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<mods:title id="ADB1CA5D6D1881F2377A343D879EF612">Revision of the carnivorous snail genus Discartemon Pfeiffer, 1856, with description of twelve new species (Pulmonata, Streptaxidae)</mods:title>
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<mods:namePart id="6500F4E7FC35FBA1EE016D9E64A81C6E">Siriboon, Thanit</mods:namePart>
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<mods:namePart id="2663F347FDF2D25C9007949D3B89242B">Naggs, Fred</mods:namePart>
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<mods:namePart id="1E4733C055DF0B44A49A7EE5682C20FC">Rowson, Ben</mods:namePart>
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<mods:namePart id="A3F040DDBF9E4FBBDF4DC35FACD4E2AB">Panha, Somsak</mods:namePart>
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<treatment id="E278A7872305646B399FD77828B54265" ID-GBIF-Taxon="152052544" LSID="urn:lsid:plazi:treatment:E278A7872305646B399FD77828B54265" httpUri="http://treatment.plazi.org/id/E278A7872305646B399FD77828B54265" lastPageId="13" lastPageNumber="57" pageId="11" pageNumber="56">
<subSubSection id="0652684EDC45BAE00C7D2B3ABA39A9A0" pageId="11" pageNumber="56" type="nomenclature">
<paragraph id="F9E34D61752B187C3ECEB9BD6C7251AB" pageId="11" pageNumber="56">
<pageBreakToken id="D45963B9D3EDEB0B122C3A63C14E6DDD" pageId="11" pageNumber="56" start="start">4</pageBreakToken>
.
<taxonomicName id="4FAC746EB395E78D5268D1D7B73E99EB" ID-CoL="872F5" LSID="http://species-id.net/wiki/Discartemon_nummus" authority="Laidlaw, 1929" authorityName="Laidlaw" authorityYear="1929" class="Gastropoda" family="Streptaxidae" genus="Discartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discartemon nummus" order="Stylommatophora" pageId="11" pageNumber="56" phylum="Mollusca" rank="species" species="nummus">Discartemon nummus (Laidlaw, 1929)</taxonomicName>
Figs 2A, 4G, 11D, E, 17
<normalizedToken id="B9B33ED30BF4558257AA39ADCDE22FED" originalValue="AE">A-E</normalizedToken>
, 22B, 23, Table 1
</paragraph>
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<subSubSection id="F8536DACA595C4AA8B104A4DCC885A04" pageId="11" pageNumber="56" type="reference_group">
<paragraph id="F1D26CC015E99AE3A1D8E318CCE88E1E" pageId="11" pageNumber="56">
<taxonomicName id="3EE4DC4EA7DEF244E0E29FE75EC25CF4" class="Gastropoda" family="Streptaxidae" genus="Odontartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Odontartemon (Discartemon) nummus" order="Stylommatophora" pageId="11" pageNumber="56" phylum="Mollusca" rank="species" species="nummus" subGenus="Discartemon">Odontartemon (Discartemon) nummus</taxonomicName>
Laidlaw, 1929: 259, 260, fig. 1. Type locality: Tale Sap, Singgora.
<bibRefCitation id="09D481F53F5E00E021CA674DE7AFD1DB" author="Laidlaw, FF" journalOrPublisher="Journal Malayan Branch Royal Asiatic Society" pageId="43" pageNumber="88" pagination="211 - 234" title="A list of the land and fresh-water Mollusca of the Malay Peninsula." volume="11" year="1933">Laidlaw 1933</bibRefCitation>
: 234.
</paragraph>
<paragraph id="7DF14F08321873C5CF0D1FAE6A5D7E38" pageId="11" pageNumber="56">
<taxonomicName id="52D8A79F158024484B18599CF1F23CB7" class="Gastropoda" family="Streptaxidae" genus="Discartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discartemon nummus" order="Stylommatophora" pageId="11" pageNumber="56" phylum="Mollusca" rank="species" species="nummus">Discartemon nummus</taxonomicName>
-
<bibRefCitation id="0C7ECF7D58CAFC6B6E838684D3DB7216" author="Benthem Jutting, WSS van" journalOrPublisher="Bulletin of the Raffles Museum" pageId="42" pageNumber="87" pagination="71 - 106" title="The Malayan Streptaxidae of the genera Discartemon and Oophana." volume="25" year="1954">Benthem Jutting 1954</bibRefCitation>
: 87, 88.
<bibRefCitation id="84E699C88FDCF74D528278B44D155887" author="Benthem Jutting, WSS van" journalOrPublisher="Beaufortia" pageId="42" pageNumber="87" pagination="41 - 191" title="Catalogue of the non-marine Mollusca of Sumatra and of its satellite islands." volume="7" year="1959">Benthem Jutting 1959</bibRefCitation>
: 168.
<bibRefCitation id="948DB89594286ABBDC8D29AC3FEC22A1" author="Richardson, L" journalOrPublisher="Tryonia" pageId="44" pageNumber="89" pagination="1 - 326" title="Streptaxacea: Catalog of species, Part I, Streptaxidae." volume="16" year="1988">Richardson 1988</bibRefCitation>
: 183.
</paragraph>
</subSubSection>
<subSubSection id="EE894949E08606C6C03C03D1172202EB" pageId="11" pageNumber="56" type="material examined">
<paragraph id="85617ACFCCFC387BEC522C63139C64F5" pageId="11" pageNumber="56">Material examined.</paragraph>
<paragraph id="0D926AA28F7CDCA598028AD68971E509" pageId="11" pageNumber="56">
Khao Ok Thalu, Phatthalung, Thailand (
<geoCoordinate id="37820374E749C5DF918B86995D30A600" direction="north" orientation="latitude" precision="1" value="7.6275277">7°37'39.1&quot;N</geoCoordinate>
,
<geoCoordinate id="6D4913A367B336AB2CD0267B988B1177" direction="east" orientation="longitude" precision="1" value="100.08864">100°5'19.1&quot;E</geoCoordinate>
): CUMZ 3594 (24 shells; Fig. 4G) and 6208 (12 specimens in ethanol; Figs 2A, 11D, E, 17
<normalizedToken id="00AD2AB6EA45F3488C3397A61306A73B" originalValue="AE">A-E</normalizedToken>
, 22B).
</paragraph>
<caption id="54E879D6931327784E1BD367F4535042" pageId="11" pageNumber="56">
<paragraph id="E17F45D805A6A4814F4177E853C74CE3" pageId="11" pageNumber="56">
Figure 2. Living snails. A
<taxonomicName id="9F4FB345E56126DB09695304D7BEFFF2" class="Gastropoda" family="Streptaxidae" genus="Discartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discartemon nummus" order="Stylommatophora" pageId="11" pageNumber="56" phylum="Mollusca" rank="species" species="nummus">Discartemon nummus</taxonomicName>
CUMZ 6208, from Patthalung (shell width about 6 mm) B
<taxonomicName id="004368DDD4437462C9EF957D639798E3" class="Gastropoda" family="Streptaxidae" genus="Discartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discartemon discadentus" order="Stylommatophora" pageId="11" pageNumber="56" phylum="Mollusca" rank="species" species="discadentus">Discartemon discadentus</taxonomicName>
sp. n. paratype CUMZ 6209 (shell width about 12 mm) C
<taxonomicName id="CF31818616A8DCF8B26EBE2098F44027" class="Gastropoda" family="Streptaxidae" genus="Discartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discartemon leptoglyphus" order="Stylommatophora" pageId="11" pageNumber="56" phylum="Mollusca" rank="species" species="leptoglyphus">Discartemon leptoglyphus</taxonomicName>
CUMZ 6007, from Ipoh, Perak, Malaysia (shell width about 6 mm) D
<taxonomicName id="CE53152502B809D9F3737A5D2D02B0E8" class="Gastropoda" family="Streptaxidae" genus="Discartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discartemon hypocrites" order="Stylommatophora" pageId="11" pageNumber="56" phylum="Mollusca" rank="species" species="hypocrites">Discartemon hypocrites</taxonomicName>
topotype CUMZ 6199 (shell width about 6 mm) E, F
<taxonomicName id="1B6BE0415CBE84FE4F582C3F552EF959" class="Gastropoda" family="Streptaxidae" genus="Discartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discartemon afthonodontia" order="Stylommatophora" pageId="11" pageNumber="56" phylum="Mollusca" rank="species" species="afthonodontia">Discartemon afthonodontia</taxonomicName>
sp. n. E paratype CUMZ 6210 (shell width about 9 mm), and F specimen CUMZ 6214, from Tam Khao Phlu, Chumphon (shell width about 8 mm).
</paragraph>
</caption>
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<subSubSection id="2EA16783C57D25985A6412AD104E2E99" pageId="11" pageNumber="56" type="description">
<paragraph id="891434069F75CE9D2D0C510D5145212B" pageId="11" pageNumber="56">Description.</paragraph>
<paragraph id="681E057E101C2C6B7D77532950155F00" pageId="11" pageNumber="56">
Shell. Shell flattened, white and semi-transparent; whorls
<normalizedToken id="30BE8B027495ACC72AB6BE60D652CD36" originalValue="5½">51/2</normalizedToken>
, spire flattened with distinct suture. Shell surface glossy with thin transverse ridges near suture and varices present. Embryonic shell about
<normalizedToken id="047E9F913D33E686B0868159942A23E4" originalValue="2½">21/2</normalizedToken>
whorls; following whorls regularly coiled. Last whorl angular with strong peripheral keel, regularly expanded; umbilicus very wide and showing all preceding whorls. Aperture triangular with sinulus; peristome continuous, thickened, expanded and reflected. Apertural dentition with a Y-shaped parietal lamella adjoining at sinulus (Fig. 4G).
</paragraph>
<paragraph id="124B172D020C5E7D552D4A7A8661101A" pageId="11" pageNumber="56">Radula. Each row consists of 39-41 teeth with formula (19-20)-1-(19-20). Central tooth very small and triangular with pointed cusp. Lateral and marginal teeth undifferentiated, unicuspid and lanceolate. Latero-marginal teeth gradually reduce in size, with outermost teeth smaller and shorter than inner teeth (Fig. 22B).</paragraph>
<paragraph id="9306D4B14D20BA36D802C1BA1C727AC1" pageId="11" pageNumber="56">Genital organs. Atrium (at) long. Proximal penis (p) long, slender; distal penis globularly enlarged. Penial sheath (ps) thin, extending about two-thirds of penis length; penial sheath retractor muscle very thin (psr), originating at genital orifice wall and inserting distally on penial sheath (Fig. 11D). Vas deferens (vd) passes through about one-fifth of penial sheath length before entering into penis distally (Fig. 11E). Penial retractor muscle (pr) thin and very long, inserting at penis and vas deferens junction.</paragraph>
<paragraph id="F9FB9BA9699113BCD12A83D2CB9D3C44" pageId="11" pageNumber="56">
Internal wall of atrium generally smooth (Fig. 17A); penial wall with scattered, short and transparent penial hooks, about 5 hooks/200
<normalizedToken id="8ED8E8B66517AE26972B15481F81A184" originalValue="µm">µm</normalizedToken>
2 (Fig. 17B); hooks located on round-ovate penial papilla. Penial hooks of small size (&lt;0.04 mm in length), slightly expanded at base, tip obtuse and directed towards genital orifice (Fig. 17C, D).
</paragraph>
<paragraph id="7329672352F939EDFB8D53167B2B835B" pageId="11" pageNumber="56">Vagina (v) short and stout, about half of penis length. Gametolytic duct (gd) a long tube extending as far as albumin gland; gametolytic sac (gs) ovate. Free oviduct (fo) long and thick; oviduct (ov) enlarged and folded; prostate gland inconspicuous and bound to oviduct. Talon (ta) small, very short and club shaped. Hermaphroditic duct (hd) bearing long seminal vesicle (sv) about one and half times longer than the length from talon to branching point of seminal vesicle (Fig. 11D).</paragraph>
<paragraph id="CCAB81E9805292633A462DDA1CB1D916" pageId="11" pageNumber="56">Vaginal wall generally with smooth surface of longitudinal vaginal folds (Fig. 17E).</paragraph>
</subSubSection>
<subSubSection id="3F369C9D15E9028F36591D568CD9455E" lastPageId="13" lastPageNumber="58" pageId="12" pageNumber="57" type="remarks">
<paragraph id="8E45E4ACB64192E5169FCBE57DA1DC0E" pageId="12" pageNumber="57">
<pageBreakToken id="D504F90D96EE792046E65F90AF13FFA3" pageId="12" pageNumber="57" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph id="94E0D9DFBE627696833F55F3FB65E673" pageId="12" pageNumber="57">
<taxonomicName id="DADC2FD58FADB7A49C875E74FB9A2F89" class="Gastropoda" family="Streptaxidae" genus="Discartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discartemon nummus" order="Stylommatophora" pageId="12" pageNumber="57" phylum="Mollusca" rank="species" species="nummus">Discartemon nummus</taxonomicName>
was described from Tale Sap (= Lake or Lagoon), Singgora (= Songkhla). In this study, living snails were found at an isolated limestone hill near the lake in Phatthalung, about 60 km north of the type locality.
</paragraph>
<paragraph id="908B88750934A127AB1F6FCB498F940B" lastPageId="13" lastPageNumber="58" pageId="12" pageNumber="57">
Having the smallest shell size clearly discriminates
<taxonomicName id="4DC5642F5B593916A4E095E3A4800231" class="Gastropoda" family="Streptaxidae" genus="Discartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discartemon nummus" order="Stylommatophora" pageId="13" pageNumber="58" phylum="Mollusca" rank="species" species="nummus">
<pageBreakToken id="40143343BC1A171E8B36350E7CFBAEDB" pageId="13" pageNumber="58" start="start">Discartemon</pageBreakToken>
nummus
</taxonomicName>
from all congeners. It is similar to
<taxonomicName id="FCCC9E809CA00A64A67ABF8AAD52EAB2" class="Gastropoda" family="Streptaxidae" genus="Discartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discartemon discus" order="Stylommatophora" pageId="13" pageNumber="58" phylum="Mollusca" rank="species" species="discus">Discartemon discus</taxonomicName>
, which has a larger shell and a peripheral keel, lacks a sinulus, and has a semi-ovate aperture with a straight parietal lamella.
<taxonomicName id="52D9B8D5A5C5E14F837E1A5B6375C7DF" class="Gastropoda" family="Streptaxidae" genus="Discartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discartemon nummus" order="Stylommatophora" pageId="13" pageNumber="58" phylum="Mollusca" rank="species" species="nummus">Discartemon nummus</taxonomicName>
can be distinguished from
<taxonomicName id="714CF6B52091006AA8482E7C8146DD58" class="Gastropoda" family="Streptaxidae" genus="Discartemon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Discartemon khaosokensis" order="Stylommatophora" pageId="13" pageNumber="58" phylum="Mollusca" rank="species" species="khaosokensis">Discartemon khaosokensis</taxonomicName>
in having a flattened spire, the last whorl regularly expanded, a triangular aperture, and a Y-shaped parietal lamella.
</paragraph>
</subSubSection>
</treatment>
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