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<document id="9CA8CD03623C068C11A9233B8CCE2566" ID-DOI="10.5281/zenodo.5341440" ID-ISSN="0374-1036" ID-Zenodo-Dep="5341440" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630190347714" checkinUser="felipe" docAuthor="Ribes, Jordi, Pagola-Carte, Santiago &amp; Heiss, Ernst" docDate="2008" docId="DC6D87A0FFE02860758BFF59FC0AF994" docLanguage="en" docName="ActaEntMusNatPra.2008.48.2.423-431.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 48 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Atomoscelis pictifrons Ribes &amp; Pagola-Carte &amp; Heiss 2008, sp. nov." docType="treatment" docVersion="4" lastPageNumber="430" masterDocId="2054FFD8FFE52867743AFF83FFB0FFA0" masterDocTitle="Two new Phylinae (Hemiptera: Heteroptera: Miridae) from the Canary Islands" masterLastPageNumber="431" masterPageNumber="423" pageNumber="428" updateTime="1699181410245" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0CA8B89577A185E3219E390772DD1F8F">Two new Phylinae (Hemiptera: Heteroptera: Miridae) from the Canary Islands</mods:title>
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<mods:name id="320BA684F6F53E2E3BC2566B4C39C27F" type="personal">
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<mods:roleTerm id="41090D7C1BE260048D44348D6E25645E">Author</mods:roleTerm>
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<mods:namePart id="EB12EF2F0AF15956E5035D8CB14259BB">Ribes, Jordi</mods:namePart>
<mods:affiliation id="8DA5EB4D69AD54FF0A4B62C5BC51A984">València 123 - 125, ent., 3 a, E- 08011 Barcelona, Catalonia, Spain; e-mail: 4354 jrr @ comb. es</mods:affiliation>
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<mods:namePart id="708DF778A9895A83C1E81A19C34223DC">Pagola-Carte, Santiago</mods:namePart>
<mods:affiliation id="11F50D41C39FF33B678C3773CDD9B72B">Azpeitia 3, 7. D, E- 20010 Donostia (Gipuzkoa), Spain; e-mail: pagolaxpc @ telefonica. net</mods:affiliation>
</mods:name>
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<mods:namePart id="2D5BAB2E54055DB9DC757EEE3FA80803">Heiss, Ernst</mods:namePart>
<mods:affiliation id="8448B941F1384BD232CB440FCBF19487">Entomologische Forschungsgruppe, Tiroler Landesmuseum, Josef-Schraffl-Strasse 2 a, A- 6020 Innsbruck, Austria; e-mail: aradus @ aon. at</mods:affiliation>
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<mods:title id="084E3D1C9899DC7B935B63F685559F7B">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="D44EEF65D33098C58B749D559B1CF2C8">2008</mods:date>
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<mods:number id="3CC2980F766DFEB34D4B808118F1DBE6">2008-12-08</mods:number>
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<mods:number id="D07311EA066653369D9827D2216395AC">48</mods:number>
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<treatment id="DC6D87A0FFE02860758BFF59FC0AF994" ID-DOI="http://doi.org/10.5281/zenodo.5444066" ID-Zenodo-Dep="5444066" LSID="urn:lsid:plazi:treatment:DC6D87A0FFE02860758BFF59FC0AF994" httpUri="http://treatment.plazi.org/id/DC6D87A0FFE02860758BFF59FC0AF994" lastPageId="7" lastPageNumber="430" pageId="5" pageNumber="428">
<subSubSection id="1CDE653DFFE02862758BFF59FCB1FF53" box="[433,769,218,243]" pageId="5" pageNumber="428" type="nomenclature">
<paragraph id="547B36B6FFE02862758BFF59FCB1FF53" blockId="5.[433,769,218,243]" box="[433,769,218,243]" pageId="5" pageNumber="428">
<heading id="0F3381DAFFE02862758BFF59FCB1FF53" bold="true" box="[433,769,218,243]" centered="true" fontSize="8" level="2" pageId="5" pageNumber="428" reason="3">
<emphasis id="66B0EAA4FFE02862758BFF59FCB1FF53" bold="true" box="[433,769,218,243]" pageId="5" pageNumber="428">
<taxonomicName id="93C44D35FFE02862758BFF59FD15FF53" ID-CoL="67WMQ" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[433,677,218,243]" class="Insecta" family="Miridae" genus="Atomoscelis" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="428" phylum="Arthropoda" rank="species" species="pictifrons" status="sp. nov.">
<emphasis id="66B0EAA4FFE02862758BFF59FD15FF53" bold="true" box="[433,677,218,243]" italics="true" pageId="5" pageNumber="428">Atomoscelis pictifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE028627696FF58FCB1FF53" box="[684,769,219,243]" pageId="5" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1CDE653DFFE028627616FF7AFD36FEAC" box="[556,646,249,268]" pageId="5" pageNumber="428" type="description">
<paragraph id="547B36B6FFE028627616FF7AFD36FEAC" blockId="5.[113,1089,249,452]" box="[556,646,249,268]" pageId="5" pageNumber="428">
(
<figureCitation id="CCFF2A33FFE028627608FF7AFD30FEAC" box="[562,640,249,268]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="5.[113,144,1539,1559]" captionStartId-1="6.[153,184,1544,1564]" captionTargetBox-0="[151,1077,710,1506]" captionTargetBox-1="[248,1058,516,1514]" captionTargetId-0="figure-159@5.[114,1089,696,1513]" captionTargetId-1="figure-122@6.[216,1062,513,1521]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Fig. 4. Atomoscelis pictifrons sp. nov., holotype. a right antenna; b head in lateral view; c head and pronotum in dorsal view; d fore leg; e claw of middle leg; f pygophore in lateral view. Different scales." captionText-1="Fig. 5. Male genitalia of Atomoscelis pictifrons sp. nov. a left paramere in different views; b right paramere; c apical portion of phallotheca; d vesica in different views. Scale bars: a, b, d = 0.2 mm; c = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5341448" figureDoi-1="http://doi.org/10.5281/zenodo.5341452" httpUri-0="https://zenodo.org/record/5341448/files/figure.png" httpUri-1="https://zenodo.org/record/5341452/files/figure.png" pageId="5" pageNumber="428">Figs. 4-5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1CDE653DFFE02862744BFE9FFCD9FE63" pageId="5" pageNumber="428" type="materials_examined">
<paragraph id="547B36B6FFE02862744BFE9FFD8DFE8F" blockId="5.[113,1089,249,452]" box="[113,573,283,303]" pageId="5" pageNumber="428">
<emphasis id="66B0EAA4FFE02862744BFE9FFF5AFE8F" bold="true" box="[113,234,283,303]" pageId="5" pageNumber="428">Type locality.</emphasis>
<collectingRegion id="9600F854FFE0286274D5FE9FFEC1FE8F" box="[239,369,284,303]" country="Spain" name="Canarias" pageId="5" pageNumber="428">Canary Islands</collectingRegion>
, Tenerife, Las Cañadas.
</paragraph>
<paragraph id="547B36B6FFE02862744BFEC5FE8CFE2E" blockId="5.[113,1089,249,452]" pageId="5" pageNumber="428">
<emphasis id="66B0EAA4FFE02862744BFEC5FF45FEF9" bold="true" box="[113,245,325,345]" pageId="5" pageNumber="428">Type material.</emphasis>
<typeStatus id="8B7F8814FFE0286274C3FEC5FEFCFEF7" box="[249,332,326,345]" pageId="5" pageNumber="428" type="holotype">HOLOTYPE</typeStatus>
: J, 25W/HE2255 / Tenerife, Las Cañadas / Fecha: 13-VII-95 / A. Camacho, leg. [white, type- and handwritten label] //
<typeStatus id="8B7F8814FFE028627542FEE3FE5DFED3" box="[376,493,352,371]" pageId="5" pageNumber="428" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="93C44D35FFE0286275C1FEE2FD08FED3" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[507,696,353,371]" class="Insecta" family="Miridae" genus="Atomoscelis" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="428" phylum="Arthropoda" rank="species" species="pictifrons" status="sp. nov.">
<emphasis id="66B0EAA4FFE0286275C1FEE2FD08FED3" box="[507,696,353,371]" italics="true" pageId="5" pageNumber="428">Atomoscelis pictifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE028627686FEE2FD5AFED4" box="[700,746,353,372]" pageId="5" pageNumber="428" rank="species">n. sp.</taxonomicNameLabel>
/ J. Ribes, Pagola-Carte / &amp; Heiss, 2008 [red, typewritten label].
</paragraph>
<paragraph id="547B36B6FFE0286274AAFE16FCD9FE63" blockId="5.[113,1089,249,452]" pageId="5" pageNumber="428">
<materialsCitation id="E4AC3CEBFFE0286274AAFE16FCD9FE63" collectorName="J. Ribes" country="Spain" location="Barcelona" pageId="5" pageNumber="428" specimenCount="1" typeStatus="holotype">
<typeStatus id="8B7F8814FFE0286274AAFE16FF52FE08" box="[144,226,405,424]" pageId="5" pageNumber="428" type="holotype">Holotype</typeStatus>
, in a rather bad condition, mounted on a card with the genitalia and some appendages or parts of them glued on a second card below. Deposited in the collection of
<collectorName id="F9315360FFE028627643FE32FD0DFE63" box="[633,701,432,452]" pageId="5" pageNumber="428">J. Ribes</collectorName>
(
<location id="511B606DFFE0286276F2FE33FC92FE63" LSID="urn:lsid:plazi:treatment:DC6D87A0FFE02860758BFF59FC0AF994:511B606DFFE0286276F2FE33FC92FE63" box="[712,802,432,451]" country="Spain" name="Barcelona" pageId="5" pageNumber="428">Barcelona</location>
,
<collectingCountry id="2CD37626FFE028627710FE33FCEEFE63" box="[810,862,432,451]" name="Spain" pageId="5" pageNumber="428">Spain</collectingCountry>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="1CDE653DFFE02860744BFE58FEE0FCF3" lastPageId="7" lastPageNumber="430" pageId="5" pageNumber="428" type="description">
<paragraph id="547B36B6FFE02862744BFE58FDE6FE53" blockId="5.[113,1089,475,660]" box="[113,598,475,500]" pageId="5" pageNumber="428">
<emphasis id="66B0EAA4FFE02862744BFE58FF4CFE53" bold="true" box="[113,252,475,499]" pageId="5" pageNumber="428">Description.</emphasis>
Length: approximately
<quantity id="933C9B53FFE0286275C7FE5FFDE2FE53" box="[509,594,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="428" unit="mm" value="2.3">2.3 mm</quantity>
.
</paragraph>
<paragraph id="547B36B6FFE0286274AAFE7FFEE4FDF4" blockId="5.[113,1089,475,660]" pageId="5" pageNumber="428">
General aspect pale greyish; frons, base of corium and cuneus faintly darker. Pronotal calli brownish red. A bright, purple-red, tear- or horn-shaped spot beside the inner margin of each eye (
<figureCitation id="CCFF2A33FFE0286274C3FDBFFEF8FDF4" box="[249,328,572,596]" captionStart="Fig" captionStartId="5.[113,144,1539,1559]" captionTargetBox="[151,1077,710,1506]" captionTargetId="figure-159@5.[114,1089,696,1513]" captionTargetPageId="5" captionText="Fig. 4. Atomoscelis pictifrons sp. nov., holotype. a right antenna; b head in lateral view; c head and pronotum in dorsal view; d fore leg; e claw of middle leg; f pygophore in lateral view. Different scales." figureDoi="http://doi.org/10.5281/zenodo.5341448" httpUri="https://zenodo.org/record/5341448/files/figure.png" pageId="5" pageNumber="428">Fig. 4c</figureCitation>
).
</paragraph>
<paragraph id="547B36B6FFE0286274AAFDDFFE11FD34" blockId="5.[113,1089,475,660]" pageId="5" pageNumber="428">Dorsal vestiture consisting of pale, shining, reclining hairs, which appear darker depending on the angle of illumination.</paragraph>
<caption id="00BB663EFFE02862744BF980FC09F991" ID-DOI="http://doi.org/10.5281/zenodo.5341448" ID-Zenodo-Dep="5341448" httpUri="https://zenodo.org/record/5341448/files/figure.png" pageId="5" pageNumber="428" startId="5.[113,144,1539,1559]" targetBox="[151,1077,710,1506]" targetPageId="5">
<paragraph id="547B36B6FFE02862744BF980FC09F991" blockId="5.[113,1089,1539,1586]" pageId="5" pageNumber="428">
<emphasis id="66B0EAA4FFE02862744BF980FF19F9B7" bold="true" box="[113,169,1539,1559]" pageId="5" pageNumber="428">Fig. 4.</emphasis>
<taxonomicName id="93C44D35FFE028627495F986FEDCF9B7" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[175,364,1541,1559]" class="Insecta" family="Miridae" genus="Atomoscelis" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="428" phylum="Arthropoda" rank="species" species="pictifrons" status="sp. nov.">
<emphasis id="66B0EAA4FFE028627495F986FEDCF9B7" box="[175,364,1541,1559]" italics="true" pageId="5" pageNumber="428">Atomoscelis pictifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE02862754BF987FE06F9B7" box="[369,438,1540,1559]" pageId="5" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
, holotype. a right antenna; b head in lateral view; c head and pronotum in dorsal view; d fore leg; e claw of middle leg; f pygophore in lateral view. Different scales.
</paragraph>
</caption>
<paragraph id="547B36B6FFE32861748FFF58FD0EFEF3" blockId="6.[151,1127,219,468]" pageId="6" pageNumber="429">
Head (
<figureCitation id="CCFF2A33FFE3286174C4FF58FEF8FF53" box="[254,328,219,243]" captionStart="Fig" captionStartId="5.[113,144,1539,1559]" captionTargetBox="[151,1077,710,1506]" captionTargetId="figure-159@5.[114,1089,696,1513]" captionTargetPageId="5" captionText="Fig. 4. Atomoscelis pictifrons sp. nov., holotype. a right antenna; b head in lateral view; c head and pronotum in dorsal view; d fore leg; e claw of middle leg; f pygophore in lateral view. Different scales." figureDoi="http://doi.org/10.5281/zenodo.5341448" httpUri="https://zenodo.org/record/5341448/files/figure.png" pageId="6" pageNumber="429">Figs. 4</figureCitation>
b-c) about 2.0 times as high as long in lateral view, 1.25 times as wide as high in frontal view, and 2.0 times as wide as long in dorsal view. Eyes extending over the upper half of head in lateral view. Vertex smooth and slightly concave; frons vertical. Clypeus nearly vertical and arising between antennal fossae.
</paragraph>
<paragraph id="547B36B6FFE32861748FFEDFFCCAFE33" blockId="6.[151,1127,219,468]" pageId="6" pageNumber="429">Rostrum reaching metacoxae. Segments I, II and III pale, segment IV partly darkened distally; segment II slightly longer than segments III and IV together.</paragraph>
<paragraph id="547B36B6FFE32860748FFE1FFD61FEB3" blockId="6.[151,1127,219,468]" lastBlockId="7.[113,1089,219,1588]" lastPageId="7" lastPageNumber="430" pageId="6" pageNumber="429">
Antennae as in
<figureCitation id="CCFF2A33FFE328617568FE18FE2CFE14" box="[338,412,411,436]" captionStart="Fig" captionStartId="5.[113,144,1539,1559]" captionTargetBox="[151,1077,710,1506]" captionTargetId="figure-159@5.[114,1089,696,1513]" captionTargetPageId="5" captionText="Fig. 4. Atomoscelis pictifrons sp. nov., holotype. a right antenna; b head in lateral view; c head and pronotum in dorsal view; d fore leg; e claw of middle leg; f pygophore in lateral view. Different scales." figureDoi="http://doi.org/10.5281/zenodo.5341448" httpUri="https://zenodo.org/record/5341448/files/figure.png" pageId="6" pageNumber="429">Fig. 4a</figureCitation>
. Segment I fusiform, darkened in its basal quarter and with two small, dark, subapical dots; segment II gradually thickened apically but not clavate, its diameter near apex approximately equal to that of segment I; segments III and IV slender. Length of antennal segments I-II-III-IV = 0.25-0.65-0.45-
<quantity id="933C9B53FFE228607653FF78FD7DFEB3" box="[617,717,251,275]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="430" unit="mm" value="0.3">0.30 mm</quantity>
.
</paragraph>
<caption id="00BB663EFFE3286174A3F98BFBB4F997" ID-DOI="http://doi.org/10.5281/zenodo.5341452" ID-Zenodo-Dep="5341452" httpUri="https://zenodo.org/record/5341452/files/figure.png" pageId="6" pageNumber="429" startId="6.[153,184,1544,1564]" targetBox="[248,1058,516,1514]" targetPageId="6">
<paragraph id="547B36B6FFE3286174A3F98BFBB4F997" blockId="6.[151,1126,1544,1591]" pageId="6" pageNumber="429">
<emphasis id="66B0EAA4FFE3286174A3F98BFF61F9BC" bold="true" box="[153,209,1544,1564]" pageId="6" pageNumber="429">Fig. 5.</emphasis>
Male genitalia of
<taxonomicName id="93C44D35FFE328617554F989FD9CF9BC" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[366,556,1546,1564]" class="Insecta" family="Miridae" genus="Atomoscelis" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="429" phylum="Arthropoda" rank="species" species="pictifrons" status="sp. nov.">
<emphasis id="66B0EAA4FFE328617554F989FD9CF9BC" box="[366,556,1546,1564]" italics="true" pageId="6" pageNumber="429">Atomoscelis pictifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE328617608F98AFDC5F9BC" box="[562,629,1545,1564]" pageId="6" pageNumber="429" rank="species">sp. nov.</taxonomicNameLabel>
a left paramere in different views; b right paramere; c apical portion of phallotheca; d vesica in different views. Scale bars: a, b, d = 0.2 mm; c = 0.1 mm.
</paragraph>
</caption>
<paragraph id="547B36B6FFE2286074AAFE98FEEFFE94" blockId="7.[113,1089,219,1588]" box="[144,351,283,308]" pageId="7" pageNumber="430">Ocular index = 1.8.</paragraph>
<paragraph id="547B36B6FFE2286074AAFEBFFF1DFE33" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="430">
Pronotum (
<figureCitation id="CCFF2A33FFE228607532FEB8FEE4FEF4" box="[264,340,315,340]" captionStart="Fig" captionStartId="5.[113,144,1539,1559]" captionTargetBox="[151,1077,710,1506]" captionTargetId="figure-159@5.[114,1089,696,1513]" captionTargetPageId="5" captionText="Fig. 4. Atomoscelis pictifrons sp. nov., holotype. a right antenna; b head in lateral view; c head and pronotum in dorsal view; d fore leg; e claw of middle leg; f pygophore in lateral view. Different scales." figureDoi="http://doi.org/10.5281/zenodo.5341448" httpUri="https://zenodo.org/record/5341448/files/figure.png" pageId="7" pageNumber="430">Fig. 4c</figureCitation>
) trapezoidal, flattened, with straight lateral sides, 2.4 times as wide as long and 1.35 times as wide as diatone. Antennal segment II 0.88 times as long as pronotum wide.
</paragraph>
<paragraph id="547B36B6FFE2286074AAFE1FFE31FDD3" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="430">
Legs pale (hind legs missing in the single specimen studied). Fore leg as in
<figureCitation id="CCFF2A33FFE22860779FFE18FC42FE13" box="[933,1010,411,435]" captionStart="Fig" captionStartId="5.[113,144,1539,1559]" captionTargetBox="[151,1077,710,1506]" captionTargetId="figure-159@5.[114,1089,696,1513]" captionTargetPageId="5" captionText="Fig. 4. Atomoscelis pictifrons sp. nov., holotype. a right antenna; b head in lateral view; c head and pronotum in dorsal view; d fore leg; e claw of middle leg; f pygophore in lateral view. Different scales." figureDoi="http://doi.org/10.5281/zenodo.5341448" httpUri="https://zenodo.org/record/5341448/files/figure.png" pageId="7" pageNumber="430">Fig. 4d</figureCitation>
. Femur with a black spot on ventral side. Tibia with scattered black spots, from which dark, nearly black, spines arise; spines longer than or as long as tibial diameter, gradually thickened apically. Tarsi pale except segment III, the latter gradually darkened on its distal half; segment III as long as segments I and II together. Claws slender (
<figureCitation id="CCFF2A33FFE2286076D0FD98FC8BFD94" box="[746,827,539,564]" captionStart="Fig" captionStartId="5.[113,144,1539,1559]" captionTargetBox="[151,1077,710,1506]" captionTargetId="figure-159@5.[114,1089,696,1513]" captionTargetPageId="5" captionText="Fig. 4. Atomoscelis pictifrons sp. nov., holotype. a right antenna; b head in lateral view; c head and pronotum in dorsal view; d fore leg; e claw of middle leg; f pygophore in lateral view. Different scales." figureDoi="http://doi.org/10.5281/zenodo.5341448" httpUri="https://zenodo.org/record/5341448/files/figure.png" pageId="7" pageNumber="430">Fig. 4e</figureCitation>
, middle leg), uniformly curved and narrowed towards apex, and provided with small but distinct pulvilli, about 0.2 times as long as the claw.
</paragraph>
<paragraph id="547B36B6FFE2286074AAFDF8FD09FD33" blockId="7.[113,1089,219,1588]" box="[144,697,635,659]" pageId="7" pageNumber="430">
Pygophore ventrally beset with stout setae (
<figureCitation id="CCFF2A33FFE22860765BFDF8FD1EFD33" box="[609,686,635,659]" captionStart="Fig" captionStartId="5.[113,144,1539,1559]" captionTargetBox="[151,1077,710,1506]" captionTargetId="figure-159@5.[114,1089,696,1513]" captionTargetPageId="5" captionText="Fig. 4. Atomoscelis pictifrons sp. nov., holotype. a right antenna; b head in lateral view; c head and pronotum in dorsal view; d fore leg; e claw of middle leg; f pygophore in lateral view. Different scales." figureDoi="http://doi.org/10.5281/zenodo.5341448" httpUri="https://zenodo.org/record/5341448/files/figure.png" pageId="7" pageNumber="430">Fig. 4f</figureCitation>
).
</paragraph>
<paragraph id="547B36B6FFE2286074AAFD18FE19FC93" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="430">
Male genitalia. Parameres small; left paramere (
<figureCitation id="CCFF2A33FFE2286076BCFD18FD60FD13" box="[646,720,667,691]" captionStart="Fig" captionStartId="6.[153,184,1544,1564]" captionTargetBox="[248,1058,516,1514]" captionTargetId="figure-122@6.[216,1062,513,1521]" captionTargetPageId="6" captionText="Fig. 5. Male genitalia of Atomoscelis pictifrons sp. nov. a left paramere in different views; b right paramere; c apical portion of phallotheca; d vesica in different views. Scale bars: a, b, d = 0.2 mm; c = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341452" httpUri="https://zenodo.org/record/5341452/files/figure.png" pageId="7" pageNumber="430">Fig. 5a</figureCitation>
) wide, with two pointed processes, posterior one longer than anterior. Right paramere (
<figureCitation id="CCFF2A33FFE2286076A9FD38FD50FD73" box="[659,736,699,723]" captionStart="Fig" captionStartId="6.[153,184,1544,1564]" captionTargetBox="[248,1058,516,1514]" captionTargetId="figure-122@6.[216,1062,513,1521]" captionTargetPageId="6" captionText="Fig. 5. Male genitalia of Atomoscelis pictifrons sp. nov. a left paramere in different views; b right paramere; c apical portion of phallotheca; d vesica in different views. Scale bars: a, b, d = 0.2 mm; c = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341452" httpUri="https://zenodo.org/record/5341452/files/figure.png" pageId="7" pageNumber="430">Fig. 5b</figureCitation>
) apically truncate and difficult to see due to its transparency. Apical portion of phallotheca as in
<figureCitation id="CCFF2A33FFE228607736FD58FCE9FD53" box="[780,857,731,755]" captionStart="Fig" captionStartId="6.[153,184,1544,1564]" captionTargetBox="[248,1058,516,1514]" captionTargetId="figure-122@6.[216,1062,513,1521]" captionTargetPageId="6" captionText="Fig. 5. Male genitalia of Atomoscelis pictifrons sp. nov. a left paramere in different views; b right paramere; c apical portion of phallotheca; d vesica in different views. Scale bars: a, b, d = 0.2 mm; c = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341452" httpUri="https://zenodo.org/record/5341452/files/figure.png" pageId="7" pageNumber="430">Fig. 5c</figureCitation>
. Vesica (
<figureCitation id="CCFF2A33FFE22860778CFD58FBB4FD53" box="[950,1028,731,755]" captionStart="Fig" captionStartId="6.[153,184,1544,1564]" captionTargetBox="[248,1058,516,1514]" captionTargetId="figure-122@6.[216,1062,513,1521]" captionTargetPageId="6" captionText="Fig. 5. Male genitalia of Atomoscelis pictifrons sp. nov. a left paramere in different views; b right paramere; c apical portion of phallotheca; d vesica in different views. Scale bars: a, b, d = 0.2 mm; c = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341452" httpUri="https://zenodo.org/record/5341452/files/figure.png" pageId="7" pageNumber="430">Fig. 5d</figureCitation>
) J- to S-shaped, with two apical, subequally long processes and with secondary gonopore located beyond their diverging point.
</paragraph>
<paragraph id="547B36B6FFE2286074AAFCB8FEE0FCF3" blockId="7.[113,1089,219,1588]" box="[144,336,827,851]" pageId="7" pageNumber="430">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="1CDE653DFFE22860744BFCDFFBBAFCD3" box="[113,1034,859,884]" pageId="7" pageNumber="430" type="etymology">
<paragraph id="547B36B6FFE22860744BFCDFFBBAFCD3" blockId="7.[113,1089,219,1588]" box="[113,1034,859,884]" pageId="7" pageNumber="430">
<emphasis id="66B0EAA4FFE22860744BFCDFFF42FCD4" bold="true" box="[113,242,860,884]" pageId="7" pageNumber="430">Etymology.</emphasis>
This species is named for the distinctive red, mask-like spots on the head.
</paragraph>
</subSubSection>
<subSubSection id="1CDE653DFFE22860744BFCF8FB89FC33" box="[113,1081,891,916]" pageId="7" pageNumber="430" type="distribution">
<paragraph id="547B36B6FFE22860744BFCF8FB89FC33" blockId="7.[113,1089,219,1588]" box="[113,1081,891,916]" pageId="7" pageNumber="430">
<emphasis id="66B0EAA4FFE22860744BFCF8FEB4FC33" bold="true" box="[113,260,891,915]" pageId="7" pageNumber="430">Distribution.</emphasis>
Up to date, only the
<typeStatus id="8B7F8814FFE2286075DEFCF8FDF1FC33" box="[484,577,891,915]" pageId="7" pageNumber="430" type="holotype">holotype</typeStatus>
has been collected in Tenerife,
<collectingRegion id="9600F854FFE2286077A9FCFFFB85FC33" box="[915,1077,891,916]" country="Spain" name="Canarias" pageId="7" pageNumber="430">Canary Islands</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="1CDE653DFFE22860744BFC18FC0AF994" pageId="7" pageNumber="430" type="discussion">
<paragraph id="547B36B6FFE22860744BFC18FDC3FBF4" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="430">
<emphasis id="66B0EAA4FFE22860744BFC18FF41FC13" bold="true" box="[113,241,923,947]" pageId="7" pageNumber="430">Discussion.</emphasis>
The new species undoubtedly belongs to the genus
<taxonomicName id="93C44D35FFE228607735FC1EFC20FC14" authorityName="Reuter" authorityYear="1875" box="[783,912,925,948]" class="Insecta" family="Miridae" genus="Atomoscelis" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="66B0EAA4FFE228607735FC1EFC20FC14" box="[783,912,925,948]" italics="true" pageId="7" pageNumber="430">Atomoscelis</emphasis>
</taxonomicName>
, since it fits well all its diagnostic characters (see
<bibRefCitation id="30554B47FFE2286075F5FC3FFDD9FC73" author="WAGNER E." box="[463,617,955,979]" pageId="7" pageNumber="430" pagination="1 - 483" refId="ref3919" refString="WAGNER E. 1975: Die Miridae Hahn, 1831, des Mittelmeerraumes und der Makaronesischen Inseln (Hemiptera, Heteroptera). Teil 3. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden 40 (Suppl.) (1975): 1 - 483." type="journal article" year="1975">WAGNER 1975</bibRefCitation>
). Particularly noticeable in this genus is the shape of the head (
<figureCitation id="CCFF2A33FFE22860750BFC58FECFFC53" box="[305,383,987,1011]" captionStart="Fig" captionStartId="5.[113,144,1539,1559]" captionTargetBox="[151,1077,710,1506]" captionTargetId="figure-159@5.[114,1089,696,1513]" captionTargetPageId="5" captionText="Fig. 4. Atomoscelis pictifrons sp. nov., holotype. a right antenna; b head in lateral view; c head and pronotum in dorsal view; d fore leg; e claw of middle leg; f pygophore in lateral view. Different scales." figureDoi="http://doi.org/10.5281/zenodo.5341448" httpUri="https://zenodo.org/record/5341448/files/figure.png" pageId="7" pageNumber="430">Fig. 4b</figureCitation>
).
<bibRefCitation id="30554B47FFE2286075B5FC5FFD84FC53" author="WAGNER E." box="[399,564,987,1011]" pageId="7" pageNumber="430" pagination="74 - 92" refId="ref3840" refString="WAGNER E. 1965: Die Gattung Atomoscelis Reuter, 1875 und ihre Verwandten in der Palaarktis (Hem. Het. Miridae). Notulae Entomologicae 45: 74 - 92." type="journal article" year="1965">WAGNER (1965)</bibRefCitation>
previously discussed and compared the characters of
<taxonomicName id="93C44D35FFE2286074B5FC7EFEA0FBB4" authorityName="Reuter" authorityYear="1875" box="[143,272,1021,1044]" class="Insecta" family="Miridae" genus="Atomoscelis" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="430" phylum="Arthropoda" rank="genus">
<emphasis id="66B0EAA4FFE2286074B5FC7EFEA0FBB4" box="[143,272,1021,1044]" italics="true" pageId="7" pageNumber="430">Atomoscelis</emphasis>
</taxonomicName>
and related genera, and
<bibRefCitation id="30554B47FFE22860762CFC7FFD77FBB3" author="LINDBERG H." box="[534,711,1019,1043]" pageId="7" pageNumber="430" pagination="1 - 304" refId="ref3638" refString="LINDBERG H. 1953: Hemiptera Insularum Canariensium. Commentationes Biologicae 14 (1): 1 - 304." type="journal article" year="1953">LINDBERG (1953)</bibRefCitation>
and
<bibRefCitation id="30554B47FFE2286076C1FC7FFC13FBB3" author="WAGNER E." box="[763,931,1019,1043]" pageId="7" pageNumber="430" pagination="74 - 92" refId="ref3840" refString="WAGNER E. 1965: Die Gattung Atomoscelis Reuter, 1875 und ihre Verwandten in der Palaarktis (Hem. Het. Miridae). Notulae Entomologicae 45: 74 - 92." type="journal article" year="1965">WAGNER (1965)</bibRefCitation>
provided keys to species and/or to the related genera; the keys are now of limited value due to subsequent taxonomic changes (KERZHNER &amp; JOSIFOV 1999).
</paragraph>
<paragraph id="547B36B6FFE2286074AAFBDEFD22FB54" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="430">
<taxonomicName id="93C44D35FFE2286074AAFBDEFEC8FBD4" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[144,376,1117,1140]" class="Insecta" family="Miridae" genus="Atomoscelis" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="430" phylum="Arthropoda" rank="species" species="pictifrons" status="sp. nov.">
<emphasis id="66B0EAA4FFE2286074AAFBDEFEC8FBD4" box="[144,376,1117,1140]" italics="true" pageId="7" pageNumber="430">Atomoscelis pictifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE228607546FBDFFE7CFBD3" box="[380,460,1116,1139]" pageId="7" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
can be easily separated from all other European and northern African species of the genus by the mask-like pair of purple-red spots on the head and the brownish-red pronotal calli (
<figureCitation id="CCFF2A33FFE22860759CFB18FE45FB14" box="[422,501,1179,1204]" captionStart="Fig" captionStartId="5.[113,144,1539,1559]" captionTargetBox="[151,1077,710,1506]" captionTargetId="figure-159@5.[114,1089,696,1513]" captionTargetPageId="5" captionText="Fig. 4. Atomoscelis pictifrons sp. nov., holotype. a right antenna; b head in lateral view; c head and pronotum in dorsal view; d fore leg; e claw of middle leg; f pygophore in lateral view. Different scales." figureDoi="http://doi.org/10.5281/zenodo.5341448" httpUri="https://zenodo.org/record/5341448/files/figure.png" pageId="7" pageNumber="430">Fig. 4c</figureCitation>
). It is also distinguished by the relatively small ocular index (
<quantity id="933C9B53FFE228607481FB3FFF49FB73" box="[187,249,1211,1236]" metricMagnitude="-2" metricUnit="m" metricValue="4.572" pageId="7" pageNumber="430" unit="in" value="1.8">1.8 in</quantity>
the male
<typeStatus id="8B7F8814FFE22860755BFB38FE73FB73" box="[353,451,1211,1235]" pageId="7" pageNumber="430" type="holotype">holotype</typeStatus>
), which is usually larger than 2.2 (males and females) in the remaining species (
<bibRefCitation id="30554B47FFE228607505FB5FFE69FB53" author="WAGNER E." box="[319,473,1243,1268]" pageId="7" pageNumber="430" pagination="1 - 483" refId="ref3919" refString="WAGNER E. 1975: Die Miridae Hahn, 1831, des Mittelmeerraumes und der Makaronesischen Inseln (Hemiptera, Heteroptera). Teil 3. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden 40 (Suppl.) (1975): 1 - 483." type="journal article" year="1975">WAGNER 1975</bibRefCitation>
,
<bibRefCitation id="30554B47FFE2286075D9FB5FFD36FB54" author="TAMANINI L." box="[483,646,1244,1268]" pageId="7" pageNumber="430" pagination="59 - 70" refId="ref3807" refString="TAMANINI L. 1978: Due nuovi Miridi endemiti della Sardegna (Heteroptera, Miridae). Bollettino della Societa Sarda di Scienze Naturali 17 (1977): 59 - 70." type="journal article" year="1978">TAMANINI 1978</bibRefCitation>
).
</paragraph>
<paragraph id="547B36B6FFE2286074AAFB78FC6EFAF4" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="430">
The most closely related taxon to
<taxonomicName id="93C44D35FFE2286075C0FB7EFDCCFAB4" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[506,636,1277,1300]" class="Insecta" family="Miridae" genus="Atomoscelis" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="430" phylum="Arthropoda" rank="species" species="pictifrons" status="sp. nov.">
<emphasis id="66B0EAA4FFE2286075C0FB7EFDCCFAB4" box="[506,636,1277,1300]" italics="true" pageId="7" pageNumber="430">A. pictifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE2286076B9FB7FFD66FAB3" box="[643,726,1276,1299]" pageId="7" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
is
<taxonomicName id="93C44D35FFE2286076CFFB7EFCEAFAB4" authorityName="AUKEMA et al." authorityYear="2006" baseAuthorityName="Fieber" baseAuthorityYear="1861" box="[757,858,1277,1300]" class="Insecta" family="Miridae" genus="Atomoscelis" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="430" phylum="Arthropoda" rank="species" species="onusta">
<emphasis id="66B0EAA4FFE2286076CFFB7EFCEAFAB4" box="[757,858,1277,1300]" italics="true" pageId="7" pageNumber="430">A. onusta</emphasis>
</taxonomicName>
. Both species share a similar habitus and the shape of the vesica. However, the apical processes of the vesica have different relative lengths (compare
<figureCitation id="CCFF2A33FFE2286075D3FAB8FD84FAF3" box="[489,564,1339,1363]" captionStart="Fig" captionStartId="6.[153,184,1544,1564]" captionTargetBox="[248,1058,516,1514]" captionTargetId="figure-122@6.[216,1062,513,1521]" captionTargetPageId="6" captionText="Fig. 5. Male genitalia of Atomoscelis pictifrons sp. nov. a left paramere in different views; b right paramere; c apical portion of phallotheca; d vesica in different views. Scale bars: a, b, d = 0.2 mm; c = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5341452" httpUri="https://zenodo.org/record/5341452/files/figure.png" pageId="7" pageNumber="430">Fig. 5d</figureCitation>
with Figs. 44b-d in
<bibRefCitation id="30554B47FFE228607737FABFFC6AFAF4" author="CARAPEZZA A." box="[781,986,1340,1364]" pageId="7" pageNumber="430" pagination="1 - 312" refId="ref3616" refString="CARAPEZZA A. 1997: Heteroptera of Tunisia. Naturalista Siciliano 21 (Suppl. A): 1 - 312." type="journal article" year="1997">CARAPEZZA (1997))</bibRefCitation>
.
</paragraph>
<paragraph id="547B36B6FFE2286074AAFADFFC0AF994" blockId="7.[113,1089,219,1588]" pageId="7" pageNumber="430">
In addition,
<taxonomicName id="93C44D35FFE228607529FADEFDF7FAD4" authority="Tamanini, 1978" authorityName="Tamanini" authorityYear="1978" box="[275,583,1371,1396]" class="Insecta" family="Miridae" genus="Atomoscelis" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="430" phylum="Arthropoda" rank="species" species="galvagnii">
<emphasis id="66B0EAA4FFE228607529FADEFE26FAD4" box="[275,406,1373,1396]" italics="true" pageId="7" pageNumber="430">A. galvagnii</emphasis>
<bibRefCitation id="30554B47FFE2286075A7FAD8FDF7FAD4" author="TAMANINI L." box="[413,583,1371,1396]" pageId="7" pageNumber="430" pagination="59 - 70" refId="ref3807" refString="TAMANINI L. 1978: Due nuovi Miridi endemiti della Sardegna (Heteroptera, Miridae). Bollettino della Societa Sarda di Scienze Naturali 17 (1977): 59 - 70." type="journal article" year="1978">Tamanini, 1978</bibRefCitation>
</taxonomicName>
, has a different colouration of the dorsal surface, one dark spot on antennal segment I, and a much more slender vesica (
<bibRefCitation id="30554B47FFE2286077A2FAFFFB8DFA34" author="TAMANINI L." box="[920,1085,1404,1428]" pageId="7" pageNumber="430" pagination="59 - 70" refId="ref3807" refString="TAMANINI L. 1978: Due nuovi Miridi endemiti della Sardegna (Heteroptera, Miridae). Bollettino della Societa Sarda di Scienze Naturali 17 (1977): 59 - 70." type="journal article" year="1978">TAMANINI 1978</bibRefCitation>
,
<bibRefCitation id="30554B47FFE22860744BFA1FFE93FA14" author="CARAPEZZA A." box="[113,291,1436,1460]" pageId="7" pageNumber="430" pagination="1 - 312" refId="ref3616" refString="CARAPEZZA A. 1997: Heteroptera of Tunisia. Naturalista Siciliano 21 (Suppl. A): 1 - 312." type="journal article" year="1997">CARAPEZZA 1997</bibRefCitation>
). The vesica is also markedly different in
<taxonomicName id="93C44D35FFE2286076E1FA1EFC44FA14" authority="Linnavuori, 1971" authorityName="Linnavuori" authorityYear="1971" box="[731,1012,1435,1460]" class="Insecta" family="Miridae" genus="Atomoscelis" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="430" phylum="Arthropoda" rank="species" species="atthis">
<emphasis id="66B0EAA4FFE2286076E1FA1EFC83FA14" box="[731,819,1437,1460]" italics="true" pageId="7" pageNumber="430">A. atthis</emphasis>
<bibRefCitation id="30554B47FFE228607703FA18FC44FA14" author="LINNAVUORI R." box="[825,1012,1435,1460]" pageId="7" pageNumber="430" pagination="27 - 33" refId="ref3658" refString="LINNAVUORI R. 1971: On some new or little known Miridae species. Miscelanea Zoologica 3 (1): 27 - 33." type="journal article" year="1971">Linnavuori, 1971</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="93C44D35FFE228607010FA1EFEC2FA74" authority="Reuter, 1902" authorityName="Reuter" authorityYear="1902" class="Insecta" family="Miridae" genus="Atomoscelis" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="430" phylum="Arthropoda" rank="species" species="noualhieri">
<emphasis id="66B0EAA4FFE228607010FA1EFF50FA74" italics="true" pageId="7" pageNumber="430">A. noualhieri</emphasis>
Reuter, 1902
</taxonomicName>
, being stouter and having more basally located secondary gonopore in both species.
<taxonomicName id="93C44D35FFE22860751BFA5EFDBFFA54" authority="Ribes &amp; Pagola-Carte &amp; Heiss, 2008" authorityName="Ribes &amp; Pagola-Carte &amp; Heiss" authorityYear="2008" box="[289,527,1501,1524]" class="Insecta" family="Miridae" genus="Atomoscelis" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="430" phylum="Arthropoda" rank="species" species="pictifrons" status="sp. nov.">
<emphasis id="66B0EAA4FFE22860751BFA5EFDBFFA54" box="[289,527,1501,1524]" italics="true" pageId="7" pageNumber="430">Atomoscelis pictifrons</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7D8357DFFFE22860762DFA5FFDDCFA53" box="[535,620,1500,1523]" pageId="7" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
also differs from these mainly Maghrebian species by the lack of longitudinal reddish stripes on the pronotum and dark dots on the hemelytra (
<bibRefCitation id="30554B47FFE2286074D0F99FFE12F994" author="LINNAVUORI R." box="[234,418,1564,1588]" pageId="7" pageNumber="430" pagination="27 - 33" refId="ref3658" refString="LINNAVUORI R. 1971: On some new or little known Miridae species. Miscelanea Zoologica 3 (1): 27 - 33." type="journal article" year="1971">LINNAVUORI 1971</bibRefCitation>
,
<bibRefCitation id="30554B47FFE228607596F99FFDF4F993" author="WAGNER E." box="[428,580,1563,1587]" pageId="7" pageNumber="430" pagination="1 - 483" refId="ref3919" refString="WAGNER E. 1975: Die Miridae Hahn, 1831, des Mittelmeerraumes und der Makaronesischen Inseln (Hemiptera, Heteroptera). Teil 3. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden 40 (Suppl.) (1975): 1 - 483." type="journal article" year="1975">WAGNER 1975</bibRefCitation>
,
<bibRefCitation id="30554B47FFE228607674F99FFD40F994" author="TAMANINI L." box="[590,752,1564,1588]" pageId="7" pageNumber="430" pagination="59 - 70" refId="ref3807" refString="TAMANINI L. 1978: Due nuovi Miridi endemiti della Sardegna (Heteroptera, Miridae). Bollettino della Societa Sarda di Scienze Naturali 17 (1977): 59 - 70." type="journal article" year="1978">TAMANINI 1978</bibRefCitation>
,
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).
</paragraph>
</subSubSection>
</treatment>
</document>