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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1004.57526" ID-GBIF-Dataset="4e633bc1-16fc-4e83-b8a0-cb9289897616" ID-PMC="PMC7758311" ID-Pensoft-Pub="1313-2970-1004-27" ID-Pensoft-UUID="BEECC18115E95A798CA5250990A350E3" ID-PubMed="33384565" ID-ZooBank="320559CF19B5423CB7FB72555290241A" ModsDocID="1313-2970-1004-27" checkinTime="1608157838683" checkinUser="pensoft" docAuthor="Maddison, Wayne P., Beattie, Imara, Marathe, Kiran, Ng, Paul Y. C., Kanesharatnam, Nilani, Benjamin, Suresh P. &amp; Kunte, Krushnamegh" docDate="2020" docId="E21D83C2CCB6588E8E429424ACF34FD6" docLanguage="en" docName="ZooKeys 1004: 27-97" docOrigin="ZooKeys 1004" docPubDate="2020-12-16" docSource="http://dx.doi.org/10.3897/zookeys.1004.57526" docTitle="Stagetillus Simon 1885" docType="treatment" docVersion="4" id="BEECC18115E95A798CA5250990A350E3" lastPageNumber="27" masterDocId="BEECC18115E95A798CA5250990A350E3" masterDocTitle="A phylogenetic and taxonomic review of baviine jumping spiders (Araneae, Salticidae, Baviini)" masterLastPageNumber="97" masterPageNumber="27" pageNumber="27" updateTime="1668146624181" updateUser="ExternalLinkService">
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<mods:title>A phylogenetic and taxonomic review of baviine jumping spiders (Araneae, Salticidae, Baviini)</mods:title>
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<mods:namePart>Maddison, Wayne P.</mods:namePart>
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<mods:affiliation>Departments of Zoology and Botany and Beaty Biodiversity Museum, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Beattie, Imara</mods:namePart>
<mods:affiliation>Department of Zoology, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Marathe, Kiran</mods:namePart>
<mods:affiliation>Department of Zoology, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada &amp; National Centre for Biological Sciences, Tata Institute of Fundamental Research, GKVK Campus, Bellary Road, Bengaluru 560065, India</mods:affiliation>
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<mods:namePart>Ng, Paul Y. C.</mods:namePart>
<mods:affiliation>205 River Valley Road, # 16 - 53, Singapore 238274, Republic of Singapore</mods:affiliation>
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<mods:namePart>Kanesharatnam, Nilani</mods:namePart>
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<mods:affiliation>National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka &amp; Department of Zoology, Faculty of Science, Eastern University, Vantharumoolai, Sri Lanka</mods:affiliation>
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<mods:namePart>Benjamin, Suresh P.</mods:namePart>
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<mods:affiliation>National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka</mods:affiliation>
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<mods:namePart>Kunte, Krushnamegh</mods:namePart>
<mods:affiliation>National Centre for Biological Sciences, Tata Institute of Fundamental Research, GKVK Campus, Bellary Road, Bengaluru 560065, India</mods:affiliation>
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<mods:date>2020</mods:date>
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<mods:number>2020-12-16</mods:number>
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<mods:number>1004</mods:number>
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<mods:start>27</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1004.57526</mods:url>
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<treatment ID-GBIF-Taxon="176088113" LSID="urn:lsid:plazi:treatment:E21D83C2CCB6588E8E429424ACF34FD6" httpUri="http://treatment.plazi.org/id/E21D83C2CCB6588E8E429424ACF34FD6" lastPageNumber="27" pageId="0" pageNumber="27">
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<taxonomicName LSID="E21D83C2-CCB6-588E-8E42-9424ACF34FD6" authority="Simon, 1885" authorityName="Simon" authorityYear="1885" class="Arachnida" family="Salticidae" genus="Stagetillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stagetillus" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">Stagetillus Simon, 1885</taxonomicName>
</paragraph>
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<paragraph pageId="0" pageNumber="27">
<taxonomicName authorityName="Simon" authorityYear="1885" class="Arachnida" family="Salticidae" genus="Stagetillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stagetillus" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">Stagetillus</taxonomicName>
Simon, 1885. Type species
<taxonomicName family="Salticidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="S. opaciceps" order="Araneae" pageId="0" pageNumber="27" rank="species" species="opaciceps">S. opaciceps</taxonomicName>
Simon, 1885.
</paragraph>
<paragraph pageId="0" pageNumber="27">
<taxonomicName authorityName="Strand" authorityYear="1911" class="Arachnida" family="Salticidae" genus="Hyctiota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyctiota" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">Hyctiota</taxonomicName>
Strand, 1911, syn. nov. Type species
<taxonomicName family="Salticidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="H. banda" order="Araneae" pageId="0" pageNumber="27" rank="species" species="banda">H. banda</taxonomicName>
Strand, 1911.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="species included">
<paragraph pageId="0" pageNumber="27">Species included.</paragraph>
<paragraph pageId="0" pageNumber="27">
<taxonomicName class="Arachnida" family="Salticidae" genus="Stagetillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stagetillus banda" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="banda">
<emphasis italics="true" pageId="0" pageNumber="27">Stagetillus banda</emphasis>
</taxonomicName>
(Strand, 1911), comb. nov.
</paragraph>
<paragraph pageId="0" pageNumber="27">
<taxonomicName authorityName="Simon" authorityYear="1885" class="Arachnida" family="Salticidae" genus="Stagetillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stagetillus opaciceps" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="opaciceps">
<emphasis italics="true" pageId="0" pageNumber="27">Stagetillus opaciceps</emphasis>
</taxonomicName>
Simon, 1885
</paragraph>
<paragraph pageId="0" pageNumber="27">
<taxonomicName class="Arachnida" family="Salticidae" genus="Stagetillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stagetillus irri" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="irri">
<emphasis italics="true" pageId="0" pageNumber="27">Stagetillus irri</emphasis>
</taxonomicName>
Maddison, sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="notes">
<paragraph pageId="0" pageNumber="27">Notes.</paragraph>
<paragraph pageId="0" pageNumber="27">
<taxonomicName authorityName="Strand" authorityYear="1911" class="Arachnida" family="Salticidae" genus="Hyctiota" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyctiota banda" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="banda">
<emphasis italics="true" pageId="0" pageNumber="27">Hyctiota banda</emphasis>
</taxonomicName>
Strand, 1911 is based on a juvenile (SMF 2511 in SMF, examined) and not easily placed. Its being a baviine is suggested by its elongate body (most like that of
<taxonomicName authorityName="Caleb &amp; Sankaran" authorityYear="2019" family="Salticidae" genus="Indopadilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Indopadilla" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Indopadilla</emphasis>
</taxonomicName>
or
<taxonomicName authorityName="Simon" authorityYear="1885" class="Arachnida" family="Salticidae" genus="Stagetillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stagetillus" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Stagetillus</emphasis>
</taxonomicName>
), plurident chelicerae, and Asian origin. Its macrosetae sockets resemble those of baviines (Figs
<figureCitation captionStart="Figures 435" captionStartId="F2" captionText="Figures 4 - 35. Variation in traits among baviines 4, 5 oblique views of female prosoma 4 Bavia nessagyna (specimen IDWM. 20004; note convex front face of chelicerae, and typical thorax) 5 Indopadilla redunca (holotype IDWM. 20011; note concave front face of chelicerae, and thoracic bulges) 6 - 12 left male endite (9 also with chelicera) 6 Bavia nessagyna (holotype IDWM. 20005) 7 Indopadilla redunca (specimen SWK 12 - M 0009) 8 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 9 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 10 Padillothorax badut (specimen SWK 12 - 4688) 11 Maripanthus jubatus (specimen AS 19.4373) 12 Piranthus planolancis (specimen AS 19.5970) 13 - 20 carapace (all females except 15) 13 Bavia nessagyna (specimen IDWM. 20004) 14 Indopadilla redunca (holotype IDWM. 20011) 15 Stagetillus opaciceps male (specimen JK. 13.02.26.0017) 16 Stagetillus cf. opaciceps female (specimen MRB 079). 17 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 18 Padillothorax mulu (from Mulu Nat. Pk.) 19 Maripanthus reinholdae (specimen SWK 12 - 1934) 20 Piranthus planolancis (specimen AS 19.5940) 21 - 28 Metatarsus of first leg, retrolateral view. All female except 24 male. All are of left legs except 23, 25, 28 which are of right leg, digitally flipped 21 Bavia nessagyna (specimen IDWM. 20004) 22 Indopadilla kahariana (specimen SWK 12 - 1876) 23 Stagetillus cf. opaciceps (specimen MRB 079) 24 Padillothorax semiostrinus (specimen SWK 12 - EP 0105) 25 Padillothorax mulu (specimen SWK 12 - EP 0105) 26 Maripanthus reinholdae (specimen SWK 12 - 1934) 27 Maripanthus draconis (from Gunung Belemut, Johor) 28 Piranthus planolancis (specimen AS 19.5940) 29 - 35 prolateral surface of male first leg femur. 29 Bavia cf. capistrata (specimen from Singapore) 30 Indopadilla kahariana (specimen from Lambir Hills Nat. Pk.) 31 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 32 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 33 Padillothorax badut (specimen SWK 12 - 4688) 34 Maripanthus draconis (specimen from Johor, Gunung Lambak) 35 Piranthus bakau (holotype). Scale bars: 0.1 mm (6 - 12, 21 - 35), 1.0 mm (4, 5, 13 - 20)." figureDoi="10.3897/zookeys.1004.57526.figures4-35" httpUri="https://binary.pensoft.net/fig/487533" pageId="0" pageNumber="27">21-23</figureCitation>
). It lacks the thoracic bulge and ridged chelicerae of
<taxonomicName authorityName="Caleb &amp; Sankaran" authorityYear="2019" family="Salticidae" genus="Indopadilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Indopadilla" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Indopadilla</emphasis>
</taxonomicName>
, and the flat carapace of
<taxonomicName authorityName="Simon" authorityYear="1901" class="Arachnida" family="Salticidae" genus="Padillothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Padillothorax" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Padillothorax</emphasis>
</taxonomicName>
. Like
<taxonomicName authorityName="Simon" authorityYear="1885" class="Arachnida" family="Salticidae" genus="Stagetillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stagetillus" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Stagetillus</emphasis>
</taxonomicName>
, the fovea is displaced slightly to the posterior. The two dark bands running through the PLE shown in
<bibRefCitation author="Strand, E" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" pageId="0" pageNumber="27" pagination="127 - 199" refId="B46" refString="Strand, E, 1911. Araneae von den Aru- und Kei-Inseln. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 34: 127 - 199" title="Araneae von den Aru- und Kei-Inseln." volume="34" year="1911">
<normalizedToken originalValue="Strands">Strand's</normalizedToken>
(1911)
</bibRefCitation>
original figure (no longer visible in specimen) resemble those of male
<taxonomicName family="Salticidae" kingdom="Animalia" lsidName="S. opaciceps" order="Araneae" pageId="0" pageNumber="27" rank="species" species="opaciceps">
<emphasis italics="true" pageId="0" pageNumber="27">S. opaciceps</emphasis>
</taxonomicName>
. The carapace is widest posteriorly, though not so clearly as in Figs
<figureCitation captionStart="Figures 435" captionStartId="F2" captionText="Figures 4 - 35. Variation in traits among baviines 4, 5 oblique views of female prosoma 4 Bavia nessagyna (specimen IDWM. 20004; note convex front face of chelicerae, and typical thorax) 5 Indopadilla redunca (holotype IDWM. 20011; note concave front face of chelicerae, and thoracic bulges) 6 - 12 left male endite (9 also with chelicera) 6 Bavia nessagyna (holotype IDWM. 20005) 7 Indopadilla redunca (specimen SWK 12 - M 0009) 8 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 9 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 10 Padillothorax badut (specimen SWK 12 - 4688) 11 Maripanthus jubatus (specimen AS 19.4373) 12 Piranthus planolancis (specimen AS 19.5970) 13 - 20 carapace (all females except 15) 13 Bavia nessagyna (specimen IDWM. 20004) 14 Indopadilla redunca (holotype IDWM. 20011) 15 Stagetillus opaciceps male (specimen JK. 13.02.26.0017) 16 Stagetillus cf. opaciceps female (specimen MRB 079). 17 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 18 Padillothorax mulu (from Mulu Nat. Pk.) 19 Maripanthus reinholdae (specimen SWK 12 - 1934) 20 Piranthus planolancis (specimen AS 19.5940) 21 - 28 Metatarsus of first leg, retrolateral view. All female except 24 male. All are of left legs except 23, 25, 28 which are of right leg, digitally flipped 21 Bavia nessagyna (specimen IDWM. 20004) 22 Indopadilla kahariana (specimen SWK 12 - 1876) 23 Stagetillus cf. opaciceps (specimen MRB 079) 24 Padillothorax semiostrinus (specimen SWK 12 - EP 0105) 25 Padillothorax mulu (specimen SWK 12 - EP 0105) 26 Maripanthus reinholdae (specimen SWK 12 - 1934) 27 Maripanthus draconis (from Gunung Belemut, Johor) 28 Piranthus planolancis (specimen AS 19.5940) 29 - 35 prolateral surface of male first leg femur. 29 Bavia cf. capistrata (specimen from Singapore) 30 Indopadilla kahariana (specimen from Lambir Hills Nat. Pk.) 31 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 32 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 33 Padillothorax badut (specimen SWK 12 - 4688) 34 Maripanthus draconis (specimen from Johor, Gunung Lambak) 35 Piranthus bakau (holotype). Scale bars: 0.1 mm (6 - 12, 21 - 35), 1.0 mm (4, 5, 13 - 20)." figureDoi="10.3897/zookeys.1004.57526.figures4-35" httpUri="https://binary.pensoft.net/fig/487533" pageId="0" pageNumber="27">15</figureCitation>
and
<figureCitation captionStart="Figures 435" captionStartId="F2" captionText="Figures 4 - 35. Variation in traits among baviines 4, 5 oblique views of female prosoma 4 Bavia nessagyna (specimen IDWM. 20004; note convex front face of chelicerae, and typical thorax) 5 Indopadilla redunca (holotype IDWM. 20011; note concave front face of chelicerae, and thoracic bulges) 6 - 12 left male endite (9 also with chelicera) 6 Bavia nessagyna (holotype IDWM. 20005) 7 Indopadilla redunca (specimen SWK 12 - M 0009) 8 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 9 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 10 Padillothorax badut (specimen SWK 12 - 4688) 11 Maripanthus jubatus (specimen AS 19.4373) 12 Piranthus planolancis (specimen AS 19.5970) 13 - 20 carapace (all females except 15) 13 Bavia nessagyna (specimen IDWM. 20004) 14 Indopadilla redunca (holotype IDWM. 20011) 15 Stagetillus opaciceps male (specimen JK. 13.02.26.0017) 16 Stagetillus cf. opaciceps female (specimen MRB 079). 17 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 18 Padillothorax mulu (from Mulu Nat. Pk.) 19 Maripanthus reinholdae (specimen SWK 12 - 1934) 20 Piranthus planolancis (specimen AS 19.5940) 21 - 28 Metatarsus of first leg, retrolateral view. All female except 24 male. All are of left legs except 23, 25, 28 which are of right leg, digitally flipped 21 Bavia nessagyna (specimen IDWM. 20004) 22 Indopadilla kahariana (specimen SWK 12 - 1876) 23 Stagetillus cf. opaciceps (specimen MRB 079) 24 Padillothorax semiostrinus (specimen SWK 12 - EP 0105) 25 Padillothorax mulu (specimen SWK 12 - EP 0105) 26 Maripanthus reinholdae (specimen SWK 12 - 1934) 27 Maripanthus draconis (from Gunung Belemut, Johor) 28 Piranthus planolancis (specimen AS 19.5940) 29 - 35 prolateral surface of male first leg femur. 29 Bavia cf. capistrata (specimen from Singapore) 30 Indopadilla kahariana (specimen from Lambir Hills Nat. Pk.) 31 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 32 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 33 Padillothorax badut (specimen SWK 12 - 4688) 34 Maripanthus draconis (specimen from Johor, Gunung Lambak) 35 Piranthus bakau (holotype). Scale bars: 0.1 mm (6 - 12, 21 - 35), 1.0 mm (4, 5, 13 - 20)." figureDoi="10.3897/zookeys.1004.57526.figures4-35" httpUri="https://binary.pensoft.net/fig/487533" pageId="0" pageNumber="27">16</figureCitation>
. Although its synonymy with
<taxonomicName authorityName="Simon" authorityYear="1885" class="Arachnida" family="Salticidae" genus="Stagetillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stagetillus" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Stagetillus</emphasis>
</taxonomicName>
is not certain, by placing it with
<taxonomicName authorityName="Simon" authorityYear="1885" class="Arachnida" family="Salticidae" genus="Stagetillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stagetillus" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Stagetillus</emphasis>
</taxonomicName>
, a salticid genus otherwise incertae sedis is provisionally settled.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="diagnosis">
<paragraph pageId="0" pageNumber="27">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="27">
Carapace distinctive in shape, widest point toward the posterior, approx. half way between the back eyes and the pedicel, and in colour, orange or yellow with the white digestive diverticulum showing beneath the transparent ocular quadrangle. Palp much like that of
<taxonomicName authorityName="Simon" authorityYear="1877" class="Arachnida" family="Pittosporaceae" genus="Bavia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bavia" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Bavia</emphasis>
</taxonomicName>
, with short blade-like embolus.
</paragraph>
</subSubSection>
</treatment>
</document>