179 lines
17 KiB
XML
179 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. & Tishechkin, Alexey K." docDate="2013" docId="53EB34910D9F2613D49398A9538B6FB5" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus hirsutipes Caterino & Tishechkin, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="202" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="200" updateTime="1668155382655" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Caterino, Michael S.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>271</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>401</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152040961" LSID="urn:lsid:zoobank.org:act:4725B183-B10E-44C4-A3E9-C8651D576EE5" httpUri="http://treatment.plazi.org/id/53EB34910D9F2613D49398A9538B6FB5" lastPageId="201" lastPageNumber="202" pageId="199" pageNumber="200">
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<subSubSection pageId="199" pageNumber="200" type="nomenclature">
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<paragraph pageId="199" pageNumber="200">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:4725B183-B10E-44C4-A3E9-C8651D576EE5" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hirsutipes" order="Coleoptera" pageId="199" pageNumber="200" phylum="Arthropoda" rank="species" species="hirsutipes">Operclipygus hirsutipes</taxonomicName>
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<taxonomicNameLabel pageId="199" pageNumber="200">sp. n.</taxonomicNameLabel>
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Figs 54
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<normalizedToken originalValue="A–C55A–D">A-C55A-D</normalizedToken>
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, GMap 19
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</paragraph>
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</subSubSection>
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<subSubSection pageId="199" pageNumber="200" type="type locality">
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<paragraph pageId="199" pageNumber="200">Type locality.</paragraph>
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<paragraph pageId="199" pageNumber="200">
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ECUADOR: Orellana:
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<normalizedToken originalValue="Yasuní">Yasuni</normalizedToken>
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Research Station [
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<geoCoordinate direction="south" orientation="latitude" precision="92" value="-0.675">0°40.5'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-76.4">76°24'W</geoCoordinate>
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].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="199" pageNumber="200" type="type material">
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<paragraph pageId="199" pageNumber="200">Type material.</paragraph>
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<paragraph pageId="199" pageNumber="200">
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Holotype male: "Ecuador: Napo, mid.Rio Tiputini,
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<normalizedToken originalValue="Yasuní">Yasuni</normalizedToken>
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Res. Stn.
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<geoCoordinate direction="south" orientation="latitude" precision="92" value="-0.675">0°40.5'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-76.4">76°24'W</geoCoordinate>
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, FIT #M1, 5-11 Jul 1999. AKT#084, C.Carlton & A.Tishechkin"/
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<normalizedToken originalValue="“LSAM0013250”">"LSAM0013250"</normalizedToken>
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(FMNH). Paratypes (5): 1: same data as type (LSAM); ECUADOR: Orellana: 1:
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<normalizedToken originalValue="Yasuní">Yasuni</normalizedToken>
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Res. Stn., mid.Rio Tiputini,
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<geoCoordinate direction="south" orientation="latitude" precision="92" value="-0.675">0°40.5'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-76.4">76°24'W</geoCoordinate>
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, 28.
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<normalizedToken originalValue="vi–">vi-</normalizedToken>
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5.vii.1999, FIT, C.E. Carlton & A.K. Tishechkin (LSAM), 1: 26.vii-4.viii.1999, FIT, A.K. Tishechkin (USFQ); 1: Tiputini Biodiversity Station,
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<geoCoordinate direction="south" orientation="latitude" precision="5" value="-0.6376">0.6376°S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-76.1499">76.1499°W</geoCoordinate>
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, 4-9.vi.2011, FIT, M.S. Caterino & A.K. Tishechkin, DNA Extract MSC-2180 (SBMNH); 1: Parque Nac.
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<normalizedToken originalValue="Yasuní">Yasuni</normalizedToken>
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, Via Maxus at Puente
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<normalizedToken originalValue="Piraña">Pirana</normalizedToken>
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,
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<geoCoordinate direction="south" orientation="latitude" precision="92" value="-0.65833336">0°39.5'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-76.433334">76°26'W</geoCoordinate>
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, 14-20.vii.2008, FIT, A.K. Tishechkin (AKTC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="199" pageNumber="200" type="other material">
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<paragraph pageId="199" pageNumber="200">Other material.</paragraph>
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<paragraph pageId="199" pageNumber="200">
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PERU: Loreto: 1: 1.5km N Teniente Lopez,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-2.5943334">2°35.66'S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="9" value="-76.11533">76°06.92'W</geoCoordinate>
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, 210-240m, 20.vii.1993, FIT, R. Leschen (SEMC).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="201" lastPageNumber="202" pageId="199" pageNumber="200" type="description">
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<paragraph pageId="199" pageNumber="200">Diagnostic description.</paragraph>
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<paragraph lastPageId="201" lastPageNumber="202" pageId="199" pageNumber="200">
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Length: 2.25-2.53 mm, width: 2.03-2.18 mm; body rufopiceous, elongate, sides subparallel to weakly rounded, moderately strongly de
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<pageBreakToken pageId="200" pageNumber="201" start="start">pressed</pageBreakToken>
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; frons broadly depressed at middle, sides of frontal stria divergent, sinuate over antennal bases, complete across front; supraorbital stria complete, strongly arched at middle, connected to sides of frontal stria; labrum short, apically emarginate; left mandible with broad, subacute tooth, right mandible with smaller acute tooth; pronotal disk with vague, linear prescutellar depression, ground punctation fine, with few (~6) coarse punctures in a cluster near sides; anterior pronotal margin weakly, bluntly projecting over head; marginal stria interrupted at middle; lateral submarginal stria complete, curved inward at front nearly to anterior stria; anterior submarginal stria transverse across middle of front, just barely recurved posterad at ends; median pronotal gland openings about halfway back on disk on each side; elytron with two complete epipleural striae, with outer subhumeral stria interrupted at middle, inner subhumeral stria present at middle, apically and especially basally abbreviated, striae 1-3 complete to apex, but distinctly abbreviated from base, 4th stria present in apical half, 5th stria present in apical third, sutural stria present in apical two-thirds, widening and diverging from suture toward front; prosternal keel rounded, projecting at base, carinal striae complete, sinuate at middle, connected in narrow anterior arch; proster
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<pageBreakToken pageId="201" pageNumber="202" start="start">nal</pageBreakToken>
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lobe with marginal stria barely abbreviated at sides; anterior mesoventral margin shallowly emarginate, with complete marginal stria; mesometaventral stria weakly and broadly arched forward to basal third of mesoventral disk, continued at sides by lateral metaventral stria extending toward middle of metacoxa; 1st abdominal ventrite with complete, arcuate inner lateral stria and weak, abbreviated outer; meso- and metatarsomeres with dense brushes of ventral setae, not just rows along ventrolateral margins; propygidium densely and uniformly covered with ovoid, ocellate punctures, separated by less than one-third their diameters; most of pygidial disk with fine, dense ground punctation only, with single row of coarse punctures along basal margin, and with sparsely scattered, inconspicuous, coarser punctures toward apex; marginal pygidial sulcus complete, very deep, crenulate mainly on inner edge.Male genitalia (Figs 55
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<normalizedToken originalValue="A–D">A-D</normalizedToken>
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, G): accessory sclerites absent; T8 with sides narrowed near base, weakly rounded to near apex, apical emargination narrow, basal emargination broadly subangulate, nearly reaching basal membrane attachment line, ventrolateral apodemes most strongly developed near base, narrowed toward apex; S8 short, with sides parallel, apical guides gradually more strongly developed to rounded apices, ventrally halves approximate at base, divergent in apical half; apices of T9 not convergent to apex, apices dorsolaterally flattened, widened and rounded on distal margin, with acute teeth on inner edges; T10 much smaller than opening in T9, with halves separate; S9 with sides more or less parallel, barely narrowed near apex, with apical emargination small, rounded, apical flanges narrow, separate; tegmen short, broad, widest just beyond midpoint, lacking medioventral process; proximal apodemes of median lobe long, three-fourths tegmen length, abruptly narrowed one-third proximad from gonopore; basal piece about one-third tegmen length.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="201" pageNumber="202" type="remarks">
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<paragraph pageId="201" pageNumber="202">Remarks.</paragraph>
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<paragraph pageId="201" pageNumber="202">The two species in this group can be separated easily by geography. This species from western Amazonia may be recognized by the generally interrupted outer subhumeral stria, the depressed frons, more conspicuous coarse pygidial punctures(Fig. 54C), and the smaller number of lateral pronotal punctures(Fig. 54B), as well as by genitalic characters, in particular by the apparently absent medioventral process of the tegmen(Fig. 55G). One specimen from Peru differs in a few minor external characters, having complete outer subhumeral striae, and having the lateral and anterior submarginal pronotal striae joined in front, and we exclude it from the type series.</paragraph>
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<caption pageId="201" pageNumber="202">
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<paragraph pageId="201" pageNumber="202">
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Figure 54.
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hirsutipes" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="hirsutipes">Operclipygus hirsutipes</taxonomicName>
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group. A Tarsus of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hirsutipes" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="hirsutipes">Operclipygus hirsutipes</taxonomicName>
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B Dorsal habitus of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hirsutipes" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="hirsutipes">Operclipygus hirsutipes</taxonomicName>
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C Pygidia of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hirsutipes" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="hirsutipes">Operclipygus hirsutipes</taxonomicName>
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D Dorsal habitus of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus guianensis" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="guianensis">Operclipygus guianensis</taxonomicName>
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E Pygidia of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus guianensis" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="guianensis">Operclipygus guianensis</taxonomicName>
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.
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</paragraph>
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</caption>
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<caption pageId="201" pageNumber="202">
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<paragraph pageId="201" pageNumber="202">
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Map 19. Records of the
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hirsutipes" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="hirsutipes">Operclipygus hirsutipes</taxonomicName>
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group.
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</paragraph>
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</caption>
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<caption pageId="201" pageNumber="202">
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<paragraph pageId="201" pageNumber="202">
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Figure 55. Male genitalia of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hirsutipes" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="hirsutipes">Operclipygus hirsutipes</taxonomicName>
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group. A T8 of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hirsutipes" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="hirsutipes">Operclipygus hirsutipes</taxonomicName>
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B S8 of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hirsutipes" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="hirsutipes">Operclipygus hirsutipes</taxonomicName>
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C T9 & T10 of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hirsutipes" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="hirsutipes">Operclipygus hirsutipes</taxonomicName>
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D S9 of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hirsutipes" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="hirsutipes">Operclipygus hirsutipes</taxonomicName>
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E S8 of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus guianensis" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="guianensis">Operclipygus guianensis</taxonomicName>
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F S9 of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus guianensis" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="guianensis">Operclipygus guianensis</taxonomicName>
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G Aedeagus, dorsal and lateral views, of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus hirsutipes" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="hirsutipes">Operclipygus hirsutipes</taxonomicName>
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H Aedeagus, dorsal and lateral views, of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus guianensis" order="Coleoptera" pageId="201" pageNumber="202" phylum="Arthropoda" rank="species" species="guianensis">Operclipygus guianensis</taxonomicName>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="201" pageNumber="202" type="etymology">
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<paragraph pageId="201" pageNumber="202">Etymology.</paragraph>
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<paragraph pageId="201" pageNumber="202">
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This
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<normalizedToken originalValue="species’">species'</normalizedToken>
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name refers to its dense tarsal setae.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |