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<document id="D93DFBFDB892549BE3E862E414CC7A62" ID-CLB-Dataset="2406" ID-DOI="10.11646/zootaxa.5093.2.2" ID-GBIF-Dataset="ffbad455-ad57-4fa5-8efa-d18cc2d12e40" ID-ISSN="1175-5326" ID-Zenodo-Dep="5905141" ID-ZooBank="31913C5B-A8D5-4A30-8D9E-4C7C9E953E1E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643182084586" checkinUser="plazi" docAuthor="Antić, Dragan &amp; Mauriès, Jean-Paul" docDate="2022" docId="03E387B4FFC9CD1956C40AC6FCA0FCDE" docLanguage="en" docName="zootaxa.5093.2.2.pdf" docOrigin="Zootaxa 5093 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Guipuzcosoma reipi Antić &amp; Mauriès 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="163" masterDocId="FFDAFFCCFFC7CD0C56530A51FFA7FFC8" masterDocTitle="Two new species of the previously monospecific genus Guipuzcosoma Vicente &amp; Mauriès, 1980 from Spain, with establishment of Guipuzcosomatidae fam. nov. (Diplopoda: Chordeumatida)" masterLastPageNumber="168" masterPageNumber="142" pageNumber="156" updateTime="1699304320109" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1DBAEAFFF2F07273232B3EC34BA99048">Two new species of the previously monospecific genus Guipuzcosoma Vicente &amp; Mauriès, 1980 from Spain, with establishment of Guipuzcosomatidae fam. nov. (Diplopoda: Chordeumatida)</mods:title>
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<mods:namePart id="4622F3BBC7890D26B06AB518CA7216C9">Antić, Dragan</mods:namePart>
<mods:affiliation id="CF75D90F517D5042E05214C203A2F7B6">University of Belgrade-Faculty of Biology, Institute of Zoology, Studentski Trg 16, 11000 Belgrade, Serbia &amp; Serbian Biospeleological Society, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia</mods:affiliation>
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<mods:namePart id="0F9463BF104BB88E04A4A27E02E36FB2">Mauriès, Jean-Paul</mods:namePart>
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<mods:affiliation id="0322AA554B1AF6A50C2B19E91D95BDB6">Muséum National d’Histoire Naturelle, Dpt. Origines &amp; Evolution, Section Arthropodes, 61 rue Buffon F- 75005 Paris, France mauries. jean-paul @ wanadoo. fr; https: // orcid. org / 0000 - 0002 - 0575 - 6937</mods:affiliation>
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<subSubSection id="C3506529FFC9CD0256C40AC6FE49FF79" box="[151,494,151,177]" pageId="14" pageNumber="156" type="nomenclature">
<paragraph id="8BF536A2FFC9CD0256C40AC6FE49FF79" blockId="14.[151,494,151,213]" box="[151,494,151,177]" pageId="14" pageNumber="156">
<heading id="D0BD81CEFFC9CD0256C40AC6FE49FF79" bold="true" box="[151,494,151,177]" fontSize="11" level="1" pageId="14" pageNumber="156" reason="1">
<emphasis id="B93EEAB0FFC9CD0256C40AC6FE49FF79" bold="true" box="[151,494,151,177]" pageId="14" pageNumber="156">
<taxonomicName id="4C4A4D21FFC9CD0256C40AC6FE2EFF79" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[151,393,151,177]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="14" pageNumber="156" phylum="Arthropoda" rank="species" species="reipi" status="sp. nov.">
<emphasis id="B93EEAB0FFC9CD0256C40AC6FE2EFF79" bold="true" box="[151,393,151,177]" italics="true" pageId="14" pageNumber="156">Guipuzcosoma reipi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20D57CBFFC9CD0257C20AC6FE49FF79" box="[401,494,151,177]" pageId="14" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C3506529FFC9CD0256C40AEAFEA1FF1D" box="[151,262,187,213]" pageId="14" pageNumber="156" type="description">
<paragraph id="8BF536A2FFC9CD0256C40AEAFEA1FF1D" blockId="14.[151,494,151,213]" box="[151,262,187,213]" pageId="14" pageNumber="156">
<figureCitation id="13712A27FFC9CD0256C40AEAFEA1FF1D" box="[151,262,187,213]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStart-2="FIGURE 9" captionStart-3="FIGURE 10" captionStart-4="FIGURE 11" captionStart-5="FIGURE 12" captionStartId-0="15.[151,250,1943,1967]" captionStartId-1="16.[151,250,1815,1839]" captionStartId-2="17.[151,250,1814,1838]" captionStartId-3="18.[151,250,1699,1723]" captionStartId-4="19.[151,250,1709,1733]" captionStartId-5="20.[151,250,1979,2003]" captionTargetBox-0="[155,1432,181,1919]" captionTargetBox-1="[151,1436,181,1791]" captionTargetBox-2="[151,1435,181,1789]" captionTargetBox-3="[151,1436,181,1675]" captionTargetBox-4="[244,1346,189,1676]" captionTargetBox-5="[151,1436,1101,1956]" captionTargetId-0="figure-19@15.[155,1432,181,1919]" captionTargetId-1="figure-17@16.[151,1436,181,1791]" captionTargetId-2="figure-19@17.[151,1435,181,1789]" captionTargetId-3="figure-17@18.[151,1436,181,1675]" captionTargetId-4="figure-19@19.[235,1353,181,1685]" captionTargetId-5="figure-470@20.[151,1436,1101,1956]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionTargetPageId-4="19" captionTargetPageId-5="20" captionText-0="FIGURE 7. Guipuzcosoma reipi sp. nov., habitus. A–D. ♂ holotype (SMNG). A. Lateral view. B. Anterior part of body, showing head and collum in dorsal view. C. Anterior part of body, lateral view. D. Midbody rings, dorsal view. E. ♂ paratype, lateral view (IZB). Scale bars: 1 mm." captionText-1="FIGURE 8. Guipuzcosoma reipi sp. nov., ♂ paratype, habitus (NHMW MY10255). A. Head, anterior view. B. Head and collum, dorso-lateral view. C. Right antenna, anterior view. D. Macrochaeta of ring 17. E. Half of ring 17, dorsal view. F. Ring 17, lateral view. Scale bars: 0.1 mm." captionText-2="FIGURE 9. Guipuzcosoma reipi sp. nov., legs and gonopods. A. ♀ paratype (NHMW MY10254), leg-pair 3, anterior view, arrow indicates lateral sternal lobe. B–D. ♂ paratype (SMNG). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Right leg 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. ♂ paratype (NHMW MY10255), anterior gonopods, anterior view. F. ♂ paratype (NHMW MY10255), posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." captionText-3="FIGURE 10. Guipuzcosoma reipi sp. nov., ♂ paratype (NHMW MY10255) and ♀ paratype (NHMW MY10254). A–F. ♂ paratype, anterior gonopods, anterior, antero-latero-distal, lateral, posterior, postero-latero-distal and distal views, respectively. G. ♂ paratype, posterior gonopod deteriorated during SEM. H. ♀ paratype, vulvae, distal view, light square indicates single sperm. I, J. Spermatozoa. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hair-like outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; S: gonopodal sternum. Scale bars: 0.1 mm (A–H), 0.005 mm (I, J)." captionText-4="FIGURE 11. Guipuzcosoma reipi sp. nov., ♂♂ paratypes and ♀ paratype. A. ♂ paratype, anterior gonopods, anterior view (NHMW MY10255). B. ♂ paratype, anterior gonopods, lateral view (SMNG). C. ♂ paratype, anterior gonopods, posterior view (NHMW MY10255). D. ♂ paratype, anterior gonopods, postero-distal view (SMNG). E, F. ♂ paratype, posterior gonopods, anterior and lateral views, respectively (SMNG). G. ♀ paratype, vulvae, distal view (NHMW MY10254). Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hair-like outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." captionText-5="FIGURE 12. Distribution of the genus Guipuzcosoma Vicente &amp; Mauriès (1980)." figureDoi-0="http://doi.org/10.5281/zenodo.5905155" figureDoi-1="http://doi.org/10.5281/zenodo.5905157" figureDoi-2="http://doi.org/10.5281/zenodo.5905159" figureDoi-3="http://doi.org/10.5281/zenodo.5905161" figureDoi-4="http://doi.org/10.5281/zenodo.5905163" figureDoi-5="http://doi.org/10.5281/zenodo.5905165" httpUri-0="https://zenodo.org/record/5905155/files/figure.png" httpUri-1="https://zenodo.org/record/5905157/files/figure.png" httpUri-2="https://zenodo.org/record/5905159/files/figure.png" httpUri-3="https://zenodo.org/record/5905161/files/figure.png" httpUri-4="https://zenodo.org/record/5905163/files/figure.png" httpUri-5="https://zenodo.org/record/5905165/files/figure.png" pageId="14" pageNumber="156">Figs 7–12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3506529FFC9CD0256C40B52FD37FDAA" pageId="14" pageNumber="156" type="diagnosis">
<paragraph id="8BF536A2FFC9CD0256C40B52FC53FDD1" blockId="14.[151,1437,259,1978]" pageId="14" pageNumber="156">
<emphasis id="B93EEAB0FFC9CD0256C40B52FEADFED5" bold="true" box="[151,266,259,285]" pageId="14" pageNumber="156">Diagnosis</emphasis>
. Differs from both congeners in the structure of angiocoxites 1, whose distal processes are sigmoid 
<emphasis id="B93EEAB0FFC9CD0256C40B79FF11FE89" box="[151,182,296,321]" italics="true" pageId="14" pageNumber="156">vs.</emphasis>
distal processes reduced, with lateral bumps in 
<taxonomicName id="4C4A4D21FFC9CD02548F0B79FCF7FE89" authorityName="Vicente &amp; Mauries" authorityYear="1980" box="[732,848,296,322]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="14" pageNumber="156" phylum="Arthropoda" rank="species" species="comasi">
<emphasis id="B93EEAB0FFC9CD02548F0B79FCF7FE89" box="[732,848,296,322]" italics="true" pageId="14" pageNumber="156">G. comasi</emphasis>
</taxonomicName>
, or almost the same width in its entire length, with an acuminate distal part bent laterad in 
<taxonomicName id="4C4A4D21FFC9CD02543A0B1DFD42FEAD" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[617,741,331,358]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="14" pageNumber="156" phylum="Arthropoda" rank="species" species="karinae" status="sp. nov.">
<emphasis id="B93EEAB0FFC9CD02543A0B1DFD42FEAD" box="[617,741,331,358]" italics="true" pageId="14" pageNumber="156">G. karinae</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFC9CD0254BC0B1AFCEBFEAD" bold="true" box="[751,844,331,357]" pageId="14" pageNumber="156">
<taxonomicNameLabel id="A20D57CBFFC9CD0254BC0B1AFCEBFEAD" box="[751,844,331,357]" pageId="14" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The anterior part of the medial bursal structure of the vulvae in 
<taxonomicName id="4C4A4D21FFC9CD0257680B21FE31FE41" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[315,406,368,394]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="14" pageNumber="156" phylum="Arthropoda" rank="species" species="reipi" status="sp. nov.">
<emphasis id="B93EEAB0FFC9CD0257680B21FE31FE41" box="[315,406,368,394]" italics="true" pageId="14" pageNumber="156">G. reipi</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFC9CD0257F20B3EFE59FE41" bold="true" box="[417,510,367,393]" pageId="14" pageNumber="156">
<taxonomicNameLabel id="A20D57CBFFC9CD0257F20B3EFE59FE41" box="[417,510,367,393]" pageId="14" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is quadrangular with straight lateral margins, 
<emphasis id="B93EEAB0FFC9CD02524C0B21FB99FE41" box="[1055,1086,368,393]" italics="true" pageId="14" pageNumber="156">vs.</emphasis>
heart-shaped in 
<taxonomicName id="4C4A4D21FFC9CD0253570B21FADEFE41" authorityName="Vicente &amp; Mauries" authorityYear="1980" box="[1284,1401,368,394]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="14" pageNumber="156" phylum="Arthropoda" rank="species" species="comasi">
<emphasis id="B93EEAB0FFC9CD0253570B21FADEFE41" box="[1284,1401,368,394]" italics="true" pageId="14" pageNumber="156">G. comasi</emphasis>
</taxonomicName>
or subquadrangular with convex lateral margins in 
<taxonomicName id="4C4A4D21FFC9CD0254E40BC5FC97FE65" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[695,816,403,430]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="14" pageNumber="156" phylum="Arthropoda" rank="species" species="karinae" status="sp. nov.">
<emphasis id="B93EEAB0FFC9CD0254E40BC5FC97FE65" box="[695,816,403,430]" italics="true" pageId="14" pageNumber="156">G. karinae</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFC9CD0255640BC2FC36FE65" bold="true" box="[823,913,403,429]" pageId="14" pageNumber="156">
<taxonomicNameLabel id="A20D57CBFFC9CD0255640BC2FC36FE65" box="[823,913,403,429]" pageId="14" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Additionally, 
<taxonomicName id="4C4A4D21FFC9CD0252600BC5FB2DFE65" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[1075,1162,404,430]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="14" pageNumber="156" phylum="Arthropoda" rank="species" species="reipi" status="sp. nov.">
<emphasis id="B93EEAB0FFC9CD0252600BC5FB2DFE65" box="[1075,1162,404,430]" italics="true" pageId="14" pageNumber="156">G. reipi</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFC9CD0252C20BC2FB4CFE65" bold="true" box="[1169,1259,403,429]" pageId="14" pageNumber="156">
<taxonomicNameLabel id="A20D57CBFFC9CD0252C20BC2FB4CFE65" box="[1169,1259,403,429]" pageId="14" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="4C4A4D21FFC9CD0253D20BC5FF41FE19" authorityName="Vicente &amp; Mauries" authorityYear="1980" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="14" pageNumber="156" phylum="Arthropoda" rank="species" species="comasi">
<emphasis id="B93EEAB0FFC9CD0253D20BC5FF41FE19" italics="true" pageId="14" pageNumber="156">G. comasi</emphasis>
</taxonomicName>
by subtriangular 
<emphasis id="B93EEAB0FFC9CD0257FF0BE9FE6CFE19" box="[428,459,440,465]" italics="true" pageId="14" pageNumber="156">vs.</emphasis>
rounded lateral parts of colpocoxites in posterior view, or by more differentiated coxal processes on leg-pairs 7 and 11 compared to 
<taxonomicName id="4C4A4D21FFC9CD0254D50B8DFD59FE3D" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[646,766,475,502]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="14" pageNumber="156" phylum="Arthropoda" rank="species" species="karinae" status="sp. nov.">
<emphasis id="B93EEAB0FFC9CD0254D50B8DFD59FE3D" box="[646,766,475,502]" italics="true" pageId="14" pageNumber="156">G. karinae</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFC9CD0255500B8AFCFBFE3D" bold="true" box="[771,860,475,501]" pageId="14" pageNumber="156">
<taxonomicNameLabel id="A20D57CBFFC9CD0255500B8AFCFBFE3D" box="[771,860,475,501]" pageId="14" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Significant differences also exist in the shape of the posterior part of the medial bursal structure of the vulvae in all three species.
</paragraph>
<paragraph id="8BF536A2FFC9CD0256940872FD37FDAA" blockId="14.[151,1437,259,1978]" pageId="14" pageNumber="156">
<emphasis id="B93EEAB0FFC9CD0256940872FEB3FDF5" bold="true" box="[199,276,547,573]" pageId="14" pageNumber="156">Name.</emphasis>
The new species is named in honour of colleague and friend, myriapodologist Hans Reip (SMNG), one of the collectors. A noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3506529FFC9CD025694083AFC0CFB6A" pageId="14" pageNumber="156" type="materials_examined">
<paragraph id="8BF536A2FFC9CD025694083AFD78FD4E" blockId="14.[151,1437,259,1978]" box="[199,735,619,646]" pageId="14" pageNumber="156">
<emphasis id="B93EEAB0FFC9CD025694083AFE0EFD4D" bold="true" box="[199,425,619,645]" pageId="14" pageNumber="156">Material examined</emphasis>
(
<specimenCount id="9D4CFD2BFFC9CD0257EA083AFE5CFD4E" box="[441,507,619,646]" pageId="14" pageNumber="156" type="male">7 ♂♂</specimenCount>
, 
<specimenCount id="9D4CFD2BFFC9CD025454083AFDE0FD4E" box="[519,583,619,646]" pageId="14" pageNumber="156" type="female">4 ♀♀</specimenCount>
, 
<specimenCount id="9D4CFD2BFFC9CD025407083AFD74FD4E" box="[596,723,619,646]" pageId="14" pageNumber="156" type="juvenile">2 juveniles</specimenCount>
).
</paragraph>
<paragraph id="8BF536A2FFC9CD02569408DEFB52FCDD" blockId="14.[151,1437,259,1978]" pageId="14" pageNumber="156">
<materialsCitation id="3B223CFFFFC9CD02569408DEFB52FCDD" ID-GBIF-Occurrence="3457637307" collectingDate="2009-04-24" collectionCode="SMNG" collectorName="de Urquilla &amp; K. Voigtlander" country="Spain" county="Zumarraundi" elevation="980" latitude="42.9241" location="Alava province" longLatPrecision="6" longitude="-2.3224" municipality="Zumarraundi" pageId="14" pageNumber="156" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93EEAB0FFC9CD02569408DEFE9AFD61" bold="true" box="[199,317,655,681]" pageId="14" pageNumber="156">
<typeStatus id="54F18800FFC9CD02569408DEFE9FFD61" box="[199,312,655,681]" pageId="14" pageNumber="156" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingCountry id="F35D7632FFC9CD02571A08DEFE30FD61" box="[329,407,655,681]" name="Spain" pageId="14" pageNumber="156">SPAIN</collectingCountry>
● 
<specimenCount id="9D4CFD2BFFC9CD0257EC08C1FE70FD62" box="[447,471,656,682]" pageId="14" pageNumber="156" type="male">♂</specimenCount>
; 
<location id="8E956079FFC9CD0257B608DEFD3FFD61" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD0257B608DEFD3FFD61" box="[485,664,655,681]" country="Spain" county="Zumarraundi" latitude="42.9241" longLatPrecision="6" longitude="-2.3224" municipality="Zumarraundi" name="Alava province" pageId="14" pageNumber="156">Alava province</location>
, 
<location id="8E956079FFC9CD0254FB08C1FC2AFD61" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD0254FB08C1FC2AFD61" box="[680,909,655,682]" country="Spain" county="Zumarraundi" latitude="42.9241" longLatPrecision="6" longitude="-2.3224" municipality="Zumarraundi" name="Montes de Altzania" pageId="14" pageNumber="156">Montes de Altzania</location>
, 
<location id="8E956079FFC9CD0255CF08DEFBD7FD61" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD0255CF08DEFBD7FD61" box="[924,1136,655,681]" country="Spain" county="Zumarraundi" latitude="42.9241" longLatPrecision="6" longitude="-2.3224" municipality="Zumarraundi" name="Sierra de Urquilla" pageId="14" pageNumber="156">
Sierra 
<collectorName id="26BF5374FFC9CD0255B908DEFBD7FD61" box="[1002,1136,655,681]" pageId="14" pageNumber="156">de Urquilla</collectorName>
</location>
, south of mountain chain 
<location id="8E956079FFC9CD0256C408E2FE36FD05" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD0256C408E2FE36FD05" box="[151,401,691,717]" country="Spain" county="Zumarraundi" latitude="42.9241" longLatPrecision="6" longitude="-2.3224" municipality="Zumarraundi" name="Aikorriko Mendikatea" pageId="14" pageNumber="156">Aikorriko Mendikatea</location>
, south west of 
<location id="8E956079FFC9CD02547F08E2FCA2FD05" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD02547F08E2FCA2FD05" box="[556,773,691,717]" country="Spain" county="Zumarraundi" latitude="42.9241" longLatPrecision="6" longitude="-2.3224" municipality="Zumarraundi" name="Mountain Aitzgorri" pageId="14" pageNumber="156">Mountain Aitzgorri</location>
, north of end of road from 
<collectingMunicipality id="6B91ACD8FFC9CD02524C08E2FE8BFD39" pageId="14" pageNumber="156">Salvatierra-Ordonana-ZalduondoZumarraundi</collectingMunicipality>
, north of 
<collectingCounty id="62944E2EFFC9CD0257C40886FD8BFD39" box="[407,556,727,753]" pageId="14" pageNumber="156">Zumarraundi</collectingCounty>
, deep gryke with stream discharge (cave), in the lower third, 
<geoCoordinate id="EE7E5065FFC9CD0252850886FA93FD39" box="[1238,1332,727,753]" degrees="42.9241" direction="north" orientation="latitude" pageId="14" pageNumber="156" precision="5" value="42.9241">42.9241</geoCoordinate>
, 
<geoCoordinate id="EE7E5065FFC9CD02536C0886FA3FFD39" box="[1343,1432,727,753]" degrees="2.3224" direction="west" orientation="longitude" pageId="14" pageNumber="156" precision="5" value="-2.3224">-2.3224</geoCoordinate>
, 
<elevation id="0067D191FFC9CD0256C408AAFEBCFCDD" box="[151,283,763,790]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="14" pageNumber="156" unit="m" value="980.0">
<quantity id="4CB29B47FFC9CD0256C408AAFF79FCDE" box="[151,222,763,790]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="14" pageNumber="156" unit="m" value="980.0">980 m</quantity>
a.s.l.
</elevation>
, 
<taxonomicName id="4C4A4D21FFC9CD02577408ADFECAFCDD" box="[295,365,764,789]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="156" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC9CD02577408ADFECAFCDD" box="[295,365,764,789]" italics="true" pageId="14" pageNumber="156">Fagus</emphasis>
</taxonomicName>
forest, thick leaf layer, in leaf litter; 
<date id="FFF41062FFC9CD02555B08AAFC09FCDD" box="[776,942,763,789]" pageId="14" pageNumber="156" value="2009-04-24">
<collectingDate id="EFB0E98AFFC9CD02555B08AAFC09FCDD" box="[776,942,763,789]" pageId="14" pageNumber="156" value="2009-04-24">24. April 2009</collectingDate>
</date>
; 
<collectorName id="26BF5374FFC9CD0255EA08ADFBFAFCDD" box="[953,1117,763,790]" pageId="14" pageNumber="156">K. Voigtländer</collectorName>
leg.; 
<collectionCode id="ED5BAE67FFC9CD0252CF08AAFB56FCDD" box="[1180,1265,763,789]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="14" pageNumber="156">SMNG</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF536A2FFC9CD025694094EFC0CFB6A" blockId="14.[151,1437,259,1978]" pageId="14" pageNumber="156">
<materialsCitation id="3B223CFFFFC9CD025694094EFC21FCF1" ID-GBIF-Occurrence="3457637302" box="[199,902,798,826]" collectingDate="2009-04-24" collectionCode="SMNG" collectorName="de Urquilla &amp; K. Voigtlander" country="Spain" county="Zumarraundi" elevation="980" latitude="42.9241" location="Alava province" longLatPrecision="6" longitude="-2.3224" municipality="Zumarraundi" pageId="14" pageNumber="156" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B93EEAB0FFC9CD025694094EFEEFFCF1" bold="true" box="[199,328,799,825]" pageId="14" pageNumber="156">
<typeStatus id="54F18800FFC9CD025694094EFEE4FCF1" box="[199,323,799,825]" pageId="14" pageNumber="156" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="F35D7632FFC9CD025702094EFE38FCF1" box="[337,415,799,825]" name="Spain" pageId="14" pageNumber="156">SPAIN</collectingCountry>
● 
<specimenCount id="9D4CFD2BFFC9CD025791094EFE56FCF2" box="[450,497,799,826]" pageId="14" pageNumber="156" type="male">1 ♂</specimenCount>
; same data as for holotype; 
<collectionCode id="ED5BAE67FFC9CD025566094EFC21FCF1" box="[821,902,799,825]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="14" pageNumber="156">SMNG</collectionCode>
</materialsCitation>
● 
<materialsCitation id="3B223CFFFFC9CD0255FA094EFDE9FC95" ID-GBIF-Occurrence="3457637303" collectingDate="2009-04-24" collectionCode="SMNG" collectorName="de Urquilla &amp; K. Voigtlander" country="Spain" county="Zumarraundi" elevation="980" latitude="42.9241" location="Alava province" longLatPrecision="6" longitude="-2.3224" municipality="Zumarraundi" pageId="14" pageNumber="156" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D4CFD2BFFC9CD0255FA094EFC71FCF2" box="[937,982,799,826]" pageId="14" pageNumber="156" type="female">1 ♀</specimenCount>
; same data as for holotype but near the entrance of cave, 
<taxonomicName id="4C4A4D21FFC9CD02570F0915FE05FC95" box="[348,418,836,861]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="156" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC9CD02570F0915FE05FC95" box="[348,418,836,861]" italics="true" pageId="14" pageNumber="156">Fagus</emphasis>
</taxonomicName>
leaves; 
<collectionCode id="ED5BAE67FFC9CD0257AE0912FDE9FC95" box="[509,590,835,861]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="14" pageNumber="156">SMNG</collectionCode>
</materialsCitation>
● 
<materialsCitation id="3B223CFFFFC9CD02543F0912FDDEFC6D" ID-GBIF-Occurrence="3457637308" collectingDate="2009-04-21" collectionCode="SMNG" collectorName="H. Reip &amp; K. Voigtlander" country="Spain" county="western Pass" elevation="550" latitude="42.9572" location="Gipuzkoa province" longLatPrecision="6" longitude="-2.1122" municipality="Etxarri-Aranaz" pageId="14" pageNumber="156" specimenCount="4" specimenCount-female="1" specimenCount-juvenile="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D4CFD2BFFC9CD02543F0912FD3FFC96" box="[620,664,835,862]" pageId="14" pageNumber="156" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D4CFD2BFFC9CD0254F70912FD6AFC96" box="[676,717,835,862]" pageId="14" pageNumber="156" type="female">1 ♀</specimenCount>
, 
<specimenCount id="9D4CFD2BFFC9CD0254880912FCFEFC95" box="[731,857,835,861]" pageId="14" pageNumber="156" type="juvenile">2 juveniles</specimenCount>
; 
<location id="8E956079FFC9CD0255300912FB9BFC95" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD0255300912FB9BFC95" box="[867,1084,835,861]" country="Spain" county="western Pass" latitude="42.9572" longLatPrecision="6" longitude="-2.1122" municipality="Etxarri-Aranaz" name="Gipuzkoa province" pageId="14" pageNumber="156">Gipuzkoa province</location>
, 
<location id="8E956079FFC9CD0252140912FB5EFC95" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD0252140912FB5EFC95" box="[1095,1273,835,861]" country="Spain" county="western Pass" latitude="42.9572" longLatPrecision="6" longitude="-2.1122" municipality="Etxarri-Aranaz" name="Sierra de Aralar" pageId="14" pageNumber="156">Sierra de Aralar</location>
, 
<location id="8E956079FFC9CD0253570912FAF8FC95" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD0253570912FAF8FC95" box="[1284,1375,835,861]" country="Spain" county="western Pass" latitude="42.9572" longLatPrecision="6" longitude="-2.1122" municipality="Etxarri-Aranaz" name="Beasain" pageId="14" pageNumber="156">Beasain</location>
, road from 
<location id="8E956079FFC9CD0256850936FE8FFC49" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD0256850936FE8FFC49" box="[214,296,871,897]" country="Spain" county="western Pass" latitude="42.9572" longLatPrecision="6" longitude="-2.1122" municipality="Etxarri-Aranaz" name="Lazkao" pageId="14" pageNumber="156">Lazkao</location>
to 
<collectingMunicipality id="6B91ACD8FFC9CD02571C0936FE5DFC49" box="[335,506,871,897]" pageId="14" pageNumber="156">Etxarri-Aranaz</collectingMunicipality>
, 
<collectingCounty id="62944E2EFFC9CD0254550939FD3DFC4A" box="[518,666,872,898]" pageId="14" pageNumber="156">western Pass</collectingCounty>
, 
<geoCoordinate id="EE7E5065FFC9CD0254F40936FCA2FC4A" box="[679,773,871,898]" degrees="42.9572" direction="north" orientation="latitude" pageId="14" pageNumber="156" precision="5" value="42.9572">42.9572</geoCoordinate>
, 
<geoCoordinate id="EE7E5065FFC9CD0255410936FCCDFC49" box="[786,874,871,897]" degrees="2.1122" direction="west" orientation="longitude" pageId="14" pageNumber="156" precision="5" value="-2.1122">-2.1122</geoCoordinate>
, 
<elevation id="0067D191FFC9CD0255250936FC5AFC49" box="[886,1021,871,898]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="14" pageNumber="156" unit="m" value="550.0">
<quantity id="4CB29B47FFC9CD0255250936FC18FC4A" box="[886,959,871,898]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="14" pageNumber="156" unit="m" value="550.0">550 m</quantity>
a.s.l.
</elevation>
, 
<taxonomicName id="4C4A4D21FFC9CD0252580939FBF6FC49" box="[1035,1105,872,897]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="156" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC9CD0252580939FBF6FC49" box="[1035,1105,872,897]" italics="true" pageId="14" pageNumber="156">Fagus</emphasis>
</taxonomicName>
forest, in leaf litter; 
<date id="FFF41062FFC9CD02536D0936FF75FC6D" pageId="14" pageNumber="156" value="2009-04-21">
<collectingDate id="EFB0E98AFFC9CD02536D0936FF75FC6D" pageId="14" pageNumber="156" value="2009-04-21">21 April 2009</collectingDate>
</date>
; 
<collectorName id="26BF5374FFC9CD02568D09DDFE9EFC6D" box="[222,313,907,934]" pageId="14" pageNumber="156">H. Reip</collectorName>
, 
<collectorName id="26BF5374FFC9CD02571709DDFE4FFC6D" box="[324,488,907,934]" pageId="14" pageNumber="156">K. Voigtländer</collectorName>
leg.; 
<collectionCode id="ED5BAE67FFC9CD02547B09DAFDDEFC6D" box="[552,633,907,933]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="14" pageNumber="156">SMNG</collectionCode>
</materialsCitation>
● 
<materialsCitation id="3B223CFFFFC9CD0254CA09DAFF67FC01" ID-GBIF-Occurrence="3457637309" collectingDate="2009-04-21" collectionCode="SEM, NHMW" collectorName="H. Reip &amp; K. Voigtlander" country="Spain" county="western Pass" elevation="550" latitude="42.9572" location="Gipuzkoa province" longLatPrecision="6" longitude="-2.1122" municipality="Etxarri-Aranaz" pageId="14" pageNumber="156" specimenCode="MY10255" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D4CFD2BFFC9CD0254CA09DAFD62FC6E" box="[665,709,907,934]" pageId="14" pageNumber="156" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D4CFD2BFFC9CD02548209DAFD5BFC6E" box="[721,764,907,934]" pageId="14" pageNumber="156" type="female">1 ♀</specimenCount>
(used for 
<collectionCode id="ED5BAE67FFC9CD02553C09DAFC0AFC6D" box="[879,941,907,933]" pageId="14" pageNumber="156">SEM</collectionCode>
); same data as previous; 
<collectionCode id="ED5BAE67FFC9CD02529509DDFA86FC6E" box="[1222,1313,908,934]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="156" type="Museum">NHMW</collectionCode>
<specimenCode id="DBEC9ED9FFC9CD02537A09DAFA3BFC6E" box="[1321,1436,907,934]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="156" type="Museum">MY10255</specimenCode>
(
<specimenCount id="9D4CFD2BFFC9CD0256F209FEFF1FFC01" box="[161,184,943,969]" pageId="14" pageNumber="156" type="male">♂</specimenCount>
)
</materialsCitation>
, MY10254 (
<specimenCount id="9D4CFD2BFFC9CD02570109FEFEC3FC01" box="[338,356,943,969]" pageId="14" pageNumber="156" type="female">♀</specimenCount>
) ● 
<materialsCitation id="3B223CFFFFC9CD0257DC09FEFC94FC02" ID-GBIF-Occurrence="3457637306" box="[399,819,943,970]" collectingDate="2009-04-21" collectionCode="IZB" collectorName="H. Reip &amp; K. Voigtlander" country="Spain" county="western Pass" elevation="550" latitude="42.9572" location="Gipuzkoa province" longLatPrecision="6" longitude="-2.1122" municipality="Etxarri-Aranaz" pageId="14" pageNumber="156" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D4CFD2BFFC9CD0257DC09FEFE1CFC02" box="[399,443,943,970]" pageId="14" pageNumber="156" type="male">1 ♂</specimenCount>
, 
<specimenCount id="9D4CFD2BFFC9CD02579B09FEFE54FC02" box="[456,499,943,970]" pageId="14" pageNumber="156" type="female">1 ♀</specimenCount>
; same data as previous; 
<collectionCode id="ED5BAE67FFC9CD02555509E1FC94FC02" box="[774,819,944,970]" pageId="14" pageNumber="156">IZB</collectionCode>
</materialsCitation>
● 
<materialsCitation id="3B223CFFFFC9CD02550009FEFC30FB91" ID-GBIF-Occurrence="3457637301" collectingDate="2009-04-24" collectionCode="SMNG" collectorName="H. Reip" country="Spain" county="Aldasbarreneta" elevation="1120" latitude="42.9427" location="Gipuzkoa province" longLatPrecision="6" longitude="-2.3346" municipality="Mountain Aitzgorri" pageId="14" pageNumber="156" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D4CFD2BFFC9CD02550009FEFC27FC02" box="[851,896,943,970]" pageId="14" pageNumber="156" type="male">1 ♂</specimenCount>
; 
<location id="8E956079FFC9CD0255DF09FEFBC2FC01" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD0255DF09FEFBC2FC01" box="[908,1125,943,969]" country="Spain" county="Aldasbarreneta" latitude="42.9427" longLatPrecision="6" longitude="-2.3346" municipality="Mountain Aitzgorri" name="Gipuzkoa province" pageId="14" pageNumber="156">Gipuzkoa province</location>
, 
<location id="8E956079FFC9CD02522209E1FAE9FC01" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD02522209E1FAE9FC01" box="[1137,1358,943,970]" country="Spain" county="Aldasbarreneta" latitude="42.9427" longLatPrecision="6" longitude="-2.3346" municipality="Mountain Aitzgorri" name="Montes de Altzania" pageId="14" pageNumber="156">Montes de Altzania</location>
, 
<location id="8E956079FFC9CD02530A09FEFEBFFC25" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD02530A09FEFEBFFC25" country="Spain" county="Aldasbarreneta" latitude="42.9427" longLatPrecision="6" longitude="-2.3346" municipality="Mountain Aitzgorri" name="Sierra de Urquilla" pageId="14" pageNumber="156">Sierra de Urquilla</location>
, south of mountain chain 
<location id="8E956079FFC9CD0254650982FC95FC25" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD0254650982FC95FC25" box="[566,818,979,1005]" country="Spain" county="Aldasbarreneta" latitude="42.9427" longLatPrecision="6" longitude="-2.3346" municipality="Mountain Aitzgorri" name="Aikorriko Mendikatea" pageId="14" pageNumber="156">Aikorriko Mendikatea</location>
, south west of 
<collectingMunicipality id="6B91ACD8FFC9CD02558B0982FB13FC25" box="[984,1204,979,1005]" pageId="14" pageNumber="156">Mountain Aitzgorri</collectingMunicipality>
, north of end of road from Salvatierra-Ordonana-Zalduondo-Zumarraundi, southern lower edge of slope to valley plain 
<collectingCounty id="62944E2EFFC9CD0252BF09A6FA3FFBD9" box="[1260,1432,1015,1041]" pageId="14" pageNumber="156">Aldasbarreneta</collectingCounty>
, 
<geoCoordinate id="EE7E5065FFC9CD0256C40E4AFF52FBFD" box="[151,245,1051,1077]" degrees="42.9427" direction="north" orientation="latitude" pageId="14" pageNumber="156" precision="5" value="42.9427">42.9427</geoCoordinate>
, 
<geoCoordinate id="EE7E5065FFC9CD0257520E4AFEFCFBFD" box="[257,347,1051,1077]" degrees="2.3346" direction="west" orientation="longitude" pageId="14" pageNumber="156" precision="5" value="-2.3346">-2.3346</geoCoordinate>
, 
<elevation id="0067D191FFC9CD0257340E4AFE5EFBFD" box="[359,505,1051,1078]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="14" pageNumber="156" unit="m" value="1120.0">
<quantity id="4CB29B47FFC9CD0257340E4AFE1CFBFE" box="[359,443,1051,1078]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="14" pageNumber="156" unit="m" value="1120.0">1120 m</quantity>
a.s.l.
</elevation>
, lower edge of doline, ca. 
<quantity id="4CB29B47FFC9CD0255740E4AFCF4FBFE" box="[807,851,1051,1078]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="156" unit="m" value="1.0">
<elevation id="0067D191FFC9CD0255740E4AFCF4FBFE" box="[807,851,1051,1078]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="156" unit="m" value="1.0">1 m</elevation>
</quantity>
above ground of doline, 
<taxonomicName id="4C4A4D21FFC9CD0252210E4DFB1FFBFD" box="[1138,1208,1052,1077]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="156" phylum="Tracheophyta" rank="genus">
<emphasis id="B93EEAB0FFC9CD0252210E4DFB1FFBFD" box="[1138,1208,1052,1077]" italics="true" pageId="14" pageNumber="156">Fagus</emphasis>
</taxonomicName>
forest, under stones in deep humus and leaf layer; 
<date id="FFF41062FFC9CD0257A40E6EFD3DFB91" box="[503,666,1087,1113]" pageId="14" pageNumber="156" value="2009-04-24">
<collectingDate id="EFB0E98AFFC9CD0257A40E6EFD3DFB91" box="[503,666,1087,1113]" pageId="14" pageNumber="156" value="2009-04-24">24 April 2009</collectingDate>
</date>
; 
<collectorName id="26BF5374FFC9CD0254FB0E11FCA5FB91" box="[680,770,1087,1114]" pageId="14" pageNumber="156">H. Reip</collectorName>
leg.; 
<collectionCode id="ED5BAE67FFC9CD0255150E6EFC30FB91" box="[838,919,1087,1113]" country="Germany" httpUri="http://grbio.org/cool/b2rf-iikk" name="Senckenberg Museum fuer Naturkunde Goerlitz" pageId="14" pageNumber="156">SMNG</collectionCode>
</materialsCitation>
● 
<materialsCitation id="3B223CFFFFC9CD0255E90E6EFC00FB6A" ID-GBIF-Occurrence="3457637305" collectingDate="2009-04-21" collectionCode="NHM" collectorName="H. Read" country="Spain" county="Goikoa" elevation="420" latitude="43.0494" location="Gipuzkoa province" longLatPrecision="6" longitude="-2.04" municipality="Bedaio" pageId="14" pageNumber="156" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D4CFD2BFFC9CD0255E90E6EFC4DFB92" box="[954,1002,1087,1114]" pageId="14" pageNumber="156" type="male">1 ♂</specimenCount>
; 
<location id="8E956079FFC9CD0255AB0E6EFB74FB91" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD0255AB0E6EFB74FB91" box="[1016,1235,1087,1113]" country="Spain" county="Goikoa" latitude="43.0494" longLatPrecision="6" longitude="-2.04" municipality="Bedaio" name="Gipuzkoa province" pageId="14" pageNumber="156">Gipuzkoa province</location>
, 
<location id="8E956079FFC9CD0252B30E6EFA3FFB91" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD0252B30E6EFA3FFB91" box="[1248,1432,1087,1113]" country="Spain" county="Goikoa" latitude="43.0494" longLatPrecision="6" longitude="-2.04" municipality="Bedaio" name="Sierra de Aralar" pageId="14" pageNumber="156">Sierra de Aralar</location>
, 
<location id="8E956079FFC9CD0256C40E32FF43FBB5" LSID="urn:lsid:plazi:treatment:03E387B4FFC9CD1956C40AC6FCA0FCDE:8E956079FFC9CD0256C40E32FF43FBB5" box="[151,228,1123,1149]" country="Spain" county="Goikoa" latitude="43.0494" longLatPrecision="6" longitude="-2.04" municipality="Bedaio" name="Tolosa" pageId="14" pageNumber="156">Tolosa</location>
, south of 
<collectingMunicipality id="6B91ACD8FFC9CD0257070E32FE03FBB5" box="[340,420,1123,1149]" pageId="14" pageNumber="156">Bedaio</collectingMunicipality>
/ 
<collectingCounty id="62944E2EFFC9CD0257FF0E32FDA7FBB5" box="[428,512,1123,1149]" pageId="14" pageNumber="156">Goikoa</collectingCounty>
, 
<geoCoordinate id="EE7E5065FFC9CD02545F0E32FDCDFBB5" box="[524,618,1123,1149]" degrees="43.0494" direction="north" orientation="latitude" pageId="14" pageNumber="156" precision="5" value="43.0494">43.0494</geoCoordinate>
, 
<geoCoordinate id="EE7E5065FFC9CD0254250E32FD68FBB5" box="[630,719,1123,1149]" degrees="2.0400" direction="west" orientation="longitude" pageId="14" pageNumber="156" precision="5" value="-2.04">-2.0400</geoCoordinate>
, 
<elevation id="0067D191FFC9CD0254880E32FCC7FBB5" box="[731,864,1123,1150]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="14" pageNumber="156" unit="m" value="420.0">
<quantity id="4CB29B47FFC9CD0254880E32FC84FBB6" box="[731,803,1123,1150]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="14" pageNumber="156" unit="m" value="420.0">420 m</quantity>
a.s.l.
</elevation>
, farm buildings and stone wall, logs, wood, stones and rubble, under stones and logs; 
<date id="FFF41062FFC9CD02544E0ED6FD1CFB69" box="[541,699,1159,1185]" pageId="14" pageNumber="156" value="2009-04-21">
<collectingDate id="EFB0E98AFFC9CD02544E0ED6FD1CFB69" box="[541,699,1159,1185]" pageId="14" pageNumber="156" value="2009-04-21">21 April 2009</collectingDate>
</date>
; 
<collectorName id="26BF5374FFC9CD0254940ED9FC84FB69" box="[711,803,1159,1186]" pageId="14" pageNumber="156">H. Read</collectorName>
leg.; 
<collectionCode id="ED5BAE67FFC9CD0255310ED9FC00FB6A" box="[866,935,1160,1186]" pageId="14" pageNumber="156">NHM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3506529FFC9CD1956940EFAFF54FE41" lastPageId="21" lastPageNumber="163" pageId="14" pageNumber="156" type="description">
<paragraph id="8BF536A2FFC9CD0256940EFAFADEFAC6" blockId="14.[151,1437,259,1978]" pageId="14" pageNumber="156">
<emphasis id="B93EEAB0FFC9CD0256940EFAFEFEFB0D" bold="true" box="[199,345,1195,1221]" pageId="14" pageNumber="156">Description.</emphasis>
SIZE AND NUMBER OF BODY RINGS. Body in adults with 30 rings (including telson). 
<typeStatus id="54F18800FFC9CD0252A60EFAFAFAFB0D" box="[1269,1373,1195,1221]" pageId="14" pageNumber="156" type="holotype">Holotype</typeStatus>
male 
<quantity id="4CB29B47FFC9CD0256C40E9EFF41FB22" box="[151,230,1231,1258]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="156" unit="mm" value="10.0">10 mm</quantity>
long, vertical diameter of the largest ring 
<quantity id="4CB29B47FFC9CD0254E90E9EFCB5FB22" box="[698,786,1231,1258]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="156" unit="mm" value="0.8">0.8 mm</quantity>
. 
<typeStatus id="54F18800FFC9CD02554F0E81FCD9FB22" box="[796,894,1232,1258]" pageId="14" pageNumber="156" type="paratype">Paratype</typeStatus>
males 
<quantity id="4CB29B47FFC9CD02559E0E9EFBF4FB22" box="[973,1107,1231,1258]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" metricValueMax="9.7" metricValueMin="7.8" pageId="14" pageNumber="156" unit="mm" value="8.75" valueMax="9.7" valueMin="7.8">7.8–9.7 mm</quantity>
long, vertical diameter of the largest ring 
<quantity id="4CB29B47FFC9CD02574F0EA2FE13FAC6" box="[284,436,1267,1294]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="14" pageNumber="156" unit="mm" value="0.675" valueMax="0.7" valueMin="0.65">0.65–0.7 mm</quantity>
. 
<typeStatus id="54F18800FFC9CD0257EC0EA5FD86FAC6" box="[447,545,1268,1294]" pageId="14" pageNumber="156" type="paratype">Paratype</typeStatus>
females 
<quantity id="4CB29B47FFC9CD0254D50EA2FCA1FAC6" box="[646,774,1267,1294]" metricMagnitude="-3" metricUnit="m" metricValue="9.25" metricValueMax="10.0" metricValueMin="8.5" pageId="14" pageNumber="156" unit="mm" value="9.25" valueMax="10.0" valueMin="8.5">8.5–10 mm</quantity>
long, vertical diameter of the largest ring 
<quantity id="4CB29B47FFC9CD02528E0EA2FAD2FAC6" box="[1245,1397,1267,1294]" metricMagnitude="-4" metricUnit="m" metricValue="7.75" metricValueMax="8.0" metricValueMin="7.5" pageId="14" pageNumber="156" unit="mm" value="0.775" valueMax="0.8" valueMin="0.75">0.75–0.8 mm</quantity>
.
</paragraph>
<paragraph id="8BF536A2FFC9CD0256C40F46FE26FA9D" blockId="14.[151,1437,259,1978]" pageId="14" pageNumber="156">
COLORATION (
<figureCitation id="13712A27FFC9CD0257780F46FECAFAFA" box="[299,365,1303,1330]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[155,1432,181,1919]" captionTargetId="figure-19@15.[155,1432,181,1919]" captionTargetPageId="15" captionText="FIGURE 7. Guipuzcosoma reipi sp. nov., habitus. A–D. ♂ holotype (SMNG). A. Lateral view. B. Anterior part of body, showing head and collum in dorsal view. C. Anterior part of body, lateral view. D. Midbody rings, dorsal view. E. ♂ paratype, lateral view (IZB). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905155" httpUri="https://zenodo.org/record/5905155/files/figure.png" pageId="14" pageNumber="156">Fig. 7</figureCitation>
). Variable. From yellowish-brown to darker brown, with greyish patterns. Ventral halves lighter. Ommatidia blackish.
</paragraph>
<paragraph id="8BF536A2FFC9CD0256940F31FCF4F9BD" blockId="14.[151,1437,259,1978]" pageId="14" pageNumber="156">
HEAD (
<figureCitation id="13712A27FFC9CD0257410F0EFE2AFAB1" box="[274,397,1375,1402]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[155,1432,181,1919]" captionTargetId="figure-19@15.[155,1432,181,1919]" captionTargetPageId="15" captionText="FIGURE 7. Guipuzcosoma reipi sp. nov., habitus. A–D. ♂ holotype (SMNG). A. Lateral view. B. Anterior part of body, showing head and collum in dorsal view. C. Anterior part of body, lateral view. D. Midbody rings, dorsal view. E. ♂ paratype, lateral view (IZB). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905155" httpUri="https://zenodo.org/record/5905155/files/figure.png" pageId="14" pageNumber="156">Figs 7B, C</figureCitation>
, 
<figureCitation id="13712A27FFC9CD0257CB0F0EFE7AFAB1" box="[408,477,1375,1401]" captionStart="FIGURE 8" captionStartId="16.[151,250,1815,1839]" captionTargetBox="[151,1436,181,1791]" captionTargetId="figure-17@16.[151,1436,181,1791]" captionTargetPageId="16" captionText="FIGURE 8. Guipuzcosoma reipi sp. nov., ♂ paratype, habitus (NHMW MY10255). A. Head, anterior view. B. Head and collum, dorso-lateral view. C. Right antenna, anterior view. D. Macrochaeta of ring 17. E. Half of ring 17, dorsal view. F. Ring 17, lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905157" httpUri="https://zenodo.org/record/5905157/files/figure.png" pageId="14" pageNumber="156">8A–C</figureCitation>
). Setose, frontal side slightly concave in males, convex in females. Labrum with three medial teeth and 5+5 labral and 2+2 supralabral setae (
<figureCitation id="13712A27FFC9CD0255560FD2FCC7FA55" box="[773,864,1411,1437]" captionStart="FIGURE 8" captionStartId="16.[151,250,1815,1839]" captionTargetBox="[151,1436,181,1791]" captionTargetId="figure-17@16.[151,1436,181,1791]" captionTargetPageId="16" captionText="FIGURE 8. Guipuzcosoma reipi sp. nov., ♂ paratype, habitus (NHMW MY10255). A. Head, anterior view. B. Head and collum, dorso-lateral view. C. Right antenna, anterior view. D. Macrochaeta of ring 17. E. Half of ring 17, dorsal view. F. Ring 17, lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905157" httpUri="https://zenodo.org/record/5905157/files/figure.png" pageId="14" pageNumber="156">Fig. 8A</figureCitation>
). Promentum subtriangular, without setae. Lingual plates with 4–6 setae arranged in one or two rows. Stipites with ca. 20 setae each.Antennae 
<quantity id="4CB29B47FFC9CD0252DE0FF6FB46FA0A" box="[1165,1249,1447,1474]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="14" pageNumber="156" unit="mm" value="1.4">1.4 mm</quantity>
long in 
<typeStatus id="54F18800FFC9CD0253690FF6FA3BFA09" box="[1338,1436,1447,1473]" pageId="14" pageNumber="156" type="holotype">holotype</typeStatus>
male. Length of antennomeres (in mm): I (0.07), II (0.17), III (0.37), IV (0.18), V (0.34), VI (0.13), VII (0.12) and VIII (0.04). Length/breadth ratios of antennomeres I–VII: I (1), II (2), III (4), IV (2), V (3.5), VI (1) and VII (1.3). Antennomeres II, IV, V, VI and VII with one, three, one, four and one sensillum trichoideum, respectively (
<figureCitation id="13712A27FFC9CD02536B0C42FA28F9E5" box="[1336,1423,1555,1581]" captionStart="FIGURE 8" captionStartId="16.[151,250,1815,1839]" captionTargetBox="[151,1436,181,1791]" captionTargetId="figure-17@16.[151,1436,181,1791]" captionTargetPageId="16" captionText="FIGURE 8. Guipuzcosoma reipi sp. nov., ♂ paratype, habitus (NHMW MY10255). A. Head, anterior view. B. Head and collum, dorso-lateral view. C. Right antenna, anterior view. D. Macrochaeta of ring 17. E. Half of ring 17, dorsal view. F. Ring 17, lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905157" httpUri="https://zenodo.org/record/5905157/files/figure.png" pageId="14" pageNumber="156">Fig. 8C</figureCitation>
). Lateral to antennal sockets a group of papillae-like outgrowths present (
<figureCitation id="13712A27FFC9CD0255E90C66FB94F99A" box="[954,1075,1591,1618]" captionStart="FIGURE 8" captionStartId="16.[151,250,1815,1839]" captionTargetBox="[151,1436,181,1791]" captionTargetId="figure-17@16.[151,1436,181,1791]" captionTargetPageId="16" captionText="FIGURE 8. Guipuzcosoma reipi sp. nov., ♂ paratype, habitus (NHMW MY10255). A. Head, anterior view. B. Head and collum, dorso-lateral view. C. Right antenna, anterior view. D. Macrochaeta of ring 17. E. Half of ring 17, dorsal view. F. Ring 17, lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905157" httpUri="https://zenodo.org/record/5905157/files/figure.png" pageId="14" pageNumber="156">Fig. 8A, B</figureCitation>
). Number of ommatidia: 
<quantity id="4CB29B47FFC9CD02531C0C66FA2FF999" box="[1359,1416,1591,1617]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="14" pageNumber="156" unit="in" value="13.0">13 in</quantity>
4 rows in both sexes, arranged in irregular subtriangle (
<figureCitation id="13712A27FFC9CD0254BD0C0AFCE2F9BD" box="[750,837,1627,1653]" captionStart="FIGURE 8" captionStartId="16.[151,250,1815,1839]" captionTargetBox="[151,1436,181,1791]" captionTargetId="figure-17@16.[151,1436,181,1791]" captionTargetPageId="16" captionText="FIGURE 8. Guipuzcosoma reipi sp. nov., ♂ paratype, habitus (NHMW MY10255). A. Head, anterior view. B. Head and collum, dorso-lateral view. C. Right antenna, anterior view. D. Macrochaeta of ring 17. E. Half of ring 17, dorsal view. F. Ring 17, lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905157" httpUri="https://zenodo.org/record/5905157/files/figure.png" pageId="14" pageNumber="156">Fig. 8B</figureCitation>
).
</paragraph>
<paragraph id="8BF536A2FFC9CD0256940C2EFE3AF976" blockId="14.[151,1437,259,1978]" pageId="14" pageNumber="156">COLLUM. Narrower than head, with six macrochaetae, as all body rings. Anterior edge semi-circular, posterior margin gently concave.</paragraph>
<paragraph id="8BF536A2FFC9CD0256940C99FB20F8E1" blockId="14.[151,1437,259,1978]" pageId="14" pageNumber="156">
BODY RINGS (
<figureCitation id="13712A27FFC9CD02570A0C96FE03F92A" box="[345,420,1735,1762]" captionStart="FIGURE 7" captionStartId="15.[151,250,1943,1967]" captionTargetBox="[155,1432,181,1919]" captionTargetId="figure-19@15.[155,1432,181,1919]" captionTargetPageId="15" captionText="FIGURE 7. Guipuzcosoma reipi sp. nov., habitus. A–D. ♂ holotype (SMNG). A. Lateral view. B. Anterior part of body, showing head and collum in dorsal view. C. Anterior part of body, lateral view. D. Midbody rings, dorsal view. E. ♂ paratype, lateral view (IZB). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905155" httpUri="https://zenodo.org/record/5905155/files/figure.png" pageId="14" pageNumber="156">Figs 7</figureCitation>
, 
<figureCitation id="13712A27FFC9CD0257E10C96FE54F929" box="[434,499,1735,1761]" captionStart="FIGURE 8" captionStartId="16.[151,250,1815,1839]" captionTargetBox="[151,1436,181,1791]" captionTargetId="figure-17@16.[151,1436,181,1791]" captionTargetPageId="16" captionText="FIGURE 8. Guipuzcosoma reipi sp. nov., ♂ paratype, habitus (NHMW MY10255). A. Head, anterior view. B. Head and collum, dorso-lateral view. C. Right antenna, anterior view. D. Macrochaeta of ring 17. E. Half of ring 17, dorsal view. F. Ring 17, lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905157" httpUri="https://zenodo.org/record/5905157/files/figure.png" pageId="14" pageNumber="156">8D–F</figureCitation>
). With lateral humps. Macrochaetae long and trichoid (
<figureCitation id="13712A27FFC9CD02523C0C96FB4EF929" box="[1135,1257,1735,1761]" captionStart="FIGURE 8" captionStartId="16.[151,250,1815,1839]" captionTargetBox="[151,1436,181,1791]" captionTargetId="figure-17@16.[151,1436,181,1791]" captionTargetPageId="16" captionText="FIGURE 8. Guipuzcosoma reipi sp. nov., ♂ paratype, habitus (NHMW MY10255). A. Head, anterior view. B. Head and collum, dorso-lateral view. C. Right antenna, anterior view. D. Macrochaeta of ring 17. E. Half of ring 17, dorsal view. F. Ring 17, lateral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5905157" httpUri="https://zenodo.org/record/5905157/files/figure.png" pageId="14" pageNumber="156">Fig. 8D–F</figureCitation>
). CIX (ring 15) = 1; MIX (ring 15) ~ 2; PIX (ring 15) = 0.8; MA (ring 15) ~ 110˚. Prozonites with hexagonal tiles. Metazonites dorsally with scale-like structures, lateroventrally striated. Lateral parts of humps smooth.
</paragraph>
<paragraph id="8BF536A2FFC9CD0256940D65FCE1F8BA" blockId="14.[151,1437,259,1978]" pageId="14" pageNumber="156">TELSON. Epiproct with a pair of spinnerets and 3+3 setae (1+1 paramedian, 2+2 marginal). Hypoproct with 1+1 distal setae. Paraprocts with 3+3 marginal setae in distal part.</paragraph>
<paragraph id="8BF536A2FFC9CD0256940D2DFCDBF872" blockId="14.[151,1437,259,1978]" pageId="14" pageNumber="156">LEG- PAIRS 1 AND 2. In both sexes with tarsal combs; femora with several long and robust setae; postfemora and tibiae each with several long and robust setae arranged in a group.</paragraph>
<caption id="DF35662AFFC8CD0356C40DC6FDBDF83E" ID-DOI="http://doi.org/10.5281/zenodo.5905155" ID-Zenodo-Dep="5905155" httpUri="https://zenodo.org/record/5905155/files/figure.png" pageId="15" pageNumber="157" startId="15.[151,250,1943,1967]" targetBox="[155,1432,181,1919]" targetPageId="15">
<paragraph id="8BF536A2FFC8CD0356C40DC6FDBDF83E" blockId="15.[151,1437,1943,2039]" pageId="15" pageNumber="157">
<emphasis id="B93EEAB0FFC8CD0356C40DC6FEB1F867" bold="true" box="[151,278,1943,1967]" pageId="15" pageNumber="157">FIGURE 7.</emphasis>
<taxonomicName id="4C4A4D21FFC8CD0357730DC6FE57F867" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[288,496,1943,1967]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="15" pageNumber="157" phylum="Arthropoda" rank="species" species="reipi" status="sp. nov.">
<emphasis id="B93EEAB0FFC8CD0357730DC6FE57F867" box="[288,496,1943,1967]" italics="true" pageId="15" pageNumber="157">Guipuzcosoma reipi</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFC8CD0357AA0DC6FDEAF867" bold="true" box="[505,589,1943,1967]" pageId="15" pageNumber="157">
<taxonomicNameLabel id="A20D57CBFFC8CD0357AA0DC6FDEAF867" box="[505,589,1943,1967]" pageId="15" pageNumber="157" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, habitus. 
<emphasis id="B93EEAB0FFC8CD0354E40DC6FD48F867" bold="true" box="[695,751,1943,1967]" pageId="15" pageNumber="157">A–D.</emphasis>
♂ holotype (SMNG). 
<emphasis id="B93EEAB0FFC8CD0355B10DC6FC5CF867" bold="true" box="[994,1019,1943,1967]" pageId="15" pageNumber="157">A.</emphasis>
Lateral view. 
<emphasis id="B93EEAB0FFC8CD0352C50DC6FB0AF867" bold="true" box="[1174,1197,1943,1967]" pageId="15" pageNumber="157">B.</emphasis>
Anterior part of body, showing head and collum in dorsal view. 
<emphasis id="B93EEAB0FFC8CD03546D0DEAFDF0F81B" bold="true" box="[574,599,1979,2003]" pageId="15" pageNumber="157">C.</emphasis>
Anterior part of body, lateral view. 
<emphasis id="B93EEAB0FFC8CD0355900DEAFC7BF81B" bold="true" box="[963,988,1979,2003]" pageId="15" pageNumber="157">D.</emphasis>
Midbody rings, dorsal view. 
<emphasis id="B93EEAB0FFC8CD0353540DEAFAB9F81B" bold="true" box="[1287,1310,1979,2003]" pageId="15" pageNumber="157">E.</emphasis>
♂ paratype, lateral view (IZB). 
<emphasis id="B93EEAB0FFC8CD0357080D8EFE75F83F" bold="true" box="[347,466,2015,2039]" pageId="15" pageNumber="157">Scale bars:</emphasis>
1 mm.
</paragraph>
</caption>
<caption id="DF35662AFFD7CD1C56C40D46FDB6F8BF" ID-DOI="http://doi.org/10.5281/zenodo.5905157" ID-Zenodo-Dep="5905157" httpUri="https://zenodo.org/record/5905157/files/figure.png" pageId="16" pageNumber="158" startId="16.[151,250,1815,1839]" targetBox="[151,1436,181,1791]" targetPageId="16">
<paragraph id="8BF536A2FFD7CD1C56C40D46FDB6F8BF" blockId="16.[151,1437,1815,1911]" pageId="16" pageNumber="158">
<emphasis id="B93EEAB0FFD7CD1C56C40D46FEB1F8E7" bold="true" box="[151,278,1815,1839]" pageId="16" pageNumber="158">FIGURE 8.</emphasis>
<taxonomicName id="4C4A4D21FFD7CD1C57730D46FE48F8E7" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[288,495,1815,1839]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="16" pageNumber="158" phylum="Arthropoda" rank="species" species="reipi" status="sp. nov.">
<emphasis id="B93EEAB0FFD7CD1C57730D46FE48F8E7" box="[288,495,1815,1839]" italics="true" pageId="16" pageNumber="158">Guipuzcosoma reipi</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFD7CD1C57AA0D46FDEBF8E7" bold="true" box="[505,588,1815,1839]" pageId="16" pageNumber="158">
<taxonomicNameLabel id="A20D57CBFFD7CD1C57AA0D46FDEBF8E7" box="[505,588,1815,1839]" pageId="16" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♂ paratype, habitus (NHMW MY10255). 
<emphasis id="B93EEAB0FFD7CD1C524B0D46FB96F8E7" bold="true" box="[1048,1073,1815,1839]" pageId="16" pageNumber="158">A.</emphasis>
Head, anterior view. 
<emphasis id="B93EEAB0FFD7CD1C534B0D46FA88F8E7" bold="true" box="[1304,1327,1815,1839]" pageId="16" pageNumber="158">B.</emphasis>
Head and collum, dorso-lateral view. 
<emphasis id="B93EEAB0FFD7CD1C57FE0D6AFE62F89B" bold="true" box="[429,453,1851,1875]" pageId="16" pageNumber="158">C.</emphasis>
Right antenna, anterior view. 
<emphasis id="B93EEAB0FFD7CD1C54A40D6AFCB7F89B" bold="true" box="[759,784,1851,1875]" pageId="16" pageNumber="158">D.</emphasis>
Macrochaeta of ring 17. 
<emphasis id="B93EEAB0FFD7CD1C525C0D6AFB87F89B" bold="true" box="[1039,1056,1851,1875]" pageId="16" pageNumber="158">E</emphasis>
. Half of ring 17, dorsal view. 
<emphasis id="B93EEAB0FFD7CD1C53020D6AFAC3F89B" bold="true" box="[1361,1380,1851,1875]" pageId="16" pageNumber="158">F.</emphasis>
Ring 17, lateral view. 
<emphasis id="B93EEAB0FFD7CD1C576C0D0EFE11F8BF" bold="true" box="[319,438,1887,1911]" pageId="16" pageNumber="158">Scale bars:</emphasis>
0.1 mm.
</paragraph>
</caption>
<caption id="DF35662AFFD6CD1D56C40D47FC20F829" ID-DOI="http://doi.org/10.5281/zenodo.5905159" ID-Zenodo-Dep="5905159" httpUri="https://zenodo.org/record/5905159/files/figure.png" pageId="17" pageNumber="159" startId="17.[151,250,1814,1838]" targetBox="[151,1435,181,1789]" targetPageId="17">
<paragraph id="8BF536A2FFD6CD1D56C40D47FC20F829" blockId="17.[151,1437,1814,2018]" pageId="17" pageNumber="159">
<emphasis id="B93EEAB0FFD6CD1D56C40D47FEB3F8E6" bold="true" box="[151,276,1814,1838]" pageId="17" pageNumber="159">FIGURE 9.</emphasis>
<taxonomicName id="4C4A4D21FFD6CD1D574E0D47FE4DF8E6" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[285,490,1814,1838]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="17" pageNumber="159" phylum="Arthropoda" rank="species" species="reipi" status="sp. nov.">
<emphasis id="B93EEAB0FFD6CD1D574E0D47FE4DF8E6" box="[285,490,1814,1838]" italics="true" pageId="17" pageNumber="159">Guipuzcosoma reipi</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFD6CD1D57A10D47FDE3F8E6" bold="true" box="[498,580,1814,1838]" pageId="17" pageNumber="159">
<taxonomicNameLabel id="A20D57CBFFD6CD1D57A10D47FDE3F8E6" box="[498,580,1814,1838]" pageId="17" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, legs and gonopods. 
<emphasis id="B93EEAB0FFD6CD1D55720D47FC9DF8E6" bold="true" box="[801,826,1814,1838]" pageId="17" pageNumber="159">A.</emphasis>
♀ paratype (NHMW MY10254), leg-pair 3, anterior view, arrow indicates lateral sternal lobe. 
<emphasis id="B93EEAB0FFD6CD1D57AF0D6BFD94F89A" bold="true" box="[508,563,1850,1874]" pageId="17" pageNumber="159">B–D.</emphasis>
♂ paratype (SMNG). 
<emphasis id="B93EEAB0FFD6CD1D55400D6BFC8DF89A" bold="true" box="[787,810,1850,1874]" pageId="17" pageNumber="159">B.</emphasis>
Leg-pair 7, posterior view, arrow indicates coxal mesal bump. 
<emphasis id="B93EEAB0FFD6CD1D56C40D0FFF17F8BE" bold="true" box="[151,176,1886,1910]" pageId="17" pageNumber="159">C.</emphasis>
Right leg 10, anterior view, arrow indicates trochanteral bump. 
<emphasis id="B93EEAB0FFD6CD1D55610D0FFCECF8BE" bold="true" box="[818,843,1886,1910]" pageId="17" pageNumber="159">D.</emphasis>
Leg-pair 11, posterior view, arrow indicates coxal process. 
<emphasis id="B93EEAB0FFD6CD1D56C40DD3FF09F852" bold="true" box="[151,174,1922,1946]" pageId="17" pageNumber="159">E.</emphasis>
♂ paratype (NHMW MY10255), anterior gonopods, anterior view. 
<emphasis id="B93EEAB0FFD6CD1D55310DD3FCD2F852" bold="true" box="[866,885,1922,1946]" pageId="17" pageNumber="159">F.</emphasis>
♂ paratype (NHMW MY10255), posterior gonopods, posterior view. 
<emphasis id="B93EEAB0FFD6CD1D57600DF7FDA7F876" bold="true" box="[307,512,1958,1982]" pageId="17" pageNumber="159">Abbreviations: A1:</emphasis>
angiocoxite 1; 
<emphasis id="B93EEAB0FFD6CD1D54C90DF7FD66F876" bold="true" box="[666,705,1958,1982]" pageId="17" pageNumber="159">A2:</emphasis>
angiocoxite 2; 
<emphasis id="B93EEAB0FFD6CD1D550E0DF7FCDFF876" bold="true" box="[861,888,1958,1982]" pageId="17" pageNumber="159">C:</emphasis>
colpocoxite; 
<emphasis id="B93EEAB0FFD6CD1D52530DF7FB84F876" bold="true" box="[1024,1059,1958,1982]" pageId="17" pageNumber="159">Ct:</emphasis>
coxotelopodite; 
<emphasis id="B93EEAB0FFD6CD1D529E0DF7FB53F876" bold="true" box="[1229,1268,1958,1982]" pageId="17" pageNumber="159">Cx:</emphasis>
coxite; 
<emphasis id="B93EEAB0FFD6CD1D53160DF7FAFBF876" bold="true" box="[1349,1372,1958,1982]" pageId="17" pageNumber="159">h:</emphasis>
mesal horns; 
<emphasis id="B93EEAB0FFD6CD1D568E0D9BFF59F82A" bold="true" box="[221,254,1994,2018]" pageId="17" pageNumber="159">pr:</emphasis>
pigment remnants; 
<emphasis id="B93EEAB0FFD6CD1D579A0D9BFE78F82A" bold="true" box="[457,479,1994,2018]" pageId="17" pageNumber="159">S:</emphasis>
gonopodal sternum. 
<emphasis id="B93EEAB0FFD6CD1D54E60D9BFC8BF82A" bold="true" box="[693,812,1994,2018]" pageId="17" pageNumber="159">Scale bars:</emphasis>
0.2 mm.
</paragraph>
</caption>
<caption id="DF35662AFFD5CD1E56C40CF2FD19F814" ID-DOI="http://doi.org/10.5281/zenodo.5905161" ID-Zenodo-Dep="5905161" httpUri="https://zenodo.org/record/5905161/files/figure.png" pageId="18" pageNumber="160" startId="18.[151,250,1699,1723]" targetBox="[151,1436,181,1675]" targetPageId="18">
<paragraph id="8BF536A2FFD5CD1E56C40CF2FD19F814" blockId="18.[151,1437,1699,2012]" pageId="18" pageNumber="160">
<emphasis id="B93EEAB0FFD5CD1E56C40CF2FE85F973" bold="true" box="[151,290,1699,1723]" pageId="18" pageNumber="160">FIGURE 10.</emphasis>
<taxonomicName id="4C4A4D21FFD5CD1E57780CF5FE5DF973" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[299,506,1700,1723]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="18" pageNumber="160" phylum="Arthropoda" rank="species" species="reipi" status="sp. nov.">
<emphasis id="B93EEAB0FFD5CD1E57780CF5FE5DF973" box="[299,506,1700,1723]" italics="true" pageId="18" pageNumber="160">Guipuzcosoma reipi</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFD5CD1E54500CF5FDF1F974" bold="true" box="[515,598,1700,1724]" pageId="18" pageNumber="160">
<taxonomicNameLabel id="A20D57CBFFD5CD1E54500CF5FDF1F974" box="[515,598,1700,1724]" pageId="18" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♂ paratype (NHMW MY10255) and ♀ paratype (NHMW MY10254). 
<emphasis id="B93EEAB0FFD5CD1E531D0CF2FA27F973" bold="true" box="[1358,1408,1699,1723]" pageId="18" pageNumber="160">A–F.</emphasis>
♂ paratype, anterior gonopods, anterior, antero-latero-distal, lateral, posterior, postero-latero-distal and distal views, respectively. 
<emphasis id="B93EEAB0FFD5CD1E56C40CBAFF16F8CB" bold="true" box="[151,177,1771,1795]" pageId="18" pageNumber="160">G.</emphasis>
♂ paratype, posterior gonopod deteriorated during SEM. 
<emphasis id="B93EEAB0FFD5CD1E55580CBDFC82F8CC" bold="true" box="[779,805,1772,1796]" pageId="18" pageNumber="160">H.</emphasis>
♀ paratype, vulvae, distal view, light square indicates single sperm. 
<emphasis id="B93EEAB0FFD5CD1E56B70D41FF49F8E0" bold="true" box="[228,238,1808,1832]" pageId="18" pageNumber="160">I</emphasis>
, 
<emphasis id="B93EEAB0FFD5CD1E56AE0D41FEB7F8E0" bold="true" box="[253,272,1808,1832]" pageId="18" pageNumber="160">J.</emphasis>
Spermatozoa. 
<emphasis id="B93EEAB0FFD5CD1E57F80D5EFDDCF8EF" bold="true" box="[427,635,1807,1831]" pageId="18" pageNumber="160">Abbreviations: A1:</emphasis>
angiocoxite 1; 
<emphasis id="B93EEAB0FFD5CD1E554D0D5EFCE1F8EF" bold="true" box="[798,838,1807,1831]" pageId="18" pageNumber="160">A2:</emphasis>
angiocoxite 2; 
<emphasis id="B93EEAB0FFD5CD1E55BA0D5EFBBAF8EF" bold="true" box="[1001,1053,1807,1831]" pageId="18" pageNumber="160">aA2:</emphasis>
anterior branch of angiocoxite 2; 
<emphasis id="B93EEAB0FFD5CD1E53D00D41FA3AF8E0" bold="true" box="[1411,1437,1808,1832]" pageId="18" pageNumber="160">B:</emphasis>
bursa; 
<emphasis id="B93EEAB0FFD5CD1E568D0D65FEA7F884" bold="true" box="[222,256,1844,1868]" pageId="18" pageNumber="160">bc:</emphasis>
basal protrusion of colpocoxite; 
<emphasis id="B93EEAB0FFD5CD1E540F0D62FDD0F883" bold="true" box="[604,631,1843,1867]" pageId="18" pageNumber="160">C:</emphasis>
colpocoxite; 
<emphasis id="B93EEAB0FFD5CD1E55550D65FC8FF884" bold="true" box="[774,808,1844,1868]" pageId="18" pageNumber="160">cb:</emphasis>
concave bulge; 
<emphasis id="B93EEAB0FFD5CD1E55860D62FC5FF883" bold="true" box="[981,1016,1843,1867]" pageId="18" pageNumber="160">Ct:</emphasis>
coxotelopodite; 
<emphasis id="B93EEAB0FFD5CD1E52FB0D62FB77F883" bold="true" box="[1192,1232,1843,1867]" pageId="18" pageNumber="160">Cx:</emphasis>
coxite; 
<emphasis id="B93EEAB0FFD5CD1E53740D62FAFBF883" bold="true" box="[1319,1372,1843,1867]" pageId="18" pageNumber="160">dA1:</emphasis>
distal process of angiocoxite 1; 
<emphasis id="B93EEAB0FFD5CD1E57F30D06FE72F8A7" bold="true" box="[416,469,1879,1903]" pageId="18" pageNumber="160">deC:</emphasis>
distal extension of colpocoxite; 
<emphasis id="B93EEAB0FFD5CD1E55760D09FC9BF8B8" bold="true" box="[805,828,1880,1904]" pageId="18" pageNumber="160">h:</emphasis>
mesal horns; 
<emphasis id="B93EEAB0FFD5CD1E559F0D09FC57F8B8" bold="true" box="[972,1008,1880,1904]" pageId="18" pageNumber="160">ho:</emphasis>
hair-like outgrowths; 
<emphasis id="B93EEAB0FFD5CD1E52860D09FAADF8B8" bold="true" box="[1237,1290,1880,1904]" pageId="18" pageNumber="160">mbs:</emphasis>
medial bursal structure; 
<emphasis id="B93EEAB0FFD5CD1E56AC0D2DFE81F85C" bold="true" box="[255,294,1916,1940]" pageId="18" pageNumber="160">ms:</emphasis>
membranous structure of colpocoxite; 
<emphasis id="B93EEAB0FFD5CD1E54EF0D2AFD70F85B" bold="true" box="[700,727,1915,1939]" pageId="18" pageNumber="160">O:</emphasis>
operculum; 
<emphasis id="B93EEAB0FFD5CD1E55080D2AFC37F85B" bold="true" box="[859,912,1915,1939]" pageId="18" pageNumber="160">pA1:</emphasis>
posterior process of angiocoxite 1; 
<emphasis id="B93EEAB0FFD5CD1E53560D2AFA9DF85B" bold="true" box="[1285,1338,1915,1939]" pageId="18" pageNumber="160">pA2:</emphasis>
posterior branch of angiocoxite 2; 
<emphasis id="B93EEAB0FFD5CD1E57C70DCEFE1BF87F" bold="true" box="[404,444,1951,1975]" pageId="18" pageNumber="160">pC:</emphasis>
posterior mesal protrusion of colpocoxite; 
<emphasis id="B93EEAB0FFD5CD1E55220DF1FC0CF870" bold="true" box="[881,939,1952,1976]" pageId="18" pageNumber="160">Pgm:</emphasis>
postgenital membranous structure; 
<emphasis id="B93EEAB0FFD5CD1E53460DCEFA8CF87F" bold="true" box="[1301,1323,1951,1975]" pageId="18" pageNumber="160">S:</emphasis>
gonopodal sternum. 
<emphasis id="B93EEAB0FFD5CD1E56A60D92FECAF814" bold="true" box="[245,365,1987,2012]" pageId="18" pageNumber="160">Scale bars:</emphasis>
0.1 mm (
<emphasis id="B93EEAB0FFD5CD1E579C0D92FDA5F814" bold="true" box="[463,514,1987,2012]" pageId="18" pageNumber="160">A–H</emphasis>
), 0.005 mm (
<emphasis id="B93EEAB0FFD5CD1E54DF0D95FD31F814" bold="true" box="[652,662,1988,2012]" pageId="18" pageNumber="160">I</emphasis>
, 
<emphasis id="B93EEAB0FFD5CD1E54F00D95FD17F814" bold="true" box="[675,688,1988,2012]" pageId="18" pageNumber="160">J</emphasis>
).
</paragraph>
</caption>
<caption id="DF35662AFFD4CD1F56C40CFCFA22F82D" ID-DOI="http://doi.org/10.5281/zenodo.5905163" ID-Zenodo-Dep="5905163" httpUri="https://zenodo.org/record/5905163/files/figure.png" pageId="19" pageNumber="161" startId="19.[151,250,1709,1733]" targetBox="[244,1346,189,1676]" targetPageId="19">
<paragraph id="8BF536A2FFD4CD1F56C40CFCFA22F82D" blockId="19.[151,1437,1709,2022]" pageId="19" pageNumber="161">
<emphasis id="B93EEAB0FFD4CD1F56C40CFCFE86F90D" bold="true" box="[151,289,1709,1733]" pageId="19" pageNumber="161">FIGURE 11.</emphasis>
<taxonomicName id="4C4A4D21FFD4CD1F577A0CFFFE50F90D" authority="Antić &amp; Mauriès, 2022" authorityName="Antić &amp; Mauriès" authorityYear="2022" box="[297,503,1710,1733]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="19" pageNumber="161" phylum="Arthropoda" rank="species" species="reipi" status="sp. nov.">
<emphasis id="B93EEAB0FFD4CD1F577A0CFFFE50F90D" box="[297,503,1710,1733]" italics="true" pageId="19" pageNumber="161">Guipuzcosoma reipi</emphasis>
</taxonomicName>
<emphasis id="B93EEAB0FFD4CD1F54530CFFFDF5F90E" bold="true" box="[512,594,1710,1734]" pageId="19" pageNumber="161">
<taxonomicNameLabel id="A20D57CBFFD4CD1F54530CFFFDF5F90E" box="[512,594,1710,1734]" pageId="19" pageNumber="161" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♂♂ paratypes and ♀ paratype. 
<emphasis id="B93EEAB0FFD4CD1F55FB0CFCFC66F90D" bold="true" box="[936,961,1709,1733]" pageId="19" pageNumber="161">A.</emphasis>
♂ paratype, anterior gonopods, anterior view (NHMW MY10255). 
<emphasis id="B93EEAB0FFD4CD1F57270C83FE2CF922" bold="true" box="[372,395,1746,1770]" pageId="19" pageNumber="161">B.</emphasis>
♂ paratype, anterior gonopods, lateral view (SMNG). 
<emphasis id="B93EEAB0FFD4CD1F55E30C80FC6FF921" bold="true" box="[944,968,1745,1769]" pageId="19" pageNumber="161">C.</emphasis>
♂ paratype, anterior gonopods, posterior view (NHMW MY10255). 
<emphasis id="B93EEAB0FFD4CD1F572A0CA7FE35F8C6" bold="true" box="[377,402,1782,1806]" pageId="19" pageNumber="161">D.</emphasis>
♂ paratype, anterior gonopods, postero-distal view (SMNG). 
<emphasis id="B93EEAB0FFD4CD1F52450CA7FB80F8C6" bold="true" box="[1046,1063,1782,1806]" pageId="19" pageNumber="161">E</emphasis>
, 
<emphasis id="B93EEAB0FFD4CD1F52660CA7FBEFF8C6" bold="true" box="[1077,1096,1782,1806]" pageId="19" pageNumber="161">F.</emphasis>
♂ paratype, posterior gonopods, anterior and lateral views, respectively (SMNG). 
<emphasis id="B93EEAB0FFD4CD1F54F30D48FD1DF8F9" bold="true" box="[672,698,1817,1841]" pageId="19" pageNumber="161">G.</emphasis>
♀ paratype, vulvae, distal view (NHMW MY10254). 
<emphasis id="B93EEAB0FFD4CD1F52A80D48FF19F89D" bold="true" pageId="19" pageNumber="161">Abbreviations: A1:</emphasis>
angiocoxite 1; 
<emphasis id="B93EEAB0FFD4CD1F570F0D6CFE24F89D" bold="true" box="[348,387,1853,1877]" pageId="19" pageNumber="161">A2:</emphasis>
angiocoxite 2; 
<emphasis id="B93EEAB0FFD4CD1F54730D6CFDF3F89D" bold="true" box="[544,596,1853,1877]" pageId="19" pageNumber="161">aA2:</emphasis>
anterior branch of angiocoxite 2; 
<emphasis id="B93EEAB0FFD4CD1F55FE0D6FFC60F89E" bold="true" box="[941,967,1854,1878]" pageId="19" pageNumber="161">B:</emphasis>
bursa; 
<emphasis id="B93EEAB0FFD4CD1F52420D6FFB94F89E" bold="true" box="[1041,1075,1854,1878]" pageId="19" pageNumber="161">bc:</emphasis>
basal protrusion of colpocoxite; 
<emphasis id="B93EEAB0FFD4CD1F53D20D6CFA3AF89D" bold="true" box="[1409,1437,1853,1877]" pageId="19" pageNumber="161">C:</emphasis>
colpocoxite; 
<emphasis id="B93EEAB0FFD4CD1F57480D33FE9BF8B2" bold="true" box="[283,316,1890,1914]" pageId="19" pageNumber="161">cb:</emphasis>
concave bulge; 
<emphasis id="B93EEAB0FFD4CD1F57B10D30FDA2F8B1" bold="true" box="[482,517,1889,1913]" pageId="19" pageNumber="161">Ct:</emphasis>
coxotelopodite; 
<emphasis id="B93EEAB0FFD4CD1F54E30D30FD70F8B1" bold="true" box="[688,727,1889,1913]" pageId="19" pageNumber="161">Cx:</emphasis>
coxite; 
<emphasis id="B93EEAB0FFD4CD1F557A0D30FCF8F8B1" bold="true" box="[809,863,1889,1913]" pageId="19" pageNumber="161">dA1:</emphasis>
distal process of angiocoxite 1; 
<emphasis id="B93EEAB0FFD4CD1F52FB0D30FB7BF8B1" bold="true" box="[1192,1244,1889,1913]" pageId="19" pageNumber="161">deC:</emphasis>
distal extension of colpocoxite; 
<emphasis id="B93EEAB0FFD4CD1F57490DD7FE96F856" bold="true" box="[282,305,1926,1950]" pageId="19" pageNumber="161">h:</emphasis>
mesal horns; 
<emphasis id="B93EEAB0FFD4CD1F57EE0DD7FE47F856" bold="true" box="[445,480,1926,1950]" pageId="19" pageNumber="161">ho:</emphasis>
hair-like outgrowths; 
<emphasis id="B93EEAB0FFD4CD1F54930DD7FD51F856" bold="true" box="[704,758,1926,1950]" pageId="19" pageNumber="161">mbs:</emphasis>
medial bursal structure; 
<emphasis id="B93EEAB0FFD4CD1F55A30DD7FBB0F856" bold="true" box="[1008,1047,1926,1950]" pageId="19" pageNumber="161">ms:</emphasis>
membranous structure of colpocoxite; 
<emphasis id="B93EEAB0FFD4CD1F56C40DF8FF14F809" bold="true" box="[151,179,1961,1985]" pageId="19" pageNumber="161">O:</emphasis>
operculum; 
<emphasis id="B93EEAB0FFD4CD1F57610DF8FECFF809" bold="true" box="[306,360,1961,1985]" pageId="19" pageNumber="161">pA1:</emphasis>
posterior process of angiocoxite 1; 
<emphasis id="B93EEAB0FFD4CD1F549C0DF8FCA2F809" bold="true" box="[719,773,1961,1985]" pageId="19" pageNumber="161">pA2:</emphasis>
posterior branch of angiocoxite 2; 
<emphasis id="B93EEAB0FFD4CD1F52360DF8FB29F809" bold="true" box="[1125,1166,1961,1985]" pageId="19" pageNumber="161">pC:</emphasis>
posterior mesal protrusion of colpocoxite; 
<emphasis id="B93EEAB0FFD4CD1F57650D9FFED7F82E" bold="true" box="[310,368,1998,2022]" pageId="19" pageNumber="161">Pgm:</emphasis>
postgenital membranous structure; 
<emphasis id="B93EEAB0FFD4CD1F54880D9FFD5AF82E" bold="true" box="[731,765,1998,2022]" pageId="19" pageNumber="161">pr:</emphasis>
pigment remnants; 
<emphasis id="B93EEAB0FFD4CD1F55940D9CFC7AF82D" bold="true" box="[967,989,1997,2021]" pageId="19" pageNumber="161">S:</emphasis>
gonopodal sternum. 
<emphasis id="B93EEAB0FFD4CD1F52E00D9CFA8DF82E" bold="true" box="[1203,1322,1997,2022]" pageId="19" pageNumber="161">Scale bars:</emphasis>
0.2 mm.
</paragraph>
</caption>
<paragraph id="8BF536A2FFD3CD1856940AC6FC46FED5" blockId="20.[151,1437,151,1077]" pageId="20" pageNumber="162">
MALE SEXUAL CHARACTERS (
<figureCitation id="13712A27FFD3CD1857AD0AC6FDDFFF79" box="[510,632,151,177]" captionStart="FIGURE 9" captionStartId="17.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1789]" captionTargetId="figure-19@17.[151,1435,181,1789]" captionTargetPageId="17" captionText="FIGURE 9. Guipuzcosoma reipi sp. nov., legs and gonopods. A. ♀ paratype (NHMW MY10254), leg-pair 3, anterior view, arrow indicates lateral sternal lobe. B–D. ♂ paratype (SMNG). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Right leg 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. ♂ paratype (NHMW MY10255), anterior gonopods, anterior view. F. ♂ paratype (NHMW MY10255), posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5905159" httpUri="https://zenodo.org/record/5905159/files/figure.png" pageId="20" pageNumber="162">Fig. 9B–C</figureCitation>
). Leg-pair 2 with genital openings on coxae. Leg-pairs 3–7 enlarged compared to leg-pairs 1 and 2. Coxa 7 with mesal bump directed posterolaterad (
<figureCitation id="13712A27FFD3CD18524F0AEAFBD3FF1D" box="[1052,1140,187,213]" captionStart="FIGURE 9" captionStartId="17.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1789]" captionTargetId="figure-19@17.[151,1435,181,1789]" captionTargetPageId="17" captionText="FIGURE 9. Guipuzcosoma reipi sp. nov., legs and gonopods. A. ♀ paratype (NHMW MY10254), leg-pair 3, anterior view, arrow indicates lateral sternal lobe. B–D. ♂ paratype (SMNG). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Right leg 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. ♂ paratype (NHMW MY10255), anterior gonopods, anterior view. F. ♂ paratype (NHMW MY10255), posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5905159" httpUri="https://zenodo.org/record/5905159/files/figure.png" pageId="20" pageNumber="162">Fig. 9B</figureCitation>
). No other peculiarities on pregonopodal legs. Leg-pair 10 with coxal sacks and with short triangular trochanteral bumps (
<figureCitation id="13712A27FFD3CD18529E0A8EFA80FF31" box="[1229,1319,223,249]" captionStart="FIGURE 9" captionStartId="17.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1789]" captionTargetId="figure-19@17.[151,1435,181,1789]" captionTargetPageId="17" captionText="FIGURE 9. Guipuzcosoma reipi sp. nov., legs and gonopods. A. ♀ paratype (NHMW MY10254), leg-pair 3, anterior view, arrow indicates lateral sternal lobe. B–D. ♂ paratype (SMNG). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Right leg 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. ♂ paratype (NHMW MY10255), anterior gonopods, anterior view. F. ♂ paratype (NHMW MY10255), posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5905159" httpUri="https://zenodo.org/record/5905159/files/figure.png" pageId="20" pageNumber="162">Fig. 9C</figureCitation>
). Leg-pair 11 with coxal sacks and with short nippled slender coxal process (
<figureCitation id="13712A27FFD3CD1855290B52FC73FED5" box="[890,980,259,285]" captionStart="FIGURE 9" captionStartId="17.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1789]" captionTargetId="figure-19@17.[151,1435,181,1789]" captionTargetPageId="17" captionText="FIGURE 9. Guipuzcosoma reipi sp. nov., legs and gonopods. A. ♀ paratype (NHMW MY10254), leg-pair 3, anterior view, arrow indicates lateral sternal lobe. B–D. ♂ paratype (SMNG). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Right leg 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. ♂ paratype (NHMW MY10255), anterior gonopods, anterior view. F. ♂ paratype (NHMW MY10255), posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5905159" httpUri="https://zenodo.org/record/5905159/files/figure.png" pageId="20" pageNumber="162">Fig. 9D</figureCitation>
).
</paragraph>
<paragraph id="8BF536A2FFD3CD1856940B76FC24FCF1" blockId="20.[151,1437,151,1077]" pageId="20" pageNumber="162">
ANTERIOR GONOPODS (
<figureCitation id="13712A27FFD3CD1857E30B76FDAEFE89" box="[432,521,295,321]" captionStart="FIGURE 9" captionStartId="17.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1789]" captionTargetId="figure-19@17.[151,1435,181,1789]" captionTargetPageId="17" captionText="FIGURE 9. Guipuzcosoma reipi sp. nov., legs and gonopods. A. ♀ paratype (NHMW MY10254), leg-pair 3, anterior view, arrow indicates lateral sternal lobe. B–D. ♂ paratype (SMNG). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Right leg 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. ♂ paratype (NHMW MY10255), anterior gonopods, anterior view. F. ♂ paratype (NHMW MY10255), posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5905159" httpUri="https://zenodo.org/record/5905159/files/figure.png" pageId="20" pageNumber="162">Figs 9E</figureCitation>
, 
<figureCitation id="13712A27FFD3CD1854470B76FDC5FE89" box="[532,610,295,321]" captionStart="FIGURE 10" captionStartId="18.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1675]" captionTargetId="figure-17@18.[151,1436,181,1675]" captionTargetPageId="18" captionText="FIGURE 10. Guipuzcosoma reipi sp. nov., ♂ paratype (NHMW MY10255) and ♀ paratype (NHMW MY10254). A–F. ♂ paratype, anterior gonopods, anterior, antero-latero-distal, lateral, posterior, postero-latero-distal and distal views, respectively. G. ♂ paratype, posterior gonopod deteriorated during SEM. H. ♀ paratype, vulvae, distal view, light square indicates single sperm. I, J. Spermatozoa. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hair-like outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; S: gonopodal sternum. Scale bars: 0.1 mm (A–H), 0.005 mm (I, J)." figureDoi="http://doi.org/10.5281/zenodo.5905161" httpUri="https://zenodo.org/record/5905161/files/figure.png" pageId="20" pageNumber="162">10A–F</figureCitation>
, 
<figureCitation id="13712A27FFD3CD18543E0B76FD18FE89" box="[621,703,295,321]" captionStart="FIGURE 11" captionStartId="19.[151,250,1709,1733]" captionTargetBox="[244,1346,189,1676]" captionTargetId="figure-19@19.[235,1353,181,1685]" captionTargetPageId="19" captionText="FIGURE 11. Guipuzcosoma reipi sp. nov., ♂♂ paratypes and ♀ paratype. A. ♂ paratype, anterior gonopods, anterior view (NHMW MY10255). B. ♂ paratype, anterior gonopods, lateral view (SMNG). C. ♂ paratype, anterior gonopods, posterior view (NHMW MY10255). D. ♂ paratype, anterior gonopods, postero-distal view (SMNG). E, F. ♂ paratype, posterior gonopods, anterior and lateral views, respectively (SMNG). G. ♀ paratype, vulvae, distal view (NHMW MY10254). Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hair-like outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5905163" httpUri="https://zenodo.org/record/5905163/files/figure.png" pageId="20" pageNumber="162">11A–D</figureCitation>
). Gonopodal sternum (
<emphasis id="B93EEAB0FFD3CD1855920B77FC76FE88" bold="true" box="[961,977,294,320]" pageId="20" pageNumber="162">S</emphasis>
) poorly developed, mainly membranous, distomedial parts forming subtriangular structures around round aperture and merging with gonopods. Coxites (
<emphasis id="B93EEAB0FFD3CD1856F20B3FFF64FE40" bold="true" box="[161,195,366,392]" pageId="20" pageNumber="162">Cx</emphasis>
) more or less spoon-shaped, wide in lateral view, supporting two pairs of angiocoxites. Anteromedial pair (= angiocoxite 1) (
<emphasis id="B93EEAB0FFD3CD18571D0BC3FED7FE64" bold="true" box="[334,368,402,428]" pageId="20" pageNumber="162">A1</emphasis>
) widest in mid-height, its mesoproximal sides accommodate shape of sternal triangle; distal process (
<emphasis id="B93EEAB0FFD3CD1856AC0BE7FE96FE18" bold="true" box="[255,305,438,464]" pageId="20" pageNumber="162">dA1</emphasis>
) sigmoid with distal parts curved laterad; posterior process (
<emphasis id="B93EEAB0FFD3CD1855A90BE7FB8BFE18" bold="true" box="[1018,1068,438,464]" pageId="20" pageNumber="162">pA1</emphasis>
) strongly developed, somewhat claw-like in lateral view with small denticles on posterior margin, skintight to lateral sides of distal extension of colpocoxite (
<emphasis id="B93EEAB0FFD3CD18577E0BAFFEFAFDD0" bold="true" box="[301,349,510,536]" pageId="20" pageNumber="162">deC</emphasis>
). Angiocoxites 2 (
<emphasis id="B93EEAB0FFD3CD18546B0BAFFDFDFDD0" bold="true" box="[568,602,510,536]" pageId="20" pageNumber="162">A2</emphasis>
) divided into two branches, anterior branch (
<emphasis id="B93EEAB0FFD3CD18523F0BAFFB3BFDD0" bold="true" box="[1132,1180,510,536]" pageId="20" pageNumber="162">aA2</emphasis>
) consisting of several flagelloid processes with denticulated posterior margins distally; posterior branch (
<emphasis id="B93EEAB0FFD3CD18520F0873FB29FDF4" bold="true" box="[1116,1166,546,572]" pageId="20" pageNumber="162">pA2</emphasis>
) long, acuminate, with longitudinal striations, leans on flagelloid processes closing a canal between; both 
<emphasis id="B93EEAB0FFD3CD18527B0817FBFDFDA8" bold="true" box="[1064,1114,582,608]" pageId="20" pageNumber="162">pA1</emphasis>
and 
<emphasis id="B93EEAB0FFD3CD1852DD0817FB67FDA8" bold="true" box="[1166,1216,582,608]" pageId="20" pageNumber="162">pA2</emphasis>
are directed mesad, and slightly posteriad with overlapping distal parts; 
<emphasis id="B93EEAB0FFD3CD185488083BFD5AFD4C" bold="true" box="[731,765,618,644]" pageId="20" pageNumber="162">A2</emphasis>
outgrows all other parts of anterior gonopods. Colpocoxites (
<emphasis id="B93EEAB0FFD3CD1856F208DFFF12FD60" bold="true" box="[161,181,654,680]" pageId="20" pageNumber="162">C</emphasis>
) massive, strongly developed, chitinized; consist of smaller basal protrusions (
<emphasis id="B93EEAB0FFD3CD18526008DEFBE8FD61" bold="true" box="[1075,1103,655,681]" pageId="20" pageNumber="162">bc</emphasis>
) roundish in lateral view and merged with main massive structures whose distal extensions (
<emphasis id="B93EEAB0FFD3CD18553008E3FC34FD04" bold="true" box="[867,915,690,716]" pageId="20" pageNumber="162">deC</emphasis>
) are in form of elongated horns strongly bent anteriad and skintight to mesal sides of 
<emphasis id="B93EEAB0FFD3CD1854060887FD20FD38" bold="true" box="[597,647,726,752]" pageId="20" pageNumber="162">pA1</emphasis>
; posterior mesal protrusions (
<emphasis id="B93EEAB0FFD3CD1855840887FC5CFD38" bold="true" box="[983,1019,726,752]" pageId="20" pageNumber="162">pC</emphasis>
) rounded and laterally flattened with circular membranous remnants (
<emphasis id="B93EEAB0FFD3CD18545408AAFD8EFCDD" bold="true" box="[519,553,763,789]" pageId="20" pageNumber="162">ms</emphasis>
) on lateral sides. Between 
<emphasis id="B93EEAB0FFD3CD18553208ABFC24FCDC" bold="true" box="[865,899,762,788]" pageId="20" pageNumber="162">A2</emphasis>
and 
<emphasis id="B93EEAB0FFD3CD1855EE08ABFC76FCDC" bold="true" box="[957,977,762,788]" pageId="20" pageNumber="162">C</emphasis>
, laterally, well-developed structure with large number of long, branched hair-like outgrowths (
<emphasis id="B93EEAB0FFD3CD1854A2094EFCA8FCF1" bold="true" box="[753,783,799,825]" pageId="20" pageNumber="162">ho</emphasis>
) presents.
</paragraph>
<paragraph id="8BF536A2FFD3CD1856940912FAF9FBD9" blockId="20.[151,1437,151,1077]" pageId="20" pageNumber="162">
POSTERIOR GONOPODS (
<figureCitation id="13712A27FFD3CD1857E90912FDB4FC95" box="[442,531,835,861]" captionStart="FIGURE 9" captionStartId="17.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1789]" captionTargetId="figure-19@17.[151,1435,181,1789]" captionTargetPageId="17" captionText="FIGURE 9. Guipuzcosoma reipi sp. nov., legs and gonopods. A. ♀ paratype (NHMW MY10254), leg-pair 3, anterior view, arrow indicates lateral sternal lobe. B–D. ♂ paratype (SMNG). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Right leg 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. ♂ paratype (NHMW MY10255), anterior gonopods, anterior view. F. ♂ paratype (NHMW MY10255), posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5905159" httpUri="https://zenodo.org/record/5905159/files/figure.png" pageId="20" pageNumber="162">Figs 9F</figureCitation>
, 
<figureCitation id="13712A27FFD3CD18544C0912FDF5FC95" box="[543,594,835,861]" captionStart="FIGURE 10" captionStartId="18.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1675]" captionTargetId="figure-17@18.[151,1436,181,1675]" captionTargetPageId="18" captionText="FIGURE 10. Guipuzcosoma reipi sp. nov., ♂ paratype (NHMW MY10255) and ♀ paratype (NHMW MY10254). A–F. ♂ paratype, anterior gonopods, anterior, antero-latero-distal, lateral, posterior, postero-latero-distal and distal views, respectively. G. ♂ paratype, posterior gonopod deteriorated during SEM. H. ♀ paratype, vulvae, distal view, light square indicates single sperm. I, J. Spermatozoa. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hair-like outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; S: gonopodal sternum. Scale bars: 0.1 mm (A–H), 0.005 mm (I, J)." figureDoi="http://doi.org/10.5281/zenodo.5905161" httpUri="https://zenodo.org/record/5905161/files/figure.png" pageId="20" pageNumber="162">10G</figureCitation>
, 
<figureCitation id="13712A27FFD3CD18540C0912FD0CFC95" box="[607,683,835,861]" captionStart="FIGURE 11" captionStartId="19.[151,250,1709,1733]" captionTargetBox="[244,1346,189,1676]" captionTargetId="figure-19@19.[235,1353,181,1685]" captionTargetPageId="19" captionText="FIGURE 11. Guipuzcosoma reipi sp. nov., ♂♂ paratypes and ♀ paratype. A. ♂ paratype, anterior gonopods, anterior view (NHMW MY10255). B. ♂ paratype, anterior gonopods, lateral view (SMNG). C. ♂ paratype, anterior gonopods, posterior view (NHMW MY10255). D. ♂ paratype, anterior gonopods, postero-distal view (SMNG). E, F. ♂ paratype, posterior gonopods, anterior and lateral views, respectively (SMNG). G. ♀ paratype, vulvae, distal view (NHMW MY10254). Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hair-like outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5905163" httpUri="https://zenodo.org/record/5905163/files/figure.png" pageId="20" pageNumber="162">11E, F</figureCitation>
). Gonopodal sternum (
<emphasis id="B93EEAB0FFD3CD1855E70913FC63FC94" bold="true" box="[948,964,834,860]" pageId="20" pageNumber="162">S</emphasis>
) poorly developed, membranous, leaving both gonopods almost free; only chitinized part is pair of anterior concave bulges (
<emphasis id="B93EEAB0FFD3CD1852380936FB20FC49" bold="true" box="[1131,1159,871,897]" pageId="20" pageNumber="162">cb</emphasis>
) for accommodation of proximal (basal) parts of posterior gonopods. Coxotelopodites (
<emphasis id="B93EEAB0FFD3CD18550F09DBFCDDFC6C" bold="true" box="[860,890,906,932]" pageId="20" pageNumber="162">Ct</emphasis>
) massive, without articulation, distally with long setae, while mesally, in upper half, with two striking acuminate horns (
<emphasis id="B93EEAB0FFD3CD18559D09FEFC79FC01" bold="true" box="[974,990,943,969]" pageId="20" pageNumber="162">h</emphasis>
); basomesally with partially chitinized colpocoxites (
<emphasis id="B93EEAB0FFD3CD1857690983FEE9FC24" bold="true" box="[314,334,978,1004]" pageId="20" pageNumber="162">C</emphasis>
) both mainly with one long distal seta and slightly serrated mesal margins. Coxotelopodites subdistally with brownish or blackish pigment remnants (
<emphasis id="B93EEAB0FFD3CD18554F09A6FC9FFBD9" bold="true" box="[796,824,1015,1041]" pageId="20" pageNumber="162">pr</emphasis>
). Distal half of 
<emphasis id="B93EEAB0FFD3CD1855B909A7FBAFFBD8" bold="true" box="[1002,1032,1014,1040]" pageId="20" pageNumber="162">Ct</emphasis>
slightly curved posteromesad.
</paragraph>
<paragraph id="8BF536A2FFD3CD1856940E4AFCEDFBFD" blockId="20.[151,1437,151,1077]" box="[199,842,1051,1077]" pageId="20" pageNumber="162">
LEG- PAIR 3 IN FEMALE (
<figureCitation id="13712A27FFD3CD1857ED0E4AFDB3FBFD" box="[446,532,1051,1077]" captionStart="FIGURE 9" captionStartId="17.[151,250,1814,1838]" captionTargetBox="[151,1435,181,1789]" captionTargetId="figure-19@17.[151,1435,181,1789]" captionTargetPageId="17" captionText="FIGURE 9. Guipuzcosoma reipi sp. nov., legs and gonopods. A. ♀ paratype (NHMW MY10254), leg-pair 3, anterior view, arrow indicates lateral sternal lobe. B–D. ♂ paratype (SMNG). B. Leg-pair 7, posterior view, arrow indicates coxal mesal bump. C. Right leg 10, anterior view, arrow indicates trochanteral bump. D. Leg-pair 11, posterior view, arrow indicates coxal process. E. ♂ paratype (NHMW MY10255), anterior gonopods, anterior view. F. ♂ paratype (NHMW MY10255), posterior gonopods, posterior view. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; C: colpocoxite; Ct: coxotelopodite; Cx: coxite; h: mesal horns; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5905159" httpUri="https://zenodo.org/record/5905159/files/figure.png" pageId="20" pageNumber="162">Fig. 9A</figureCitation>
). With lateral sternal lobes.
</paragraph>
<caption id="DF35662AFFD3CD1856C40DEAFC70F81B" ID-DOI="http://doi.org/10.5281/zenodo.5905165" ID-Zenodo-Dep="5905165" box="[151,983,1979,2003]" httpUri="https://zenodo.org/record/5905165/files/figure.png" pageId="20" pageNumber="162" startId="20.[151,250,1979,2003]" targetBox="[151,1436,1101,1956]" targetPageId="20">
<paragraph id="8BF536A2FFD3CD1856C40DEAFC70F81B" blockId="20.[151,983,1979,2003]" box="[151,983,1979,2003]" pageId="20" pageNumber="162">
<emphasis id="B93EEAB0FFD3CD1856C40DEAFE87F81B" bold="true" box="[151,288,1979,2003]" pageId="20" pageNumber="162">FIGURE 12.</emphasis>
Distribution of the genus 
<taxonomicName id="4C4A4D21FFD3CD18547A0DEAFC73F81B" authority="Vicente &amp; Mauries (1980)" authorityName=", Vicente &amp; Mauries" authorityYear="1980" box="[553,980,1979,2003]" class="Diplopoda" family="Vandeleumatidae" genus="Guipuzcosoma" kingdom="Animalia" order="Chordeumatida" pageId="20" pageNumber="162" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEAB0FFD3CD18547A0DEAFD67F81A" box="[553,704,1979,2002]" italics="true" pageId="20" pageNumber="162">Guipuzcosoma</emphasis>
<bibRefCitation id="EFDB4B53FFD3CD1854950DEAFC73F81B" author="Vicente, M. C. &amp; Mauries, J. - P." box="[710,980,1979,2003]" pageId="20" pageNumber="162" pagination="9 - 13" refId="ref15742" refString="Vicente, M. C. &amp; Mauries, J. - P. (1980) Un genero y una especie nuevos de Diplopodos Cavernicolas de Guipuzcoa, Espana (Myriapoda - Diplopoda). Speleon, Barcelona, 25, 9 - 13." type="journal article" year="1980">Vicente &amp; Mauriès (1980)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BF536A2FFD2CD1956940AC6FF54FE41" blockId="21.[151,1437,151,790]" pageId="21" pageNumber="163">
VULVAE (
<figureCitation id="13712A27FFD2CD19577C0AC6FE38FF79" box="[303,415,151,177]" captionStart="FIGURE 10" captionStartId="18.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1675]" captionTargetId="figure-17@18.[151,1436,181,1675]" captionTargetPageId="18" captionText="FIGURE 10. Guipuzcosoma reipi sp. nov., ♂ paratype (NHMW MY10255) and ♀ paratype (NHMW MY10254). A–F. ♂ paratype, anterior gonopods, anterior, antero-latero-distal, lateral, posterior, postero-latero-distal and distal views, respectively. G. ♂ paratype, posterior gonopod deteriorated during SEM. H. ♀ paratype, vulvae, distal view, light square indicates single sperm. I, J. Spermatozoa. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hair-like outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; S: gonopodal sternum. Scale bars: 0.1 mm (A–H), 0.005 mm (I, J)." figureDoi="http://doi.org/10.5281/zenodo.5905161" httpUri="https://zenodo.org/record/5905161/files/figure.png" pageId="21" pageNumber="163">Figs 10H</figureCitation>
, 
<figureCitation id="13712A27FFD2CD1957FC0AC6FE79FF79" box="[431,478,151,177]" captionStart="FIGURE 11" captionStartId="19.[151,250,1709,1733]" captionTargetBox="[244,1346,189,1676]" captionTargetId="figure-19@19.[235,1353,181,1685]" captionTargetPageId="19" captionText="FIGURE 11. Guipuzcosoma reipi sp. nov., ♂♂ paratypes and ♀ paratype. A. ♂ paratype, anterior gonopods, anterior view (NHMW MY10255). B. ♂ paratype, anterior gonopods, lateral view (SMNG). C. ♂ paratype, anterior gonopods, posterior view (NHMW MY10255). D. ♂ paratype, anterior gonopods, postero-distal view (SMNG). E, F. ♂ paratype, posterior gonopods, anterior and lateral views, respectively (SMNG). G. ♀ paratype, vulvae, distal view (NHMW MY10254). Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hair-like outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; pr: pigment remnants; S: gonopodal sternum. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5905163" httpUri="https://zenodo.org/record/5905163/files/figure.png" pageId="21" pageNumber="163">11G</figureCitation>
). With a strongly-developed, massive, wrinkled postgenital membranous structure (
<emphasis id="B93EEAB0FFD2CD1956F20AEAFF70FF1D" bold="true" box="[161,215,187,213]" pageId="21" pageNumber="163">Pgm</emphasis>
) covering posterior and partially lateral parts of bursae (
<emphasis id="B93EEAB0FFD2CD1955060AEAFCCFFF1D" bold="true" box="[853,872,187,213]" pageId="21" pageNumber="163">B</emphasis>
). Lateral valves of 
<emphasis id="B93EEAB0FFD2CD1952100AEAFBF1FF1D" bold="true" box="[1091,1110,187,213]" pageId="21" pageNumber="163">B</emphasis>
rounded posteriorly, without denticles and spicule-like outgrowths in distoposterior ⅔; distoanterior ⅓ wrinkled with long setae. Mesal valves of 
<emphasis id="B93EEAB0FFD2CD1956E40B52FF6DFED5" bold="true" box="[183,202,259,285]" pageId="21" pageNumber="163">B</emphasis>
completely merged, only with anterior fissure as evidence of former separation; anterior ⅔ wrinkled distally with long setae and straight lateral margins with sharp posterior corners; posterior ⅓ separated from rest by deep depression and represents striking unpaired medial structure (
<emphasis id="B93EEAB0FFD2CD1955340B1AFC3EFEAD" bold="true" box="[871,921,331,357]" pageId="21" pageNumber="163">mbs</emphasis>
) in the form of mushroom. Operculum (
<emphasis id="B93EEAB0FFD2CD19532E0B1BFA34FEAC" bold="true" box="[1405,1427,330,356]" pageId="21" pageNumber="163">O</emphasis>
) bilobed.
</paragraph>
</subSubSection>
<subSubSection id="C3506529FFD2CD1956940BC2FB8BFD05" pageId="21" pageNumber="163" type="discussion">
<paragraph id="8BF536A2FFD2CD1956940BC2FB8BFD05" blockId="21.[151,1437,151,790]" pageId="21" pageNumber="163">
<emphasis id="B93EEAB0FFD2CD1956940BC2FE9CFE65" bold="true" box="[199,315,403,429]" pageId="21" pageNumber="163">Remarks.</emphasis>
Interestingly, on the SEM micrograph of the vulvae (
<figureCitation id="13712A27FFD2CD1955F90BC2FBB5FE65" box="[938,1042,403,429]" captionStart="FIGURE 10" captionStartId="18.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1675]" captionTargetId="figure-17@18.[151,1436,181,1675]" captionTargetPageId="18" captionText="FIGURE 10. Guipuzcosoma reipi sp. nov., ♂ paratype (NHMW MY10255) and ♀ paratype (NHMW MY10254). A–F. ♂ paratype, anterior gonopods, anterior, antero-latero-distal, lateral, posterior, postero-latero-distal and distal views, respectively. G. ♂ paratype, posterior gonopod deteriorated during SEM. H. ♀ paratype, vulvae, distal view, light square indicates single sperm. I, J. Spermatozoa. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hair-like outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; S: gonopodal sternum. Scale bars: 0.1 mm (A–H), 0.005 mm (I, J)." figureDoi="http://doi.org/10.5281/zenodo.5905161" httpUri="https://zenodo.org/record/5905161/files/figure.png" pageId="21" pageNumber="163">Fig. 10H</figureCitation>
) in the fissure between the lateral valves and the medial structures (mesal valve), as well as on the surface of the left lateral valve (highlighted square), structures that we consider to be spermatozoa can be seen. These structures (
<figureCitation id="13712A27FFD2CD1955B20B8AFBF5FE3D" box="[993,1106,475,501]" captionStart="FIGURE 10" captionStartId="18.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1675]" captionTargetId="figure-17@18.[151,1436,181,1675]" captionTargetPageId="18" captionText="FIGURE 10. Guipuzcosoma reipi sp. nov., ♂ paratype (NHMW MY10255) and ♀ paratype (NHMW MY10254). A–F. ♂ paratype, anterior gonopods, anterior, antero-latero-distal, lateral, posterior, postero-latero-distal and distal views, respectively. G. ♂ paratype, posterior gonopod deteriorated during SEM. H. ♀ paratype, vulvae, distal view, light square indicates single sperm. I, J. Spermatozoa. Abbreviations: A1: angiocoxite 1; A2: angiocoxite 2; aA2: anterior branch of angiocoxite 2; B: bursa; bc: basal protrusion of colpocoxite; C: colpocoxite; cb: concave bulge; Ct: coxotelopodite; Cx: coxite; dA1: distal process of angiocoxite 1; deC: distal extension of colpocoxite; h: mesal horns; ho: hair-like outgrowths; mbs: medial bursal structure; ms: membranous structure of colpocoxite; O: operculum; pA1: posterior process of angiocoxite 1; pA2: posterior branch of angiocoxite 2; pC: posterior mesal protrusion of colpocoxite; Pgm: postgenital membranous structure; S: gonopodal sternum. Scale bars: 0.1 mm (A–H), 0.005 mm (I, J)." figureDoi="http://doi.org/10.5281/zenodo.5905161" httpUri="https://zenodo.org/record/5905161/files/figure.png" pageId="21" pageNumber="163">Fig. 10I, J</figureCitation>
) are about 12 µm in diameter, and are characterized by a narrow medial protuberance, which rises from the coin-like structure. The medial protuberance could be an acrosome-containing structure, while the basal structure would contain a nucleus. If we are right, these could be the first photographs of spermatozoa in the entire order 
<taxonomicName id="4C4A4D21FFD2CD19525E0816FB1AFDA9" authorityName="Pocock" authorityYear="1894" box="[1037,1213,583,609]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="21" pageNumber="163" phylum="Arthropoda" rank="order">Chordeumatida</taxonomicName>
. These spermatozoa nicely fit the general appearance of spermatozoa in other 
<taxonomicName id="4C4A4D21FFD2CD195572083AFC3DFD4D" box="[801,922,619,645]" class="Diplopoda" kingdom="Animalia" pageId="21" pageNumber="163" phylum="Arthropoda" rank="class">Diplopoda</taxonomicName>
, whose hypothetical evolution is presented in Bacceti 
<emphasis id="B93EEAB0FFD2CD1956A008DEFE8AFD61" box="[243,301,655,681]" italics="true" pageId="21" pageNumber="163">et al.</emphasis>
(1979: 102, fig. 14). It is clear that only cytological methods will show whether the spermatozoa of 
<taxonomicName id="4C4A4D21FFD2CD1956C408E2FEE3FD05" authorityName="Pocock" authorityYear="1894" box="[151,324,691,717]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="21" pageNumber="163" phylum="Arthropoda" rank="order">Chordeumatida</taxonomicName>
belong to the so-called monolayered or bilayered acrosome clade.
</paragraph>
</subSubSection>
<subSubSection id="C3506529FFD2CD1956940887FCA0FCDE" pageId="21" pageNumber="163" type="distribution">
<paragraph id="8BF536A2FFD2CD1956940887FCA0FCDE" blockId="21.[151,1437,151,790]" pageId="21" pageNumber="163">
<emphasis id="B93EEAB0FFD2CD1956940887FEC6FD38" bold="true" box="[199,353,726,752]" pageId="21" pageNumber="163">Distribution.</emphasis>
So far, known only from a few localities on the borders between Alava and Gipuzkoa, and Gipuzkoa and 
<collectingRegion id="498EF840FFD2CD19576908AAFE33FCDD" box="[314,404,763,789]" country="Spain" name="Navarra" pageId="21" pageNumber="163">Navarre</collectingRegion>
provinces (
<figureCitation id="13712A27FFD2CD19544B08AAFDCDFCDD" box="[536,618,763,789]" captionStart="FIGURE 12" captionStartId="20.[151,250,1979,2003]" captionTargetBox="[151,1436,1101,1956]" captionTargetId="figure-470@20.[151,1436,1101,1956]" captionTargetPageId="20" captionText="FIGURE 12. Distribution of the genus Guipuzcosoma Vicente &amp; Mauriès (1980)." figureDoi="http://doi.org/10.5281/zenodo.5905165" httpUri="https://zenodo.org/record/5905165/files/figure.png" pageId="21" pageNumber="163">Fig. 12</figureCitation>
, blue circles).
</paragraph>
</subSubSection>
</treatment>
</document>