae" genus="Gourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B94C074CA242FFA7FBCFFE57FB7A39F7" box="[1124,1235,475,501]" italics="true" pageId="7" pageNumber="560">Gourretia</emphasis>
de
<bibRefCitation id="EFA9A6AFA242FFA7FB56FE57FF663A1B" author="Saint Laurent, M. de" pageId="7" pageNumber="560" pagination="513 - 516" refId="ref6940" refString="Saint Laurent, M. de (1973) Sur la systematique et la phylogenie des Thalassinidea: definition des familles des Callianassidae et des Upogebiidae et diagnose de cinq genres noveaux (Crustacea Decapoda). Comptes Rendus Hebdomadaires des Seances de l'Academie des Sciences, Paris, 277 D, 513 - 516. [http: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 5474901 n / f 589. image]" type="journal article" year="1973">Saint Laurent, 1973</bibRefCitation>
</taxonomicName>
are separated from
<taxonomicName id="4C38A0DDA242FFA7FE18FE73FD863A1A" authorityName="Manning &amp; Felder" authorityYear="1991" box="[435,559,511,536]" class="Malacostraca" family="Ctenochelidae" genus="Dawsonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B94C074CA242FFA7FE18FE73FD863A1A" box="[435,559,511,536]" italics="true" pageId="7" pageNumber="560">Dawsonius</emphasis>
</taxonomicName>
, the only other ctenochelid genus known to occur in the western Atlantic, by the presence of an exopod on the third maxilliped. Species of these genera also differ from
<taxonomicName id="4C38A0DDA242FFA7FB0CFDAFFA8A3A3E" authorityName="Manning &amp; Felder" authorityYear="1991" box="[1191,1315,547,572]" class="Malacostraca" family="Ctenochelidae" genus="Dawsonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B94C074CA242FFA7FB0CFDAFFA8A3A3E" box="[1191,1315,547,572]" italics="true" pageId="7" pageNumber="560">Dawsonius</emphasis>
</taxonomicName>
in lacking a lateral projection or keel in the anterior half of the sixth pleomere, except for
<taxonomicName id="4C38A0DDA242FFA7FB83FDCBFAB13A63" baseAuthorityName="Blanco Rambla &amp; Linero Arana" baseAuthorityYear="1994" box="[1064,1304,583,609]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="biffari">
<emphasis id="B94C074CA242FFA7FB83FDCBFAB13A63" box="[1064,1304,583,609]" italics="true" pageId="7" pageNumber="560">Paragourretia biffari</emphasis>
</taxonomicName>
in which it occurs in a somewhat ventrolateral position (
<figureCitation id="1303C7DBA242FFA7FD3AFDE7FD463A87" box="[657,751,619,645]" captionStart="FIGURE 3" captionStartId="6.[151,250,1148,1172]" captionTargetBox="[151,1436,181,1124]" captionTargetId="figure-29@6.[151,1436,181,1124]" captionTargetPageId="6" captionText="FIGURE 3. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304): A, right lateral surface; B, dorsal surface. Paragourretia biffari (Blanco Rambla &amp; Liñero Arana, 1994), male, cl 10.6 mm, northwestern Gulf of Mexico, USNM 1540482 (ULLZ 4673): C, dorsal surface. P. biffari, female cl 6.0 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541735 (ULLZ 7305): D, dorsal surface. P. biffari, female holotype, cl 17.3 mm, Venezuela, USNM 259410: E, major (left) cheliped, internal surface; F, minor (right) cheliped, internal surface; G, posterior pleon and uropods, right ventrolateral, arrow indicating angular submarginal keel." figureDoi="http://doi.org/10.5281/zenodo.7929719" httpUri="https://zenodo.org/record/7929719/files/figure.png" pageId="7" pageNumber="560">Figs 3G</figureCitation>
,
<figureCitation id="1303C7DBA242FFA7FD50FDE7FCB43A87" box="[763,797,619,645]" captionStart="FIGURE 4" captionStartId="8.[151,250,982,1006]" captionTargetBox="[152,1434,182,957]" captionTargetId="figure-29@8.[151,1436,181,958]" captionTargetPageId="8" captionText="FIGURE 4. Paragourretia biffari (Blanco Rambla &amp; Liñero Arana, 1994), male, cl 10.6 mm, northwestern Gulf of Mexico, USNM 1540482 (ULLZ 4673). A, carapace front, eyestalks, and anterior appendages, dorsal surface; B, right third maxilliped, external surface; C, same, internal surface; D, major (right) cheliped, external surface; E, minor (left) cheliped, external surface; F, sixth pleomere, telson, and uropods, right lateral surface; G, same, dorsal surface; H, left first pleopod, lateral surface. Scale bars = 3 mm (AG); 1 mm (H)." figureDoi="http://doi.org/10.5281/zenodo.7929721" httpUri="https://zenodo.org/record/7929721/files/figure.png" pageId="7" pageNumber="560">4F</figureCitation>
). All members of
<taxonomicName id="4C38A0DDA242FFA7FC43FDE7FB203A86" authorityName="Sakai" authorityYear="2004" box="[1000,1161,619,644]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B94C074CA242FFA7FC43FDE7FB203A86" box="[1000,1161,619,644]" italics="true" pageId="7" pageNumber="560">Paragourretia</emphasis>
</taxonomicName>
can in turn be separated from
<taxonomicName id="4C38A0DDA242FFA7FF73FD03FEEE3AAB" authorityName="de Saint Laurent" authorityYear="1973" box="[216,327,655,681]" class="Malacostraca" family="Ctenochelidae" genus="Gourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B94C074CA242FFA7FF73FD03FEEE3AAB" box="[216,327,655,681]" italics="true" pageId="7" pageNumber="560">Gourretia</emphasis>
</taxonomicName>
by their lacking sharp proximally directed teeth along the cutting edges of the minor cheliped fingers characteristic of that genus (
<bibRefCitation id="EFA9A6AFA242FFA7FD88FD3FFD4C3ACF" author="Poore, G. C. B. &amp; Dworschak, P. C. &amp; Robles, R. &amp; Mantelatto, F. &amp; Felder, D. L." box="[547,741,691,717]" pageId="7" pageNumber="560" pagination="73 - 146" refId="ref6503" refString="Poore, G. C. B., Dworschak, P. C., Robles, R., Mantelatto, F. &amp; Felder, D. L. (2019) A new classification of families and genera of Callianassoidea (Crustacea: Decapoda: Axiidea) derived from a molecular phylogeny with morphological support. Memoirs of Museum Victoria, 78, 73 - 146. https: // doi. org / 10.24199 / j. mmv. 2019.78.05" type="journal article" year="2019">
Poore
<emphasis id="B94C074CA242FFA7FDC7FD3FFD353ACF" box="[620,668,691,717]" italics="true" pageId="7" pageNumber="560">et al</emphasis>
. 2019
</bibRefCitation>
: fig. 17i). As in most other members of the genus, the carapace in
<taxonomicName id="4C38A0DDA242FFA7FF1FFD5BFE833AF3" authority="Felder &amp; Dworschak, 2023" authorityName="Felder &amp; Dworschak" authorityYear="2023" box="[180,298,727,753]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="sandrae" status="sp. nov.">
<emphasis id="B94C074CA242FFA7FF1FFD5BFE833AF3" box="[180,298,727,753]" italics="true" pageId="7" pageNumber="560">P. sandrae</emphasis>
</taxonomicName>
<emphasis id="B94C074CA242FFA7FE99FD5BFED83AF3" bold="true" box="[306,369,727,753]" pageId="7" pageNumber="560">
<taxonomicNameLabel id="A27FBA37A242FFA7FE99FD5BFED83AF3" box="[306,369,727,753]" pageId="7" pageNumber="560" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
appears to have a dorsal cardiac prominence of its carapace, albeit weakly expressed. As in its congeners, the antennular peduncle is both heavier and shorter than the antennal peduncle, and the antenna bears a short bladelike dorsal scaphocerite where its second article joins to the third. Typical of the genus, its second maxilliped is comparatively small, with a straplike exopod carried against the internal surface of the slightly longer endopodal merus, and a small epipod originating at its base.
</paragraph>
<paragraph id="8B87DB5EA242FFA7FF6CFC07FDA53D0F" blockId="7.[151,1437,151,1941]" pageId="7" pageNumber="560">
The description of
<taxonomicName id="4C38A0DDA242FFA7FE37FC07FDB83BA7" authority="Felder &amp; Dworschak, 2023" authorityName="Felder &amp; Dworschak" authorityYear="2023" box="[412,529,907,933]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="sandrae" status="sp. nov.">
<emphasis id="B94C074CA242FFA7FE37FC07FDB83BA7" box="[412,529,907,933]" italics="true" pageId="7" pageNumber="560">P. sandrae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27FBA37A242FFA7FDB3FC07FDFE3BA7" box="[536,599,907,933]" pageId="7" pageNumber="560" rank="species">
<emphasis id="B94C074CA242FFA7FDB3FC07FDF93BA7" bold="true" box="[536,592,907,933]" pageId="7" pageNumber="560">n. sp</emphasis>
.
</taxonomicNameLabel>
brings known world membership of the genus
<taxonomicName id="4C38A0DDA242FFA7FBC3FC07FAA03BA6" authorityName="Sakai" authorityYear="2004" box="[1128,1289,907,932]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B94C074CA242FFA7FBC3FC07FAA03BA6" box="[1128,1289,907,932]" italics="true" pageId="7" pageNumber="560">Paragourretia</emphasis>
</taxonomicName>
to ten, six of which are found only in the Indo-West Pacific. Among the known species, the majority differ from
<taxonomicName id="4C38A0DDA242FFA7FAACFC23FAD43BCB" authority="Felder &amp; Dworschak, 2023" authorityName="Felder &amp; Dworschak" authorityYear="2023" box="[1287,1405,943,969]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="sandrae" status="sp. nov.">
<emphasis id="B94C074CA242FFA7FAACFC23FAD43BCB" box="[1287,1405,943,969]" italics="true" pageId="7" pageNumber="560">P. sandrae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27FBA37A242FFA7FA2DFC23FF103BEF" pageId="7" pageNumber="560" rank="species">
<emphasis id="B94C074CA242FFA7FA2DFC23FF183BEF" bold="true" pageId="7" pageNumber="560">n. sp</emphasis>
.
</taxonomicNameLabel>
and resemble
<taxonomicName id="4C38A0DDA242FFA7FEF0FC5FFE143BEF" baseAuthorityName="Blanco Rambla &amp; Linero Arana" baseAuthorityYear="1994" box="[347,445,979,1005]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="biffari">
<emphasis id="B94C074CA242FFA7FEF0FC5FFE143BEF" box="[347,445,979,1005]" italics="true" pageId="7" pageNumber="560">P. biffari</emphasis>
</taxonomicName>
in having a distinct marginal notch or indentation in the margin of the uropodal exopod. From illustrations accompanying descriptions, this feature appears to be conspicuous in at least
<taxonomicName id="4C38A0DDA242FFA7FB51FC7BFE993C37" authority="(Sakai, 2002)" baseAuthorityName="Sakai" baseAuthorityYear="2002" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="aungtonyae">
<emphasis id="B94C074CA242FFA7FB51FC7BFA343C12" box="[1274,1437,1015,1040]" italics="true" pageId="7" pageNumber="560">P. aungtonyae</emphasis>
(
<bibRefCitation id="EFA9A6AFA242FFA7FF34FB97FE813C37" author="Sakai, K." box="[159,296,1051,1077]" pageId="7" pageNumber="560" pagination="461 - 532" refId="ref7059" refString="Sakai, K. (2002) Callianassidae (Decapoda, Thalassinidea) in the Andaman Sea, Thailand. Phuket Marine Biological Center Special Publication, 23, 461 - 532." type="journal article" year="2002">Sakai, 2002</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C38A0DDA242FFA7FE91FB97FD6E3C37" authority="(Poore &amp; Griffin, 1979)" baseAuthorityName="Poore &amp; Griffin" baseAuthorityYear="1979" box="[314,711,1051,1077]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="coolibah">
<emphasis id="B94C074CA242FFA7FE91FB97FE1F3C37" box="[314,438,1051,1077]" italics="true" pageId="7" pageNumber="560">P. coolibah</emphasis>
(
<bibRefCitation id="EFA9A6AFA242FFA7FE6FFB97FD163C37" author="Poore, G. C. B. &amp; Griffin, D. J. G." box="[452,703,1051,1077]" pageId="7" pageNumber="560" pagination="217 - 321" refId="ref6588" refString="Poore, G. C. B. &amp; Griffin, D. J. G. (1979) The Thalassinidea (Crustacea: Decapoda) of Australia. Records of the Australian Museum, 32, 217 - 321. https: // doi. org / 10.3853 / j. 0067 - 1975.32.1979.457" type="journal article" year="1979">Poore &amp; Griffin, 1979</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C38A0DDA242FFA7FD7AFB97FC453C37" authority="(Sakai, 2017)" baseAuthorityName="Sakai" baseAuthorityYear="2017" box="[721,1004,1051,1077]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="galathea">
<emphasis id="B94C074CA242FFA7FD7AFB97FCE73C37" box="[721,846,1051,1077]" italics="true" pageId="7" pageNumber="560">P. galathea</emphasis>
(
<bibRefCitation id="EFA9A6AFA242FFA7FCF7FB97FC4D3C37" author="Sakai, K." box="[860,996,1051,1077]" pageId="7" pageNumber="560" pagination="177 - 197" refId="ref7169" refString="Sakai, K. (2017) Descriptions of eight species from the superfamilies Axioidea Huxley, 1879 and Callianassoidea Dana, 1852, with a revised key to the species of the genus Acanthaxius Sakai &amp; de Saint Laurent, 1989 (Decapoda, Callianassidea). Crustaceana, 90, 177 - 197. https: // doi. org / 10.1163 / 15685403 - 00003621" type="journal article" year="2017">Sakai, 2017</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="4C38A0DDA242FFA7FB8EFB97FA303C37" authority="(Liu &amp; Liu, 2010)" baseAuthorityName="Liu &amp; Liu" baseAuthorityYear="2010" box="[1061,1433,1051,1077]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="laevidactyla">
<emphasis id="B94C074CA242FFA7FB8EFB97FB613C37" box="[1061,1224,1051,1077]" italics="true" pageId="7" pageNumber="560">P. laevidactyla</emphasis>
(
<bibRefCitation id="EFA9A6AFA242FFA7FB7DFB97FA383C37" author="Liu, W. L. &amp; Liu, R. Y." box="[1238,1425,1051,1077]" pageId="7" pageNumber="560" pagination="745 - 756" refId="ref6305" refString="Liu, W. L. &amp; Liu, R. Y. (2010) Two new species of the axiidean genus Gourretia de Saint Laurent, 1973 (Decapoda: Ctenochelidae) from the South China Sea. Journal of Crustacean Biology, 30, 745 - 756. https: // doi. org / 10.1651 / 10 - 3282.1" type="journal article" year="2010">Liu &amp; Liu, 2010</bibRefCitation>
)
</taxonomicName>
. However, it appears to be weakly evident in
<taxonomicName id="4C38A0DDA242FFA7FD09FBB3FC8D3C5A" baseAuthorityName="Ngoc-Ho" baseAuthorityYear="1991" box="[674,804,1087,1112]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="crosnieri">
<emphasis id="B94C074CA242FFA7FD09FBB3FC8D3C5A" box="[674,804,1087,1112]" italics="true" pageId="7" pageNumber="560">P. crosnieri</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFA9A6AFA242FFA7FC9DFBB3FC433C5B" author="Ngoc-Ho, N." box="[822,1002,1087,1113]" pageId="7" pageNumber="560" pagination="281 - 311" refId="ref6425" refString="Ngoc-Ho, N. (1991) Sur quelques Callianassidae et Upogebiidae de Nouvelle-Caledonie (Crustacea, Thalassinidea). In: Richer de Forges, B. (Ed.), Le benthos des fonds meubles des lagons de Nouvelle-Caledonie. ORSTOM Editions, Paris, pp. 281 - 311, figs. 281 - 211. [https: // www. documentation. ird. fr / hor / fdi: 35618]" type="book chapter" year="1991">Ngoc-Ho, 1991</bibRefCitation>
) and not evident in illustrations of
<taxonomicName id="4C38A0DDA242FFA7FA23FBB3FE1F3C7F" authority="(Sakai, 2002)" baseAuthorityName="Sakai" baseAuthorityYear="2002" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="phuketensis">
<emphasis id="B94C074CA242FFA7FA23FBB3FEB33C7F" italics="true" pageId="7" pageNumber="560">P. phuketensis</emphasis>
(
<bibRefCitation id="EFA9A6AFA242FFA7FE83FBEFFE063C7F" author="Sakai, K." box="[296,431,1123,1149]" pageId="7" pageNumber="560" pagination="461 - 532" refId="ref7059" refString="Sakai, K. (2002) Callianassidae (Decapoda, Thalassinidea) in the Andaman Sea, Thailand. Phuket Marine Biological Center Special Publication, 23, 461 - 532." type="journal article" year="2002">Sakai, 2002</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C38A0DDA242FFA7FE41FBEFFC0F3C7F" authority="(Le Loeuff &amp; Intes, 1974)" baseAuthorityName="Le Loeuff &amp; Intes" baseAuthorityYear="1974" box="[490,934,1123,1149]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="lahouensis">
<emphasis id="B94C074CA242FFA7FE41FBEFFDD43C7F" box="[490,637,1123,1149]" italics="true" pageId="7" pageNumber="560">P. lahouensis</emphasis>
(
<bibRefCitation id="EFA9A6AFA242FFA7FD20FBEFFC343C7F" author="Le Loeuff, P. &amp; Intes, A." box="[651,925,1123,1149]" pageId="7" pageNumber="560" pagination="17 - 69" refId="ref6240" refString="Le Loeuff, P. &amp; Intes, A. (1974) Les Thalassinidea (Crustacea, Decapoda) du Golfe de Guinee systematique - ecologie. Cahiers de l'Office de Recherches Scientifiques et Techniques Outre-Mer, serie Oceanographique, 12, 17 - 69. [http: // www. documentation. ird. fr / hor / fdi: 19672]" type="journal article" year="1974">Le Loeuff &amp; Intès, 1974</bibRefCitation>
)
</taxonomicName>
. In
<taxonomicName id="4C38A0DDA242FFA7FC66FBEFFBEB3C7F" authority="Felder &amp; Dworschak, 2023" authorityName="Felder &amp; Dworschak" authorityYear="2023" box="[973,1090,1123,1149]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="sandrae" status="sp. nov.">
<emphasis id="B94C074CA242FFA7FC66FBEFFBEB3C7F" box="[973,1090,1123,1149]" italics="true" pageId="7" pageNumber="560">P. sandrae</emphasis>
</taxonomicName>
<emphasis id="B94C074CA242FFA7FBE3FBEFFB2F3C7F" bold="true" box="[1096,1158,1123,1149]" pageId="7" pageNumber="560">
<taxonomicNameLabel id="A27FBA37A242FFA7FBE3FBEFFB2F3C7F" box="[1096,1158,1123,1149]" pageId="7" pageNumber="560" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, the anterolateral margin of the uropodal exopod is intersected by the anteriormost of two dorsal longitudinal ridges producing a very slight vertical offset in the margin, most evident when viewed obliquely under magnification. The ridge itself appears to demarcate the edge of a very slightly offset dorsal plate on the exopod that is found in most members of the genus (
<bibRefCitation id="EFA9A6AFA242FFA7FF34FB7FFE333D0F" author="Poore, G. C. B. &amp; Dworschak, P. C. &amp; Robles, R. &amp; Mantelatto, F. &amp; Felder, D. L." box="[159,410,1267,1293]" pageId="7" pageNumber="560" pagination="73 - 146" refId="ref6503" refString="Poore, G. C. B., Dworschak, P. C., Robles, R., Mantelatto, F. &amp; Felder, D. L. (2019) A new classification of families and genera of Callianassoidea (Crustacea: Decapoda: Axiidea) derived from a molecular phylogeny with morphological support. Memoirs of Museum Victoria, 78, 73 - 146. https: // doi. org / 10.24199 / j. mmv. 2019.78.05" type="journal article" year="2019">
Poore
<emphasis id="B94C074CA242FFA7FF42FB7FFEB33D0F" box="[233,282,1267,1293]" italics="true" pageId="7" pageNumber="560">et al</emphasis>
. 2019: 118
</bibRefCitation>
, fig. 16e).
</paragraph>
<paragraph id="8B87DB5EA242FFA7FF6CFA9BFBF83F97" blockId="7.[151,1437,151,1941]" pageId="7" pageNumber="560">
The absence of an obvious marginal notch in the uropodal exopod of
<taxonomicName id="4C38A0DDA242FFA7FC4FFA9BFBF23D33" authorityName="Felder &amp; Dworschak" authorityYear="2023" box="[996,1115,1303,1329]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="sandrae">
<emphasis id="B94C074CA242FFA7FC4FFA9BFBF23D33" box="[996,1115,1303,1329]" italics="true" pageId="7" pageNumber="560">P. sandrae</emphasis>
</taxonomicName>
<emphasis id="B94C074CA242FFA7FBCFFA9BFB343D33" bold="true" box="[1124,1181,1303,1329]" pageId="7" pageNumber="560">n. sp</emphasis>
, a feature shared with regional ctenochelids of the genera
<taxonomicName id="4C38A0DDA242FFA7FD98FAB7FD063D56" authorityName="Manning &amp; Felder" authorityYear="1991" box="[563,687,1339,1364]" class="Malacostraca" family="Ctenochelidae" genus="Dawsonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B94C074CA242FFA7FD98FAB7FD063D56" box="[563,687,1339,1364]" italics="true" pageId="7" pageNumber="560">Dawsonius</emphasis>
</taxonomicName>
and
<taxonomicName id="4C38A0DDA242FFA7FD40FAB7FCF33D57" authorityName="de Saint Laurent" authorityYear="1973" box="[747,858,1339,1365]" class="Malacostraca" family="Ctenochelidae" genus="Gourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="genus">
<emphasis id="B94C074CA242FFA7FD40FAB7FCF33D57" box="[747,858,1339,1365]" italics="true" pageId="7" pageNumber="560">Gourretia</emphasis>
</taxonomicName>
, readily separates the new species from
<taxonomicName id="4C38A0DDA242FFA7FA9AFAB7FA3C3D57" baseAuthorityName="Blanco Rambla &amp; Linero Arana" baseAuthorityYear="1994" box="[1329,1429,1339,1365]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="biffari">
<emphasis id="B94C074CA242FFA7FA9AFAB7FA3C3D57" box="[1329,1429,1339,1365]" italics="true" pageId="7" pageNumber="560">P. biffari</emphasis>
</taxonomicName>
. Like
<taxonomicName id="4C38A0DDA242FFA7FF78FAD3FE133D7B" baseAuthorityName="Dawson" baseAuthorityYear="1967" box="[211,442,1375,1401]" class="Malacostraca" family="Ctenochelidae" genus="Dawsonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="latispina">
<emphasis id="B94C074CA242FFA7FF78FAD3FE133D7B" box="[211,442,1375,1401]" italics="true" pageId="7" pageNumber="560">Dawsonius latispina</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFA9A6AFA242FFA7FE53FAD3FD763D7B" author="Rabalais, N. N. &amp; Holt, S. A. &amp; Flint, R. W." box="[504,735,1375,1401]" pageId="7" pageNumber="560" pagination="96 - 115" refId="ref6699" refString="Rabalais, N. N., Holt, S. A. &amp; Flint, R. W. (1981) Mud shrimps (Crustacea, Decapoda, Thalassinidea) of the northwestern Gulf of Mexico. Bulletin of Marine Science, 31, 96 - 115." type="journal article" year="1981">
Rabalais
<emphasis id="B94C074CA242FFA7FDCAFAD3FD323D7B" box="[609,667,1375,1401]" italics="true" pageId="7" pageNumber="560">et al.</emphasis>
1981
</bibRefCitation>
: fig. 3I),
<taxonomicName id="4C38A0DDA242FFA7FCE0FAD3FC6B3D7B" authority="Felder &amp; Dworschak, 2023" authorityName="Felder &amp; Dworschak" authorityYear="2023" box="[843,962,1375,1401]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="sandrae" status="sp. nov.">
<emphasis id="B94C074CA242FFA7FCE0FAD3FC6B3D7B" box="[843,962,1375,1401]" italics="true" pageId="7" pageNumber="560">P. sandrae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27FBA37A242FFA7FC61FAD3FBA23D7B" box="[970,1035,1375,1401]" pageId="7" pageNumber="560" rank="species">
<emphasis id="B94C074CA242FFA7FC61FAD3FBAD3D7B" bold="true" box="[970,1028,1375,1401]" pageId="7" pageNumber="560">n. sp</emphasis>
.
</taxonomicNameLabel>
also lacks a median terminal spine on the telson (
<figureCitation id="1303C7DBA242FFA7FE9EFA0FFE253D9F" box="[309,396,1411,1437]" captionStart="FIGURE 1" captionStartId="4.[151,250,1756,1780]" captionTargetBox="[160,1424,182,1723]" captionTargetId="figure-29@4.[151,1436,181,1732]" captionTargetPageId="4" captionText="FIGURE 1. Paragourretia sandrae n. sp., male holotype, cl 4.4 mm, Campeche Banks, southwestern Gulf of Mexico, USNM 1541885 (= ULLZ 7304). A, carapace front, eyestalks, antennular and antennal peduncles, dorsal surface; B, same, right lateral surface; C, right mandible, external surface; D, same, internal surface; E, right maxillule, external surface; F, posterior carapace and pleomeres 16, dorsal surface; G, sixth pleomere, telson, and uropods, right lateral surface; H, same, dorsal surface. Scale bars = 1 mm (AE, G, H); 3 mm (F)." figureDoi="http://doi.org/10.5281/zenodo.7929711" httpUri="https://zenodo.org/record/7929711/files/figure.png" pageId="7" pageNumber="560">Fig. 1H</figureCitation>
), a character now documented to occur in
<taxonomicName id="4C38A0DDA242FFA7FCCCFA0FFC603D9F" baseAuthorityName="Blanco Rambla &amp; Linero Arana" baseAuthorityYear="1994" box="[871,969,1411,1437]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="biffari">
<emphasis id="B94C074CA242FFA7FCCCFA0FFC603D9F" box="[871,969,1411,1437]" italics="true" pageId="7" pageNumber="560">P. biffari</emphasis>
</taxonomicName>
following re-examination of the
<typeStatus id="548365FCA242FFA7FA91FA0FFA353D9F" box="[1338,1436,1411,1437]" pageId="7" pageNumber="560" type="holotype">holotype</typeStatus>
and study of additional specimens. Specimens from the western Gulf of Mexico and the Caribbean coast of
<collectingCountry id="F32F9BCEA242FFA7FAEFFA2BFA343DC3" box="[1348,1437,1447,1473]" name="Panama" pageId="7" pageNumber="560">Panama</collectingCountry>
(see Materials and methods, above) were assigned to
<taxonomicName id="4C38A0DDA242FFA7FCAEFA47FC5F3DE7" baseAuthorityName="Blanco Rambla &amp; Linero Arana" baseAuthorityYear="1994" box="[773,1014,1483,1509]" class="Malacostraca" family="Ctenochelidae" genus="Paragourretia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="biffari">
<emphasis id="B94C074CA242FFA7FCAEFA47FC5F3DE7" box="[773,1014,1483,1509]" italics="true" pageId="7" pageNumber="560">Paragourretia biffari</emphasis>
</taxonomicName>
only after direct comparisons to the female
<typeStatus id="548365FCA242FFA7FF40FA63FEE43E0B" box="[235,333,1519,1545]" pageId="7" pageNumber="560" type="holotype">holotype</typeStatus>
(USNM 259410). These add to the original report from
<collectingCountry id="F32F9BCEA242FFA7FC67FA63FB963E0B" box="[972,1087,1519,1545]" name="Venezuela" pageId="7" pageNumber="560">Venezuela</collectingCountry>
and
<collectingCountry id="F32F9BCEA242FFA7FBDCFA63FB4E3E0B" box="[1143,1255,1519,1545]" name="Honduras" pageId="7" pageNumber="560">Honduras</collectingCountry>
, suggesting that the species is widely distributed in offshore tropical to subtropical waters of the western Atlantic. Illustrations accompanying the original description (
<bibRefCitation id="EFA9A6AFA242FFA7FDF0F9BBFBA23E53" author="Blanco Rambla, J. P. &amp; Linero Arana, I." box="[603,1035,1591,1617]" pageId="7" pageNumber="560" pagination="16 - 29" refId="ref5558" refString="Blanco Rambla, J. P. &amp; Linero Arana, I. (1994) New records and new species of ghost shrimps (Crustacea: Thalassinidea) from Venezuela. Bulletin of Marine Science, 55, 16 - 29." type="journal article" year="1994">Blanco Rambla &amp; Liñero Arana 1994</bibRefCitation>
) were of limited detail and did not show the now confirmed characteristic posterior median spine on the telson (
<figureCitation id="1303C7DBA242FFA7FC4FF9D7FB953E77" box="[996,1084,1627,1653]" captionStart="FIGURE 4" captionStartId="8.[151,250,982,1006]" captionTargetBox="[152,1434,182,957]" captionTargetId="figure-29@8.[151,1436,181,958]" captionTargetPageId="8" captionText="FIGURE 4. Paragourretia biffari (Blanco Rambla &amp; Liñero Arana, 1994), male, cl 10.6 mm, northwestern Gulf of Mexico, USNM 1540482 (ULLZ 4673). A, carapace front, eyestalks, and anterior appendages, dorsal surface; B, right third maxilliped, external surface; C, same, internal surface; D, major (right) cheliped, external surface; E, minor (left) cheliped, external surface; F, sixth pleomere, telson, and uropods, right lateral surface; G, same, dorsal surface; H, left first pleopod, lateral surface. Scale bars = 3 mm (AG); 1 mm (H)." figureDoi="http://doi.org/10.5281/zenodo.7929721" httpUri="https://zenodo.org/record/7929721/files/figure.png" pageId="7" pageNumber="560">Fig. 4G</figureCitation>
), which was also not mentioned in the accompanying text. The authors did specifically mention the lack of a sharp lateral projection on the sixth pleomere, which, along with the presence of an exopod on the third maxilliped, was postulated to distinguish the species from
<taxonomicName id="4C38A0DDA242FFA7FE81F94BFDA73EE3" baseAuthorityName="Dawson" baseAuthorityYear="1967" box="[298,526,1735,1761]" class="Malacostraca" family="Ctenochelidae" genus="Dawsonius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="560" phylum="Arthropoda" rank="species" species="latispina">
<emphasis id="B94C074CA242FFA7FE81F94BFDA73EE3" box="[298,526,1735,1761]" italics="true" pageId="7" pageNumber="560">Dawsonius latispina</emphasis>
</taxonomicName>
. However, on close study of the
<typeStatus id="548365FCA242FFA7FCDDF94BFC713EE3" box="[886,984,1735,1761]" pageId="7" pageNumber="560" type="holotype">holotype</typeStatus>
this sharp ridge-like projection or keel is present, albeit in a somewhat ventrolateral position, as is also the case in some Gulf of Mexico specimens. Varying in the degree to which it projects laterally, perhaps due to effects of preservation, it is not always evident in dorsal view. Gulf of Mexico and Panamanian materials share this keel, the median telson spine, dentition of the chelae, and other morphology with the Venezuelan
<typeStatus id="548365FCA242FFA7FD2DF8DBFD413F73" box="[646,744,1879,1905]" pageId="7" pageNumber="560" type="holotype">holotype</typeStatus>
of this species. However, the female
<typeStatus id="548365FCA242FFA7FB3AF8DBFB5A3F73" box="[1169,1267,1879,1905]" pageId="7" pageNumber="560" type="holotype">holotype</typeStatus>
is much larger (cl
<quantity id="4CC076BBA242FFA7FF17F8F7FE883F97" box="[188,289,1915,1941]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" pageId="7" pageNumber="560" unit="mm" value="17.3">17.3 mm</quantity>
) than known Gulf of Mexico and Panamanian materials (cl
<quantity id="4CC076BBA242FFA7FC6AF8F6FBEC3F97" box="[961,1093,1914,1941]" metricMagnitude="-3" metricUnit="m" metricValue="9.35" metricValueMax="13.0" metricValueMin="5.7" pageId="7" pageNumber="560" unit="mm" value="9.35" valueMax="13.0" valueMin="5.7">5.713 mm</quantity>
).
</paragraph>
</subSubSection>
</treatment>
</document>