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<document id="7B84CF79F3779046E8D6BE6FDA9BBC8A" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629797405601" checkinUser="felipe" docAuthor="Carranza, Salvador &amp; Arnold, Edwin Nicholas" docDate="2012" docId="03E36252C528FFFAF39BFBC9FA0BF8EF" docLanguage="en" docName="zt03378p095.pdf" docOrigin="Zootaxa 3378" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hemidactylus luqueorum Carranza &amp; Arnold 2012, sp. nov." docType="treatment" docVersion="2" lastPageNumber="28" masterDocId="FFDA1A2AC53FFFE1F30CFFADFF85FF9A" masterDocTitle="3378" masterLastPageNumber="95" masterPageNumber="1" pageNumber="24" updateTime="1699113039157" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:namePart id="4628653C72F125E17954C8141B833D66">Carranza, Salvador</mods:namePart>
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<mods:namePart id="9CBF98ED846827AD2288937EAEFAC5AB">Arnold, Edwin Nicholas</mods:namePart>
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<mods:date id="AEF8207E68F74264414672DAB563454F">2012</mods:date>
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<mods:number id="7CDCF184CA010810488ABCDCD1EB5D61">2012-07-04</mods:number>
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<mods:number id="207D45002C603D62E64B2E8C173E4785">3378</mods:number>
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<treatment id="03E36252C528FFFAF39BFBC9FA0BF8EF" LSID="urn:lsid:plazi:treatment:03E36252C528FFFAF39BFBC9FA0BF8EF" httpUri="http://treatment.plazi.org/id/03E36252C528FFFAF39BFBC9FA0BF8EF" lastPageId="27" lastPageNumber="28" pageId="23" pageNumber="24">
<subSubSection id="C35080CFC528FFF6F39BFBC9FDABFBE7" box="[151,558,1123,1150]" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8BF5D344C528FFF6F39BFBC9FDABFBE7" blockId="23.[151,960,1123,1218]" box="[151,558,1123,1150]" pageId="23" pageNumber="24">
<heading id="D0BD6428C528FFF6F39BFBC9FDABFBE7" bold="true" box="[151,558,1123,1150]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="B93E0F56C528FFF6F39BFBC9FDABFBE7" bold="true" box="[151,558,1123,1150]" pageId="23" pageNumber="24">
<taxonomicName id="4C4AA8C7C528FFF6F39BFBC9FE4CFBE4" authority="Carranza &amp; Arnold, 2012" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[151,457,1124,1150]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="luqueorum" status="sp. nov.">
<emphasis id="B93E0F56C528FFF6F39BFBC9FE4CFBE4" bold="true" box="[151,457,1124,1150]" italics="true" pageId="23" pageNumber="24">Hemidactylus luqueorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DB22DC528FFF6F2DDFBCEFDABFBE7" box="[465,558,1123,1149]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35080CFC528FFF6F39BFB2AFC45FB58" pageId="23" pageNumber="24" type="description">
<paragraph id="8BF5D344C528FFF6F39BFB2AFCAFFB05" blockId="23.[151,960,1123,1218]" box="[151,810,1159,1183]" pageId="23" pageNumber="24">
(
<figureCitation id="1371CFC1C528FFF6F393FB2AFF6AFB05" box="[159,239,1159,1183]" captionStart="FIGURE 2" captionStartId="13.[151,244,1580,1601]" captionTargetBox="[160,1436,197,1542]" captionTargetId="figure-120@13.[151,1436,191,1544]" captionTargetPageId="13" captionText="FIGURE 2. Distribution map of Hemidactylus luqueorum sp. nov., H. hajarensis sp. nov., H. alkiyumii sp. nov. and H. festivus sp. nov. Color dots indicate specimens included in the molecular analyses and listed in Table 1. Stars indicate Museum specimens included in the morphological analyses only and listed in Appendix I: 1) BMNH1975.916; 2) BMNH1975.41, BMNH1980.558; 3) BMNH1976.1404; 4) BMNH1977.35; 5) BMNH1983.706; 6) BMNH1977.975; 7) BMNH1977.976–981; 8) BMNH1977.963–966; 9) BMNH1977.956–959; 10) BMNH1977.972–973 and BMNH1976.1409. Dashed lines delimit the different genetic lineages found within H. hajarensis (B1 and B2) and H. alkiyumii (C1, C2 and C3) and shown in Fig. 5 and Appendix III." pageId="23" pageNumber="24">Figs. 2</figureCitation>
,
<figureCitation id="1371CFC1C528FFF6F3F6FB2AFE9AFB05" box="[250,287,1159,1183]" captionStart="FIGURE 5" captionStartId="19.[151,244,1802,1823]" captionTargetBox="[294,1293,193,1791]" captionTargetId="figure-180@19.[294,1314,185,1793]" captionTargetPageId="19" captionText="FIGURE 5. Maximum likelihood (ML) phylogenetic tree of 131 Hemidactylus specimens of the Arid clade (Carranza &amp; Arnold, 2006) based on 2866 bp of concatenated sequences of four mitochondrial (12S, cyb, nd4 and tRNAs) and three nuclear (c-mos, mc1r and rag2) genes. Two specimens of H. flaviviridis were used to root the tree and have not been included in the figure. Tree topology and branch lengths are for the sampled tree with the highest likelihood by RaxML (100 searches, log likelihood = -20540.526926). Each sequence is labeled with the specimen code and taxa name (see Table 1). Maximum-likelihood bootstrap support values above 70% are indicated above branches and black dots by the nodes indicate a posterior probability value ≥ 0.95 in the Bayesian analysis. Age estimates inferred with BEAST are indicated in italics below some relevant nodes and include the mean and, between brackets, the HPD 95% confidence interval. Silhouettes of Hemidactylus taxa are not size-scaled." pageId="23" pageNumber="24">5A</figureCitation>
,
<figureCitation id="1371CFC1C528FFF6F227FB2AFEBEFB05" box="[299,315,1159,1183]" captionStart="FIGURE 6" captionStartId="20.[151,244,1945,1966]" captionTargetBox="[192,1390,199,1918]" captionTargetId="figure-83@20.[191,1396,191,1928]" captionTargetPageId="20" captionText="FIGURE 6. Statistical parsimony nuclear allele networks of c-mos, mc1r and rag2 loci with colors corresponding to species H. persicus, H. luqueorum sp. nov. and H. hajarensis sp. nov.. Circle sizes are proportional to the number of alleles. White circles represent mutational steps. B1 and B2 refer to the different genetic lineages found within H. hajarensis (see Fig. 5 and Appendix III)." pageId="23" pageNumber="24">6</figureCitation>
,
<figureCitation id="1371CFC1C528FFF6F24BFB2AFE04FB05" box="[327,385,1159,1183]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStartId-0="25.[151,244,1730,1751]" captionStartId-1="26.[151,244,1650,1671]" captionStartId-2="28.[151,244,1906,1927]" captionTargetBox-0="[151,1436,191,1708]" captionTargetBox-1="[151,1436,191,1628]" captionTargetBox-2="[228,1358,191,1884]" captionTargetId-0="figure-78@25.[151,1436,191,1708]" captionTargetId-1="figure-92@26.[151,1436,191,1628]" captionTargetId-2="figure-118@28.[228,1358,191,1884]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="28" captionText-0="FIGURE 9. A) male, Holotype of H. luqueorum sp. nov. from Sayq, Jebel Akhdar (BMNH2005.1660); B) left: male of H. luqueorum Wadi Sayq, Jebel Akhdar (BMNH1980.558); right: female of H. hajarensis sp. nov. from Wadi Sabt, Jebel Akhdar (BMNH1977.35); C) above, H. hajarensis (BMNH1977.35); below H. luqueorum (BMNH1980.558); D) underside of right hind feet; left: H. luqueorum (BMNH1980.558), right: H. hajarensis (BMNH1977.35)." captionText-1="FIGURE 10. Live specimens of H. luqueorum sp. nov. A) and B) male from Wadi al Khahafa, Jebel Akhdar (IBES8068); C) unvouchered specimen photographed in its natural habitat in a cave at approximately 1 km E of Hat, Jebel Akhdar (photograph by Felix Amat); D) detail of an area of regenerated skin on the back of an unvouchered specimen, probably as a results of fights with conspecifics or attacks from predators." captionText-2="FIGURE 11. Different localities where H. luqueorum sp. nov. has been found A) 1 km E of Hat, Jebel Akhdar. In this locality H. luqueorum was found out during the day close to crevices and in small caves on the large boulders at the sides of the narrow Wadi; B) Wadi Bani Habib, Jebel Akhdar. In this locality H. luqueorum was found at night in man made constructions and on boulders; C) Wadi al Khahafa, Jebel Akhdar, where two specimens of H. luqueorum have been found at just 492 m altitude on the rocky sides of the wadi." pageId="23" pageNumber="24">9–11</figureCitation>
; Table 1; Appendix I; Appendix IIIA)
</paragraph>
<paragraph id="8BF5D344C528FFF6F39BFB07FC45FB58" blockId="23.[151,960,1123,1218]" box="[151,960,1194,1218]" pageId="23" pageNumber="24">
<materialsCitation id="3B22D919C528FFF6F39BFB07FE17FB58" accessionNumber="M94288" box="[151,402,1194,1218]" pageId="23" pageNumber="24" specimenCount="1">MorphoBank M94288</materialsCitation>
–
<materialsCitation id="3B22D919C528FFF6F2ACFB07FDE1FB58" accessionNumber="M94372 M94378" box="[416,612,1194,1218]" isTailEnumeration="true" pageId="23" pageNumber="24" specimenCount="1">
M94372
<accessionNumber id="94194EA7C528FFF6F10AFB07FDE1FB58" box="[518,612,1194,1218]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M94378" pageId="23" pageNumber="24" type="EnaNcbi">M94378</accessionNumber>
</materialsCitation>
–
<materialsCitation id="3B22D919C528FFF6F17FFB07FD57FB58" accessionNumber="M94393" box="[627,722,1194,1218]" pageId="23" pageNumber="24" specimenCount="1">M94393</materialsCitation>
M100049–M100093
</paragraph>
</subSubSection>
<subSubSection id="C35080CFC528FFF6F39BFB5EFB57FABD" pageId="23" pageNumber="24" type="reference_group">
<paragraph id="8BF5D344C528FFF6F39BFB5EFB57FABD" blockId="23.[151,1436,1266,1319]" pageId="23" pageNumber="24">
<treatmentCitationGroup id="AB5AF46AC528FFF6F39BFB5EFB57FABD" pageId="23" pageNumber="24">
<treatmentCitation id="0AEBF555C528FFF6F39BFB5EFD49FA92" author="Arnold, E. N. &amp; Gallagher, M. D." box="[151,716,1266,1289]" page="65" pageId="23" pageNumber="24" year="1977">
<taxonomicName id="4C4AA8C7C528FFF6F39BFB5EFD49FA92" authority="Arnold and Gallagher, 1977: 65" authorityName="Arnold and Gallagher" authorityPageNumber="65" authorityYear="1977" box="[151,716,1266,1289]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B93E0F56C528FFF6F39BFB5EFEF8FA92" box="[151,381,1267,1289]" italics="true" pageId="23" pageNumber="24">Hemidactylus persicus</emphasis>
<bibRefCitation id="EFDBAEB5C528FFF6F288FB5FFD49FA92" author="Arnold, E. N. &amp; Gallagher, M. D." box="[388,716,1266,1288]" pageId="23" pageNumber="24" pagination="59 - 80" refId="ref41599" refString="Arnold, E. N. &amp; Gallagher, M. D. (1977) The scientific results of the Oman flora and fauna survey 1975. Reptiles and amphibians from the mountains of northern Oman with special reference to the Jebel Akhdar region. Journal of Oman studies special report, 1, 59 - 80." type="journal article" year="1977">Arnold and Gallagher, 1977: 65</bibRefCitation>
</taxonomicName>
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; Arnold, 1977: 102;
<treatmentCitation id="0AEBF555C528FFF6F0ADFB5FFBE5FA92" author="Arnold, E. N." box="[929,1120,1266,1288]" page="419" pageId="23" pageNumber="24" year="1986">
<bibRefCitation id="EFDBAEB5C528FFF6F0ADFB5FFBE5FA92" author="Arnold, E. N." box="[929,1120,1266,1288]" pageId="23" pageNumber="24" pagination="385 - 435" refId="ref41530" refString="Arnold, E. N. (1986) A key and annotated checklist to the lizards and amphisbaenians of Arabia. Fauna of Saudi Arabia, 8, 385 - 435." type="journal article" year="1986">Arnold, 1986: 419</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AEBF555C528FFF6F767FB5FFEF2FABD" author="Leviton, A. E. &amp; Anderson, S. C. &amp; Adler, K. &amp; Minton, S. A." page="38" pageId="23" pageNumber="24" year="1992">
<bibRefCitation id="EFDBAEB5C528FFF6F767FB5FFEF2FABD" author="Leviton, A. E. &amp; Anderson, S. C. &amp; Adler, K. &amp; Minton, S. A." pageId="23" pageNumber="24" refId="ref45006" refString="Leviton, A. E., Anderson, S. C., Adler, K. &amp; Minton, S. A. (1992) Handbook to Middle East amphibians and reptiles. Society for the Study of Amphibians and Reptiles, Oxford, Ohio, 252 pp." type="book" year="1992">Leviton, Anderson, Adler and Minton, 1992: 38</bibRefCitation>
</treatmentCitation>
(part.);
<treatmentCitation id="0AEBF555C528FFF6F2C6FABFFD43FABD" author="van der Kooij, J." box="[458,710,1297,1319]" page="112" pageId="23" pageNumber="24" year="2000">
<bibRefCitation id="EFDBAEB5C528FFF6F2C6FABFFD43FABD" author="van der Kooij, J." box="[458,710,1297,1319]" pageId="23" pageNumber="24" pagination="105 - 120" refId="ref46617" refString="van der Kooij, J. (2000) The herpetofauna of the Sultanate of Oman. Part 2: the geckos. Podarcis, 1, 105 - 120." type="journal article" year="2000">van der Kooij, 2000: 112</bibRefCitation>
</treatmentCitation>
(part.);
<treatmentCitation id="0AEBF555C528FFF6F015FABCFB03FABD" author="Sindaco, R. &amp; Jeremcenko, V. K." box="[793,1158,1297,1319]" page="115" pageId="23" pageNumber="24" year="2008">
<bibRefCitation id="EFDBAEB5C528FFF6F015FABCFB03FABD" author="Sindaco, R. &amp; Jeremcenko, V. K." box="[793,1158,1297,1319]" pageId="23" pageNumber="24" refId="ref45930" refString="Sindaco, R. &amp; Jeremcenko, V. K. (2008) The reptiles of the Western Palearctic. Annotated checklist and distributional atlas of the turtles, crocodiles, amphisbaenians and lizards of Europe, North Africa, Middle East and Central Asia. Monografie della Societas Herpetologica Italica - I. 579 pp." type="book" year="2008">Sindaco and Jeremcenko, 2008: 115</bibRefCitation>
</treatmentCitation>
(part.).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35080CFC528FFFAF39BFAF9FA0BF8EF" lastPageId="27" lastPageNumber="28" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BF5D344C528FFF6F39BFAF9FF7BFAF6" blockId="23.[151,1437,1364,2005]" box="[151,254,1364,1388]" pageId="23" pageNumber="24">
<emphasis id="B93E0F56C528FFF6F39BFAF9FF7BFAF6" box="[151,254,1364,1388]" italics="true" pageId="23" pageNumber="24">
<typeStatus id="54F16DE6C528FFF6F39BFAF9FF7BFAF6" box="[151,254,1364,1388]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BF5D344C528FFF9F3CBFAD5FD19FF4E" blockId="23.[151,1437,1364,2005]" lastBlockId="24.[151,1436,151,212]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<materialsCitation id="3B22D919C528FFF9F3CBFAD5FD19FF4E" accessionNumber="M94288, M94303, M94304, M94312, M94313, M94350, M94378, M94384, M94363, M94372, M00093" collectingDate="2005-10" collectingDateMax="2011-05" collectingDateMin="2005-10" collectionCode="BMNH, IBES, ONHM" collectorName="S. Carranza &amp; E. N. Arnold &amp; D. Donaire &amp; M. D. Gallagher &amp; D. L. Harrison &amp; F. Amat &amp; MorphoBank M &amp; Carranza, E &amp; Jebel Akhdar &amp; MorphoBank" country="Oman" county="North" elevation="1961" lastPageId="24" lastPageNumber="25" latitude="57.60417" location="Sayq" municipality="Wadi Bani Habib" pageId="23" pageNumber="24" specimenCode="BMNH2005.1660, BMNH1971.41, BMNH1980.558, BMNH1975.916, BMNH2005.1661, BMNH2005.1658, BMNH2005.1659, IBES8068, IBES7771, IBES6085, ONHM3705" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBEC7B3FC528FFF6F3CBFAD5FE11FA0A" box="[199,404,1400,1424]" collectionCode="BMNH" pageId="23" pageNumber="24">BMNH2005.1660</specimenCode>
, male from
<location id="8E95859FC528FFF6F112FAD5FDDCFA0A" LSID="urn:lsid:plazi:treatment:03E36252C528FFFAF39BFBC9FA0BF8EF:8E95859FC528FFF6F112FAD5FDDCFA0A" box="[542,601,1400,1424]" country="Oman" county="North" latitude="57.60417" municipality="Wadi Bani Habib" name="Sayq" pageId="23" pageNumber="24">Sayq</location>
,
<quantity id="4CB27EA1C528FFF6F16AFAD5FD3BFA0A" box="[614,702,1400,1424]" metricMagnitude="3" metricUnit="m" metricValue="1.9609999999999999" pageId="23" pageNumber="24" unit="m" value="1961.0">
<elevation id="00673477C528FFF6F16AFAD5FD3BFA0A" box="[614,702,1400,1424]" metricMagnitude="3" metricUnit="m" metricValue="1.9609999999999999" pageId="23" pageNumber="24" unit="m" value="1961.0">1961 m</elevation>
</quantity>
,
<location id="8E95859FC528FFF6F1C7FAD5FCE5FA0A" LSID="urn:lsid:plazi:treatment:03E36252C528FFFAF39BFBC9FA0BF8EF:8E95859FC528FFF6F1C7FAD5FCE5FA0A" box="[715,864,1400,1424]" country="Oman" county="North" latitude="57.60417" municipality="Wadi Bani Habib" name="Jebel Akhdar" pageId="23" pageNumber="24">Jebel Akhdar</location>
(
<location id="8E95859FC528FFF6F07CFAD5FC36FA0A" LSID="urn:lsid:plazi:treatment:03E36252C528FFFAF39BFBC9FA0BF8EF:8E95859FC528FFF6F07CFAD5FC36FA0A" box="[880,947,1400,1424]" country="Oman" county="North" latitude="57.60417" municipality="Wadi Bani Habib" name="North" pageId="23" pageNumber="24">North</location>
<collectingCountry id="F35D93D4C528FFF6F0B7FAD5FB81FA0A" box="[955,1028,1400,1424]" name="Oman" pageId="23" pageNumber="24">Oman</collectingCountry>
), 23.07639’
<geoCoordinate id="EE7EB583C528FFF6F786FAD5FA98FA0A" box="[1162,1309,1400,1424]" degrees="57.62861" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="57.62861">N 57.62861</geoCoordinate>
’E WGS84, collected in
<date id="FFF4F584C528FFF6F223FA31FE55FA2E" box="[303,464,1436,1460]" pageId="23" pageNumber="24" value="2005-10">
<collectingDate id="EFB00C6CC528FFF6F223FA31FE55FA2E" box="[303,464,1436,1460]" pageId="23" pageNumber="24" value="2005-10">October 2005</collectingDate>
</date>
by
<collectorName id="26BFB692C528FFF6F105FA31FD13FA2E" box="[521,662,1436,1460]" pageId="23" pageNumber="24">S. Carranza</collectorName>
,
<collectorName id="26BFB692C528FFF6F1A5FA31FCBFFA2E" box="[681,826,1436,1460]" pageId="23" pageNumber="24">E.N. Arnold</collectorName>
and
<collectorName id="26BFB692C528FFF6F08CFA31FB81FA2E" box="[896,1028,1436,1460]" pageId="23" pageNumber="24">D. Donaire</collectorName>
(
<location id="8E95859FC528FFF6F716FA31FB34FA2E" LSID="urn:lsid:plazi:treatment:03E36252C528FFFAF39BFBC9FA0BF8EF:8E95859FC528FFF6F716FA31FB34FA2E" box="[1050,1201,1436,1460]" country="Oman" county="North" latitude="57.60417" municipality="Wadi Bani Habib" name="MorphoBank" pageId="23" pageNumber="24">MorphoBank</location>
M94288–M94303).
<emphasis id="B93E0F56C528FFF6F39BFA6DFE8AFA42" bold="true" box="[151,271,1472,1496]" pageId="23" pageNumber="24">
<typeStatus id="54F16DE6C528FFF6F39BFA6DFE8AFA42" box="[151,271,1472,1496]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCode id="DBEC7B3FC528FFF6F22DFA6DFE56FA42" box="[289,467,1472,1496]" collectionCode="BMNH" pageId="23" pageNumber="24">BMNH1971.41</specimenCode>
, female from
<location id="8E95859FC528FFF6F176FA6DFD7FFA42" LSID="urn:lsid:plazi:treatment:03E36252C528FFFAF39BFBC9FA0BF8EF:8E95859FC528FFF6F176FA6DFD7FFA42" box="[634,762,1472,1496]" country="Oman" county="North" latitude="57.60417" municipality="Wadi Bani Habib" name="Wadi Sayq" pageId="23" pageNumber="24">Wadi Sayq</location>
,
<quantity id="4CB27EA1C528FFF6F004FA6DFCE6FA42" box="[776,867,1472,1496]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="23" pageNumber="24" unit="m" value="1900.0">
<elevation id="00673477C528FFF6F004FA6DFCE6FA42" box="[776,867,1472,1496]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="23" pageNumber="24" unit="m" value="1900.0">1900 m</elevation>
</quantity>
, Jebel Akhdar (North
<collectingCountry id="F35D93D4C528FFF6F767FA6DFB31FA42" box="[1131,1204,1472,1496]" name="Oman" pageId="23" pageNumber="24">Oman</collectingCountry>
), collected by
<collectorName id="26BFB692C528FFF6F66CFA6CFE80FA66" pageId="23" pageNumber="24">M.D. Gallagher</collectorName>
(MorphoBank M94304–M94312);
<specimenCode id="DBEC7B3FC528FFF6F1ABFA49FCE2FA66" box="[679,871,1508,1532]" collectionCode="BMNH" pageId="23" pageNumber="24">BMNH1980.558</specimenCode>
, male from
<location id="8E95859FC528FFF6F0F3FA49FB07FA66" LSID="urn:lsid:plazi:treatment:03E36252C528FFFAF39BFBC9FA0BF8EF:8E95859FC528FFF6F0F3FA49FB07FA66" box="[1023,1154,1508,1532]" country="Oman" county="North" latitude="57.60417" municipality="Wadi Bani Habib" name="Wadi Sayq" pageId="23" pageNumber="24">Wadi Sayq</location>
,
<quantity id="4CB27EA1C528FFF6F79EFA49FB75FA61" box="[1170,1264,1508,1532]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="23" pageNumber="24" unit="m" value="1900.0">
<elevation id="00673477C528FFF6F79EFA49FB75FA61" box="[1170,1264,1508,1532]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="23" pageNumber="24" unit="m" value="1900.0">1900 m</elevation>
</quantity>
, Jebel Akhdar (North
<collectingCountry id="F35D93D4C528FFF6F3E1F9A5FEB3F9BA" box="[237,310,1544,1568]" name="Oman" pageId="23" pageNumber="24">Oman</collectingCountry>
), collected by
<collectorName id="26BFB692C528FFF6F2E4F9A4FD18F9BA" box="[488,669,1544,1568]" pageId="23" pageNumber="24">M.D. Gallagher</collectorName>
(MorphoBank
<accessionNumber id="94194EA7C528FFF6F058F9A5FC30F9BA" box="[852,949,1544,1568]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M94313" pageId="23" pageNumber="24" type="EnaNcbi">M94313</accessionNumber>
–M94350);
<specimenCode id="DBEC7B3FC528FFF6F731F9A5FB78F9BA" box="[1085,1277,1544,1568]" collectionCode="BMNH" pageId="23" pageNumber="24">BMNH1975.916</specimenCode>
, female from Birkat Sahfan, Jebel Akhdar (
<collectingCountry id="F35D93D4C528FFF6F12AF981FDF5F9DE" box="[550,624,1580,1604]" name="Oman" pageId="23" pageNumber="24">Oman</collectingCountry>
), collected by
<collectorName id="26BFB692C528FFF6F049F981FC77F9DE" box="[837,1010,1580,1604]" pageId="23" pageNumber="24">D.L. Harrison</collectorName>
(MorphoBank
<accessionNumber id="94194EA7C528FFF6F7CCF981FAA4F9DE" box="[1216,1313,1580,1604]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M94378" pageId="23" pageNumber="24" type="EnaNcbi">M94378</accessionNumber>
–M94384).
<specimenCode id="DBEC7B3FC528FFF6F39BF9FDFEE1F9F2" box="[151,356,1616,1640]" collectionCode="BMNH" pageId="23" pageNumber="24">BMNH2005.1661</specimenCode>
, juvenile from Sayq,
<quantity id="4CB27EA1C528FFF6F16CF9FDFD3FF9F2" box="[608,698,1616,1640]" metricMagnitude="3" metricUnit="m" metricValue="1.9609999999999999" pageId="23" pageNumber="24" unit="m" value="1961.0">
<elevation id="00673477C528FFF6F16CF9FDFD3FF9F2" box="[608,698,1616,1640]" metricMagnitude="3" metricUnit="m" metricValue="1.9609999999999999" pageId="23" pageNumber="24" unit="m" value="1961.0">1961 m</elevation>
</quantity>
, Jebel Akhdar (North
<collectingCountry id="F35D93D4C528FFF6F0B2F9FDFB82F9F2" box="[958,1031,1616,1640]" name="Oman" pageId="23" pageNumber="24">Oman</collectingCountry>
), 23.07639’
<geoCoordinate id="EE7EB583C528FFF6F783F9FDFA9EF9F2" box="[1167,1307,1616,1640]" degrees="57.62861" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="57.62861">N 57.62861</geoCoordinate>
’E WGS84, collected in
<date id="FFF4F584C528FFF6F213F9D9FE3DF916" box="[287,440,1652,1676]" pageId="23" pageNumber="24" value="2005-10">
<collectingDate id="EFB00C6CC528FFF6F213F9D9FE3DF916" box="[287,440,1652,1676]" pageId="23" pageNumber="24" value="2005-10">October 2005</collectingDate>
</date>
by
<collectorName id="26BFB692C528FFF6F2EEF9D9FDE2F916" box="[482,615,1652,1676]" pageId="23" pageNumber="24">S. Carranza</collectorName>
,
<collectorName id="26BFB692C528FFF6F17EF9D9FD7EF916" box="[626,763,1652,1676]" pageId="23" pageNumber="24">E.N. Arnold</collectorName>
and
<collectorName id="26BFB692C528FFF6F03EF9D9FC37F916" box="[818,946,1652,1676]" pageId="23" pageNumber="24">D. Donaire</collectorName>
;
<specimenCode id="DBEC7B3FC528FFF6F0B1F9D9FB0EF916" box="[957,1163,1652,1676]" collectionCode="BMNH" pageId="23" pageNumber="24">BMNH2005.1658</specimenCode>
, female from
<location id="8E95859FC528FFF6F62AF9D9FF5BF92A" LSID="urn:lsid:plazi:treatment:03E36252C528FFFAF39BFBC9FA0BF8EF:8E95859FC528FFF6F62AF9D9FF5BF92A" country="Oman" county="North" latitude="57.60417" municipality="Wadi Bani Habib" name="Wadi Bani Habib" pageId="23" pageNumber="24">Wadi Bani Habib</location>
,
<quantity id="4CB27EA1C528FFF6F3E1F935FEC2F92A" box="[237,327,1688,1712]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="23" pageNumber="24" unit="m" value="2200.0">
<elevation id="00673477C528FFF6F3E1F935FEC2F92A" box="[237,327,1688,1712]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="23" pageNumber="24" unit="m" value="2200.0">2200 m</elevation>
</quantity>
, Jebel Akhdar (North
<collectingCountry id="F35D93D4C528FFF6F142F935FD13F92A" box="[590,662,1688,1712]" name="Oman" pageId="23" pageNumber="24">Oman</collectingCountry>
), 23.0711’
<geoCoordinate id="EE7EB583C528FFF6F003F935FC18F92A" box="[783,925,1688,1712]" degrees="57.60417" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="57.60417">N 57.60417</geoCoordinate>
’E WGS84, collected in
<date id="FFF4F584C528FFF6F7B4F935FAD0F92A" box="[1208,1365,1688,1712]" pageId="23" pageNumber="24" value="2005-10">
<collectingDate id="EFB00C6CC528FFF6F7B4F935FAD0F92A" box="[1208,1365,1688,1712]" pageId="23" pageNumber="24" value="2005-10">October 2005</collectingDate>
</date>
by
<collectorName id="26BFB692C528FFF6F68AF935FF7AF94E" pageId="23" pageNumber="24">S. Carranza</collectorName>
,
<collectorName id="26BFB692C528FFF6F201F911FE1CF94E" box="[269,409,1724,1748]" pageId="23" pageNumber="24">E.N. Arnold</collectorName>
and
<collectorName id="26BFB692C528FFF6F2DAF911FDD0F94E" box="[470,597,1724,1748]" pageId="23" pageNumber="24">D. Donaire</collectorName>
(MorphoBank M94363–M94372);
<specimenCode id="DBEC7B3FC528FFF6F0E3F911FB38F94E" box="[1007,1213,1724,1748]" collectionCode="BMNH" pageId="23" pageNumber="24">BMNH2005.1659</specimenCode>
, female from
<location id="8E95859FC528FFF6F66EF911FE9CF962" LSID="urn:lsid:plazi:treatment:03E36252C528FFFAF39BFBC9FA0BF8EF:8E95859FC528FFF6F66EF911FE9CF962" country="Oman" county="North" latitude="57.60417" municipality="Wadi Bani Habib" name="Wadi Bani Habib" pageId="23" pageNumber="24">Wadi Bani Habib</location>
,
<quantity id="4CB27EA1C528FFF6F228F94DFEFEF962" box="[292,379,1760,1784]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="23" pageNumber="24" unit="m" value="2200.0">
<elevation id="00673477C528FFF6F228F94DFEFEF962" box="[292,379,1760,1784]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="23" pageNumber="24" unit="m" value="2200.0">2200 m</elevation>
</quantity>
, Jebel Akhdar (North
<collectingCountry id="F35D93D4C528FFF6F17EF94DFD3EF962" box="[626,699,1760,1784]" name="Oman" pageId="23" pageNumber="24">Oman</collectingCountry>
), 23.0711’
<geoCoordinate id="EE7EB583C528FFF6F023F94DFC3DF962" box="[815,952,1760,1784]" degrees="57.60417" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="57.60417">N 57.60417</geoCoordinate>
’E WGS84, collected in
<date id="FFF4F584C528FFF6F7C8F94DFAD8F962" box="[1220,1373,1760,1784]" pageId="23" pageNumber="24" value="2005-10">
<collectingDate id="EFB00C6CC528FFF6F7C8F94DFAD8F962" box="[1220,1373,1760,1784]" pageId="23" pageNumber="24" value="2005-10">October 2005</collectingDate>
</date>
by
<collectorName id="26BFB692C528FFF6F68AF94DFF7AF886" pageId="23" pageNumber="24">S. Carranza</collectorName>
,
<collectorName id="26BFB692C528FFF6F21EF8A9FE26F886" box="[274,419,1796,1820]" pageId="23" pageNumber="24">E.N. Arnold</collectorName>
and
<collectorName id="26BFB692C528FFF6F2E5F8A9FDE8F886" box="[489,621,1796,1820]" pageId="23" pageNumber="24">D. Donaire</collectorName>
(MorphoBank M94363–M94372);
<specimenCode id="DBEC7B3FC528FFF6F718F8A9FB09F886" box="[1044,1164,1796,1820]" collectionCode="IBES" pageId="23" pageNumber="24">IBES8068</specimenCode>
, female from Wadi al Khahafa,
<quantity id="4CB27EA1C528FFF6F20AF886FED5F8D8" box="[262,336,1835,1859]" metricMagnitude="2" metricUnit="m" metricValue="4.92" pageId="23" pageNumber="24" unit="m" value="492.0">
<elevation id="00673477C528FFF6F20AF886FED5F8D8" box="[262,336,1835,1859]" metricMagnitude="2" metricUnit="m" metricValue="4.92" pageId="23" pageNumber="24" unit="m" value="492.0">492 m</elevation>
</quantity>
, Jebel Akhdar (North
<collectingCountry id="F35D93D4C528FFF6F15CF886FD1DF8D9" box="[592,664,1835,1859]" name="Oman" pageId="23" pageNumber="24">Oman</collectingCountry>
), 23.07419’
<geoCoordinate id="EE7EB583C528FFF6F02CF886FC2FF8D9" box="[800,938,1835,1859]" degrees="57.12208" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="57.12208">N 57.12208</geoCoordinate>
’E WGS84, collected on the 10
<superScript id="7C3F7E0CC528FFF6F601F885FA9CF8AC" attach="left" box="[1293,1305,1832,1846]" fontSize="6" pageId="23" pageNumber="24">th</superScript>
of
<date id="FFF4F584C528FFF6F64DF886FF4AF8FD" pageId="23" pageNumber="24" value="2010-10">
<collectingDate id="EFB00C6CC528FFF6F64DF886FF4AF8FD" pageId="23" pageNumber="24" value="2010-10">October 2010</collectingDate>
</date>
by
<collectorName id="26BFB692C528FFF6F3F6F8E2FEFBF8FD" box="[250,382,1871,1895]" pageId="23" pageNumber="24">S. Carranza</collectorName>
and
<collectorName id="26BFB692C528FFF6F2BAF8E2FD95F8FD" box="[438,528,1871,1895]" pageId="23" pageNumber="24">F. Amat</collectorName>
(
<collectorName id="26BFB692C528FFF6F113F8E2FD52F8FD" box="[543,727,1871,1895]" pageId="23" pageNumber="24">MorphoBank M</collectorName>
100056–M100064);
<specimenCode id="DBEC7B3FC528FFF6F0B3F8E2FBB3F8FD" box="[959,1078,1871,1895]" collectionCode="IBES" pageId="23" pageNumber="24">IBES7771</specimenCode>
, female from
<locationDeviation id="9A47CA66C528FFF6F7D8F8E2FA1DF8FC" box="[1236,1432,1871,1895]" location="East of Hat" pageId="23" pageNumber="24">
<quantity id="4CB27EA1C528FFF6F7D8F8E2FA8BF8FD" box="[1236,1294,1871,1895]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="km" value="1.0">1 km</quantity>
East of Hat
</locationDeviation>
,
<quantity id="4CB27EA1C528FFF6F39BF8DEFF6BF810" box="[151,238,1907,1931]" metricMagnitude="3" metricUnit="m" metricValue="1.124" pageId="23" pageNumber="24" unit="m" value="1124.0">
<elevation id="00673477C528FFF6F39BF8DEFF6BF810" box="[151,238,1907,1931]" metricMagnitude="3" metricUnit="m" metricValue="1.124" pageId="23" pageNumber="24" unit="m" value="1124.0">1124 m</elevation>
</quantity>
, Jebel Akhdar (North
<collectingCountry id="F35D93D4C528FFF6F2E6F8DEFDB7F811" box="[490,562,1907,1931]" name="Oman" pageId="23" pageNumber="24">Oman</collectingCountry>
), 23.18292’
<geoCoordinate id="EE7EB583C528FFF6F1B5F8DEFCC6F811" box="[697,835,1907,1931]" degrees="57.41627" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="57.41627">N 57.41627</geoCoordinate>
’E WGS84, collected by
<collectorName id="26BFB692C528FFF6F757F8DEFB64F811" box="[1115,1249,1907,1931]" pageId="23" pageNumber="24">S. Carranza</collectorName>
, E. Gómez-Díaz and
<collectorName id="26BFB692C528FFF6F3CBF837FEA4F82B" box="[199,289,1945,1969]" pageId="23" pageNumber="24">F. Amat</collectorName>
on the 9
<superScript id="7C3F7E0CC528FFF6F288F83BFE15F83E" attach="left" box="[388,400,1942,1956]" fontSize="6" pageId="23" pageNumber="24">th</superScript>
of
<date id="FFF4F584C528FFF6F2BBF837FDACF82B" box="[439,553,1945,1970]" pageId="23" pageNumber="24" value="2011-05">
<collectingDate id="EFB00C6CC528FFF6F2BBF837FDACF82B" box="[439,553,1945,1970]" pageId="23" pageNumber="24" value="2011-05">May 2011</collectingDate>
</date>
(
<collectorName id="26BFB692C528FFF6F134F834FD74F82B" box="[568,753,1945,1969]" pageId="23" pageNumber="24">MorphoBank M</collectorName>
100065–M100073);
<specimenCode id="DBEC7B3FC528FFF6F0D4F834FBCAF82B" box="[984,1103,1945,1969]" collectionCode="IBES" pageId="23" pageNumber="24">IBES6085</specimenCode>
, female, same collecting data as IBES7771 (
<collectorName id="26BFB692C528FFF6F24EF810FE7BF84F" box="[322,510,1981,2005]" pageId="23" pageNumber="24">MorphoBank M</collectorName>
100083–
<accessionNumber id="94194EA7C528FFF6F16EF810FD41F84F" box="[610,708,1981,2005]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/M00093" pageId="23" pageNumber="24" type="EnaNcbi">M00093</accessionNumber>
);
<specimenCode id="DBEC7B3FC528FFF6F1D0F810FCE9F84F" box="[732,876,1981,2005]" collectionCode="ONHM" pageId="23" pageNumber="24">ONHM3705</specimenCode>
, female from
<collectingMunicipality id="6B91493EC528FFF6F71BF810FB63F84F" box="[1047,1254,1981,2005]" pageId="23" pageNumber="24">Wadi Bani Habib</collectingMunicipality>
,
<quantity id="4CB27EA1C528FFF6F7FAF810FAD7F84F" box="[1270,1362,1981,2005]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="23" pageNumber="24" unit="m" value="2200.0">
<elevation id="00673477C528FFF6F7FAF810FAD7F84F" box="[1270,1362,1981,2005]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="23" pageNumber="24" unit="m" value="2200.0">2200 m</elevation>
</quantity>
,
<collectorName id="26BFB692C528FFF9F66EF810FF6EFF35" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">Jebel Akhdar</collectorName>
(
<collectingCounty id="6294ABC8C527FFF9F3F4FF3AFEBEFF35" box="[248,315,151,175]" pageId="24" pageNumber="25">North</collectingCounty>
<collectingCountry id="F35D93D4C527FFF9F24EFF3AFE0EFF35" box="[322,395,151,175]" name="Oman" pageId="24" pageNumber="25">Oman</collectingCountry>
), 23.0711’
<geoCoordinate id="EE7EB583C527FFF9F2F3FF3AFD0CFF35" box="[511,649,151,175]" degrees="57.60417" direction="north" orientation="latitude" pageId="24" pageNumber="25" precision="1" value="57.60417">N 57.60417</geoCoordinate>
’E WGS84, collected in
<date id="FFF4F584C527FFF9F098FF3AFBA9FF35" box="[916,1068,151,175]" pageId="24" pageNumber="25" value="2005-10">
<collectingDate id="EFB00C6CC527FFF9F098FF3AFBA9FF35" box="[916,1068,151,175]" pageId="24" pageNumber="25" value="2005-10">October 2005</collectingDate>
</date>
by
<collectorName id="26BFB692C527FFF9F759FF3AFB5FFF35" box="[1109,1242,151,175]" pageId="24" pageNumber="25">S. Carranza</collectorName>
,
<collectorName id="26BFB692C527FFF9F7E8FF35FAE8FF35" box="[1252,1389,151,175]" pageId="24" pageNumber="25">E.N. Arnold</collectorName>
and
<collectorName id="26BFB692C527FFF9F39BFF11FE96FF4E" box="[151,275,188,212]" pageId="24" pageNumber="25">D. Donaire</collectorName>
(
<collectorName id="26BFB692C527FFF9F22EFF11FE3CFF4E" box="[290,441,188,212]" pageId="24" pageNumber="25">MorphoBank</collectorName>
M94363–M94372).
</materialsCitation>
</paragraph>
<paragraph id="8BF5D344C527FFF9F39BFEA9FE35FE86" blockId="24.[151,1435,260,356]" box="[151,432,260,284]" pageId="24" pageNumber="25">
<emphasis id="B93E0F56C527FFF9F39BFEA9FE35FE86" box="[151,432,260,284]" italics="true" pageId="24" pageNumber="25">Other material examined</emphasis>
</paragraph>
<paragraph id="8BF5D344C527FFF9F3CBFE85FB2AFEFE" blockId="24.[151,1435,260,356]" pageId="24" pageNumber="25">
Two vouchers listed in Appendix I under
<taxonomicName id="4C4AA8C7C527FFF9F1ACFE84FCB8FEDA" authority="Carranza &amp; Arnold, 2012" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[672,829,296,320]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="luqueorum" status="sp. nov.">
<emphasis id="B93E0F56C527FFF9F1ACFE84FCB8FEDA" box="[672,829,296,320]" italics="true" pageId="24" pageNumber="25">H. luqueorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DB22DC527FFF9F049FE85FC18FEA5" box="[837,925,296,319]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
and not mentioned above. Two unvouchered specimens (tissue codes S6080 and S7843) included in the molecular analyses only (Table 1).
</paragraph>
<paragraph id="8BF5D344C527FFF9F39BFE38FE8CFE36" blockId="24.[151,1436,405,937]" box="[151,265,405,428]" pageId="24" pageNumber="25">
<emphasis id="B93E0F56C527FFF9F39BFE38FE8CFE36" box="[151,265,405,428]" italics="true" pageId="24" pageNumber="25">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BF5D344C527FFF9F3CBFE1AFBA8FD6C" blockId="24.[151,1436,405,937]" pageId="24" pageNumber="25">
A large-sized
<taxonomicName id="4C4AA8C7C527FFF9F27EFE15FD89FE4A" authorityName="Oken" authorityYear="1817" box="[370,524,440,464]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="B93E0F56C527FFF9F27EFE15FD89FE4A" box="[370,524,440,464]" italics="true" pageId="24" pageNumber="25">Hemidactylus</emphasis>
</taxonomicName>
with a maximum recorded SVL of
<quantity id="4CB27EA1C527FFF9F0C1FE1AFBA3FE55" box="[973,1062,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="8.8" pageId="24" pageNumber="25" unit="mm" value="88.0">88 mm</quantity>
; with a mean of 14.2 (13–15) longitudinal rows of enlarged dorsal tubercles at mid-body; adhesive pads broad, in adults maximum width of pad on fourth toe of the pes more than half its length; lamellae under the 1
<superScript id="7C3F7E0CC527FFF9F0CAFE52FC55FD97" attach="left" box="[966,976,511,525]" fontSize="6" pageId="24" pageNumber="25">st</superScript>
toe of pes mean 10.3 (10–11); lamellae under the 4
<superScript id="7C3F7E0CC527FFF9F215FD8BFEA0FDAE" attach="left" box="[281,293,550,564]" fontSize="6" pageId="24" pageNumber="25">th</superScript>
toe of pes mean 13.6 (13–14); preanal pores mean 5.3 (5–6); expanded subcaudal scales extending proximally as far as the second whorl after the vent and starting just after the hemipenial bulge in males; dorsum grey–buff with irregular small spots; a dark stripe from the nostril, through the eye, on to cheek above ear and often on to neck; tail with small irregular dark blotches basally and numerous transverse dark bands more distally, the total number being around 17. Underside of tail pale but large subcaudals suffused with grey formed by dark chromatophores that increase in intensity distally; underside of toe pads also grey.
</paragraph>
<paragraph id="8BF5D344C527FFF9F3CBFCACFA02FC33" blockId="24.[151,1436,405,937]" pageId="24" pageNumber="25">
<taxonomicName id="4C4AA8C7C527FFF9F3CBFCACFE64FC83" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[199,481,769,793]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="luqueorum">
<emphasis id="B93E0F56C527FFF9F3CBFCACFE64FC83" box="[199,481,769,793]" italics="true" pageId="24" pageNumber="25">Hemidactylus luqueorum</emphasis>
</taxonomicName>
is generally similar to
<taxonomicName id="4C4AA8C7C527FFF9F1EFFCAFFCE7FC83" authorityName="Arnold and Gallagher" authorityYear="1977" box="[739,866,770,793]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B93E0F56C527FFF9F1EFFCAFFCE7FC83" box="[739,866,770,793]" italics="true" pageId="24" pageNumber="25">H. persicus</emphasis>
</taxonomicName>
in number of its moderately-sized dorsal tubercles across mid-body, and large adhesive pads on toes but differs from it in its much larger size (SVL mean
<quantity id="4CB27EA1C527FFF9F63CFC88FA12FCA7" box="[1328,1431,805,829]" metricMagnitude="-2" metricUnit="m" metricValue="7.68" pageId="24" pageNumber="25" unit="mm" value="76.8">76.8 mm</quantity>
, max.
<quantity id="4CB27EA1C527FFF9F3D9FCE4FEA2FCFB" box="[213,295,841,865]" metricMagnitude="-2" metricUnit="m" metricValue="8.8" pageId="24" pageNumber="25" unit="mm" value="88.0">88 mm</quantity>
, compared with mean.
<quantity id="4CB27EA1C527FFF9F121FCE4FD16FCFB" box="[557,659,841,865]" metricMagnitude="-2" metricUnit="m" metricValue="5.64" pageId="24" pageNumber="25" unit="mm" value="56.4">56.4 mm</quantity>
, max.
<quantity id="4CB27EA1C527FFF9F1D2FCE4FCB7FCFB" box="[734,818,841,865]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" pageId="24" pageNumber="25" unit="mm" value="67.0">67 mm</quantity>
), reduced number of preanal pores in males (mean 5.3, 5–6, compared with mean 9.2, 8–11), and presence of more lamellae under the first toe of pes (mean 10.3, 10–11, compared with mean 8.8, 8–9). For differences from the second North
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species of
<taxonomicName id="4C4AA8C7C527FFF9F77DFC3CFA8EFC33" authorityName="Oken" authorityYear="1817" box="[1137,1291,913,937]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="B93E0F56C527FFF9F77DFC3CFA8EFC33" box="[1137,1291,913,937]" italics="true" pageId="24" pageNumber="25">Hemidactylus</emphasis>
</taxonomicName>
see below.
</paragraph>
<paragraph id="8BF5D344C527FFF9F39BFC74FE8AFC6B" blockId="24.[151,1436,985,1117]" box="[151,271,985,1009]" pageId="24" pageNumber="25">
<emphasis id="B93E0F56C527FFF9F39BFC74FE8AFC6B" box="[151,271,985,1009]" italics="true" pageId="24" pageNumber="25">Etymology</emphasis>
</paragraph>
<paragraph id="8BF5D344C527FFF9F3CBFC50FD26FBC7" blockId="24.[151,1436,985,1117]" pageId="24" pageNumber="25">
The species epithet &quot;
<taxonomicName id="4C4AA8C7C527FFF9F2BCFC53FDACFB8C" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[432,553,1022,1046]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="luqueorum">
<emphasis id="B93E0F56C527FFF9F2BCFC53FDACFB8C" box="[432,553,1022,1046]" italics="true" pageId="24" pageNumber="25">luqueorum</emphasis>
</taxonomicName>
&quot; is a collective genitive plural Latin noun to honour Salvador Carranza’s wife, Maria Teresa Luque, and her family for all their love and support. Without their encouragement and help it would have been impossible to accomplish this work.
</paragraph>
<paragraph id="8BF5D344C527FFF9F39BFB2AFDC0FB05" blockId="24.[151,1436,1159,1651]" box="[151,581,1159,1183]" pageId="24" pageNumber="25">
<emphasis id="B93E0F56C527FFF9F39BFB2AFDC0FB05" box="[151,581,1159,1183]" italics="true" pageId="24" pageNumber="25">Genetic and phylogeographic remarks</emphasis>
</paragraph>
<paragraph id="8BF5D344C527FFF9F3CBFB06FDE1F9E9" blockId="24.[151,1436,1159,1651]" pageId="24" pageNumber="25">
<taxonomicName id="4C4AA8C7C527FFF9F3CBFB06FE66FB59" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[199,483,1195,1219]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="luqueorum">
<emphasis id="B93E0F56C527FFF9F3CBFB06FE66FB59" box="[199,483,1195,1219]" italics="true" pageId="24" pageNumber="25">Hemidactylus luqueorum</emphasis>
</taxonomicName>
is monophyletic in the phylogenetic analyses of Dataset 1 (
<figureCitation id="1371CFC1C527FFF9F798FB06FB6BFB59" box="[1172,1262,1195,1219]" captionStart="FIGURE 5" captionStartId="19.[151,244,1802,1823]" captionTargetBox="[294,1293,193,1791]" captionTargetId="figure-180@19.[294,1314,185,1793]" captionTargetPageId="19" captionText="FIGURE 5. Maximum likelihood (ML) phylogenetic tree of 131 Hemidactylus specimens of the Arid clade (Carranza &amp; Arnold, 2006) based on 2866 bp of concatenated sequences of four mitochondrial (12S, cyb, nd4 and tRNAs) and three nuclear (c-mos, mc1r and rag2) genes. Two specimens of H. flaviviridis were used to root the tree and have not been included in the figure. Tree topology and branch lengths are for the sampled tree with the highest likelihood by RaxML (100 searches, log likelihood = -20540.526926). Each sequence is labeled with the specimen code and taxa name (see Table 1). Maximum-likelihood bootstrap support values above 70% are indicated above branches and black dots by the nodes indicate a posterior probability value ≥ 0.95 in the Bayesian analysis. Age estimates inferred with BEAST are indicated in italics below some relevant nodes and include the mean and, between brackets, the HPD 95% confidence interval. Silhouettes of Hemidactylus taxa are not size-scaled." pageId="24" pageNumber="25">Fig. 5A</figureCitation>
) and Dataset 3 (Appendix
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). In both phylogenetic trees it forms a clade together with
<taxonomicName id="4C4AA8C7C527FFF9F0D6FB7DFBDFFB7D" authorityName="Arnold and Gallagher" authorityYear="1977" box="[986,1114,1232,1255]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B93E0F56C527FFF9F0D6FB7DFBDFFB7D" box="[986,1114,1232,1255]" italics="true" pageId="24" pageNumber="25">H. persicus</emphasis>
</taxonomicName>
and the second North
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species of
<taxonomicName id="4C4AA8C7C527FFF9F200FB5EFE23FA91" authorityName="Oken" authorityYear="1817" box="[268,422,1267,1291]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="B93E0F56C527FFF9F200FB5EFE23FA91" box="[268,422,1267,1291]" italics="true" pageId="24" pageNumber="25">Hemidactylus</emphasis>
</taxonomicName>
described below, although the bootstrap support and pp values are very low (see
<figureCitation id="1371CFC1C527FFF9F620FB5EFAE8FA91" box="[1324,1389,1267,1291]" captionStart="FIGURE 5" captionStartId="19.[151,244,1802,1823]" captionTargetBox="[294,1293,193,1791]" captionTargetId="figure-180@19.[294,1314,185,1793]" captionTargetPageId="19" captionText="FIGURE 5. Maximum likelihood (ML) phylogenetic tree of 131 Hemidactylus specimens of the Arid clade (Carranza &amp; Arnold, 2006) based on 2866 bp of concatenated sequences of four mitochondrial (12S, cyb, nd4 and tRNAs) and three nuclear (c-mos, mc1r and rag2) genes. Two specimens of H. flaviviridis were used to root the tree and have not been included in the figure. Tree topology and branch lengths are for the sampled tree with the highest likelihood by RaxML (100 searches, log likelihood = -20540.526926). Each sequence is labeled with the specimen code and taxa name (see Table 1). Maximum-likelihood bootstrap support values above 70% are indicated above branches and black dots by the nodes indicate a posterior probability value ≥ 0.95 in the Bayesian analysis. Age estimates inferred with BEAST are indicated in italics below some relevant nodes and include the mean and, between brackets, the HPD 95% confidence interval. Silhouettes of Hemidactylus taxa are not size-scaled." pageId="24" pageNumber="25">Fig. 5</figureCitation>
and Appendix
<collectionCode id="ED5B4B81C527FFF9F200FABAFEA9FAB5" box="[268,300,1303,1327]" pageId="24" pageNumber="25">III</collectionCode>
). According to the results of the dating analysis inferred with Dataset 2, these three species split about 12.6 mya (95% HPD: 7.7–17.9). According to
<figureCitation id="1371CFC1C527FFF9F1A1FA96FD75FAC9" box="[685,752,1339,1363]" captionStart="FIGURE 5" captionStartId="19.[151,244,1802,1823]" captionTargetBox="[294,1293,193,1791]" captionTargetId="figure-180@19.[294,1314,185,1793]" captionTargetPageId="19" captionText="FIGURE 5. Maximum likelihood (ML) phylogenetic tree of 131 Hemidactylus specimens of the Arid clade (Carranza &amp; Arnold, 2006) based on 2866 bp of concatenated sequences of four mitochondrial (12S, cyb, nd4 and tRNAs) and three nuclear (c-mos, mc1r and rag2) genes. Two specimens of H. flaviviridis were used to root the tree and have not been included in the figure. Tree topology and branch lengths are for the sampled tree with the highest likelihood by RaxML (100 searches, log likelihood = -20540.526926). Each sequence is labeled with the specimen code and taxa name (see Table 1). Maximum-likelihood bootstrap support values above 70% are indicated above branches and black dots by the nodes indicate a posterior probability value ≥ 0.95 in the Bayesian analysis. Age estimates inferred with BEAST are indicated in italics below some relevant nodes and include the mean and, between brackets, the HPD 95% confidence interval. Silhouettes of Hemidactylus taxa are not size-scaled." pageId="24" pageNumber="25">Fig. 5</figureCitation>
and Appendix
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,
<taxonomicName id="4C4AA8C7C527FFF9F0C0FA96FB62FAC9" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[972,1255,1339,1363]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="luqueorum">
<emphasis id="B93E0F56C527FFF9F0C0FA96FB62FAC9" box="[972,1255,1339,1363]" italics="true" pageId="24" pageNumber="25">Hemidactylus luqueorum</emphasis>
</taxonomicName>
is more closely related to the second North
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species of
<emphasis id="B93E0F56C527FFF9F19EFAF2FCA9FAED" box="[658,812,1375,1399]" italics="true" pageId="24" pageNumber="25">Hemidacytlus</emphasis>
described below (
<figureCitation id="1371CFC1C527FFF9F0F6FAF2FBD4FAED" box="[1018,1105,1375,1399]" captionStart="FIGURE 5" captionStartId="19.[151,244,1802,1823]" captionTargetBox="[294,1293,193,1791]" captionTargetId="figure-180@19.[294,1314,185,1793]" captionTargetPageId="19" captionText="FIGURE 5. Maximum likelihood (ML) phylogenetic tree of 131 Hemidactylus specimens of the Arid clade (Carranza &amp; Arnold, 2006) based on 2866 bp of concatenated sequences of four mitochondrial (12S, cyb, nd4 and tRNAs) and three nuclear (c-mos, mc1r and rag2) genes. Two specimens of H. flaviviridis were used to root the tree and have not been included in the figure. Tree topology and branch lengths are for the sampled tree with the highest likelihood by RaxML (100 searches, log likelihood = -20540.526926). Each sequence is labeled with the specimen code and taxa name (see Table 1). Maximum-likelihood bootstrap support values above 70% are indicated above branches and black dots by the nodes indicate a posterior probability value ≥ 0.95 in the Bayesian analysis. Age estimates inferred with BEAST are indicated in italics below some relevant nodes and include the mean and, between brackets, the HPD 95% confidence interval. Silhouettes of Hemidactylus taxa are not size-scaled." pageId="24" pageNumber="25">Fig. 5B</figureCitation>
, Appendix
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), from which it split approximately 9.6 mya (95% HPD: 5.7–13.8). The level of genetic variability within
<taxonomicName id="4C4AA8C7C527FFF9F792FA29FABCFA01" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[1182,1337,1411,1435]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="luqueorum">
<emphasis id="B93E0F56C527FFF9F792FA29FABCFA01" box="[1182,1337,1411,1435]" italics="true" pageId="24" pageNumber="25">H. luqueorum</emphasis>
</taxonomicName>
is 2% in the
<emphasis id="B93E0F56C527FFF9F3B3FA0AFF6BFA25" box="[191,238,1447,1471]" italics="true" pageId="24" pageNumber="25">cytb</emphasis>
and 0.2% in the
<emphasis id="B93E0F56C527FFF9F2A4FA0AFE57FA25" box="[424,466,1447,1471]" italics="true" pageId="24" pageNumber="25">12S</emphasis>
. The uncorrected genetic distances between
<taxonomicName id="4C4AA8C7C527FFF9F0C8FA05FBDBFA25" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[964,1118,1447,1471]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="luqueorum">
<emphasis id="B93E0F56C527FFF9F0C8FA05FBDBFA25" box="[964,1118,1447,1471]" italics="true" pageId="24" pageNumber="25">H. luqueorum</emphasis>
</taxonomicName>
and the second North
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species of
<taxonomicName id="4C4AA8C7C527FFF9F21CFA66FE2FFA79" authorityName="Oken" authorityYear="1817" box="[272,426,1483,1507]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="genus">
<emphasis id="B93E0F56C527FFF9F21CFA66FE2FFA79" box="[272,426,1483,1507]" italics="true" pageId="24" pageNumber="25">Hemidactylus</emphasis>
</taxonomicName>
described below are 15.6% in the
<emphasis id="B93E0F56C527FFF9F035FA66FCEDFA79" box="[825,872,1483,1507]" italics="true" pageId="24" pageNumber="25">cytb</emphasis>
and 6.9% in the
<emphasis id="B93E0F56C527FFF9F721FA66FBD2FA79" box="[1069,1111,1483,1507]" italics="true" pageId="24" pageNumber="25">12S</emphasis>
; and between
<taxonomicName id="4C4AA8C7C527FFF9F7F3FA61FA19FA79" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[1279,1436,1483,1507]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="luqueorum">
<emphasis id="B93E0F56C527FFF9F7F3FA61FA19FA79" box="[1279,1436,1483,1507]" italics="true" pageId="24" pageNumber="25">H. luqueorum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4AA8C7C527FFF9F3CBFA5DFEC2F99D" authorityName="Arnold and Gallagher" authorityYear="1977" box="[199,327,1520,1543]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B93E0F56C527FFF9F3CBFA5DFEC2F99D" box="[199,327,1520,1543]" italics="true" pageId="24" pageNumber="25">H. persicus</emphasis>
</taxonomicName>
are 14.6% in the
<emphasis id="B93E0F56C527FFF9F11DFA42FDC5F99D" box="[529,576,1519,1543]" italics="true" pageId="24" pageNumber="25">cytb</emphasis>
and 9.8% in the
<emphasis id="B93E0F56C527FFF9F00DFA42FCAEF99D" box="[769,811,1519,1543]" italics="true" pageId="24" pageNumber="25">12S</emphasis>
. The results of the nuclear networks presented in
<figureCitation id="1371CFC1C527FFF9F66DFA42FA19F99D" box="[1377,1436,1519,1543]" captionStart="FIGURE 6" captionStartId="20.[151,244,1945,1966]" captionTargetBox="[192,1390,199,1918]" captionTargetId="figure-83@20.[191,1396,191,1928]" captionTargetPageId="20" captionText="FIGURE 6. Statistical parsimony nuclear allele networks of c-mos, mc1r and rag2 loci with colors corresponding to species H. persicus, H. luqueorum sp. nov. and H. hajarensis sp. nov.. Circle sizes are proportional to the number of alleles. White circles represent mutational steps. B1 and B2 refer to the different genetic lineages found within H. hajarensis (see Fig. 5 and Appendix III)." pageId="24" pageNumber="25">Fig 6</figureCitation>
and a network analysis including all specimens from Dataset 1 (data not shown) indicate that all alleles of
<taxonomicName id="4C4AA8C7C527FFF9F68DF9B9FE95F9D5" authorityName="Carranza &amp; Arnold" authorityYear="2012" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="luqueorum">
<emphasis id="B93E0F56C527FFF9F68DF9B9FE95F9D5" italics="true" pageId="24" pageNumber="25">H. luqueorum</emphasis>
</taxonomicName>
for all three independent loci analyzed (
<emphasis id="B93E0F56C527FFF9F1EDF995FCA1F9D5" box="[737,804,1592,1615]" italics="true" pageId="24" pageNumber="25">c-mos</emphasis>
,
<emphasis id="B93E0F56C527FFF9F038F99AFCE9F9D5" box="[820,876,1591,1615]" italics="true" pageId="24" pageNumber="25">mc1r</emphasis>
and
<emphasis id="B93E0F56C527FFF9F0ABF99AFC5EF9D5" box="[935,987,1591,1615]" italics="true" pageId="24" pageNumber="25">rag2</emphasis>
) are private (not shared with any other species included in the present analyses).
</paragraph>
<paragraph id="8BF5D344C527FFF9F39BF931FEA5F92E" blockId="24.[151,1436,1692,1860]" box="[151,288,1692,1716]" pageId="24" pageNumber="25">
<emphasis id="B93E0F56C527FFF9F39BF931FEA5F92E" box="[151,288,1692,1716]" italics="true" pageId="24" pageNumber="25">Distribution</emphasis>
</paragraph>
<paragraph id="8BF5D344C527FFF9F3CBF96DFF69F8DE" blockId="24.[151,1436,1692,1860]" pageId="24" pageNumber="25">
Despite intensive surveys across the Hajar Mountain range and especially the Eastern Hajars,
<taxonomicName id="4C4AA8C7C527FFF9F7F2F96CFA19F942" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[1278,1436,1728,1752]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="luqueorum">
<emphasis id="B93E0F56C527FFF9F7F2F96CFA19F942" box="[1278,1436,1728,1752]" italics="true" pageId="24" pageNumber="25">H. luqueorum</emphasis>
</taxonomicName>
has only been found in the Jebel Akhdar, the largest structural domain of the Western Hajar Mountains in North
<collectingCountry id="F35D93D4C527FFF9F39BF8A5FF5EF8BA" box="[151,219,1800,1824]" name="Oman" pageId="24" pageNumber="25">Oman</collectingCountry>
(
<figureCitation id="1371CFC1C527FFF9F3E2F8A5FEBBF8BA" box="[238,318,1800,1824]" captionStart="FIGURE 1" captionStartId="5.[151,244,1755,1776]" captionTargetBox="[153,1433,205,1733]" captionTargetId="figure-148@5.[151,1436,203,1735]" captionTargetPageId="5" captionText="FIGURE 1. Schematic map of the study region showing the position of the Northern and Southern Oman mountains with the name of some relevant massifs." pageId="24" pageNumber="25">Figs. 1</figureCitation>
and
<figureCitation id="1371CFC1C527FFF9F277F8A5FE09F8BA" box="[379,396,1800,1824]" captionStart="FIGURE 2" captionStartId="13.[151,244,1580,1601]" captionTargetBox="[160,1436,197,1542]" captionTargetId="figure-120@13.[151,1436,191,1544]" captionTargetPageId="13" captionText="FIGURE 2. Distribution map of Hemidactylus luqueorum sp. nov., H. hajarensis sp. nov., H. alkiyumii sp. nov. and H. festivus sp. nov. Color dots indicate specimens included in the molecular analyses and listed in Table 1. Stars indicate Museum specimens included in the morphological analyses only and listed in Appendix I: 1) BMNH1975.916; 2) BMNH1975.41, BMNH1980.558; 3) BMNH1976.1404; 4) BMNH1977.35; 5) BMNH1983.706; 6) BMNH1977.975; 7) BMNH1977.976–981; 8) BMNH1977.963–966; 9) BMNH1977.956–959; 10) BMNH1977.972–973 and BMNH1976.1409. Dashed lines delimit the different genetic lineages found within H. hajarensis (B1 and B2) and H. alkiyumii (C1, C2 and C3) and shown in Fig. 5 and Appendix III." pageId="24" pageNumber="25">2</figureCitation>
). It has been recorded from
<quantity id="4CB27EA1C527FFF9F1D5F8A5FCA6F8BA" box="[729,803,1800,1824]" metricMagnitude="2" metricUnit="m" metricValue="4.92" pageId="24" pageNumber="25" unit="m" value="492.0">492 m</quantity>
altitude (Wadi Al Khahafa) up to
<quantity id="4CB27EA1C527FFF9F7BBF8A5FA8AF8BA" box="[1207,1295,1800,1824]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="24" pageNumber="25" unit="m" value="2200.0">2200 m</quantity>
(Wadi Bani Habib).
</paragraph>
<paragraph id="8BF5D344C527FFF9F39BF8C3FF67F81C" blockId="24.[151,1436,1902,2033]" box="[151,226,1902,1926]" pageId="24" pageNumber="25">
<emphasis id="B93E0F56C527FFF9F39BF8C3FF67F81C" box="[151,226,1902,1926]" italics="true" pageId="24" pageNumber="25">Habits</emphasis>
</paragraph>
<paragraph id="8BF5D344C527FFFAF3CBF83CFE06FE1D" blockId="24.[151,1436,1902,2033]" lastBlockId="27.[151,1436,151,391]" lastPageId="27" lastPageNumber="28" pageId="24" pageNumber="25">
The species occurs on rocky sides of wadis and on buildings and occasionally on gravely wadi floors. Mainly nocturnal, several specimens were active during the day in a narrow wadi
<quantity id="4CB27EA1C527FFF9F0D6F818FB91F857" box="[986,1044,1973,1997]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="25" unit="km" value="1.0">1 km</quantity>
East of Hat (
<figureCitation id="1371CFC1C527FFF9F7A6F818FA97F857" box="[1194,1298,1973,1997]" captionStart="FIGURE 11" captionStartId="28.[151,244,1906,1927]" captionTargetBox="[228,1358,191,1884]" captionTargetId="figure-118@28.[228,1358,191,1884]" captionTargetPageId="28" captionText="FIGURE 11. Different localities where H. luqueorum sp. nov. has been found A) 1 km E of Hat, Jebel Akhdar. In this locality H. luqueorum was found out during the day close to crevices and in small caves on the large boulders at the sides of the narrow Wadi; B) Wadi Bani Habib, Jebel Akhdar. In this locality H. luqueorum was found at night in man made constructions and on boulders; C) Wadi al Khahafa, Jebel Akhdar, where two specimens of H. luqueorum have been found at just 492 m altitude on the rocky sides of the wadi." pageId="24" pageNumber="25">Fig. 11A</figureCitation>
). According to
<bibRefCitation id="EFDBAEB5C527FFF9F3BFF874FE7EF86B" author="Arnold, E. N. &amp; Gallagher, M. D." box="[179,507,2009,2033]" pageId="24" pageNumber="25" pagination="59 - 80" refId="ref41599" refString="Arnold, E. N. &amp; Gallagher, M. D. (1977) The scientific results of the Oman flora and fauna survey 1975. Reptiles and amphibians from the mountains of northern Oman with special reference to the Jebel Akhdar region. Journal of Oman studies special report, 1, 59 - 80." type="journal article" year="1977">Arnold and Gallagher (1977)</bibRefCitation>
, specimens
<collectionCode id="ED5B4B81C527FFF9F18FF874FD5CF86B" box="[643,729,2009,2033]" pageId="24" pageNumber="25">BMNH</collectionCode>
1971.41 and
<collectionCode id="ED5B4B81C527FFF9F064F874FC38F86B" box="[872,957,2009,2033]" pageId="24" pageNumber="25">BMNH</collectionCode>
1975.916 were caught during the day, one on an overhanging rock-face and the other in a shallow cave. It can be locally abundant inside large caves and share habitat with
<taxonomicName id="4C4AA8C7C524FFFAF226FF11FDAAFF4E" authorityName="Arnold and Gardner" authorityYear="1994" box="[298,559,188,212]" class="Reptilia" family="Phyllodactylidae" genus="Asaccus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="platyrhynchus">
<emphasis id="B93E0F56C524FFFAF226FF11FDAAFF4E" box="[298,559,188,212]" italics="true" pageId="27" pageNumber="28">Asaccus platyrhynchus</emphasis>
</taxonomicName>
. In Wadi Bani Habib, at
<quantity id="4CB27EA1C524FFFAF051FF11FC32FF49" box="[861,951,188,212]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="27" pageNumber="28" unit="m" value="2000.0">2000 m</quantity>
, it has been found together with
<taxonomicName id="4C4AA8C7C524FFFAF64CFF11FE81FF62" authorityName="Gardner" authorityYear="1994" class="Reptilia" family="Phyllodactylidae" genus="Asaccus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="montanus">
<emphasis id="B93E0F56C524FFFAF64CFF11FE81FF62" italics="true" pageId="27" pageNumber="28">Asaccus montanus</emphasis>
</taxonomicName>
and at Wadi al Kahafa, at much lower altitude, it shares habitat with both
<taxonomicName id="4C4AA8C7C524FFFAF7AEFF4DFAE3FF62" authorityName="Arnold and Gardner" authorityYear="1994" box="[1186,1382,224,248]" class="Reptilia" family="Phyllodactylidae" genus="Asaccus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="platyrhynchus">
<emphasis id="B93E0F56C524FFFAF7AEFF4DFAE3FF62" box="[1186,1382,224,248]" italics="true" pageId="27" pageNumber="28">A. platyrhynchus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4AA8C7C524FFFAF39BFEA9FE3EFE86" baseAuthorityName="Donndorf" baseAuthorityYear="1798" box="[151,443,260,284]" class="Reptilia" family="Phyllodactylidae" genus="Ptyodactylus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="hasselquistii">
<emphasis id="B93E0F56C524FFFAF39BFEA9FE3EFE86" box="[151,443,260,284]" italics="true" pageId="27" pageNumber="28">Ptyodactylus hasselquistii</emphasis>
</taxonomicName>
. It moves relatively slowly and is quite confident, sometimes allowing one to approach quite closely to take pictures. It losses the skin very easily when being handled and sometimes specimens have large scars of regenerated skin on the back, probably as a results of fights with conspecifics or attacks from predators (
<figureCitation id="1371CFC1C524FFFAF202FEC2FEF2FE1D" box="[270,375,367,391]" captionStart="FIGURE 10" captionStartId="26.[151,244,1650,1671]" captionTargetBox="[151,1436,191,1628]" captionTargetId="figure-92@26.[151,1436,191,1628]" captionTargetPageId="26" captionText="FIGURE 10. Live specimens of H. luqueorum sp. nov. A) and B) male from Wadi al Khahafa, Jebel Akhdar (IBES8068); C) unvouchered specimen photographed in its natural habitat in a cave at approximately 1 km E of Hat, Jebel Akhdar (photograph by Felix Amat); D) detail of an area of regenerated skin on the back of an unvouchered specimen, probably as a results of fights with conspecifics or attacks from predators." pageId="27" pageNumber="28">Fig. 10D</figureCitation>
).
</paragraph>
<caption id="DF3583CCC526FFF8F39BF96FFD47F8A0" pageId="25" pageNumber="26" startId="25.[151,244,1730,1751]" targetBox="[151,1436,191,1708]" targetPageId="25">
<paragraph id="8BF5D344C526FFF8F39BF96FFD47F8A0" blockId="25.[151,1436,1730,1850]" pageId="25" pageNumber="26">
<emphasis id="B93E0F56C526FFF8F39BF96FFE89F94D" bold="true" box="[151,268,1730,1751]" pageId="25" pageNumber="26">FIGURE 9.</emphasis>
A) male, Holotype of
<taxonomicName id="4C4AA8C7C526FFF8F2E8F96EFDECF942" authority="Carranza &amp; Arnold, 2012" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[484,617,1731,1752]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="luqueorum" status="sp. nov.">
<emphasis id="B93E0F56C526FFF8F2E8F96EFDECF942" box="[484,617,1731,1752]" italics="true" pageId="25" pageNumber="26">H. luqueorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DB22DC526FFF8F163F96FFD3FF94D" box="[623,698,1730,1751]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
from Sayq, Jebel Akhdar (BMNH2005.1660); B) left: male of
<taxonomicName id="4C4AA8C7C526FFF8F61BF96EFE0EF96D" authority="Wadi Sayq, Jebel Akhdar" authorityName="Wadi Sayq, Jebel Akhdar" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="luqueorum">
<emphasis id="B93E0F56C526FFF8F61BF96EFA19F942" box="[1303,1436,1731,1752]" italics="true" pageId="25" pageNumber="26">H. luqueorum</emphasis>
Wadi Sayq, Jebel Akhdar
</taxonomicName>
(BMNH1980.558); right: female of
<taxonomicName id="4C4AA8C7C526FFF8F1E1F94EFCEBF962" authority="Carranza &amp; Arnold, 2012" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[749,878,1763,1784]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="hajarensis" status="sp. nov.">
<emphasis id="B93E0F56C526FFF8F1E1F94EFCEBF962" box="[749,878,1763,1784]" italics="true" pageId="25" pageNumber="26">H. hajarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DB22DC526FFF8F078F94FFC45F96D" box="[884,960,1762,1783]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
from Wadi Sabt, Jebel Akhdar (BMNH1977.35); C) above,
<taxonomicName id="4C4AA8C7C526FFF8F3F6F8AEFEFFF882" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[250,378,1795,1816]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="hajarensis">
<emphasis id="B93E0F56C526FFF8F3F6F8AEFEFFF882" box="[250,378,1795,1816]" italics="true" pageId="25" pageNumber="26">H. hajarensis</emphasis>
</taxonomicName>
(BMNH1977.35); below
<taxonomicName id="4C4AA8C7C526FFF8F17EF8AEFD72F882" box="[626,759,1795,1816]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="luqueorum">
<emphasis id="B93E0F56C526FFF8F17EF8AEFD72F882" box="[626,759,1795,1816]" italics="true" pageId="25" pageNumber="26">H. luqueorum</emphasis>
</taxonomicName>
(BMNH1980.558); D) underside of right hind feet; left:
<taxonomicName id="4C4AA8C7C526FFF8F61BF8AEFA19F882" box="[1303,1436,1795,1816]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="luqueorum">
<emphasis id="B93E0F56C526FFF8F61BF8AEFA19F882" box="[1303,1436,1795,1816]" italics="true" pageId="25" pageNumber="26">H. luqueorum</emphasis>
</taxonomicName>
(BMNH1980.558), right:
<taxonomicName id="4C4AA8C7C526FFF8F282F888FD8AF8A0" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[398,527,1829,1850]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="hajarensis">
<emphasis id="B93E0F56C526FFF8F282F888FD8AF8A0" box="[398,527,1829,1850]" italics="true" pageId="25" pageNumber="26">H. hajarensis</emphasis>
</taxonomicName>
(BMNH1977.35).
</paragraph>
</caption>
<caption id="DF3583CCC525FFFBF39BF9DFFD8EF945" pageId="26" pageNumber="27" startId="26.[151,244,1650,1671]" targetBox="[151,1436,191,1628]" targetPageId="26">
<paragraph id="8BF5D344C525FFFBF39BF9DFFD8EF945" blockId="26.[151,1436,1650,1759]" pageId="26" pageNumber="27">
<emphasis id="B93E0F56C525FFFBF39BF9DFFE99F91D" bold="true" box="[151,284,1650,1671]" pageId="26" pageNumber="27">FIGURE 10.</emphasis>
Live specimens of
<taxonomicName id="4C4AA8C7C525FFFBF2E5F9DEFDF7F912" authority="Carranza &amp; Arnold, 2012" authorityName="Carranza &amp; Arnold" authorityYear="2012" box="[489,626,1651,1672]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="luqueorum" status="sp. nov.">
<emphasis id="B93E0F56C525FFFBF2E5F9DEFDF7F912" box="[489,626,1651,1672]" italics="true" pageId="26" pageNumber="27">H. luqueorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DB22DC525FFFBF170F9DFFD4EF91D" box="[636,715,1650,1671]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
A) and B) male from Wadi al Khahafa, Jebel Akhdar (IBES8068); C) unvouchered specimen photographed in its natural habitat in a cave at approximately 1 km E of Hat, Jebel Akhdar (photograph by Felix Amat); D) detail of an area of regenerated skin on the back of an unvouchered specimen, probably as a results of fights with conspecifics or attacks from predators.
</paragraph>
</caption>
<paragraph id="8BF5D344C524FFFAF39BFE15FE9EFE55" blockId="27.[151,1436,440,1763]" box="[151,283,440,463]" pageId="27" pageNumber="28">
<emphasis id="B93E0F56C524FFFAF39BFE15FE9EFE55" box="[151,283,440,463]" italics="true" pageId="27" pageNumber="28">Description</emphasis>
</paragraph>
<paragraph id="8BF5D344C524FFFAF3CBFE71FB68FDC5" blockId="27.[151,1436,440,1763]" pageId="27" pageNumber="28">Head and body markedly depressed; head broad, especially posteriorly and neck well defined. Head length about 24–28% of SVL (mean males 25%, mean females 26%), head width 70–78% of head length (mean males 75%, mean females 74%), and head height 36–51% of head length (mean males 46%, mean females 44%). Adhesive pads broad; in adults maximum width of pad on fourth hind toe more than half its length.</paragraph>
<paragraph id="8BF5D344C524FFFAF3CBFDC1FB04FC3E" blockId="27.[151,1436,440,1763]" pageId="27" pageNumber="28">Nostril between rostral, supranasal and two superposed postnasals, with the first supralabial scale usually also entering narrowly into its border. One, occasionally 2 scales separating supranasals on midline. About 14–19 scales in a straight line from postnasal to edge of orbit. Small conical tubercles scattered in orbital area, crown of head and temporal area above the level of ear opening and immediately in front of the upper part of this. Ear opening with its longest axis running upwards and backwards, smooth-edged, usually half or more of eye diameter. Supralabial scales mean 11.8 (10–14), infralabials mean 9.1 (8–10). Mental scale broadly triangular posteriorly bordered by two large postmentals making contact behind it, a second pair of more lateral postmentals also present, all four with a smooth transverse posterior border, the postmentals contacting the first and second supralabials; second and more posterior infralabials bordered by more irregular and smaller enlarged scales. Gulars fine.</paragraph>
<paragraph id="8BF5D344C524FFFAF3CBFC02FCE8FA95" blockId="27.[151,1436,440,1763]" pageId="27" pageNumber="28">
Enlarged tubercles present on back, arranged in 14.2 (mean) (13–15) longitudinal rows at mid-body, which also form backwardly directed oblique rows from near midline to flank, 12–16 across mid-body, and
<quantity id="4CB27EA1C524FFFAF62EFC79FA02FC76" box="[1314,1415,980,1004]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="4.572" metricValueMin="4.064" pageId="27" pageNumber="28" unit="in" value="17.0" valueMax="18.0" valueMin="16.0">16–18 in</quantity>
a paravertebral row from the level of the axilla to that of the groin, where they are separated by spaces of about their own length. Enlarged tubercles keeled and trihedral but becoming smaller and more pointed on flanks. Ventrals small, but larger than dorsals and more imbricate, about
<quantity id="4CB27EA1C524FFFAF029FB92FC0EFBCD" box="[805,907,1087,1111]" metricMagnitude="0" metricUnit="m" metricValue="1.2446" metricValueMax="1.27" metricValueMin="1.2192" pageId="27" pageNumber="28" unit="in" value="49.0" valueMax="50.0" valueMin="48.0">48–50 in</quantity>
a transverse row at mid body between lateral folds. Males with 5–6 preanal pores (mean 5.3) separated by one or two scales giving a formula of 2+3, 3+2 or 3+3. Scales on upper forelimb small and imbricate, interspersed with enlarged tubercles on distal section. Scales on front of thigh and beneath about same size as belly scales and imbricate, rather larger under tibia, enlarged tubercles present on upper surface of both femur and tibia and also on posterior edge of foot. Lamellae under the toes of pes: 1
<superScript id="7C3F7E0CC524FFFAF227FB59FEB0FA98" attach="left" box="[299,309,1268,1282]" fontSize="6" pageId="27" pageNumber="28">st</superScript>
toe mean 10.3 (10–11); 4
<superScript id="7C3F7E0CC524FFFAF15AFB59FDE7FA98" attach="left" box="[598,610,1268,1282]" fontSize="6" pageId="27" pageNumber="28">th</superScript>
toe mean 13.6 (13–14).
</paragraph>
<paragraph id="8BF5D344C524FFFAF3CBFAB7FE40FA7D" blockId="27.[151,1436,440,1763]" pageId="27" pageNumber="28">Tail relatively slender, although sometimes thickened proximally; six enlarged, keeled and pointed tubercles on each whorl proximally, dropping to four around whorl 8 or 10. Tubercles about one third the length of basal whorls, becoming smaller and placed more posteriorly on whorls distally. About 10–11 small scales in longitudinal row on fourth whorl after vent, around seven small scales between tubercles on fourth and fifth whorls. Subcaudal scales enlarged and broad, extending proximally as far as the second whorl after the vent and starting just after the hemipenial bulge in males.</paragraph>
<paragraph id="8BF5D344C524FFFAF3CBFA5EFCC1F979" blockId="27.[151,1436,440,1763]" pageId="27" pageNumber="28">In spirit pale grey-buff; a dark stripe from the nostril, through the eye, on to cheek above ear and often on to neck; body with irregular small spots; some tubercles on forebody and vertebral area have opaque white coloring on one side and dark coloring on the other. Belly pale but there may be a slight stipple at the sides, the dark punctate spots being smaller than the scales. Tail with small irregular dark blotches basally and numerous transverse dark bands more distally, initially on every other whorl and then on each one, the total number being around 17. Underside of tail pale but large subcaudals suffused with grey formed by dark chromatophores that increase in intensity distally. Underside of toe pads also grey.</paragraph>
<paragraph id="8BF5D344C524FFFAF39BF8BEFE7EF8B1" blockId="27.[151,1435,1811,1909]" box="[151,507,1811,1835]" pageId="27" pageNumber="28">
<emphasis id="B93E0F56C524FFFAF39BF8BEFE7EF8B1" box="[151,507,1811,1835]" italics="true" pageId="27" pageNumber="28">
Distinctive features of
<typeStatus id="54F16DE6C524FFFAF298F8BEFE7EF8B1" box="[404,507,1811,1835]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BF5D344C524FFFAF3CBF89AFA0BF8EF" blockId="27.[151,1435,1811,1909]" pageId="27" pageNumber="28">
Male,
<quantity id="4CB27EA1C524FFFAF218F89AFEF8F8D4" box="[276,381,1847,1871]" metricMagnitude="-2" metricUnit="m" metricValue="8.04" pageId="27" pageNumber="28" unit="mm" value="80.4">80.4 mm</quantity>
SVL; tail truncated,
<quantity id="4CB27EA1C524FFFAF18CF89AFD56F8D4" box="[640,723,1847,1871]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" pageId="27" pageNumber="28" unit="mm" value="59.0">59 mm</quantity>
long. Supralabial scales 13/12; infralabials 9/9; 15 rows of enlarged tubercles at mid-back; 6 (3+3) preanal pores; lamellae under the 1
<superScript id="7C3F7E0CC524FFFAF0ECF8F7FC6FF8F2" attach="left" box="[992,1002,1882,1896]" fontSize="6" pageId="27" pageNumber="28">st</superScript>
toe of pes 10/11, 4
<superScript id="7C3F7E0CC524FFFAF7CCF8F7FB49F8F2" attach="left" box="[1216,1228,1882,1896]" fontSize="6" pageId="27" pageNumber="28">th</superScript>
toe of pes 14/13.
</paragraph>
</subSubSection>
</treatment>
</document>
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