215 lines
24 KiB
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215 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1045.63810" ID-GBIF-Dataset="e3ab2700-b888-4721-b7c1-fcb2d0bb38b2" ID-PMC="PMC8233300" ID-Pensoft-Pub="1313-2970-1045-1" ID-Pensoft-UUID="CDADD9D94DEB5471834AF0F1AB8B48AA" ID-PubMed="34228772" ID-ZooBank="2C3076FD13FB4842A7F6B0EBE9B23795" ModsDocID="1313-2970-1045-1" checkinTime="1624048151249" checkinUser="pensoft" docAuthor="Giron, Jennifer C. & Short, Andrew Edward Z." docDate="2021" docId="B8136D6F704C9A23DBC94DEE71C425A1" docLanguage="en" docName="ZooKeys 1045: 1-236" docOrigin="ZooKeys 1045" docPubDate="2021-06-18" docSource="http://dx.doi.org/10.3897/zookeys.1045.63810" docTitle="Troglochares Spangler 1981" docType="treatment" docVersion="4" id="CDADD9D94DEB5471834AF0F1AB8B48AA" lastPageNumber="1" masterDocId="CDADD9D94DEB5471834AF0F1AB8B48AA" masterDocTitle="The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species" masterLastPageNumber="236" masterPageNumber="1" pageNumber="1" updateTime="1668150528805" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species</mods:title>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Giron, Jennifer C.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0851-6883</mods:nameIdentifier>
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<mods:affiliation>Department of Entomology, Purdue University, West Lafayette, Indiana, USA & Natural Science Research Laboratory, Museum of Texas Tech University, Lubbock, Texas, USA</mods:affiliation>
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<mods:nameIdentifier type="email">entiminae@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Short, Andrew Edward Z.</mods:namePart>
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<mods:affiliation>Department of Ecology and Evolutionary Biology, and Division of Entomology, Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-06-18</mods:number>
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<mods:detail type="volume">
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<mods:number>1045</mods:number>
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<mods:start>1</mods:start>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1045.63810</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1045.63810</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1045-1</mods:identifier>
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<subSection pageId="0" pageNumber="1" type="results">
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<treatment ID-GBIF-Taxon="182959269" LSID="urn:lsid:plazi:treatment:B8136D6F704C9A23DBC94DEE71C425A1" httpUri="http://treatment.plazi.org/id/B8136D6F704C9A23DBC94DEE71C425A1" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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Genus
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<taxonomicName authority="Spangler, 1981" authorityName="Spangler" authorityYear="1981" class="Insecta" family="Hydrophilidae" genus="Troglochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Troglochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Troglochares Spangler, 1981</taxonomicName>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Known distribution of genera of Acidocerinae: Novochares, Peltochares, Primocerus, Quadriops, Radicitus, Sindolus, Tobochares, and Troglochares." figureDoi="10.3897/zookeys.1045.63810.figure6" httpUri="https://binary.pensoft.net/fig/557405" pageId="0" pageNumber="1">Figs 6</figureCitation>
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Holotype and labels of Troglochares ashmolei A mount of holotype B head, dorsal view C labels." figureDoi="10.3897/zookeys.1045.63810.figure56" httpUri="https://binary.pensoft.net/fig/557455" pageId="0" pageNumber="1">, 56</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Spangler" authorityYear="1981" class="Insecta" family="Hydrophilidae" genus="Troglochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Troglochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Troglochares</taxonomicName>
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Spangler, 1981a: 316.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="gender">
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<paragraph pageId="0" pageNumber="1">Gender.</paragraph>
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<paragraph pageId="0" pageNumber="1">Masculine.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type species">
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<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Spangler" authorityYear="1981" class="Insecta" family="Hydrophilidae" genus="Troglochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Troglochares ashmolei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ashmolei">
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<emphasis italics="true" pageId="0" pageNumber="1">Troglochares ashmolei</emphasis>
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</taxonomicName>
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Spangler, 1981a: 318; by original designation and monotypy.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Small beetles, body length 1.9 mm. Body shape oval in dorsal view; moderately convex in lateral view (
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<bibRefCitation author="Hansen, M" journalOrPublisher="Biologiske Skrifter" pageId="0" pageNumber="1" pagination="1 - 367" refId="B139" refString="Hansen, M, 1991. The hydrophiloid beetles. Phylogeny, classification and a revision of the genera (Coleoptera: Hydrophilidae). Biologiske Skrifter 40: 1 - 367" title="The hydrophiloid beetles. Phylogeny, classification and a revision of the genera (Coleoptera: Hydrophilidae)." volume="40" year="1991">Hansen 1991</bibRefCitation>
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: fig. 39). Color yellowish light brown; ground punctation extremely shallowly marked. Shape of head somewhat oval. Eyes absent (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Holotype and labels of Troglochares ashmolei A mount of holotype B head, dorsal view C labels." figureDoi="10.3897/zookeys.1045.63810.figure56" httpUri="https://binary.pensoft.net/fig/557455" pageId="0" pageNumber="1">56B</figureCitation>
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). Clypeus trapezoidal, with anterior margin broadly emarginate, with medial region of emargination nearly straight (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Holotype and labels of Troglochares ashmolei A mount of holotype B head, dorsal view C labels." figureDoi="10.3897/zookeys.1045.63810.figure56" httpUri="https://binary.pensoft.net/fig/557455" pageId="0" pageNumber="1">56B</figureCitation>
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). Labrum fully exposed, convex. Mentum rather smooth and antero-medially depressed; median anterior depression broad. Antennae with nine antennomeres (
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<bibRefCitation author="Spangler, PJ" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="1" pagination="316 - 323" publicationUrl="https://biodiversitylibrary.org/page/16364657" refId="B279" refString="Spangler, PJ, 1981a. A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae). Proceedings of the Entomological Society of Washington 83 (2): 316 - 323, https://biodiversitylibrary.org/page/16364657" title="A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae)." url="https://biodiversitylibrary.org/page/16364657" volume="83" year="1981 a">Spangler 1981a</bibRefCitation>
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: fig. 3); cupule slightly asymmetric, with rounded outline. Maxillary palps slender, nearly as long as width of head; inner margin of maxillary palpomere 2 nearly straight, outer margin curved along apical third; maxillary palpomere 3 slightly shorter than 4. Prosternum non carinate, slightly convex. Elytra without sutural striae; ground punctation fine, shallow; outer margins slightly flared (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Holotype and labels of Troglochares ashmolei A mount of holotype B head, dorsal view C labels." figureDoi="10.3897/zookeys.1045.63810.figure56" httpUri="https://binary.pensoft.net/fig/557455" pageId="0" pageNumber="1">56A</figureCitation>
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). Posterior elevation of mesoventrite with curved, transverse ridge (
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<bibRefCitation author="Spangler, PJ" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="1" pagination="316 - 323" publicationUrl="https://biodiversitylibrary.org/page/16364657" refId="B279" refString="Spangler, PJ, 1981a. A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae). Proceedings of the Entomological Society of Washington 83 (2): 316 - 323, https://biodiversitylibrary.org/page/16364657" title="A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae)." url="https://biodiversitylibrary.org/page/16364657" volume="83" year="1981 a">Spangler 1981a</bibRefCitation>
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: fig. 8); anapleural sutures concave, separated at anterior margin by distance 0.7
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<normalizedToken originalValue="×">x</normalizedToken>
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width of anterior margin of mesepisternum. Metaventrite densely pubescent except for median short and narrow posterior glabrous patch; metaventrite short (nearly as long as first abdominal ventrite;
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<bibRefCitation author="Spangler, PJ" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="1" pagination="316 - 323" publicationUrl="https://biodiversitylibrary.org/page/16364657" refId="B279" refString="Spangler, PJ, 1981a. A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae). Proceedings of the Entomological Society of Washington 83 (2): 316 - 323, https://biodiversitylibrary.org/page/16364657" title="A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae)." url="https://biodiversitylibrary.org/page/16364657" volume="83" year="1981 a">Spangler 1981a</bibRefCitation>
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: fig. 8). Protibiae with spines of anterior row long; apical spurs of protibiae moderately slender, reaching apex of protarsomere 2; metatarsomeres 2-4 slightly decreasing in size; metatarsomere 5 nearly as long as 2-4 combined. Posterior femora densely covered by hydrofuge pubescence along basal 2/3 (
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<bibRefCitation author="Spangler, PJ" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="1" pagination="316 - 323" publicationUrl="https://biodiversitylibrary.org/page/16364657" refId="B279" refString="Spangler, PJ, 1981a. A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae). Proceedings of the Entomological Society of Washington 83 (2): 316 - 323, https://biodiversitylibrary.org/page/16364657" title="A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae)." url="https://biodiversitylibrary.org/page/16364657" volume="83" year="1981 a">Spangler 1981a</bibRefCitation>
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: fig. 8). Fifth abdominal ventrite apically truncate, without stout setae (
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<bibRefCitation author="Spangler, PJ" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="1" pagination="316 - 323" publicationUrl="https://biodiversitylibrary.org/page/16364657" refId="B279" refString="Spangler, PJ, 1981a. A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae). Proceedings of the Entomological Society of Washington 83 (2): 316 - 323, https://biodiversitylibrary.org/page/16364657" title="A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae)." url="https://biodiversitylibrary.org/page/16364657" volume="83" year="1981 a">Spangler 1981a</bibRefCitation>
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: fig. 9).
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</paragraph>
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<caption doi="10.3897/zookeys.1045.63810.figure56" httpUri="https://binary.pensoft.net/fig/557455" pageId="0" pageNumber="1" start="Figure 56" startId="F56">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 56.</emphasis>
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Holotype and labels of
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<taxonomicName authorityName="Spangler" authorityYear="1981" class="Insecta" family="Hydrophilidae" genus="Troglochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Troglochares ashmolei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ashmolei">
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<emphasis italics="true" pageId="0" pageNumber="1">Troglochares ashmolei</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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mount of holotype
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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head, dorsal view
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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labels.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Spangler" authorityYear="1981" class="Insecta" family="Hydrophilidae" genus="Troglochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Troglochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Troglochares</emphasis>
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</taxonomicName>
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is the only genus of acidocerines (and Hydrophilids) lacking eyes.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Neotropical</emphasis>
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: Ecuador; Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Known distribution of genera of Acidocerinae: Novochares, Peltochares, Primocerus, Quadriops, Radicitus, Sindolus, Tobochares, and Troglochares." figureDoi="10.3897/zookeys.1045.63810.figure6" httpUri="https://binary.pensoft.net/fig/557405" pageId="0" pageNumber="1">6</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="natural history">
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<paragraph pageId="0" pageNumber="1">Natural history.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The only known specimen was collected in a cave on calcite formations and is presumably aquatic (
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<bibRefCitation author="Spangler, PJ" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="1" pagination="316 - 323" publicationUrl="https://biodiversitylibrary.org/page/16364657" refId="B279" refString="Spangler, PJ, 1981a. A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae). Proceedings of the Entomological Society of Washington 83 (2): 316 - 323, https://biodiversitylibrary.org/page/16364657" title="A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae)." url="https://biodiversitylibrary.org/page/16364657" volume="83" year="1981 a">Spangler 1981a</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="larvae">
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<paragraph pageId="0" pageNumber="1">Larvae.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The immature stages are unknown for
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<taxonomicName authorityName="Spangler" authorityYear="1981" class="Insecta" family="Hydrophilidae" genus="Troglochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Troglochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Troglochares</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="taxonomic history">
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<paragraph pageId="0" pageNumber="1">Taxonomic history.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The genus and its only known species were described by
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<bibRefCitation author="Spangler, PJ" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="1" pagination="316 - 323" publicationUrl="https://biodiversitylibrary.org/page/16364657" refId="B279" refString="Spangler, PJ, 1981a. A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae). Proceedings of the Entomological Society of Washington 83 (2): 316 - 323, https://biodiversitylibrary.org/page/16364657" title="A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae)." url="https://biodiversitylibrary.org/page/16364657" volume="83" year="1981 a">Spangler (1981a)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The genus is only known from a single female specimen, which is pin-mounted in pieces (Fig.
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<figureCitation captionStart="Figure 56" captionStartId="F56" captionText="Figure 56. Holotype and labels of Troglochares ashmolei A mount of holotype B head, dorsal view C labels." figureDoi="10.3897/zookeys.1045.63810.figure56" httpUri="https://binary.pensoft.net/fig/557455" pageId="0" pageNumber="1">56</figureCitation>
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). This species was not included in the molecular phylogeny by
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<bibRefCitation DOI="https://doi.org/10.1111/syen.12467" author="Short, AEZ" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="1" pagination="380 - 395" refId="B274" refString="Short, AEZ, Giron, JC, Toussaint, EFA, 2021. Evolution and biogeography of acidocerine water scavenger beetles (Coleoptera: Hydrophilidae) shaped by Gondwanan vicariance and Cenozoic isolation of South America. Systematic Entomology 46 (2): 380 - 395, DOI: https://doi.org/10.1111/syen.12467" title="Evolution and biogeography of acidocerine water scavenger beetles (Coleoptera: Hydrophilidae) shaped by Gondwanan vicariance and Cenozoic isolation of South America." url="https://doi.org/10.1111/syen.12467" volume="46" year="2021">Short et al. (2021)</bibRefCitation>
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. Its assignment to the
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<taxonomicName authorityName="Short & Garcia" authorityYear="2007" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Tobochares</emphasis>
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</taxonomicName>
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group is based primarily on its tiny size (excluding the
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<taxonomicName authorityName="Girón & Short" authorityYear="2021" class="Insecta" family="Hydrophilidae" genus="Helochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Helochares" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Helochares</emphasis>
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</taxonomicName>
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group), presence in the Neotropical region (excluding the
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<taxonomicName authorityName="Regimbart" authorityYear="1903" class="Insecta" family="Hydrophilidae" genus="Agraphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agraphydrus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Agraphydrus</emphasis>
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</taxonomicName>
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group), and lack of a sutural stria (excluding the
|
||
<taxonomicName authorityName="Giron & Short" authorityYear="2019" class="Insecta" family="Hydrophilidae" genus="Primocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Primocerus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Primocerus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Sharp" authorityYear="1882" class="Insecta" family="Hydrophilidae" genus="Chasmogenus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chasmogenus" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Chasmogenus</emphasis>
|
||
</taxonomicName>
|
||
groups).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="species examined">
|
||
<paragraph pageId="0" pageNumber="1">Species examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
The holotype specimen of
|
||
<taxonomicName authorityName="Spangler" authorityYear="1981" class="Insecta" family="Hydrophilidae" genus="Troglochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Troglochares ashmolei" order="Coleoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ashmolei">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Troglochares ashmolei</emphasis>
|
||
</taxonomicName>
|
||
Spangler was examined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="selected references">
|
||
<paragraph pageId="0" pageNumber="1">Selected references.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<bibRefCitation author="Spangler, PJ" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="1" pagination="316 - 323" publicationUrl="https://biodiversitylibrary.org/page/16364657" refId="B279" refString="Spangler, PJ, 1981a. A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae). Proceedings of the Entomological Society of Washington 83 (2): 316 - 323, https://biodiversitylibrary.org/page/16364657" title="A new water beetle, Troglochares ashmolei, n. gen., n. sp., from Ecuador; the first known eyeless cavernicolous hydrophilid beetle (Coleoptera: Hydrophilidae)." url="https://biodiversitylibrary.org/page/16364657" volume="83" year="1981 a">Spangler 1981a</bibRefCitation>
|
||
: original description;
|
||
<bibRefCitation DOI="https://doi.org/10.1111/syen.12467" author="Short, AEZ" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="1" pagination="380 - 395" refId="B274" refString="Short, AEZ, Giron, JC, Toussaint, EFA, 2021. Evolution and biogeography of acidocerine water scavenger beetles (Coleoptera: Hydrophilidae) shaped by Gondwanan vicariance and Cenozoic isolation of South America. Systematic Entomology 46 (2): 380 - 395, DOI: https://doi.org/10.1111/syen.12467" title="Evolution and biogeography of acidocerine water scavenger beetles (Coleoptera: Hydrophilidae) shaped by Gondwanan vicariance and Cenozoic isolation of South America." url="https://doi.org/10.1111/syen.12467" volume="46" year="2021">Short et al. 2021</bibRefCitation>
|
||
: morphological affinities discussed in a phylogenetic context.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |