treatments-xml/data/03/E3/29/03E3293EFFAEFF97FE7E6AED43618824.xml

<document id="564C4B8CD52090C56472BABFF0B5833D" ID-DOI="10.1080/00222933.2010.535914" ID-ISSN="1464-5262" ID-Zenodo-Dep="5203342" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629048039960" checkinUser="felipe" docAuthor="Zacharda, Miloslav, Isaia, Marco &amp; Piva, Erminio" docDate="2011" docId="03E3293EFFAEFF97FE7E6AED43618824" docLanguage="en" docName="JNatHist.45.11-12.641-666.pdf" docOrigin="Journal of Natural History 45 (11 - 12)" docSource="http://dx.doi.org/10.1080/00222933.2010.535914" docStyle="DocumentStyle:DE8A86AD4006EFA56F95B2AAEFBF2813.5:JNatHist.2008-2012.journal_article.2cover" docStyleId="DE8A86AD4006EFA56F95B2AAEFBF2813" docStyleName="JNatHist.2008-2012.journal_article.2cover" docStyleVersion="5" docTitle="Troglocheles lanai Zacharda &amp; Isaia &amp; Piva 2011, sp. nov." docType="treatment" docVersion="3" lastPageNumber="656" masterDocId="FFDA5146FFA4FF86FF9B6E15477E8D0F" masterDocTitle="New troglobitic species of the genus Troglocheles (Acari: Prostigmata: Rhagidiidae) from caves in northern Italy and Austria, with a key to adult species of the genus" masterLastPageNumber="666" masterPageNumber="641" pageNumber="649" updateTime="1699057451977" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DE6C341F1E2377BCEA5ABDAADC8D8289">New troglobitic species of the genus Troglocheles (Acari: Prostigmata: Rhagidiidae) from caves in northern Italy and Austria, with a key to adult species of the genus</mods:title>
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<mods:namePart id="5B6C704B8291080C00645D6047622B15">Zacharda, Miloslav</mods:namePart>
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<mods:namePart id="12EDB7A6E44B472D7E9F0B9C0406592D">Isaia, Marco</mods:namePart>
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<mods:title id="AF25761C0244C592C3E3970E74A9847A">Journal of Natural History</mods:title>
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<mods:date id="57F26D826347C27ACB760413158C7C22">2011</mods:date>
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<subSubSection id="C350CBA3FFAEFF8CFE7E6AED44FC881F" box="[485,898,1272,1296]" pageId="10" pageNumber="649" type="nomenclature">
<paragraph id="8BF59828FFAEFF8CFE7E6AED44FC881F" blockId="10.[485,898,1272,1328]" box="[485,898,1272,1296]" pageId="10" pageNumber="649">
<heading id="D0BD2F44FFAEFF8CFE7E6AED44FC881F" box="[485,898,1272,1296]" centered="true" fontSize="36" level="2" pageId="10" pageNumber="649" reason="7">
<taxonomicName id="4C4AE3ABFFAEFF8CFE7E6AED445D881F" ID-CoL="608dcb3b-a4aa-4ad8-84c5-621d003bd2a5" authority="Zacharda" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[485,803,1272,1296]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="10" pageNumber="649" phylum="Arthropoda" rank="species" species="lanai" status="sp. nov.">
<emphasis id="B93E443AFFAEFF8CFE7E6AED45D2881F" bold="true" box="[485,684,1272,1296]" italics="true" pageId="10" pageNumber="649">Troglocheles lanai</emphasis>
Zacharda
</taxonomicName>
, 
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<subSubSection id="C350CBA3FFAEFF97FDF26B0D43618824" lastPageId="17" lastPageNumber="656" pageId="10" pageNumber="649" type="description">
<paragraph id="8BF59828FFAEFF8CFDF26B0D4580883F" blockId="10.[485,898,1272,1328]" box="[617,766,1304,1328]" pageId="10" pageNumber="649">
(
<figureCitation id="137184ADFFAEFF8CFDEA6B0D4588883F" box="[625,758,1304,1328]" captionStart-0="Figure 6" captionStart-1="Figure 7" captionStart-2="Figure 8" captionStart-3="Figure 9" captionStartId-0="11.[144,211,1244,1266]" captionStartId-1="12.[213,280,1484,1506]" captionStartId-2="13.[144,211,1638,1660]" captionStartId-3="14.[213,280,1630,1652]" captionTargetBox-0="[142,1102,160,1171]" captionTargetBox-1="[210,1170,161,1448]" captionTargetBox-2="[142,1102,160,1567]" captionTargetBox-3="[209,1172,162,1596]" captionTargetId-0="figure-7@11.[142,1102,160,1171]" captionTargetId-1="figure-6@12.[210,1138,160,1441]" captionTargetId-2="figure-7@13.[142,1102,160,1567]" captionTargetId-3="figure-6@14.[209,1135,161,1597]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionText-0="Figure 6. Troglocheles lanai sp. nov., adult male; (A) dorsum; (B) venter (for setal notation see Description)." captionText-1="Figure 7. Troglocheles lanai sp. nov., adult female; (A,B) rhagidial organ I; (C) rhagidial organ II; (D) chelicera, lateral aspect; (E) palpus, lateral aspect; (F) subcapitulum, ventral aspect; (G) anomalous neotrichy on trochanter III; (H) elongated naso." captionText-2="Figure 8. Troglocheles lanai sp. nov., adult female, setal arrangement on legs, lateral aspect; (A) leg I – genu, tibia, tarsus; (B) leg I – trochanter, basi- and telofemur; (C,D) variations in positions of dorsodistal rhagidial Φ1 and spiniform Φ2 solenidia on tibia I, dorsal aspect; (E) leg II; (F,G) variations in positions of dorsodistal lanceolate Φ1 and spiniform Φ2 solenidia on tibia II, dorsal and lateral aspect, respectively; (ω, Φ, σ and ε designations for solenidia on tarsus, tibia, genu and for famulus, respectively)." captionText-3="Figure 9. Troglocheles lanai sp. nov., adult female, setal arrangement on legs, lateral aspect; (A) leg III, (B) apex of tarsus III with oar-like iteral (it) setae and two rhagidial solenidia, lateral aspect; (C) leg IV (ω, Φ, σ designations for solenidia on tarsus, tibia, genu, respectively)." figureDoi-0="http://doi.org/10.5281/zenodo.5203355" figureDoi-1="http://doi.org/10.5281/zenodo.5203357" figureDoi-2="http://doi.org/10.5281/zenodo.5203359" figureDoi-3="http://doi.org/10.5281/zenodo.5203361" httpUri-0="https://zenodo.org/record/5203355/files/figure.png" httpUri-1="https://zenodo.org/record/5203357/files/figure.png" httpUri-2="https://zenodo.org/record/5203359/files/figure.png" httpUri-3="https://zenodo.org/record/5203361/files/figure.png" pageId="10" pageNumber="649">Figures 6–9</figureCitation>
)
</paragraph>
<paragraph id="8BF59828FFAEFF8CFF4E6B6246D98880" blockId="10.[213,423,1399,1423]" box="[213,423,1399,1423]" pageId="10" pageNumber="649">
<emphasis id="B93E443AFFAEFF8CFF4E6B6246D98880" box="[213,423,1399,1423]" italics="true" pageId="10" pageNumber="649">Material examined</emphasis>
</paragraph>
<paragraph id="8BF59828FFAEFF8CFF4E6B8A45198B39" blockId="10.[213,1170,1439,1591]" pageId="10" pageNumber="649">
<materialsCitation id="3B229275FFAEFF8CFF4E6B8A45198B39" collectingDate="2004-08-21" collectionCode="CN, UTM" collectorName="E. Lana &amp; Ohio State University &amp; Type No." country="Italy" county="Adult" elevation="1050" latitude="44.263054" location="Valdieri" longLatPrecision="19" longitude="7.410278" municipality="Cuneo" pageId="10" pageNumber="649" specimenCode="T 373117, OSAL007450" specimenCount="1" stateProvince="Piemonte" typeStatus="holotype">
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<typeStatus id="54F1268AFFAEFF8CFF4E6B8A464088B8" box="[213,318,1439,1463]" pageId="10" pageNumber="649" type="holotype">Holotype</typeStatus>
.
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<collectingCounty id="6294E0A4FFAEFF8CFED16B8A46F288B8" box="[330,396,1439,1463]" pageId="10" pageNumber="649">Adult</collectingCounty>
female, 
<collectingCountry id="F35DD8B8FFAEFF8CFE636B8A455388B8" box="[504,557,1439,1463]" name="Italy" pageId="10" pageNumber="649">Italy</collectingCountry>
, 
<collectingRegion id="498E56CAFFAEFF8CFDA56B8A45D188B8" box="[574,687,1439,1463]" country="Italy" name="Piemonte" pageId="10" pageNumber="649">Piedmont</collectingRegion>
, 
<collectingMunicipality id="6B910252FFAEFF8CFD5A6B8A447088B8" box="[705,782,1439,1463]" pageId="10" pageNumber="649">Cuneo</collectingMunicipality>
, 
<location id="8E95CEF3FFAEFF8CFCBB6B8A440788B8" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFAEFF8CFCBB6B8A440788B8" box="[800,889,1439,1463]" country="Italy" county="Adult" latitude="44.263054" longLatPrecision="19" longitude="7.410278" municipality="Cuneo" name="Valdieri" pageId="10" pageNumber="649" stateProvince="Piemonte">Valdieri</location>
(1214 Pi 
<emphasis id="B93E443AFFAEFF8CFC726B8A448B88B8" box="[1001,1013,1439,1463]" italics="true" pageId="10" pageNumber="649">/</emphasis>
<collectionCode id="ED5B00EDFFAEFF8CFC6E6B8A435F88B8" box="[1013,1057,1439,1463]" country="United Kingdom" name="Wellcome Collection of Bacteria, Burroughs Wellcome Research Laboratories" pageId="10" pageNumber="649">CN</collectionCode>
), 
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dell’Infernotto, 
<location id="8E95CEF3FFAEFF8CFE106BAA450288D8" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFAEFF8CFE106BAA450288D8" box="[395,636,1471,1495]" country="Italy" county="Adult" latitude="44.263054" longLatPrecision="19" longitude="7.410278" municipality="Cuneo" name="Baron Litron-Maissa" pageId="10" pageNumber="649" stateProvince="Piemonte">Barôn Litrôn-Maissa</location>
6 
<location id="8E95CEF3FFAEFF8CFD066BAA45A888D8" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFAEFF8CFD066BAA45A888D8" box="[669,726,1471,1495]" country="Italy" county="Adult" latitude="44.263054" longLatPrecision="19" longitude="7.410278" municipality="Cuneo" name="Cave" pageId="10" pageNumber="649" stateProvince="Piemonte">Cave</location>
, universal transverse mercator (
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) coordinates: 32 
<specimenCode id="DBEC3053FFAEFF8CFE1E6BCA468E88F8" box="[389,496,1503,1527]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="649">T 373117</specimenCode>
4902365 (
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44 
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15 
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47 
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N
</geoCoordinate>
, 
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7 
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24 
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37 
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E
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), 
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<quantity id="4CB235CDFFAEFF8CFC006BCA448E88F8" box="[923,1008,1503,1527]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="10" pageNumber="649" unit="m" value="1050.0">1050 m</quantity>
a.s.l.
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, coll. 
<collectorName id="26BFFDFEFFAEFF8CFBE26BF5466D8B19" pageId="10" pageNumber="649">E. Lana</collectorName>
, 
<date id="FFF4BEE8FFAEFF8CFEBB6BEA46AA8B18" box="[288,468,1535,1559]" pageId="10" pageNumber="649" value="2004-08-21">
<collectingDate id="EFB04700FFAEFF8CFEBB6BEA46AA8B18" box="[288,468,1535,1559]" pageId="10" pageNumber="649" value="2004-08-21">21 August 2004</collectingDate>
</date>
, deposited in the 
<location id="8E95CEF3FFAEFF8CFD056BEA43728B18" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFAEFF8CFD056BEA43728B18" box="[670,1036,1535,1559]" country="Italy" county="Adult" latitude="44.263054" longLatPrecision="19" longitude="7.410278" municipality="Cuneo" name="Museum of Biological Diversity" pageId="10" pageNumber="649" stateProvince="Piemonte">Museum of Biological Diversity</location>
, 
<collectorName id="26BFFDFEFFAEFF8CFB836BEA46328B39" pageId="10" pageNumber="649">Ohio State University</collectorName>
, 
<collectorName id="26BFFDFEFFAEFF8CFECD680A46C18B39" box="[342,447,1567,1591]" pageId="10" pageNumber="649">Type No.</collectorName>
<specimenCode id="DBEC3053FFAEFF8CFE5E680B451C8B39" box="[453,610,1566,1590]" collectionCode="OSAL" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34094" lsid="urn:lsid:biocol.org:col:34094" name="Ohio State University Acarology Laboratory" pageId="10" pageNumber="649">OSAL007450</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF59828FFAEFF8CFF4E687743EF8B97" blockId="10.[213,1170,1632,1688]" pageId="10" pageNumber="649">
<materialsCitation id="3B229275FFAEFF8CFF4E687743F38B77" box="[213,1165,1632,1657]" collectingDate="2004-08-21" collectionCode="CN, UTM" collectorName="E. Lana &amp; Ohio State University &amp; Type No." country="Italy" county="Adult" elevation="1050" latitude="44.263054" location="Valdieri" longLatPrecision="19" longitude="7.410278" municipality="Cuneo" pageId="10" pageNumber="649" specimenCount="1" specimenCount-adult="1" stateProvince="Piemonte" typeStatus="paratype">
<emphasis id="B93E443AFFAEFF8CFF4E687746348B76" box="[213,330,1634,1657]" italics="true" pageId="10" pageNumber="649">
<typeStatus id="54F1268AFFAEFF8CFF4E6877463B8B76" box="[213,325,1634,1657]" pageId="10" pageNumber="649" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Deposited with the 
<typeStatus id="54F1268AFFAEFF8CFDAC687545E08B77" box="[567,670,1632,1656]" pageId="10" pageNumber="649" type="holotype">holotype</typeStatus>
; 
<specimenCount id="9D4C53A1FFAEFF8CFD37687444648B77" box="[684,794,1632,1656]" pageId="10" pageNumber="649" type="adult">two adult</specimenCount>
females, same data as holotype
</materialsCitation>
, 
<materialsCitation id="3B229275FFAEFF8CFF4E689543EF8B97" box="[213,1169,1664,1688]" collectingDate="2003-04-13" collectionCode="OSAL" collectorName="E. Lana" country="Italy" county="Adult" elevation="1050" latitude="44.263054" location="Valdieri" longLatPrecision="19" longitude="7.410278" municipality="Cuneo" pageId="10" pageNumber="649" specimenCode="OSAL007448, OSAL007449" specimenCount="1" stateProvince="Piemonte" typeStatus="paratype">
coll. 
<collectorName id="26BFFDFEFFAEFF8CFE90689446168B97" box="[267,360,1665,1688]" pageId="10" pageNumber="649">E. Lana</collectorName>
, 
<date id="FFF4BEE8FFAEFF8CFEE9689545758B97" box="[370,523,1664,1688]" pageId="10" pageNumber="649" value="2003-04-13">
<collectingDate id="EFB04700FFAEFF8CFEE9689545758B97" box="[370,523,1664,1688]" pageId="10" pageNumber="649" value="2003-04-13">13 April 2003</collectingDate>
</date>
, 
<specimenCode id="DBEC3053FFAEFF8CFD8E689545D18B97" box="[533,687,1664,1688]" collectionCode="OSAL" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34094" lsid="urn:lsid:biocol.org:col:34094" name="Ohio State University Acarology Laboratory" pageId="10" pageNumber="649">OSAL007448</specimenCode>
and 
<specimenCode id="DBEC3053FFAEFF8CFD7E689544FF8B97" box="[741,897,1664,1688]" collectionCode="OSAL" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34094" lsid="urn:lsid:biocol.org:col:34094" name="Ohio State University Acarology Laboratory" pageId="10" pageNumber="649">OSAL007449</specimenCode>
; adult female, same data
</materialsCitation>
</paragraph>
<paragraph id="8BF59828FFAFFF8DFE236A8B469689B6" blockId="11.[440,488,1182,1209]" box="[440,488,1182,1209]" pageId="11" pageNumber="650">A,B</paragraph>
<caption id="DF35C8A0FFAFFF8DFF0B6AC9466F8800" ID-DOI="http://doi.org/10.5281/zenodo.5203355" ID-Zenodo-Dep="5203355" httpUri="https://zenodo.org/record/5203355/files/figure.png" pageId="11" pageNumber="650" startId="11.[144,211,1244,1266]" targetBox="[142,1102,160,1171]" targetPageId="11">
<paragraph id="8BF59828FFAFFF8DFF0B6AC9466F8800" blockId="11.[144,1101,1244,1295]" pageId="11" pageNumber="650">
Figure 6. 
<taxonomicName id="4C4AE3ABFFAFFF8DFF636AC946D589FD" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[248,427,1244,1266]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="11" pageNumber="650" phylum="Arthropoda" rank="species" species="lanai" status="sp. nov.">
<emphasis id="B93E443AFFAFFF8DFF636AC946D589FD" box="[248,427,1244,1266]" italics="true" pageId="11" pageNumber="650">Troglocheles lanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFAFFF8DFE296AC8457F89FD" box="[434,513,1245,1266]" pageId="11" pageNumber="650" rank="species">sp. nov.</taxonomicNameLabel>
, adult male; (A) dorsum; (B) venter (for setal notation see 
<emphasis id="B93E443AFFAFFF8DFF0B6AEF467D8800" box="[144,259,1274,1295]" italics="true" pageId="11" pageNumber="650">Description</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BF59828FFAFFF8AFF0B6B7743EC8B97" blockId="11.[144,1101,1377,1688]" lastBlockId="12.[213,1170,1601,1688]" lastPageId="12" lastPageNumber="651" pageId="11" pageNumber="650">
<materialsCitation id="3B229275FFAFFF8DFF0B6B7744BA8876" box="[144,964,1377,1401]" collectingDate="2006-12-03" collectionCode="OSAL" collectorName="M. Isaia &amp; E. Lana" pageId="11" pageNumber="650" specimenCode="OSAL007447" specimenCount="1" typeStatus="holotype">
as 
<typeStatus id="54F1268AFFAFFF8DFF346B74466A8876" box="[175,276,1377,1401]" pageId="11" pageNumber="650" type="holotype">holotype</typeStatus>
, coll. 
<collectorName id="26BFFDFEFFAFFF8DFECD6B7746CA8876" box="[342,436,1377,1401]" pageId="11" pageNumber="650">M. Isaia</collectorName>
and 
<collectorName id="26BFFDFEFFAFFF8DFE766B7745358876" box="[493,587,1378,1401]" pageId="11" pageNumber="650">E. Lana</collectorName>
, 
<date id="FFF4BEE8FFAFFF8DFDCD6B7444638876" box="[598,797,1377,1401]" pageId="11" pageNumber="650" value="2006-12-03">
<collectingDate id="EFB04700FFAFFF8DFDCD6B7444638876" box="[598,797,1377,1401]" pageId="11" pageNumber="650" value="2006-12-03">3 December 2006</collectingDate>
</date>
, 
<specimenCode id="DBEC3053FFAFFF8DFCB36B7444BA8876" box="[808,964,1377,1401]" collectionCode="OSAL" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34094" lsid="urn:lsid:biocol.org:col:34094" name="Ohio State University Acarology Laboratory" pageId="11" pageNumber="650">OSAL007447</specimenCode>
</materialsCitation>
; 
<materialsCitation id="3B229275FFAFFF8DFC546B7443378896" collectingDate="2009-05-14" collectionCode="OSAL" collectorName="M. Isaia &amp; M. Paschetta" pageId="11" pageNumber="650" specimenCode="OSAL007446" specimenCount="1" typeStatus="holotype">
adult male, same data as holotype, coll. 
<collectorName id="26BFFDFEFFAFFF8DFE4B6B9745518896" box="[464,559,1409,1433]" pageId="11" pageNumber="650">M. Isaia</collectorName>
and 
<collectorName id="26BFFDFEFFAFFF8DFDF36B97447E8896" box="[616,768,1409,1433]" pageId="11" pageNumber="650">M. Paschetta</collectorName>
, 
<date id="FFF4BEE8FFAFFF8DFC976B9444DF8896" box="[780,929,1409,1434]" pageId="11" pageNumber="650" value="2009-05-14">
<collectingDate id="EFB04700FFAFFF8DFC976B9444DF8896" box="[780,929,1409,1434]" pageId="11" pageNumber="650" value="2009-05-14">14 May 2009</collectingDate>
</date>
, 
<specimenCode id="DBEC3053FFAFFF8DFC376B9443378896" box="[940,1097,1409,1433]" collectionCode="OSAL" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34094" lsid="urn:lsid:biocol.org:col:34094" name="Ohio State University Acarology Laboratory" pageId="11" pageNumber="650">OSAL007446</specimenCode>
</materialsCitation>
; 
<materialsCitation id="3B229275FFAFFF8DFF0B6BB4468388D6" collectionCode="CN, UTM" collectorName="Valle Corsaglia" county="Cuneo" location="Pi" municipality="Frabosa Soprana" pageId="11" pageNumber="650" specimenCount="1" stateProvince="Piemonte" typeStatus="holotype">
adult female, 
<collectingRegion id="498E56CAFFAFFF8DFEAD6BB446D988B6" box="[310,423,1441,1465]" country="Italy" name="Piemonte" pageId="11" pageNumber="650">Piedmont</collectingRegion>
, 
<collectingCounty id="6294E0A4FFAFFF8DFE236BB4457B88B6" box="[440,517,1441,1465]" pageId="11" pageNumber="650">Cuneo</collectingCounty>
, 
<collectingMunicipality id="6B910252FFAFFF8DFD8D6BB445A088B6" box="[534,734,1441,1465]" pageId="11" pageNumber="650">Frabosa Soprana</collectingMunicipality>
(108 
<location id="8E95CEF3FFAFFF8DFCB16BB4443F88B6" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFAFFF8DFCB16BB4443F88B6" box="[810,833,1441,1465]" county="Cuneo" municipality="Frabosa Soprana" name="Pi" pageId="11" pageNumber="650" stateProvince="Piemonte">Pi</location>
<emphasis id="B93E443AFFAFFF8DFCDA6BB7443388B5" box="[833,845,1442,1466]" italics="true" pageId="11" pageNumber="650">/</emphasis>
<collectionCode id="ED5B00EDFFAFFF8DFCD56BB4440488B6" box="[846,890,1441,1465]" country="United Kingdom" name="Wellcome Collection of Bacteria, Burroughs Wellcome Research Laboratories" pageId="11" pageNumber="650">CN</collectionCode>
), 
<collectorName id="26BFFDFEFFAFFF8DFC0F6BB4433788B6" box="[916,1097,1441,1465]" pageId="11" pageNumber="650">Valle Corsaglia</collectorName>
, 
<location id="8E95CEF3FFAFFF8DFF0B6BD446D688D6" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFAFFF8DFF0B6BD446D688D6" box="[144,424,1473,1497]" county="Cuneo" municipality="Frabosa Soprana" name="Grotta di Bossea Cave" pageId="11" pageNumber="650" stateProvince="Piemonte">Grotta di Bossea Cave</location>
, 
<collectionCode id="ED5B00EDFFAFFF8DFE206BD7468388D6" box="[443,509,1474,1497]" pageId="11" pageNumber="650">UTM</collectionCode>
</materialsCitation>
<materialsCitation id="3B229275FFAFFF8DFD976BD4444F88D6" box="[524,817,1473,1497]" collectionCode="T" pageId="11" pageNumber="650" specimenCode="T 407464" specimenCount="1" stateProvince="Piemonte" typeStatus="holotype">
coordinates: 32 
<specimenCode id="DBEC3053FFAFFF8DFD5A6BD4444F88D6" box="[705,817,1473,1497]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="650">T 407464</specimenCode>
</materialsCitation>
<materialsCitation id="3B229275FFAFFF8DFCDB6BD4437F88F6" collectingDate="2001-09-23" collectionCode="OSAL" collectorName="E. Lana" elevation="836" latitude="44.24278" longLatPrecision="19" longitude="7.8408337" pageId="11" pageNumber="650" specimenCode="OSAL007445" specimenCount="1" stateProvince="Piemonte" typeStatus="holotype">
4899559 (
<geoCoordinate id="EE7EFEEFFFAFFF8DFC2F6BD4433688D6" box="[948,1096,1469,1497]" degrees="44" direction="north" minutes="14" orientation="latitude" pageId="11" pageNumber="650" precision="15" seconds="34" value="44.24278">
44 
<superScript id="7C3F3560FFAFFF8DFC4B6BA844A788C1" attach="none" box="[976,985,1469,1486]" fontSize="7" pageId="11" pageNumber="650">◦</superScript>
14 
<superScript id="7C3F3560FFAFFF8DFC6E6BA8448488C1" attach="left" box="[1013,1018,1469,1486]" fontSize="7" pageId="11" pageNumber="650">′</superScript>
34 
<superScript id="7C3F3560FFAFFF8DFB8D6BA8435E88C1" attach="left" box="[1046,1056,1469,1486]" fontSize="7" pageId="11" pageNumber="650">′′</superScript>
N
</geoCoordinate>
, 
<geoCoordinate id="EE7EFEEFFFAFFF8DFF0B6BF7467188F6" box="[144,271,1501,1529]" degrees="7" direction="east" minutes="50" orientation="longitude" pageId="11" pageNumber="650" precision="15" seconds="27" value="7.8408337">
7 
<superScript id="7C3F3560FFAFFF8DFF066BC847D888E1" attach="none" box="[157,166,1501,1518]" fontSize="7" pageId="11" pageNumber="650">◦</superScript>
50 
<superScript id="7C3F3560FFAFFF8DFF586BC847B688E1" attach="none" box="[195,200,1501,1518]" fontSize="7" pageId="11" pageNumber="650">′</superScript>
27 
<superScript id="7C3F3560FFAFFF8DFF7F6BC8479088E1" attach="left" box="[228,238,1501,1518]" fontSize="7" pageId="11" pageNumber="650">′′</superScript>
E
</geoCoordinate>
), 
<elevation id="00677F1BFFAFFF8DFEB26BF446CD88F6" box="[297,435,1505,1529]" metricMagnitude="2" metricUnit="m" metricValue="8.36" pageId="11" pageNumber="650" unit="m" value="836.0">
<quantity id="4CB235CDFFAFFF8DFEB26BF4460D88F6" box="[297,371,1505,1529]" metricMagnitude="2" metricUnit="m" metricValue="8.36" pageId="11" pageNumber="650" unit="m" value="836.0">836 m</quantity>
a.s.l.
</elevation>
, coll. 
<collectorName id="26BFFDFEFFAFFF8DFD9B6BF7451D88F6" box="[512,611,1506,1529]" pageId="11" pageNumber="650">E. Lana</collectorName>
, 
<date id="FFF4BEE8FFAFFF8DFDE86BF4442B88F6" box="[627,853,1505,1529]" pageId="11" pageNumber="650" value="2001-09-23">
<collectingDate id="EFB04700FFAFFF8DFDE86BF4442B88F6" box="[627,853,1505,1529]" pageId="11" pageNumber="650" value="2001-09-23">23 September 2001</collectingDate>
</date>
, 
<specimenCode id="DBEC3053FFAFFF8DFCFD6BF4437F88F6" box="[870,1025,1505,1529]" collectionCode="OSAL" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34094" lsid="urn:lsid:biocol.org:col:34094" name="Ohio State University Acarology Laboratory" pageId="11" pageNumber="650">OSAL007445</specimenCode>
</materialsCitation>
; 
<materialsCitation id="3B229275FFAFFF8DFB896BF4461F8B36" collectionCode="CN" county="Cuneo" location="Pi" municipality="Frabosa Soprana" pageId="11" pageNumber="650" specimenCode="UTM 32, T 403600" specimenCount="1" stateProvince="Piemonte" typeStatus="holotype">
adult female, 
<collectingRegion id="498E56CAFFAFFF8DFF75681446218B16" box="[238,351,1537,1561]" country="Italy" name="Piemonte" pageId="11" pageNumber="650">Piedmont</collectingRegion>
, 
<collectingCounty id="6294E0A4FFAFFF8DFEF4681446C28B16" box="[367,444,1537,1561]" pageId="11" pageNumber="650">Cuneo</collectingCounty>
, 
<collectingMunicipality id="6B910252FFAFFF8DFE56681445E68B16" box="[461,664,1537,1561]" pageId="11" pageNumber="650">Frabosa Soprana</collectingMunicipality>
, (837 
<location id="8E95CEF3FFAFFF8DFD7C681445808B16" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFAFFF8DFD7C681445808B16" box="[743,766,1537,1561]" county="Cuneo" municipality="Frabosa Soprana" name="Pi" pageId="11" pageNumber="650" stateProvince="Piemonte">Pi</location>
<emphasis id="B93E443AFFAFFF8DFD65681444748B16" box="[766,778,1537,1561]" italics="true" pageId="11" pageNumber="650">/</emphasis>
<collectionCode id="ED5B00EDFFAFFF8DFC90681444498B16" box="[779,823,1537,1561]" country="United Kingdom" name="Wellcome Collection of Bacteria, Burroughs Wellcome Research Laboratories" pageId="11" pageNumber="650">CN</collectionCode>
), 
<location id="8E95CEF3FFAFFF8DFCCA681443378B16" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFAFFF8DFCCA681443378B16" box="[849,1097,1537,1561]" county="Cuneo" municipality="Frabosa Soprana" name="Abisso Bacardi Cave" pageId="11" pageNumber="650" stateProvince="Piemonte">Abisso Bacardi Cave</location>
, 
<specimenCode id="DBEC3053FFAFFF8DFF0B683747898B36" box="[144,247,1569,1593]" collectionCode="UTM" pageId="11" pageNumber="650">UTM 32</specimenCode>
<specimenCode id="DBEC3053FFAFFF8DFF6C6834461F8B36" box="[247,353,1569,1593]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="650">T 403600</specimenCode>
</materialsCitation>
<materialsCitation id="3B229275FFAFFF8DFEF1683446538B56" collectingDate="1995-05-14" collectionCode="OSAL" collectorName="E. Lana" latitude="44.234444" longLatPrecision="19" longitude="7.7930555" pageId="11" pageNumber="650" specimenCode="OSAL007444" specimenCount="1" stateProvince="Piemonte" typeStatus="holotype">
4898640 (
<geoCoordinate id="EE7EFEEFFFAFFF8DFE43683445188B36" box="[472,614,1565,1593]" degrees="44" direction="north" minutes="14" orientation="latitude" pageId="11" pageNumber="650" precision="15" seconds="04" value="44.234444">
44 
<superScript id="7C3F3560FFAFFF8DFE6F680846838B21" attach="none" box="[500,509,1565,1582]" fontSize="7" pageId="11" pageNumber="650">◦</superScript>
14 
<superScript id="7C3F3560FFAFFF8DFD82680845608B21" attach="left" box="[537,542,1565,1582]" fontSize="7" pageId="11" pageNumber="650">′</superScript>
04 
<superScript id="7C3F3560FFAFFF8DFDA16808453A8B21" attach="left" box="[570,580,1565,1582]" fontSize="7" pageId="11" pageNumber="650">′′</superScript>
N
</geoCoordinate>
, 
<geoCoordinate id="EE7EFEEFFFAFFF8DFDEF6837458E8B36" box="[628,752,1565,1593]" degrees="7" direction="east" minutes="47" orientation="longitude" pageId="11" pageNumber="650" precision="15" seconds="35" value="7.7930555">
7 
<superScript id="7C3F3560FFAFFF8DFD19680845F58B21" attach="none" box="[642,651,1565,1582]" fontSize="7" pageId="11" pageNumber="650">◦</superScript>
47 
<superScript id="7C3F3560FFAFFF8DFD3C680845D28B21" attach="none" box="[679,684,1565,1582]" fontSize="7" pageId="11" pageNumber="650">′</superScript>
35 
<superScript id="7C3F3560FFAFFF8DFD53680845AC8B21" attach="left" box="[712,722,1565,1582]" fontSize="7" pageId="11" pageNumber="650">′′</superScript>
E
</geoCoordinate>
), coll. 
<collectorName id="26BFFDFEFFAFFF8DFCDA683744DF8B36" box="[833,929,1570,1593]" pageId="11" pageNumber="650">E. Lana</collectorName>
, 
<date id="FFF4BEE8FFAFFF8DFC34683443368B36" box="[943,1096,1569,1594]" pageId="11" pageNumber="650" value="1995-05-14">
<collectingDate id="EFB04700FFAFFF8DFC34683443368B36" box="[943,1096,1569,1594]" pageId="11" pageNumber="650" value="1995-05-14">14 May 1995</collectingDate>
</date>
, 
<specimenCode id="DBEC3053FFAFFF8DFF0B685446538B56" box="[144,301,1601,1625]" collectionCode="OSAL" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34094" lsid="urn:lsid:biocol.org:col:34094" name="Ohio State University Acarology Laboratory" pageId="11" pageNumber="650">OSAL007444</specimenCode>
</materialsCitation>
; 
<materialsCitation id="3B229275FFAFFF8DFEA26854463D8B97" collectingDate="1998-12-26" collectionCode="CN" collectorName="Valle Corsaglia &amp; E. Lana" county="Cuneo" location="Pi" municipality="Frabosa Soprana" pageId="11" pageNumber="650" specimenCount="1" stateProvince="Piemonte">
additional material examined: adult male, 
<collectingRegion id="498E56CAFFAFFF8DFC80685444F28B56" box="[795,908,1601,1625]" country="Italy" name="Piemonte" pageId="11" pageNumber="650">Piedmont</collectingRegion>
, 
<collectingCounty id="6294E0A4FFAFFF8DFC036854449B8B56" box="[920,997,1601,1625]" pageId="11" pageNumber="650">Cuneo</collectingCounty>
, 
<collectingMunicipality id="6B910252FFAFFF8DFC6A685447918B77" pageId="11" pageNumber="650">Frabosa Soprana</collectingMunicipality>
(108 
<location id="8E95CEF3FFAFFF8DFEA3687546318B77" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFAFFF8DFEA3687546318B77" box="[312,335,1632,1656]" county="Cuneo" municipality="Frabosa Soprana" name="Pi" pageId="11" pageNumber="650" stateProvince="Piemonte">Pi</location>
<emphasis id="B93E443AFFAFFF8DFECB687446228B76" box="[336,348,1633,1657]" italics="true" pageId="11" pageNumber="650">/</emphasis>
<collectionCode id="ED5B00EDFFAFFF8DFEC7687546F68B77" box="[348,392,1632,1656]" country="United Kingdom" name="Wellcome Collection of Bacteria, Burroughs Wellcome Research Laboratories" pageId="11" pageNumber="650">CN</collectionCode>
), 
<collectorName id="26BFFDFEFFAFFF8DFE3A6875452B8B77" box="[417,597,1632,1656]" pageId="11" pageNumber="650">Valle Corsaglia</collectorName>
, 
<location id="8E95CEF3FFAFFF8DFDFE6875440D8B77" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFAFFF8DFDFE6875440D8B77" box="[613,883,1632,1656]" county="Cuneo" municipality="Frabosa Soprana" name="Grotta di Bossea Cave" pageId="11" pageNumber="650" stateProvince="Piemonte">Grotta di Bossea Cave</location>
, coll. 
<collectorName id="26BFFDFEFFAFFF8DFC246874435C8B77" box="[959,1058,1633,1656]" pageId="11" pageNumber="650">E. Lana</collectorName>
, 
<date id="FFF4BEE8FFAFFF8DFBA96875463D8B97" pageId="11" pageNumber="650" value="1998-12-26">
<collectingDate id="EFB04700FFAFFF8DFBA96875463D8B97" pageId="11" pageNumber="650" value="1998-12-26">26 December 1998</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B229275FFAFFF8DFED4689543378B97" box="[335,1097,1664,1688]" collectingDate="2009-12-16" collectionCode="CN" collectorName="M. Isaia &amp; M. Paschetta" county="Cuneo" location="Pi" municipality="Frabosa Soprana" pageId="11" pageNumber="650" specimenCount="1" stateProvince="Piemonte">
adult male, same data, coll. 
<collectorName id="26BFFDFEFFAFFF8DFD15689445908B97" box="[654,750,1664,1688]" pageId="11" pageNumber="650">M. Isaia</collectorName>
and 
<collectorName id="26BFFDFEFFAFFF8DFCB3689444BF8B97" box="[808,961,1664,1688]" pageId="11" pageNumber="650">M. Paschetta</collectorName>
, 
<date id="FFF4BEE8FFAFFF8DFC56689543378B97" box="[973,1097,1664,1688]" pageId="11" pageNumber="650" value="2009-12-16">
<collectingDate id="EFB04700FFAFFF8DFC56689543378B97" box="[973,1097,1664,1688]" pageId="11" pageNumber="650" value="2009-12-16">16.12.2009</collectingDate>
</date>
</materialsCitation>
; 
<materialsCitation id="3B229275FFA8FF8AFF4E6854468C8B97" collectingDate="1997-04-15" collectionCode="CN" collectorName="E. Lana &amp; M. Zacharda" county="Cuneo" location="Pi" municipality="Frabosa Soprana" pageId="12" pageNumber="651" specimenCount="2" specimenCount-adult="2" stateProvince="Piemonte">
<specimenCount id="9D4C53A1FFA8FF8AFF4E6854462D8B56" box="[213,339,1601,1625]" pageId="12" pageNumber="651" type="adult">three adult</specimenCount>
females and 
<specimenCount id="9D4C53A1FFA8FF8AFE6A6854451E8B56" box="[497,608,1601,1625]" pageId="12" pageNumber="651" type="adult">one adult</specimenCount>
male, 
<collectingRegion id="498E56CAFFA8FF8AFD2B6854445F8B56" box="[688,801,1601,1625]" country="Italy" name="Piemonte" pageId="12" pageNumber="651">Piedmont</collectingRegion>
, 
<collectingCounty id="6294E0A4FFA8FF8AFCB4685444028B56" box="[815,892,1601,1625]" pageId="12" pageNumber="651">Cuneo</collectingCounty>
, 
<collectingMunicipality id="6B910252FFA8FF8AFC116854432C8B56" box="[906,1106,1601,1625]" pageId="12" pageNumber="651">Frabosa Soprana</collectingMunicipality>
, (837 
<location id="8E95CEF3FFA8FF8AFF4E687547928B77" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFA8FF8AFF4E687547928B77" box="[213,236,1632,1656]" county="Cuneo" municipality="Frabosa Soprana" name="Pi" pageId="12" pageNumber="651" stateProvince="Piemonte">Pi</location>
<emphasis id="B93E443AFFA8FF8AFF76687447878B76" box="[237,249,1633,1657]" italics="true" pageId="12" pageNumber="651">/</emphasis>
<collectionCode id="ED5B00EDFFA8FF8AFF626875465B8B77" box="[249,293,1632,1656]" country="United Kingdom" name="Wellcome Collection of Bacteria, Burroughs Wellcome Research Laboratories" pageId="12" pageNumber="651">CN</collectionCode>
), 
<location id="8E95CEF3FFA8FF8AFEDA687545438B77" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFA8FF8AFEDA687545438B77" box="[321,573,1632,1656]" county="Cuneo" municipality="Frabosa Soprana" name="Abisso Bacardi Cave" pageId="12" pageNumber="651" stateProvince="Piemonte">Abisso Bacardi Cave</location>
, coll. 
<collectorName id="26BFFDFEFFA8FF8AFD146874458B8B77" box="[655,757,1633,1656]" pageId="12" pageNumber="651">E. Lana</collectorName>
, 
<date id="FFF4BEE8FFA8FF8AFC93687544CA8B77" box="[776,948,1632,1656]" pageId="12" pageNumber="651" value="1997-04-15">
<collectingDate id="EFB04700FFA8FF8AFC93687544CA8B77" box="[776,948,1632,1656]" pageId="12" pageNumber="651" value="1997-04-15">15 April 1997</collectingDate>
</date>
, deposited in collection of 
<collectorName id="26BFFDFEFFA8FF8AFECF6894468C8B97" box="[340,498,1664,1688]" pageId="12" pageNumber="651">M. Zacharda</collectorName>
</materialsCitation>
; 
<materialsCitation id="3B229275FFA8FF8AFD9F689543EC8B97" box="[516,1170,1664,1688]" collectorName="Frabosa Soprana" location="Cuneo" pageId="12" pageNumber="651" specimenCount="1" stateProvince="Piemonte">
adult female, 
<collectingRegion id="498E56CAFFA8FF8AFD31689544658B97" box="[682,795,1664,1688]" country="Italy" name="Piemonte" pageId="12" pageNumber="651">Piedmont</collectingRegion>
, 
<location id="8E95CEF3FFA8FF8AFCB7689544078B97" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFA8FF8AFCB7689544078B97" box="[812,889,1664,1688]" name="Cuneo" pageId="12" pageNumber="651" stateProvince="Piemonte">Cuneo</location>
, 
<collectorName id="26BFFDFEFFA8FF8AFC106895432D8B97" box="[907,1107,1664,1688]" pageId="12" pageNumber="651">Frabosa Soprana</collectorName>
(108
</materialsCitation>
</paragraph>
<caption id="DF35C8A0FFA8FF8AFF4E6BD9444E8B13" ID-DOI="http://doi.org/10.5281/zenodo.5203357" ID-Zenodo-Dep="5203357" httpUri="https://zenodo.org/record/5203357/files/figure.png" pageId="12" pageNumber="651" startId="12.[213,280,1484,1506]" targetBox="[210,1170,161,1448]" targetPageId="12">
<paragraph id="8BF59828FFA8FF8AFF4E6BD9444E8B13" blockId="12.[213,1170,1484,1564]" pageId="12" pageNumber="651">
Figure 7. 
<taxonomicName id="4C4AE3ABFFA8FF8AFEA66BD9468E88ED" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[317,496,1484,1506]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="12" pageNumber="651" phylum="Arthropoda" rank="species" species="lanai" status="sp. nov.">
<emphasis id="B93E443AFFA8FF8AFEA66BD9468E88ED" box="[317,496,1484,1506]" italics="true" pageId="12" pageNumber="651">Troglocheles lanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFA8FF8AFE6D6BD8453B88ED" box="[502,581,1485,1506]" pageId="12" pageNumber="651" rank="species">sp. nov.</taxonomicNameLabel>
, adult female; (A,B) rhagidial organ I; (C) rhagidial organ II; (D) chelicera, lateral aspect; (E) palpus, lateral aspect; (F) subcapitulum, ventral aspect; (G) anomalous neotrichy on trochanter III; (H) elongated naso.
</paragraph>
</caption>
<paragraph id="8BF59828FFA9FF8BFD43683345928B4B" blockId="13.[728,748,1574,1604]" box="[728,748,1574,1604]" pageId="13" pageNumber="652">F</paragraph>
<caption id="DF35C8A0FFA9FF8BFF0B6873454F8A01" ID-DOI="http://doi.org/10.5281/zenodo.5203359" ID-Zenodo-Dep="5203359" httpUri="https://zenodo.org/record/5203359/files/figure.png" pageId="13" pageNumber="652" startId="13.[144,211,1638,1660]" targetBox="[142,1102,160,1567]" targetPageId="13">
<paragraph id="8BF59828FFA9FF8BFF0B6873454F8A01" blockId="13.[144,1101,1638,1806]" pageId="13" pageNumber="652">
Figure 8. 
<taxonomicName id="4C4AE3ABFFA9FF8BFF61687246CE8B72" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[250,432,1639,1661]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="13" pageNumber="652" phylum="Arthropoda" rank="species" species="lanai" status="sp. nov.">
<emphasis id="B93E443AFFA9FF8BFF61687246CE8B72" box="[250,432,1639,1661]" italics="true" pageId="13" pageNumber="652">Troglocheles lanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFA9FF8BFE22687245758B73" box="[441,523,1639,1660]" pageId="13" pageNumber="652" rank="species">sp. nov.</taxonomicNameLabel>
, adult female, setal arrangement on legs, lateral aspect; (A) leg I – genu, tibia, tarsus; (B) leg I – trochanter, basi- and telofemur; (C,D) variations in positions of dorsodistal rhagidial 
<emphasis id="B93E443AFFA9FF8BFE7868B446848BB7" box="[483,506,1697,1720]" italics="true" pageId="13" pageNumber="652">
Φ 
<subScript id="17CE9A6DFFA9FF8BFE6968BF46848BB7" attach="left" box="[498,506,1706,1720]" fontSize="6" pageId="13" pageNumber="652">1</subScript>
</emphasis>
and spiniform 
<emphasis id="B93E443AFFA9FF8BFD0F68B445D58BB7" box="[660,683,1697,1720]" italics="true" pageId="13" pageNumber="652">
Φ 
<subScript id="17CE9A6DFFA9FF8BFD3868BF45D58BB7" attach="left" box="[675,683,1706,1720]" fontSize="6" pageId="13" pageNumber="652">2</subScript>
</emphasis>
solenidia on tibia I, dorsal aspect; (E) leg II; (F,G) variations in positions of dorsodistal lanceolate 
<emphasis id="B93E443AFFA9FF8BFD5568AB459B8BDA" box="[718,741,1726,1749]" italics="true" pageId="13" pageNumber="652">
Φ 
<subScript id="17CE9A6DFFA9FF8BFD4668D2459B8BDA" attach="left" box="[733,741,1735,1749]" fontSize="6" pageId="13" pageNumber="652">1</subScript>
</emphasis>
and spiniform 
<emphasis id="B93E443AFFA9FF8BFC1B68AB44E98BDA" box="[896,919,1726,1749]" italics="true" pageId="13" pageNumber="652">
Φ 
<subScript id="17CE9A6DFFA9FF8BFC1468D244E98BDA" attach="left" box="[911,919,1735,1749]" fontSize="6" pageId="13" pageNumber="652">2</subScript>
</emphasis>
solenidia on tibia II, dorsal and lateral aspect, respectively; (
<emphasis id="B93E443AFFA9FF8BFDDD68C945288BFD" box="[582,598,1756,1778]" italics="true" pageId="13" pageNumber="652">ω</emphasis>
, 
<emphasis id="B93E443AFFA9FF8BFDFE68C9450D8BFD" box="[613,627,1756,1778]" italics="true" pageId="13" pageNumber="652">Φ</emphasis>
, 
<emphasis id="B93E443AFFA9FF8BFD1968C945F18BFD" box="[642,655,1756,1778]" italics="true" pageId="13" pageNumber="652">σ</emphasis>
and 
<emphasis id="B93E443AFFA9FF8BFD5368C945AD8BFD" box="[712,723,1756,1778]" italics="true" pageId="13" pageNumber="652">ε</emphasis>
designations for solenidia on tarsus, tibia, genu and for famulus, respectively).
</paragraph>
</caption>
<caption id="DF35C8A0FFAAFF88FF4E684B43338BA0" ID-DOI="http://doi.org/10.5281/zenodo.5203361" ID-Zenodo-Dep="5203361" httpUri="https://zenodo.org/record/5203361/files/figure.png" pageId="14" pageNumber="653" startId="14.[213,280,1630,1652]" targetBox="[209,1172,162,1596]" targetPageId="14">
<paragraph id="8BF59828FFAAFF88FF4E684B43338BA0" blockId="14.[213,1170,1630,1711]" pageId="14" pageNumber="653">
Figure 9. 
<taxonomicName id="4C4AE3ABFFAAFF88FEA7684A46908B7A" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[316,494,1631,1653]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="14" pageNumber="653" phylum="Arthropoda" rank="species" species="lanai" status="sp. nov.">
<emphasis id="B93E443AFFAAFF88FEA7684A46908B7A" box="[316,494,1631,1653]" italics="true" pageId="14" pageNumber="653">Troglocheles lanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFAAFF88FE6F684A453C8B7B" box="[500,578,1631,1652]" pageId="14" pageNumber="653" rank="species">sp. nov.</taxonomicNameLabel>
, adult female, setal arrangement on legs, lateral aspect; (A) leg III, (B) apex of tarsus III with oar-like iteral (
<emphasis id="B93E443AFFAAFF88FD4F6868459C8B9D" box="[724,738,1661,1682]" italics="true" pageId="14" pageNumber="653">it</emphasis>
) setae and two rhagidial solenidia, lateral aspect; (C) leg IV (
<emphasis id="B93E443AFFAAFF88FE08688C46DD8BA0" box="[403,419,1689,1711]" italics="true" pageId="14" pageNumber="653">ω</emphasis>
, 
<emphasis id="B93E443AFFAAFF88FE2B688C46C08BA0" box="[432,446,1689,1711]" italics="true" pageId="14" pageNumber="653">Φ</emphasis>
, 
<emphasis id="B93E443AFFAAFF88FE50688C46A68BA0" box="[459,472,1689,1711]" italics="true" pageId="14" pageNumber="653">σ</emphasis>
designations for solenidia on tarsus, tibia, genu, respectively).
</paragraph>
</caption>
<paragraph id="8BF59828FFABFF89FF0B6EB346F48C92" blockId="15.[144,1101,166,414]" pageId="15" pageNumber="654">
<materialsCitation id="3B229275FFABFF89FF0B6EB3479E8DB1" box="[144,224,166,190]" collectionCode="CN" collectorName="Pi" pageId="15" pageNumber="654" specimenCount="1">
<collectorName id="26BFFDFEFFABFF89FF0B6EB347D98DB1" box="[144,167,166,190]" pageId="15" pageNumber="654">Pi</collectorName>
<emphasis id="B93E443AFFABFF89FF3C6EB347CD8DB1" box="[167,179,166,190]" italics="true" pageId="15" pageNumber="654">/</emphasis>
<collectionCode id="ED5B00EDFFABFF89FF2F6EB3479E8DB1" box="[180,224,166,190]" country="United Kingdom" name="Wellcome Collection of Bacteria, Burroughs Wellcome Research Laboratories" pageId="15" pageNumber="654">CN</collectionCode>
</materialsCitation>
), 
<materialsCitation id="3B229275FFABFF89FF616EB3445F8DB1" box="[250,801,166,190]" collectionCode="UTM" collectorName="Valle Corsaglia" location="Grotta di Bossea Cave" pageId="15" pageNumber="654" specimenCount="1">
<collectorName id="26BFFDFEFFABFF89FF616EB346D08DB1" box="[250,430,166,190]" pageId="15" pageNumber="654">Valle Corsaglia</collectorName>
, 
<location id="8E95CEF3FFABFF89FE246EB345B08DB1" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFABFF89FE246EB345B08DB1" box="[447,718,166,190]" name="Grotta di Bossea Cave" pageId="15" pageNumber="654">Grotta di Bossea Cave</location>
, 
<collectionCode id="ED5B00EDFFABFF89FD446EB2445F8DB1" box="[735,801,167,190]" pageId="15" pageNumber="654">UTM</collectionCode>
</materialsCitation>
<materialsCitation id="3B229275FFABFF89FCB66EB343338DB1" box="[813,1101,166,190]" collectionCode="T" pageId="15" pageNumber="654" specimenCode="T 407464" specimenCount="1">
coordinates: 32 
<specimenCode id="DBEC3053FFABFF89FC446EB343338DB1" box="[991,1101,166,190]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="654">T 407464</specimenCode>
</materialsCitation>
<materialsCitation id="3B229275FFABFF89FF0B6ED345588C12" collectingDate="2009-12-20" collectionCode="CN, UTM" collectorName="E. Lana" county="Cuneo" elevation="836" latitude="44.24278" location="Vallone" longLatPrecision="19" longitude="7.8408337" municipality="Valdieri" pageId="15" pageNumber="654" specimenCount="1" stateProvince="Piemonte">
4899559 (
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44 
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14 
<superScript id="7C3F3560FFABFF89FEDB6ED7463B8DDC" attach="none" box="[320,325,194,211]" fontSize="7" pageId="15" pageNumber="654">′</superScript>
34 
<superScript id="7C3F3560FFABFF89FEFA6ED746158DDC" attach="left" box="[353,363,194,211]" fontSize="7" pageId="15" pageNumber="654">′′</superScript>
N
</geoCoordinate>
, 
<geoCoordinate id="EE7EFEEFFFABFF89FE076ED245668DD1" box="[412,536,194,222]" degrees="7" direction="east" minutes="50" orientation="longitude" pageId="15" pageNumber="654" precision="15" seconds="27" value="7.8408337">
7 
<superScript id="7C3F3560FFABFF89FE326ED746CC8DDC" attach="none" box="[425,434,194,211]" fontSize="7" pageId="15" pageNumber="654">◦</superScript>
50 
<superScript id="7C3F3560FFABFF89FE556ED746AD8DDC" attach="none" box="[462,467,194,211]" fontSize="7" pageId="15" pageNumber="654">′</superScript>
27 
<superScript id="7C3F3560FFABFF89FE746ED746878DDC" attach="left" box="[495,505,194,211]" fontSize="7" pageId="15" pageNumber="654">′′</superScript>
E
</geoCoordinate>
), 
<elevation id="00677F1BFFABFF89FDAB6ED345C88DD1" box="[560,694,198,222]" metricMagnitude="2" metricUnit="m" metricValue="8.36" pageId="15" pageNumber="654" unit="m" value="836.0">
<quantity id="4CB235CDFFABFF89FDAB6ED345068DD1" box="[560,632,198,222]" metricMagnitude="2" metricUnit="m" metricValue="8.36" pageId="15" pageNumber="654" unit="m" value="836.0">836 m</quantity>
a.s.l.
</elevation>
, coll. 
<collectorName id="26BFFDFEFFABFF89FC9B6ED2441E8DD1" box="[768,864,199,222]" pageId="15" pageNumber="654">E. Lana</collectorName>
, 
<date id="FFF4BEE8FFABFF89FCF46ED343368DD1" box="[879,1096,198,222]" pageId="15" pageNumber="654" value="2009-12-20">
<collectingDate id="EFB04700FFABFF89FCF46ED343368DD1" box="[879,1096,198,222]" pageId="15" pageNumber="654" value="2009-12-20">20 December 2009</collectingDate>
</date>
; one tritonymph, 
<collectingRegion id="498E56CAFFABFF89FECE6EF346B88DF1" box="[341,454,230,254]" country="Italy" name="Piemonte" pageId="15" pageNumber="654">Piedmont</collectingRegion>
, 
<collectingCounty id="6294E0A4FFABFF89FE4F6EF3455F8DF1" box="[468,545,230,254]" pageId="15" pageNumber="654">Cuneo</collectingCounty>
, 
<collectingMunicipality id="6B910252FFABFF89FDB46EF345F68DF1" box="[559,648,230,254]" pageId="15" pageNumber="654">Valdieri</collectingMunicipality>
(1214 Pi 
<emphasis id="B93E443AFFABFF89FD6A6EF345838DF1" box="[753,765,230,254]" italics="true" pageId="15" pageNumber="654">/</emphasis>
<collectionCode id="ED5B00EDFFABFF89FD656EF344548DF1" box="[766,810,230,254]" country="United Kingdom" name="Wellcome Collection of Bacteria, Burroughs Wellcome Research Laboratories" pageId="15" pageNumber="654">CN</collectionCode>
), 
<location id="8E95CEF3FFABFF89FCDA6EF344E98DF1" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFABFF89FCDA6EF344E98DF1" box="[833,919,230,254]" county="Cuneo" latitude="44.24278" longLatPrecision="19" longitude="7.8408337" municipality="Valdieri" name="Vallone" pageId="15" pageNumber="654" stateProvince="Piemonte">Vallone</location>
dell’Infernotto, 
<location id="8E95CEF3FFABFF89FF0B6F1346FE8C11" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFABFF89FF0B6F1346FE8C11" box="[144,384,262,286]" county="Cuneo" latitude="44.24278" longLatPrecision="19" longitude="7.8408337" municipality="Valdieri" name="Baron Litron-Maissa" pageId="15" pageNumber="654" stateProvince="Piemonte">Barôn Litrôn-Maissa</location>
6 
<location id="8E95CEF3FFABFF89FE046F1346A68C11" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFABFF89FE046F1346A68C11" box="[415,472,262,286]" county="Cuneo" latitude="44.24278" longLatPrecision="19" longitude="7.8408337" municipality="Valdieri" name="Cave" pageId="15" pageNumber="654" stateProvince="Piemonte">Cave</location>
, 
<collectionCode id="ED5B00EDFFABFF89FE7F6F1345588C12" box="[484,550,262,285]" pageId="15" pageNumber="654">UTM</collectionCode>
</materialsCitation>
<materialsCitation id="3B229275FFABFF89FDB46F1344388C11" box="[559,838,262,286]" collectionCode="T" pageId="15" pageNumber="654" specimenCode="T 373117" specimenCount="1" stateProvince="Piemonte">
coordinates: 32 
<specimenCode id="DBEC3053FFABFF89FD476F1344388C11" box="[732,838,262,286]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="654">T 373117</specimenCode>
</materialsCitation>
<materialsCitation id="3B229275FFABFF89FCD56F1346F88C92" collectingDate="2010-04-24" collectionCode="EDIT" collectorName="M. Isaia" county="Natural Museum of Torino" elevation="1050" latitude="44.263054" location="Monitoring Project" longLatPrecision="19" longitude="7.410278" municipality="All Taxa Biodiversity Inventory" pageId="15" pageNumber="654" specimenCount="1" stateProvince="Piemonte">
4902365 (
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44 
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15 
<superScript id="7C3F3560FFABFF89FC676F17437F8C1C" attach="left" box="[1020,1025,258,275]" fontSize="7" pageId="15" pageNumber="654">′</superScript>
47 
<superScript id="7C3F3560FFABFF89FB866F1743598C1C" attach="left" box="[1053,1063,258,275]" fontSize="7" pageId="15" pageNumber="654">′′</superScript>
N
</geoCoordinate>
, 
<geoCoordinate id="EE7EFEEFFFABFF89FF0B6F3346758C32" box="[144,267,290,317]" degrees="7" direction="east" minutes="24" orientation="longitude" pageId="15" pageNumber="654" precision="15" seconds="37" value="7.410278">
7 
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24 
<superScript id="7C3F3560FFABFF89FF586F3747B68C3C" attach="none" box="[195,200,290,307]" fontSize="7" pageId="15" pageNumber="654">′</superScript>
37 
<superScript id="7C3F3560FFABFF89FF7F6F3747908C3C" attach="left" box="[228,238,290,307]" fontSize="7" pageId="15" pageNumber="654">′′</superScript>
E
</geoCoordinate>
), 
<elevation id="00677F1BFFABFF89FEBA6F3046CF8C32" box="[289,433,293,317]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="15" pageNumber="654" unit="m" value="1050.0">
<quantity id="4CB235CDFFABFF89FEBA6F30460A8C32" box="[289,372,293,317]" metricMagnitude="3" metricUnit="m" metricValue="1.05" pageId="15" pageNumber="654" unit="m" value="1050.0">1050 m</quantity>
a.s.l.
</elevation>
, coll. 
<collectorName id="26BFFDFEFFABFF89FE6D6F3345278C32" box="[502,601,293,317]" pageId="15" pageNumber="654">M. Isaia</collectorName>
, 
<date id="FFF4BEE8FFABFF89FDFE6F30447D8C32" box="[613,771,293,317]" pageId="15" pageNumber="654" value="2010-04-24">
<collectingDate id="EFB04700FFABFF89FDFE6F30447D8C32" box="[613,771,293,317]" pageId="15" pageNumber="654" value="2010-04-24">24 April 2010</collectingDate>
</date>
; the latter 
<specimenCount id="9D4C53A1FFABFF89FCE56F33435C8C32" box="[894,1058,293,317]" pageId="15" pageNumber="654" type="generic">two specimens</specimenCount>
are deposited in the 
<collectingCounty id="6294E0A4FFABFF89FED26F5045028C52" box="[329,636,325,349]" pageId="15" pageNumber="654">Natural Museum of Torino</collectingCounty>
as the voucher specimens of the 
<collectingMunicipality id="6B910252FFABFF89FC776F5046F38C71" pageId="15" pageNumber="654">All Taxa Biodiversity Inventory</collectingMunicipality>
and 
<location id="8E95CEF3FFABFF89FE5E6F7045E18C72" LSID="urn:lsid:plazi:treatment:03E3293EFFAEFF97FE7E6AED43618824:8E95CEF3FFABFF89FE5E6F7045E18C72" box="[453,671,357,381]" county="Natural Museum of Torino" latitude="44.263054" longLatPrecision="19" longitude="7.410278" municipality="All Taxa Biodiversity Inventory" name="Monitoring Project" pageId="15" pageNumber="654" stateProvince="Piemonte">Monitoring Project</location>
of the 
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(
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)
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.
</paragraph>
<paragraph id="8BF59828FFABFF89FF0B6FF647828CF5" blockId="15.[144,252,483,506]" box="[144,252,483,506]" pageId="15" pageNumber="654">
<emphasis id="B93E443AFFABFF89FF0B6FF647828CF5" box="[144,252,483,506]" italics="true" pageId="15" pageNumber="654">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BF59828FFABFF89FF0B6C1C44658E6F" blockId="15.[144,1101,521,864]" pageId="15" pageNumber="654">
Rhagidial organ I comprises 12, 13 or 14 rhagidial solenidia lying symmetrically or asymmetrically, in separate oblique depressions; rhagidial organ II with four rhagidial solenidia lying in separate oblique depressions, and small spiniform famulus (
<emphasis id="B93E443AFFABFF89FBAC6C5F433D8F6D" box="[1079,1091,586,610]" italics="true" pageId="15" pageNumber="654">ε</emphasis>
) subtending proximal rhagidial solenidion. Tarsus III with two adjacent dorsodistal rhagidial solenidia lying in separate depressions axially, famulus absent. Spiniform solenidion on tibia I and II long, positioned dorsodistally just proximad of dorsodistal rhagidial or lanceolate solenidion, respectively. Tibia IV with one long spiniform dorsal medioproximal solenidion. Ratio of tarsus I length to dorsoventral width 6.25– 9.33; ratio of palpal tarsus length to width 4.23–5.25. Bothridial setae 
<emphasis id="B93E443AFFABFF89FC4B6D1F44918E2C" box="[976,1007,778,803]" italics="true" pageId="15" pageNumber="654">
sc 
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</emphasis>
slightly overlapping insertions of opisthosomal setae 
<emphasis id="B93E443AFFABFF89FD006D3C45D98E4F" box="[667,679,809,832]" italics="true" pageId="15" pageNumber="654">c</emphasis>
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<emphasis id="B93E443AFFABFF89FD3C6D2745CE8E4C" box="[679,688,818,835]" italics="true" pageId="15" pageNumber="654">1</emphasis>
.
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Movable digit of chelicera striated along proximal two-thirds of length – see in lateral aspect.
</paragraph>
<paragraph id="8BF59828FFABFF89FF0B6DB345188EB2" blockId="15.[144,614,933,957]" box="[144,614,933,957]" pageId="15" pageNumber="654">
<emphasis id="B93E443AFFABFF89FF0B6DB345188EB2" box="[144,614,933,957]" italics="true" pageId="15" pageNumber="654">Description – adult females (10 examined)</emphasis>
</paragraph>
<paragraph id="8BF59828FFABFF89FF0B6DD8467D890B" blockId="15.[144,1100,973,1028]" pageId="15" pageNumber="654">Length of idiosoma 1056–1280 µm. Ratio of leg I length to idiosomal length 1.98–2.43.</paragraph>
<paragraph id="8BF59828FFABFF89FF0B6A3946F68BCE" blockId="15.[144,1101,1068,1729]" pageId="15" pageNumber="654">
<emphasis id="B93E443AFFABFF89FF0B6A39465C894B" box="[144,290,1068,1092]" italics="true" pageId="15" pageNumber="654">
<taxonomicName id="4C4AE3ABFFABFF89FF0B6A394660894B" authorityName="Krefft" authorityYear="1871" box="[144,286,1068,1092]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="15" pageNumber="654" phylum="Nematoda" rank="genus">Gnathosoma</taxonomicName>
.
</emphasis>
Subcapitulum slenderly elongated, cylindrical, ratio of length to breadth 1.65–1.71; distal hypostomal lips with spiniform internal and serrate external malar processes; adoral setae nude and overlapping apex of subcapitulum, proximal subcapitular setae fine, pointed, pubescent, external pair slightly longer than internal pair (
<figureCitation id="137184ADFFABFF89FF036ABE467389CC" box="[152,269,1195,1219]" captionStart="Figure 7" captionStartId="12.[213,280,1484,1506]" captionTargetBox="[210,1170,161,1448]" captionTargetId="figure-6@12.[210,1138,160,1441]" captionTargetPageId="12" captionText="Figure 7. Troglocheles lanai sp. nov., adult female; (A,B) rhagidial organ I; (C) rhagidial organ II; (D) chelicera, lateral aspect; (E) palpus, lateral aspect; (F) subcapitulum, ventral aspect; (G) anomalous neotrichy on trochanter III; (H) elongated naso." figureDoi="http://doi.org/10.5281/zenodo.5203357" httpUri="https://zenodo.org/record/5203357/files/figure.png" pageId="15" pageNumber="654">Figure 7F</figureCitation>
). Dorsal surface of chelicera with saddle-shaped depression above level of bases of digits; cheliceral digits strikingly long, slender; fixed digit terminates in two large cusps, smooth along masticatory surface and with small lateral vertical fissure at level of bases of digits (
<figureCitation id="137184ADFFABFF89FE276B1E4548882C" box="[444,566,1291,1315]" captionStart="Figure 7" captionStartId="12.[213,280,1484,1506]" captionTargetBox="[210,1170,161,1448]" captionTargetId="figure-6@12.[210,1138,160,1441]" captionTargetPageId="12" captionText="Figure 7. Troglocheles lanai sp. nov., adult female; (A,B) rhagidial organ I; (C) rhagidial organ II; (D) chelicera, lateral aspect; (E) palpus, lateral aspect; (F) subcapitulum, ventral aspect; (G) anomalous neotrichy on trochanter III; (H) elongated naso." figureDoi="http://doi.org/10.5281/zenodo.5203357" httpUri="https://zenodo.org/record/5203357/files/figure.png" pageId="15" pageNumber="654">Figure 7D</figureCitation>
); movable digit smooth along masticatory surface and strikingly striated along proximal two-thirds (see in lateral aspect). Chelicera with two setae inserted distad of articulation of movable digit; tip of proximal seta almost reaching insertion of distal seta; tip of distal seta overlaps apex of fixed digit. Length of chelicera 320–429 µm, dorsoventral width 79–145 µm, length of movable digit 162–221 µm, length of proximal and distal cheliceral setae 23–33 and 66–86 µm, respectively, distance between their insertions 23–46 µm. Ratios: cheliceral length to dorsoventral width 2.86–4.05; length of movable digit to length of chelicera 0.50–0.53; length of movable digit to dorsoventral width of chelicera 1.50–2.05. Palpal tarsus shorter than femorogenu (
<figureCitation id="137184ADFFABFF89FE24683F45498B4D" box="[447,567,1578,1602]" captionStart="Figure 7" captionStartId="12.[213,280,1484,1506]" captionTargetBox="[210,1170,161,1448]" captionTargetId="figure-6@12.[210,1138,160,1441]" captionTargetPageId="12" captionText="Figure 7. Troglocheles lanai sp. nov., adult female; (A,B) rhagidial organ I; (C) rhagidial organ II; (D) chelicera, lateral aspect; (E) palpus, lateral aspect; (F) subcapitulum, ventral aspect; (G) anomalous neotrichy on trochanter III; (H) elongated naso." figureDoi="http://doi.org/10.5281/zenodo.5203357" httpUri="https://zenodo.org/record/5203357/files/figure.png" pageId="15" pageNumber="654">Figure 7E</figureCitation>
), ratio of length to width of tarsus 4.16–5.25. Length of palpal trochanter, femorogenu, tibia and tarsus 59–66, 247–264, 76-102 and 181–208 µm, respectively. Number of setae and solenidia (in brackets) on palpal trochanter, femorogenu, tibia and tarsus 0-2-3-10(1), respectively; tarsal solenidion long, spiniform, erect.
</paragraph>
<paragraph id="8BF59828FFB4FF96FF4E6EB346EE8C12" blockId="16.[213,1170,166,288]" pageId="16" pageNumber="655">
<emphasis id="B93E443AFFB4FF96FF4E6EB346288DB1" box="[213,342,166,190]" italics="true" pageId="16" pageNumber="655">Prodorsum.</emphasis>
Naso well-developed (
<figureCitation id="137184ADFFB4FF96FDF96EB3459F8DB1" box="[610,737,166,190]" captionStart="Figure 7" captionStartId="12.[213,280,1484,1506]" captionTargetBox="[210,1170,161,1448]" captionTargetId="figure-6@12.[210,1138,160,1441]" captionTargetPageId="12" captionText="Figure 7. Troglocheles lanai sp. nov., adult female; (A,B) rhagidial organ I; (C) rhagidial organ II; (D) chelicera, lateral aspect; (E) palpus, lateral aspect; (F) subcapitulum, ventral aspect; (G) anomalous neotrichy on trochanter III; (H) elongated naso." figureDoi="http://doi.org/10.5281/zenodo.5203357" httpUri="https://zenodo.org/record/5203357/files/figure.png" pageId="16" pageNumber="655">Figure 7H</figureCitation>
), with pair of long internal vertical setae 
<emphasis id="B93E443AFFB4FF96FE826ED246538DEE" box="[281,301,199,225]" italics="true" pageId="16" pageNumber="655">
v 
<subScript id="17CE9A6DFFB4FF96FEBF6EC546538DEE" attach="left" box="[292,301,208,225]" fontSize="7" pageId="16" pageNumber="655">1</subScript>
</emphasis>
. Bothridial setae 
<emphasis id="B93E443AFFB4FF96FD9E6ED2455A8DEE" box="[517,548,199,225]" italics="true" pageId="16" pageNumber="655">
sc 
<subScript id="17CE9A6DFFB4FF96FD806EC5455A8DEE" attach="left" box="[539,548,208,225]" fontSize="7" pageId="16" pageNumber="655">1</subScript>
</emphasis>
filiform, finely pubescent, slightly overlapping insertions of opisthosomal setae 
<emphasis id="B93E443AFFB4FF96FD856EF2454C8C0E" box="[542,562,231,257]" italics="true" pageId="16" pageNumber="655">
c 
<subScript id="17CE9A6DFFB4FF96FDB26EE5454C8C0E" attach="left" box="[553,562,240,257]" fontSize="7" pageId="16" pageNumber="655">1</subScript>
</emphasis>
. Length of setae: 
<emphasis id="B93E443AFFB4FF96FC976EF2445F8C0E" box="[780,801,231,257]" italics="true" pageId="16" pageNumber="655">
v 
<subScript id="17CE9A6DFFB4FF96FC836EE5445F8C0E" attach="left" box="[792,801,240,257]" fontSize="7" pageId="16" pageNumber="655">1</subScript>
</emphasis>
99–115, 
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v 
<subScript id="17CE9A6DFFB4FF96FC016EE544DD8C0E" attach="left" box="[922,931,240,257]" fontSize="7" pageId="16" pageNumber="655">2</subScript>
</emphasis>
72–79, 
<emphasis id="B93E443AFFB4FF96FB986EF2435F8C0E" box="[1027,1057,231,257]" italics="true" pageId="16" pageNumber="655">
sc 
<subScript id="17CE9A6DFFB4FF96FB836EE5435F8C0E" attach="left" box="[1048,1057,240,257]" fontSize="7" pageId="16" pageNumber="655">1</subScript>
</emphasis>
138–188, 
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sc 
<subScript id="17CE9A6DFFB4FF96FF706F1A478A8C2F" attach="left" box="[235,244,271,288]" fontSize="7" pageId="16" pageNumber="655">2</subScript>
</emphasis>
155–188 µm.
</paragraph>
<paragraph id="8BF59828FFB4FF96FF4E6F5E43F48CED" blockId="16.[213,1170,331,485]" pageId="16" pageNumber="655">
<emphasis id="B93E443AFFB4FF96FF4E6F5E45FA8C6C" box="[213,644,331,355]" italics="true" pageId="16" pageNumber="655">Opisthosomal dorsum and anal region.</emphasis>
Cupules 
<emphasis id="B93E443AFFB4FF96FD6A6F59447B8C6C" box="[753,773,332,355]" italics="true" pageId="16" pageNumber="655">ia</emphasis>
positioned proximad of insertions of setae 
<emphasis id="B93E443AFFB4FF96FEAC6F7946328C8A" box="[311,332,364,389]" italics="true" pageId="16" pageNumber="655">
c 
<subScript id="17CE9A6DFFB4FF96FED86F6146328C8A" attach="left" box="[323,332,372,389]" fontSize="7" pageId="16" pageNumber="655">2</subScript>
</emphasis>
, 
<emphasis id="B93E443AFFB4FF96FEC56F7946048C8C" box="[350,378,364,387]" italics="true" pageId="16" pageNumber="655">im</emphasis>
laterad and slightly distad of insertions of 
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e 
<subScript id="17CE9A6DFFB4FF96FCE76F6144FB8C8A" attach="left" box="[892,901,372,389]" fontSize="7" pageId="16" pageNumber="655">1</subScript>
</emphasis>
, 
<emphasis id="B93E443AFFB4FF96FC0C6F7944D58C8C" box="[919,939,364,387]" italics="true" pageId="16" pageNumber="655">ip</emphasis>
laterad and slightly distad of insertions of 
<emphasis id="B93E443AFFB4FF96FE436F9E46978CAA" box="[472,489,395,421]" italics="true" pageId="16" pageNumber="655">
f 
<subScript id="17CE9A6DFFB4FF96FE7B6F8146978CAA" attach="left" box="[480,489,404,421]" fontSize="7" pageId="16" pageNumber="655">1</subScript>
</emphasis>
, 
<emphasis id="B93E443AFFB4FF96FE616F9E45708CAC" box="[506,526,395,419]" italics="true" pageId="16" pageNumber="655">ih</emphasis>
positioned laterodistad of insertions of adanal setae 
<emphasis id="B93E443AFFB4FF96FBF66F9E43EE8CAA" box="[1133,1168,395,421]" italics="true" pageId="16" pageNumber="655">
ad 
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</emphasis>
(
<figureCitation id="137184ADFFB4FF96FF466FBF462A8CCD" box="[221,340,426,450]" captionStart="Figure 6" captionStartId="11.[144,211,1244,1266]" captionTargetBox="[142,1102,160,1171]" captionTargetId="figure-7@11.[142,1102,160,1171]" captionTargetPageId="11" captionText="Figure 6. Troglocheles lanai sp. nov., adult male; (A) dorsum; (B) venter (for setal notation see Description)." figureDoi="http://doi.org/10.5281/zenodo.5203355" httpUri="https://zenodo.org/record/5203355/files/figure.png" pageId="16" pageNumber="655">Figure 6A</figureCitation>
). Length of setae: 
<emphasis id="B93E443AFFB4FF96FDBB6FBE454A8CCA" box="[544,564,427,453]" italics="true" pageId="16" pageNumber="655">
c 
<subScript id="17CE9A6DFFB4FF96FDB06FA1454A8CCA" attach="left" box="[555,564,436,453]" fontSize="7" pageId="16" pageNumber="655">1</subScript>
</emphasis>
79–132, 
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c 
<subScript id="17CE9A6DFFB4FF96FD3E6FA145D08CCA" attach="left" box="[677,686,436,453]" fontSize="7" pageId="16" pageNumber="655">2</subScript>
</emphasis>
168–198, 
<emphasis id="B93E443AFFB4FF96FCBB6FBE44488CCA" box="[800,822,427,453]" italics="true" pageId="16" pageNumber="655">
d 
<subScript id="17CE9A6DFFB4FF96FCB66FA144488CCA" attach="left" box="[813,822,436,453]" fontSize="7" pageId="16" pageNumber="655">1</subScript>
</emphasis>
92–122, 
<emphasis id="B93E443AFFB4FF96FC006FBE44D18CCA" box="[923,943,427,453]" italics="true" pageId="16" pageNumber="655">
e 
<subScript id="17CE9A6DFFB4FF96FC3D6FA144D18CCA" attach="left" box="[934,943,436,453]" fontSize="7" pageId="16" pageNumber="655">1</subScript>
</emphasis>
92–115, 
<emphasis id="B93E443AFFB4FF96FB8F6FBE43588CCA" box="[1044,1062,427,453]" italics="true" pageId="16" pageNumber="655">
f 
<subScript id="17CE9A6DFFB4FF96FB866FA143588CCA" attach="left" box="[1053,1062,436,453]" fontSize="7" pageId="16" pageNumber="655">1</subScript>
</emphasis>
145–175, 
<emphasis id="B93E443AFFB4FF96FF4E6FDE47998CEA" box="[213,231,459,485]" italics="true" pageId="16" pageNumber="655">
f 
<subScript id="17CE9A6DFFB4FF96FF456FC147998CEA" attach="left" box="[222,231,468,485]" fontSize="7" pageId="16" pageNumber="655">2</subScript>
</emphasis>
63–105, 
<emphasis id="B93E443AFFB4FF96FED66FDE461D8CEA" box="[333,355,459,485]" italics="true" pageId="16" pageNumber="655">
h 
<subScript id="17CE9A6DFFB4FF96FEC16FC1461D8CEA" attach="left" box="[346,355,468,485]" fontSize="7" pageId="16" pageNumber="655">1</subScript>
</emphasis>
165–208, 
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h 
<subScript id="17CE9A6DFFB4FF96FE7F6FC146938CEA" attach="left" box="[484,493,468,485]" fontSize="7" pageId="16" pageNumber="655">2</subScript>
</emphasis>
99–138, 
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ps 
<subScript id="17CE9A6DFFB4FF96FDF16FC1450D8CEA" attach="left" box="[618,627,468,485]" fontSize="7" pageId="16" pageNumber="655">1</subScript>
</emphasis>
138–175, 
<emphasis id="B93E443AFFB4FF96FD7D6FDE44798CEA" box="[742,775,459,485]" italics="true" pageId="16" pageNumber="655">
ps 
<subScript id="17CE9A6DFFB4FF96FD656FC144798CEA" attach="left" box="[766,775,468,485]" fontSize="7" pageId="16" pageNumber="655">2</subScript>
</emphasis>
89–115, 
<emphasis id="B93E443AFFB4FF96FCF76FDE44F38CEA" box="[876,909,459,485]" italics="true" pageId="16" pageNumber="655">
ps 
<subScript id="17CE9A6DFFB4FF96FC1F6FC144F38CEA" attach="left" box="[900,909,468,485]" fontSize="7" pageId="16" pageNumber="655">3</subScript>
</emphasis>
49–69, 
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ad 
<subScript id="17CE9A6DFFB4FF96FB9B6FC143778CEA" attach="left" box="[1024,1033,468,485]" fontSize="7" pageId="16" pageNumber="655">1</subScript>
</emphasis>
60–67 µm.
</paragraph>
<paragraph id="8BF59828FFB4FF96FF4E6C0545DC8F48" blockId="16.[213,1170,527,583]" pageId="16" pageNumber="655">
<emphasis id="B93E443AFFB4FF96FF4E6C0546318F27" box="[213,335,528,552]" italics="true" pageId="16" pageNumber="655">Podosoma.</emphasis>
Coxae I-II-III-IV with 3-1-5-3, exceptionally 4-1-5-3 or 4 
<emphasis id="B93E443AFFB4FF96FC756C0544848F27" box="[1006,1018,528,552]" italics="true" pageId="16" pageNumber="655">/</emphasis>
3-1-5-3 finely pubescent setae, respectively (
<figureCitation id="137184ADFFB4FF96FDBA6C3A45E88F48" box="[545,662,559,583]" captionStart="Figure 6" captionStartId="11.[144,211,1244,1266]" captionTargetBox="[142,1102,160,1171]" captionTargetId="figure-7@11.[142,1102,160,1171]" captionTargetPageId="11" captionText="Figure 6. Troglocheles lanai sp. nov., adult male; (A) dorsum; (B) venter (for setal notation see Description)." figureDoi="http://doi.org/10.5281/zenodo.5203355" httpUri="https://zenodo.org/record/5203355/files/figure.png" pageId="16" pageNumber="655">Figure 6B</figureCitation>
).
</paragraph>
<paragraph id="8BF59828FFB4FF96FF4E6C6046CD8FE3" blockId="16.[213,1170,628,748]" pageId="16" pageNumber="655">
<emphasis id="B93E443AFFB4FF96FF4E6C6046078F83" box="[213,377,629,653]" italics="true" pageId="16" pageNumber="655">Genital region.</emphasis>
Genital valves each with five finely pubescent genital setae (
<emphasis id="B93E443AFFB4FF96FBBF6C60434F8F83" box="[1060,1073,629,652]" italics="true" pageId="16" pageNumber="655">g</emphasis>
) of similar length, about 26–46 µm, arranged evenly along medial edge of valve. Five pairs of pointed aggenital setae (
<emphasis id="B93E443AFFB4FF96FD906CA045588FC3" box="[523,550,693,716]" italics="true" pageId="16" pageNumber="655">ag</emphasis>
) of similar length, about 59–86 µm. Length of genital valves 115–165 µm.
</paragraph>
<paragraph id="8BF59828FFB4FF96FF4E6D0F46D388E1" blockId="16.[213,1170,793,1518]" pageId="16" pageNumber="655">
<emphasis id="B93E443AFFB4FF96FF4E6D0F466F8E3E" box="[213,273,794,817]" italics="true" pageId="16" pageNumber="655">Legs.</emphasis>
Leg 
<date id="FFF4BEE8FFB4FF96FED16D0F46F18E3E" box="[330,399,793,817]" pageId="16" pageNumber="655" value="2224-01">I 2224</date>
–2720 µm long, about 1.98–2.43 times as long as idiosoma. Empodia of all legs setulose, longer than claws, no ventrobasal clawlets on claws. Trochanter I strikingly enlarged. Tarsi II, III, IV each with oar-like subapical setae typical of 
<taxonomicName id="4C4AE3ABFFB4FF96FF4E6D6C46208E9E" authorityName="Zacharda" authorityYear="1980" box="[213,350,889,913]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="16" pageNumber="655" phylum="Arthropoda" rank="genus">
<emphasis id="B93E443AFFB4FF96FF4E6D6C46208E9E" box="[213,350,889,913]" italics="true" pageId="16" pageNumber="655">Troglocheles</emphasis>
</taxonomicName>
(
<figureCitation id="137184ADFFB4FF96FEF46D6C46998E9E" box="[367,487,889,913]" captionStart="Figure 9" captionStartId="14.[213,280,1630,1652]" captionTargetBox="[209,1172,162,1596]" captionTargetId="figure-6@14.[209,1135,161,1597]" captionTargetPageId="14" captionText="Figure 9. Troglocheles lanai sp. nov., adult female, setal arrangement on legs, lateral aspect; (A) leg III, (B) apex of tarsus III with oar-like iteral (it) setae and two rhagidial solenidia, lateral aspect; (C) leg IV (ω, Φ, σ designations for solenidia on tarsus, tibia, genu, respectively)." figureDoi="http://doi.org/10.5281/zenodo.5203361" httpUri="https://zenodo.org/record/5203361/files/figure.png" pageId="16" pageNumber="655">Figure 9C</figureCitation>
). Number of setae and solenidia (solenidia and famulus (
<emphasis id="B93E443AFFB4FF96FBE76D6C43F68E9E" box="[1148,1160,889,913]" italics="true" pageId="16" pageNumber="655">ε</emphasis>
) bracketed), respectively, on legs I-II-III-IV (
<figureCitation id="137184ADFFB4FF96FD5D6D8D444C8EBF" box="[710,818,920,944]" captionStart="Figure 8" captionStartId="13.[144,211,1638,1660]" captionTargetBox="[142,1102,160,1567]" captionTargetId="figure-7@13.[142,1102,160,1567]" captionTargetPageId="13" captionText="Figure 8. Troglocheles lanai sp. nov., adult female, setal arrangement on legs, lateral aspect; (A) leg I – genu, tibia, tarsus; (B) leg I – trochanter, basi- and telofemur; (C,D) variations in positions of dorsodistal rhagidial Φ1 and spiniform Φ2 solenidia on tibia I, dorsal aspect; (E) leg II; (F,G) variations in positions of dorsodistal lanceolate Φ1 and spiniform Φ2 solenidia on tibia II, dorsal and lateral aspect, respectively; (ω, Φ, σ and ε designations for solenidia on tarsus, tibia, genu and for famulus, respectively)." figureDoi="http://doi.org/10.5281/zenodo.5203359" httpUri="https://zenodo.org/record/5203359/files/figure.png" pageId="16" pageNumber="655">Figures 8</figureCitation>
, 
<figureCitation id="137184ADFFB4FF96FCA46D8D44318EBF" box="[831,847,920,944]" captionStart="Figure 9" captionStartId="14.[213,280,1630,1652]" captionTargetBox="[209,1172,162,1596]" captionTargetId="figure-6@14.[209,1135,161,1597]" captionTargetPageId="14" captionText="Figure 9. Troglocheles lanai sp. nov., adult female, setal arrangement on legs, lateral aspect; (A) leg III, (B) apex of tarsus III with oar-like iteral (it) setae and two rhagidial solenidia, lateral aspect; (C) leg IV (ω, Φ, σ designations for solenidia on tarsus, tibia, genu, respectively)." figureDoi="http://doi.org/10.5281/zenodo.5203361" httpUri="https://zenodo.org/record/5203361/files/figure.png" pageId="16" pageNumber="655">9</figureCitation>
): trochanters 1-1-2-2, exceptionally three setae on trochanter III; basifemora + telofemora 5+5-6+5-4+4-3+4; genua 17(1)-9(1)-9(1)-7; tibiae 12(2)-8(2)-7(2)-6(1); tarsi 18(13 or 14+ 
<emphasis id="B93E443AFFB4FF96FB946DCC43658EFE" box="[1039,1051,985,1009]" italics="true" pageId="16" pageNumber="655">ε</emphasis>
)-17(4+ 
<emphasis id="B93E443AFFB4FF96FBE86DCC43018EFE" box="[1139,1151,985,1009]" italics="true" pageId="16" pageNumber="655">ε</emphasis>
)- 16-15. Spiniform solenidia on leg segments conspicuously long. Genua I and II each with one erect spiniform distoventral solenidion (
<emphasis id="B93E443AFFB4FF96FCCC6A0D441B893F" box="[855,869,1048,1072]" italics="true" pageId="16" pageNumber="655">σ</emphasis>
); genu III with one long spiniform dorso-mediodistal solenidion (
<figureCitation id="137184ADFFB4FF96FD356A2D4455895F" box="[686,811,1080,1104]" captionStart="Figure 9" captionStartId="14.[213,280,1630,1652]" captionTargetBox="[209,1172,162,1596]" captionTargetId="figure-6@14.[209,1135,161,1597]" captionTargetPageId="14" captionText="Figure 9. Troglocheles lanai sp. nov., adult female, setal arrangement on legs, lateral aspect; (A) leg III, (B) apex of tarsus III with oar-like iteral (it) setae and two rhagidial solenidia, lateral aspect; (C) leg IV (ω, Φ, σ designations for solenidia on tarsus, tibia, genu, respectively)." figureDoi="http://doi.org/10.5281/zenodo.5203361" httpUri="https://zenodo.org/record/5203361/files/figure.png" pageId="16" pageNumber="655">Figure 9A</figureCitation>
). Tibia I with one erect spiniform dorsodistal solenidion (
<emphasis id="B93E443AFFB4FF96FDB76A4D4545897F" box="[556,571,1112,1136]" italics="true" pageId="16" pageNumber="655">Φ</emphasis>
) positioned just proximad of dorsodistal rhagidial solenidion (
<figureCitation id="137184ADFFB4FF96FEC66A6D468A899F" box="[349,500,1144,1168]" captionStart="Figure 8" captionStartId="13.[144,211,1638,1660]" captionTargetBox="[142,1102,160,1567]" captionTargetId="figure-7@13.[142,1102,160,1567]" captionTargetPageId="13" captionText="Figure 8. Troglocheles lanai sp. nov., adult female, setal arrangement on legs, lateral aspect; (A) leg I – genu, tibia, tarsus; (B) leg I – trochanter, basi- and telofemur; (C,D) variations in positions of dorsodistal rhagidial Φ1 and spiniform Φ2 solenidia on tibia I, dorsal aspect; (E) leg II; (F,G) variations in positions of dorsodistal lanceolate Φ1 and spiniform Φ2 solenidia on tibia II, dorsal and lateral aspect, respectively; (ω, Φ, σ and ε designations for solenidia on tarsus, tibia, genu and for famulus, respectively)." figureDoi="http://doi.org/10.5281/zenodo.5203359" httpUri="https://zenodo.org/record/5203359/files/figure.png" pageId="16" pageNumber="655">Figure 8C,D</figureCitation>
); tibia II with one spiniform erect dorsodistal solenidion positioned just proximad of dorsodistal lanceolate solenidion recessed in deep pit with broadly open surface pore (
<figureCitation id="137184ADFFB4FF96FDC46AA2458B89C0" box="[607,757,1207,1231]" captionStart="Figure 8" captionStartId="13.[144,211,1638,1660]" captionTargetBox="[142,1102,160,1567]" captionTargetId="figure-7@13.[142,1102,160,1567]" captionTargetPageId="13" captionText="Figure 8. Troglocheles lanai sp. nov., adult female, setal arrangement on legs, lateral aspect; (A) leg I – genu, tibia, tarsus; (B) leg I – trochanter, basi- and telofemur; (C,D) variations in positions of dorsodistal rhagidial Φ1 and spiniform Φ2 solenidia on tibia I, dorsal aspect; (E) leg II; (F,G) variations in positions of dorsodistal lanceolate Φ1 and spiniform Φ2 solenidia on tibia II, dorsal and lateral aspect, respectively; (ω, Φ, σ and ε designations for solenidia on tarsus, tibia, genu and for famulus, respectively)." figureDoi="http://doi.org/10.5281/zenodo.5203359" httpUri="https://zenodo.org/record/5203359/files/figure.png" pageId="16" pageNumber="655">Figure 8F,G</figureCitation>
); tibia III with two adjacent erect spiniform dorsomedial solenidia (
<figureCitation id="137184ADFFB4FF96FDFE6AC2459A89E0" box="[613,740,1239,1263]" captionStart="Figure 9" captionStartId="14.[213,280,1630,1652]" captionTargetBox="[209,1172,162,1596]" captionTargetId="figure-6@14.[209,1135,161,1597]" captionTargetPageId="14" captionText="Figure 9. Troglocheles lanai sp. nov., adult female, setal arrangement on legs, lateral aspect; (A) leg III, (B) apex of tarsus III with oar-like iteral (it) setae and two rhagidial solenidia, lateral aspect; (C) leg IV (ω, Φ, σ designations for solenidia on tarsus, tibia, genu, respectively)." figureDoi="http://doi.org/10.5281/zenodo.5203361" httpUri="https://zenodo.org/record/5203361/files/figure.png" pageId="16" pageNumber="655">Figure 9A</figureCitation>
); tibia IV with one erect spiniform dorsoproximal solenidion (
<figureCitation id="137184ADFFB4FF96FD926AE245FF8800" box="[521,641,1271,1295]" captionStart="Figure 9" captionStartId="14.[213,280,1630,1652]" captionTargetBox="[209,1172,162,1596]" captionTargetId="figure-6@14.[209,1135,161,1597]" captionTargetPageId="14" captionText="Figure 9. Troglocheles lanai sp. nov., adult female, setal arrangement on legs, lateral aspect; (A) leg III, (B) apex of tarsus III with oar-like iteral (it) setae and two rhagidial solenidia, lateral aspect; (C) leg IV (ω, Φ, σ designations for solenidia on tarsus, tibia, genu, respectively)." figureDoi="http://doi.org/10.5281/zenodo.5203361" httpUri="https://zenodo.org/record/5203361/files/figure.png" pageId="16" pageNumber="655">Figure 9C</figureCitation>
). Tarsus I slender, its tip slightly tapers in lateral view, ratio length to width 8.66–9.33, with 12–14 rhagidial solenidia (
<emphasis id="B93E443AFFB4FF96FBAD6B0243398820" box="[1078,1095,1303,1327]" italics="true" pageId="16" pageNumber="655">ω</emphasis>
) lying obliquely in separate depressions dorsodistally (
<figureCitation id="137184ADFFB4FF96FC9D6B2244E48840" box="[774,922,1335,1359]" captionStart="Figure 7" captionStartId="12.[213,280,1484,1506]" captionTargetBox="[210,1170,161,1448]" captionTargetId="figure-6@12.[210,1138,160,1441]" captionTargetPageId="12" captionText="Figure 7. Troglocheles lanai sp. nov., adult female; (A,B) rhagidial organ I; (C) rhagidial organ II; (D) chelicera, lateral aspect; (E) palpus, lateral aspect; (F) subcapitulum, ventral aspect; (G) anomalous neotrichy on trochanter III; (H) elongated naso." figureDoi="http://doi.org/10.5281/zenodo.5203357" httpUri="https://zenodo.org/record/5203357/files/figure.png" pageId="16" pageNumber="655">Figure 7A,B</figureCitation>
), stellate famulus (
<emphasis id="B93E443AFFB4FF96FBE76B2243F68840" box="[1148,1160,1335,1359]" italics="true" pageId="16" pageNumber="655">ε</emphasis>
) inserted between first and second proximal rhagidial solenidia antiaxially; tarsus II with four rhagidial solenidia lying in separate oblique depressions, and small spiniform famulus (
<emphasis id="B93E443AFFB4FF96FE186B8246F188A0" box="[387,399,1431,1455]" italics="true" pageId="16" pageNumber="655">ε</emphasis>
) subtending proximal rhagidial solenidion antiaxially (
<figureCitation id="137184ADFFB4FF96FB976B8243FB88A0" box="[1036,1157,1431,1455]" captionStart="Figure 7" captionStartId="12.[213,280,1484,1506]" captionTargetBox="[210,1170,161,1448]" captionTargetId="figure-6@12.[210,1138,160,1441]" captionTargetPageId="12" captionText="Figure 7. Troglocheles lanai sp. nov., adult female; (A,B) rhagidial organ I; (C) rhagidial organ II; (D) chelicera, lateral aspect; (E) palpus, lateral aspect; (F) subcapitulum, ventral aspect; (G) anomalous neotrichy on trochanter III; (H) elongated naso." figureDoi="http://doi.org/10.5281/zenodo.5203357" httpUri="https://zenodo.org/record/5203357/files/figure.png" pageId="16" pageNumber="655">Figure 7C</figureCitation>
), tarsus III with two adjacent solenidia arranged in separate depressions axially, famulus absent (
<figureCitation id="137184ADFFB4FF96FEB76BC346DF88E1" box="[300,417,1494,1518]" captionStart="Figure 9" captionStartId="14.[213,280,1630,1652]" captionTargetBox="[209,1172,162,1596]" captionTargetId="figure-6@14.[209,1135,161,1597]" captionTargetPageId="14" captionText="Figure 9. Troglocheles lanai sp. nov., adult female, setal arrangement on legs, lateral aspect; (A) leg III, (B) apex of tarsus III with oar-like iteral (it) setae and two rhagidial solenidia, lateral aspect; (C) leg IV (ω, Φ, σ designations for solenidia on tarsus, tibia, genu, respectively)." figureDoi="http://doi.org/10.5281/zenodo.5203361" httpUri="https://zenodo.org/record/5203361/files/figure.png" pageId="16" pageNumber="655">Figure 9B</figureCitation>
).
</paragraph>
<paragraph id="8BF59828FFB4FF96FF4E682F45DE8B5E" blockId="16.[213,672,1593,1617]" box="[213,672,1593,1617]" pageId="16" pageNumber="655">
<emphasis id="B93E443AFFB4FF96FF4E682F45DE8B5E" box="[213,672,1593,1617]" italics="true" pageId="16" pageNumber="655">Description – adult male (four examined)</emphasis>
</paragraph>
<paragraph id="8BF59828FFB4FF96FF4E687543EC8B97" blockId="16.[213,1170,1632,1688]" pageId="16" pageNumber="655">
A club-shaped sperm sac (
<emphasis id="B93E443AFFB4FF96FD91687745608B76" box="[522,542,1634,1657]" italics="true" pageId="16" pageNumber="655">ss</emphasis>
) showing through the opisthosomal tegument; genital valves each with five and six setae asymmetrically; otherwise as in female (
<figureCitation id="137184ADFFB4FF96FB8A689543F88B97" box="[1041,1158,1664,1688]" captionStart="Figure 6" captionStartId="11.[144,211,1244,1266]" captionTargetBox="[142,1102,160,1171]" captionTargetId="figure-7@11.[142,1102,160,1171]" captionTargetPageId="11" captionText="Figure 6. Troglocheles lanai sp. nov., adult male; (A) dorsum; (B) venter (for setal notation see Description)." figureDoi="http://doi.org/10.5281/zenodo.5203355" httpUri="https://zenodo.org/record/5203355/files/figure.png" pageId="16" pageNumber="655">Figure 6B</figureCitation>
).
</paragraph>
<paragraph id="8BF59828FFB5FF97FF0B6EB3478B8DB1" blockId="17.[144,245,166,190]" box="[144,245,166,190]" pageId="17" pageNumber="656">
<emphasis id="B93E443AFFB5FF97FF0B6EB3478B8DB1" box="[144,245,166,190]" italics="true" pageId="17" pageNumber="656">Affinities</emphasis>
</paragraph>
<paragraph id="8BF59828FFB5FF97FF0B6EDB46838FEB" blockId="17.[144,1101,206,740]" pageId="17" pageNumber="656">
Among other representatives of the genus 
<taxonomicName id="4C4AE3ABFFB5FF97FDF76EDB458B8DE9" authorityName="Zacharda" authorityYear="1980" box="[620,757,206,230]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="656" phylum="Arthropoda" rank="genus">
<emphasis id="B93E443AFFB5FF97FDF76EDB458B8DE9" box="[620,757,206,230]" italics="true" pageId="17" pageNumber="656">Troglocheles</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4AE3ABFFB5FF97FC9F6EDA44298DE9" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[772,855,206,230]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="656" phylum="Arthropoda" rank="species" species="lanai" status="sp. nov.">
<emphasis id="B93E443AFFB5FF97FC9F6EDA44298DE9" box="[772,855,206,230]" italics="true" pageId="17" pageNumber="656">T. lanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFB5FF97FCC46EDA44CA8DE9" box="[863,948,207,230]" pageId="17" pageNumber="656" rank="species">sp. nov.</taxonomicNameLabel>
possesses the most expressed adaptive apomorphic troglomorphisms, i.e. 12–14 rhagidial solenidia in the rhagidial organ I (both tarsi examined in the same specimen, i.e. the number of solenidia in the rhagidial organ I may be asymmetrical), and two rhagidial solenidia on tarsus III, i.e. the rhagidial organ III. 
<taxonomicName id="4C4AE3ABFFB5FF97FDEF6F5B44438C69" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[628,829,334,358]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="656" phylum="Arthropoda" rank="species" species="lanai" status="sp. nov.">
<emphasis id="B93E443AFFB5FF97FDEF6F5B44438C69" box="[628,829,334,358]" italics="true" pageId="17" pageNumber="656">Troglocheles lanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFB5FF97FCDC6F5B44E18C6A" box="[839,927,334,357]" pageId="17" pageNumber="656" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the 
<taxonomicName id="4C4AE3ABFFB5FF97FF0B6F7B46688C89" baseAuthorityName="Willmann" baseAuthorityYear="1953" box="[144,278,366,390]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="656" phylum="Arthropoda" rank="species" species="vornatscheri">
<emphasis id="B93E443AFFB5FF97FF0B6F7B46688C89" box="[144,278,366,390]" italics="true" pageId="17" pageNumber="656">vornatscheri</emphasis>
</taxonomicName>
species group because its coxal formula is 3-1-5-3, the distance between insertions of the cheliceral setae on the fixed digits is almost the same as the length of the proximal cheliceral seta, and the spiniform solenidion on tibia I is positioned dorsodistally (cf. 
<bibRefCitation id="EFDBE5D9FFB5FF97FEC96FD846838CEA" author="Zacharda M" box="[338,509,461,485]" pageId="17" pageNumber="656" pagination="463 - 478" refId="ref11291" refString="Zacharda M. 2000. New species of the genus Troglocheles (Acari: Prostigmata: Rhagidiidae) from Oetztal Alps, with a key to adult species of the genus. J Nat Hist. 34: 463 - 478." type="journal article" year="2000">Zacharda 2000</bibRefCitation>
, Table 1: 470). However, 
<taxonomicName id="4C4AE3ABFFB5FF97FC8E6FDB44198CEA" authority="Zacharda &amp; Isaia &amp; Piva, 2011" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[789,871,461,485]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="656" phylum="Arthropoda" rank="species" species="lanai" status="sp. nov.">
<emphasis id="B93E443AFFB5FF97FC8E6FDB44198CEA" box="[789,871,461,485]" italics="true" pageId="17" pageNumber="656">T. lanai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20DF941FFB5FF97FCF46FDB44BD8CEA" box="[879,963,462,485]" pageId="17" pageNumber="656" rank="species">sp. nov.</taxonomicNameLabel>
differs from 
<taxonomicName id="4C4AE3ABFFB5FF97FF0B6FFB464C8F0A" baseAuthorityName="Willmann" baseAuthorityYear="1953" box="[144,306,493,517]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="656" phylum="Arthropoda" rank="species" species="vornatscheri">
<emphasis id="B93E443AFFB5FF97FF0B6FFB464C8F0A" box="[144,306,493,517]" italics="true" pageId="17" pageNumber="656">T. vornatscheri</emphasis>
</taxonomicName>
in the following characteristics: (1) The number of rhagidial solenidia in rhagidial organ I is 12, 13 or 14 (examine both tarsi I); in 
<taxonomicName id="4C4AE3ABFFB5FF97FC8E6C1B44C68F2A" baseAuthorityName="Willmann" baseAuthorityYear="1953" box="[789,952,525,549]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="656" phylum="Arthropoda" rank="species" species="vornatscheri">
<emphasis id="B93E443AFFB5FF97FC8E6C1B44C68F2A" box="[789,952,525,549]" italics="true" pageId="17" pageNumber="656">T. vornatscheri</emphasis>
</taxonomicName>
the rhagidial organ I comprises eight rhagidial solenidia. (2) Two adjacent rhagidial solenidia are on tarsus III (rhagidial organ III); in 
<taxonomicName id="4C4AE3ABFFB5FF97FDAC6C5B459E8F6A" baseAuthorityName="Willmann" baseAuthorityYear="1953" box="[567,736,589,613]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="656" phylum="Arthropoda" rank="species" species="vornatscheri">
<emphasis id="B93E443AFFB5FF97FDAC6C5B45378F6A" box="[567,585,590,613]" italics="true" pageId="17" pageNumber="656">T</emphasis>
. 
<emphasis id="B93E443AFFB5FF97FDC16C58459E8F6A" box="[602,736,589,613]" italics="true" pageId="17" pageNumber="656">vornatscheri</emphasis>
</taxonomicName>
no rhagidial solenidia on tarsus III. (3) The position of the spiniform solenidion on tibia II is dorsodistal, just behind the dorsodistal lanceolate solenidion; in 
<taxonomicName id="4C4AE3ABFFB5FF97FDCF6C9845888FAB" baseAuthorityName="Willmann" baseAuthorityYear="1953" box="[596,758,652,676]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="656" phylum="Arthropoda" rank="species" species="vornatscheri">
<emphasis id="B93E443AFFB5FF97FDCF6C9845888FAB" box="[596,758,652,676]" italics="true" pageId="17" pageNumber="656">T. vornatscheri</emphasis>
</taxonomicName>
this solenidion is laterodorsal, proximal. (4) The position of the spiniform solenidion on genu III is dorsodistal; in 
<taxonomicName id="4C4AE3ABFFB5FF97FF0B6CD8464D8FEB" baseAuthorityName="Willmann" baseAuthorityYear="1953" box="[144,307,716,740]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="656" phylum="Arthropoda" rank="species" species="vornatscheri">
<emphasis id="B93E443AFFB5FF97FF0B6CD8464D8FEB" box="[144,307,716,740]" italics="true" pageId="17" pageNumber="656">T. vornatscheri</emphasis>
</taxonomicName>
it is lateromedial.
</paragraph>
<paragraph id="8BF59828FFB5FF97FF0B6D3C46768E4E" blockId="17.[144,264,809,833]" box="[144,264,809,833]" pageId="17" pageNumber="656">
<emphasis id="B93E443AFFB5FF97FF0B6D3C46768E4E" box="[144,264,809,833]" italics="true" pageId="17" pageNumber="656">Etymology</emphasis>
</paragraph>
<paragraph id="8BF59828FFB5FF97FF0B6D4546C48EC7" blockId="17.[144,1101,848,968]" pageId="17" pageNumber="656">
The epithet 
<taxonomicName id="4C4AE3ABFFB5FF97FE8C6D4446338E66" authorityName="Zacharda &amp; Isaia &amp; Piva" authorityYear="2011" box="[279,333,849,873]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="656" phylum="Arthropoda" rank="species" species="lanai">
<emphasis id="B93E443AFFB5FF97FE8C6D4446338E66" box="[279,333,849,873]" italics="true" pageId="17" pageNumber="656">lanai</emphasis>
</taxonomicName>
was used in honour of Enrico Lana, the Italian biospeleologist, collector and photographer of the cave invertebrates in the region of 
<collectingRegion id="498E56CAFFB5FF97FC096D65437E8E87" box="[914,1024,880,904]" country="Italy" name="Piemonte" pageId="17" pageNumber="656">Piedmont</collectingRegion>
(
<bibRefCitation id="EFDBE5D9FFB5FF97FB8A6D6547B48EA7" author="Lana E" pageId="17" pageNumber="656" refId="ref10849" refString="Lana E. 2001. Biospeleologia del Piemonte. Atlante fotografico sistematico. Torino: Associazione Gruppi Speleologici Piemontesi, La Grafica Nuova." type="book" year="2001">Lana 2001</bibRefCitation>
). He first discovered this new troglobitic species in the Barôn Litrôn cave (locus typicus). Feminine gender.
</paragraph>
<paragraph id="8BF59828FFB5FF97FF0B6A18478C892A" blockId="17.[144,242,1037,1061]" box="[144,242,1037,1061]" pageId="17" pageNumber="656">
<emphasis id="B93E443AFFB5FF97FF0B6A18478C892A" box="[144,242,1037,1061]" italics="true" pageId="17" pageNumber="656">Remarks</emphasis>
</paragraph>
<paragraph id="8BF59828FFB5FF97FF0B6A21443689E3" blockId="17.[144,1101,1076,1323]" pageId="17" pageNumber="656">
Hitherto the rhagidial organ III has been known only in the troglomorphic 
<taxonomicName id="4C4AE3ABFFB5FF97FF0B6A4145618963" authority="Elliott, 1976" authorityName="Elliott" authorityYear="1976" box="[144,543,1108,1132]" class="Arachnida" family="Rhagidiidae" genus="Flabellorhagidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="17" pageNumber="656" phylum="Arthropoda" rank="species" species="pecki">
<emphasis id="B93E443AFFB5FF97FF0B6A4146F98963" box="[144,391,1108,1132]" italics="true" pageId="17" pageNumber="656">Flabellorhagidia pecki</emphasis>
<bibRefCitation id="EFDBE5D9FFB5FF97FE0A6A4145618963" author="Elliott W" box="[401,543,1108,1132]" pageId="17" pageNumber="656" pagination="1 - 15" refId="ref10717" refString="Elliott W. 1976. New cavernicolous Rhagidiidae from Idaho, Washington, and Utah (Prostigmata: Acari: Arachnida). Occ Pap Mus Texas Tech Univ. 43: 1 - 15." type="journal article" year="1976">Elliott, 1976</bibRefCitation>
</taxonomicName>
from the Boy Scout Cave, Craters of the Moon National Monument, Butte Co., ID, 
<collectingCountry id="F35DD8B8FFB5FF97FDCA6A6145F48983" box="[593,650,1140,1164]" name="United States of America" pageId="17" pageNumber="656">USA</collectingCountry>
(
<bibRefCitation id="EFDBE5D9FFB5FF97FD046A61444E8983" author="Elliott W" box="[671,816,1140,1164]" pageId="17" pageNumber="656" pagination="1 - 15" refId="ref10717" refString="Elliott W. 1976. New cavernicolous Rhagidiidae from Idaho, Washington, and Utah (Prostigmata: Acari: Arachnida). Occ Pap Mus Texas Tech Univ. 43: 1 - 15." type="journal article" year="1976">Elliott 1976</bibRefCitation>
). The chelicerae of this rhagidiid mite are strikingly similar to these in the 
<taxonomicName id="4C4AE3ABFFB5FF97FD436A8144C389A3" box="[728,957,1172,1196]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="656" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93E443AFFB5FF97FD436A81441F89A3" box="[728,865,1172,1196]" italics="true" pageId="17" pageNumber="656">Troglocheles</emphasis>
species.
</taxonomicName>
However, in 
<taxonomicName id="4C4AE3ABFFB5FF97FF0B6AA0479789C3" authorityName="Elliott" authorityYear="1976" box="[144,233,1204,1228]" class="Arachnida" family="Rhagidiidae" genus="Flabellorhagidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Acari" pageId="17" pageNumber="656" phylum="Arthropoda" rank="species" species="pecki">
<emphasis id="B93E443AFFB5FF97FF0B6AA047DE89C3" box="[144,160,1205,1228]" italics="true" pageId="17" pageNumber="656">F</emphasis>
. 
<emphasis id="B93E443AFFB5FF97FF2B6AA1479789C3" box="[176,233,1204,1228]" italics="true" pageId="17" pageNumber="656">pecki</emphasis>
</taxonomicName>
the tarsi II, III and IV are with the normal flagelliform iteral setae, i.e. not the oar-like iteral setae that are typical of the 
<taxonomicName id="4C4AE3ABFFB5FF97FDFF6AC1443689E3" box="[612,840,1236,1260]" class="Arachnida" family="Rhagidiidae" genus="Troglocheles" kingdom="Animalia" order="Trombidiformes" pageId="17" pageNumber="656" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93E443AFFB5FF97FDFF6AC1459389E3" box="[612,749,1236,1260]" italics="true" pageId="17" pageNumber="656">Troglocheles</emphasis>
species.
</taxonomicName>
</paragraph>
<paragraph id="8BF59828FFB5FF97FF206AE143618824" blockId="17.[144,1101,1076,1323]" pageId="17" pageNumber="656">
More detailed information on the Piedmontese caves can be found elsewhere (
<bibRefCitation id="EFDBE5D9FFB5FF97FF036B0646DF8824" author="Peano G &amp; Morisi A" box="[152,417,1299,1323]" pageId="17" pageNumber="656" pagination="317 - 336" refId="ref10979" refString="Peano G, Morisi A. 1982. Importanza naturalistica e valorizzazione scientifica della grotta di Bossea. Le Grotte d'Italia 1981 (4): 317 - 336." type="journal article" year="1982">Peano and Morisi 1982</bibRefCitation>
; 
<bibRefCitation id="EFDBE5D9FFB5FF97FE376B0145588824" author="Lana E" box="[428,550,1299,1323]" pageId="17" pageNumber="656" refId="ref10849" refString="Lana E. 2001. Biospeleologia del Piemonte. Atlante fotografico sistematico. Torino: Associazione Gruppi Speleologici Piemontesi, La Grafica Nuova." type="book" year="2001">Lana 2001</bibRefCitation>
; Gruppo Speleologico Alpi Marittime 2005).
</paragraph>
</subSubSection>
</treatment>
</document>