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<mods:title id="F2764B2C93C6B2E9167234281FF9D384">Review of the genus Engyprosopon Günther, 1862 (Pleuronectiformes: Bothidae) from waters off Taiwan, with descriptions of two new species</mods:title>
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<taxonomicName id="FDF2F6ECFFA7FF8930E1FA55FD70994B" authority="Hensley, 2003" authorityName="Hensley" authorityYear="2003" box="[151,706,1534,1560]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="21" pageNumber="470" phylum="Chordata" rank="species" species="mozambiquense">
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<emphasis id="0886517DFFA7FF8930E1FA55FDA2994B" bold="true" box="[151,528,1534,1560]" italics="true" pageId="21" pageNumber="470">Engyprosopon mozambiquense</emphasis>
<bibRefCitation id="5E63F09EFFA7FF893261FA55FD70994B" author="Hensley, D. A." box="[535,706,1534,1560]" journalOrPublisher="Copeia, 2003" pageId="21" pageNumber="470" pagination="833 - 837" part="4" publicationUrl="https://doi.org/10.1643/IA02-266.1" refId="ref14895" refString="Hensley, D. A. (2003) On the status of Rhombus cocosensis Bleeker, and descrption of a new species of Engyprosopon based on misidentification of this species (Pleuronectiformes: Bothidae). Copeia, 2003 (4), 833 - 837. https: // doi. org / 10.1643 / IA 02 - 266.1" title="On the status of Rhombus cocosensis Bleeker, and descrption of a new species of Engyprosopon based on misidentification of this species (Pleuronectiformes: Bothidae)" type="journal article" year="2003">Hensley, 2003</bibRefCitation>
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<figureCitation id="A2C991EAFFA7FF8930E1F98AFE889969" box="[151,314,1569,1594]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStart-2="FIGURE 15" captionStartId-0="22.[151,250,1841,1863]" captionStartId-1="23.[151,250,1480,1502]" captionStartId-2="24.[151,250,699,721]" captionTargetBox-0="[216,1425,617,1793]" captionTargetBox-1="[151,1436,193,1459]" captionTargetBox-2="[355,1229,200,679]" captionTargetId-0="figure@22.[174,1425,617,1826]" captionTargetId-1="figure@23.[151,1436,193,1459]" captionTargetId-2="figure@24.[346,1246,193,685]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionTargetPageId-2="24" captionText-0="FIGURE 13. Engyprosopon mozambiqense Hensley, 2003. Ocular side, A. NMMB-P22218, male, 58.8 mm SL. B. NMMB- P22218, female, 47.6 mm SL." captionText-1="FIGURE 14. Engyprosopon mozambiquense Hensley, 2003. Blind side, same specimens as those in Figure 12." captionText-2="FIGURE 15. Engyprosopon mozambiquense Hensley, 2003. Gill rakers on first arch from NMMB-P22218." httpUri-0="https://zenodo.org/record/1227817/files/figure.png" httpUri-1="https://zenodo.org/record/1227819/files/figure.png" httpUri-2="https://zenodo.org/record/1227821/files/figure.png" pageId="21" pageNumber="470">Figures 1315</figureCitation>
,
<figureCitation id="A2C991EAFFA7FF893130F989FED79969" box="[326,357,1570,1594]" captionStart="FIGURE 19" captionStartId="29.[151,250,1905,1927]" captionTargetBox="[198,1387,223,1845]" captionTargetId="figure@29.[151,1436,193,1884]" captionTargetPageId="29" captionText="FIGURE 19. Relationship between SL and interorbital width (A), and ocular-side pectoral fin length (B) in Engyprosopon mozambiquense (trangles) and Engyprosopon parvipectorale sp. nov. (circles)." httpUri="https://zenodo.org/record/1227829/files/figure.png" pageId="21" pageNumber="470">19</figureCitation>
;
<tableCitation id="7770B8D4FFA7FF893107F98AFE749969" box="[369,454,1569,1594]" captionStart="TABLE 2" captionStartId="21.[151,239,368,391]" captionText="TABLE 2. Morphometric data (% of SL) of four Engyprosopon species in Taiwan. M = male(s); F = female(s); O = ocular side; B = blind side." pageId="21" pageNumber="470">Table 2</tableCitation>
,
<tableCitation id="7770B8D4FFA7FF8931A7F989FE6D9969" box="[465,479,1570,1594]" captionStart="TABLE 4" captionStartId="30.[151,239,152,175]" captionText="TABLE 4. Comparison of proportional measurements and counts between Engyprosopon parvipectorale sp. nov. and E. mozambiquense. Numbers in bold numbers show differences between Engyprosopon parvipectorale sp. nov. and E. mozambiqensis. M = male(s); F = female(s); O = ocular side; B = blind side." pageId="21" pageNumber="470">4</tableCitation>
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<emphasis id="0886517DFFA7FF8930E1F9C2FECC992C" box="[151,382,1641,1663]" italics="true" pageId="21" pageNumber="470">Arnoglossus annulatus</emphasis>
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(not of
<bibRefCitation id="5E63F09EFFA7FF8931B9F9C2FDE1992C" author="Weber, M." bookContentInfo="710 pp." box="[463,595,1641,1663]" journalOrPublisher="Brill, E. J., Leiden" pageId="21" pageNumber="470" refId="ref16199" refString="Weber, M. (1913) Die Fische der Siboga-Expedition. Brill, E. J., Leiden, 710 pp." title="Die Fische der Siboga-Expedition" type="book" year="1913">Weber, 1913</bibRefCitation>
):
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<bibRefCitation id="5E63F09EFFA7FF893212F9C1FD48992C" author="Norman, J. R." box="[612,762,1641,1663]" journalOrPublisher="Records of the Indian Museum" pageId="21" pageNumber="470" pagination="7 - 48" part="29" refId="ref15589" refString="Norman, J. R. (1927) The flatfishes (Heterosomata) of India, with a list of the specimens in the Indian Museum. Records of the Indian Museum, 29 (1), 7 - 48." title="The flatfishes (Heterosomata) of India, with a list of the specimens in the Indian Museum" type="journal article" year="1927">Norman, 1927</bibRefCitation>
: 19
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<taxonomicName id="FDF2F6ECFFA7FF8930E1F922FE2E99CD" ID-CoL="39TQV" baseAuthorityName="non Bleeker" baseAuthorityYear="1855" box="[151,412,1673,1694]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="21" pageNumber="470" phylum="Chordata" rank="species" species="cocosensis">
<emphasis id="0886517DFFA7FF8930E1F922FE2E99CD" box="[151,412,1673,1694]" italics="true" pageId="21" pageNumber="470">Engyprosopon cocosensis</emphasis>
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(not of
<bibRefCitation id="5E63F09EFFA7FF893199F923FD3399CD" author="Bleeker, P." box="[495,641,1672,1694]" journalOrPublisher="Natuurkundig Tijdschrift Nederlandsch Indie" pageId="21" pageNumber="470" pagination="169 - 180" part="8" refId="ref14465" refString="Bleeker, P. (1855) Derde bijdrage tot de kennis der ichthyologische fauna van de Kokos-eilanden. Natuurkundig Tijdschrift Nederlandsch Indie, 8, 169 - 180." title="Derde bijdrage tot de kennis der ichthyologische fauna van de Kokos-eilanden" type="journal article" year="1855">Bleeker, 1855</bibRefCitation>
):
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<bibRefCitation id="5E63F09EFFA7FF8932E5F922FC9B99CD" author="Norman, J. R." box="[659,809,1672,1694]" journalOrPublisher="Records of the Indian Museum" pageId="21" pageNumber="470" pagination="7 - 48" part="29" refId="ref15589" refString="Norman, J. R. (1927) The flatfishes (Heterosomata) of India, with a list of the specimens in the Indian Museum. Records of the Indian Museum, 29 (1), 7 - 48." title="The flatfishes (Heterosomata) of India, with a list of the specimens in the Indian Museum" type="journal article" year="1927">Norman, 1927</bibRefCitation>
: 24
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;
<treatmentCitation id="BB53AB7EFFA7FF89332CF922FB9199CD" author="" box="[858,1059,1672,1694]" page="204" pageId="21" pageNumber="470">
<bibRefCitation id="5E63F09EFFA7FF89332CF922FC4299CD" author="Norman, J. R." bookContentInfo="459 pp." box="[858,1008,1672,1694]" journalOrPublisher="British Museum, London" pageId="21" pageNumber="470" publicationUrl="https://doi.org/10.5962/bhl.title.8585" refId="ref15632" refString="Norman, J. R. (1934) A systematic monograph of the flatfishes (Heterosomata). 1. Psettodidae, Bothidae, Pleuronectidae. British Museum, London, 459 pp. https: // doi. org / 10.5962 / bhl. title. 8585" title="A systematic monograph of the flatfishes (Heterosomata). 1. Psettodidae, Bothidae, Pleuronectidae" type="book" year="1934">Norman, 1934</bibRefCitation>
: 204
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, fig. 151;
<treatmentCitation id="BB53AB7EFFA7FF8934FDF923FF4F99EF" author="" page="3830" pageId="21" pageNumber="470">
<bibRefCitation id="5E63F09EFFA7FF8934FDF923FA2A99CD" author="Hensley, D. A. &amp; Amaoka, K." box="[1163,1432,1672,1694]" editor="Carpenter, K. E." journalOrPublisher="FAO, Rome" pageId="21" pageNumber="470" pagination="3799 - 3841" refId="ref15048" refString="Hensley, D. A. &amp; Amaoka, K. (2001) Family Bothidae. In: Carpenter, K. E. &amp; Niem, V. E. (Eds.), FAO species identification guide for fishery Purposes. The living marine resources of the western central Pacific. Bony fishes part 4 (Labridae to Latimeriidae), estuarine crocodiles, sea turtles, sea snakes and marine mammals. FAO, Rome, pp. 3799 - 3841." title="Family Bothidae" type="book chapter" volumeTitle="FAO species identification guide for fishery Purposes. The living marine resources of the western central Pacific. Bony fishes part 4 (Labridae to Latimeriidae), estuarine crocodiles, sea turtles, sea snakes and marine mammals" year="2001">Hensley &amp; Amaoka, 2001</bibRefCitation>
: 3830
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<taxonomicName id="FDF2F6ECFFA7FF8930E1F96EFC8D9988" ID-CoL="MP5D" authority=": Weber &amp; de Beaufort, 1929: 428" authorityName=": Weber &amp; de Beaufort" authorityPageNumber="428" authorityYear="1929" box="[151,831,1733,1755]" class="Actinopterygii" family="Bothidae" genus="Bothus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="21" pageNumber="470" phylum="Chordata" rank="species" species="cocosensis" subGenus="Arnoglossus">
<emphasis id="0886517DFFA7FF8930E1F96EFE569988" box="[151,484,1733,1755]" italics="true" pageId="21" pageNumber="470">Bothus (Arnoglossus) cocosensis</emphasis>
: Weber &amp; de
<bibRefCitation id="5E63F09EFFA7FF893207F96EFCBE9988" author="Weber, M. &amp; de Beaufort, L. F." bookContentInfo="458 pp." box="[625,780,1733,1755]" journalOrPublisher="Brill, E. J., Leiden" pageId="21" pageNumber="470" refId="ref16223" refString="Weber, M. &amp; de Beaufort, L. F. (1929) The fishes of the Indo-Australian Archipelago. V. Anacanthini, Allotriognathi, Heterostomata, Berycomorphi, Percomorphi: families Kuhliidae, Apogonidae, Plesiopidae, Pseudoplesiopidae, Priacanthidae, Centropomidae. Brill, E. J., Leiden, 458 pp." title="The fishes of the Indo-Australian Archipelago. V. Anacanthini, Allotriognathi, Heterostomata, Berycomorphi, Percomorphi: families Kuhliidae, Apogonidae, Plesiopidae, Pseudoplesiopidae, Priacanthidae, Centropomidae" type="book" year="1929">Beaufort, 1929</bibRefCitation>
: 428
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.
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<emphasis id="0886517DFFA7FF8930E1F94EFE9899A9" box="[151,298,1765,1786]" italics="true" pageId="21" pageNumber="470">Engyprosopon</emphasis>
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sp.:
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: 858
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<emphasis id="0886517DFFA7FF8930E1F8A8FE6A984B" box="[151,472,1794,1816]" italics="true" pageId="21" pageNumber="470">Engyprosopon mozambiquensis</emphasis>
<treatmentCitation id="BB53AB7EFFA7FF8931ABF8A9FD10984B" author="" box="[477,674,1794,1816]" page="834" pageId="21" pageNumber="470">
<bibRefCitation id="5E63F09EFFA7FF8931ABF8A9FDC3984B" author="Hensley, D. A." box="[477,625,1794,1816]" journalOrPublisher="Copeia, 2003" pageId="21" pageNumber="470" pagination="833 - 837" part="4" publicationUrl="https://doi.org/10.1643/IA02-266.1" refId="ref14895" refString="Hensley, D. A. (2003) On the status of Rhombus cocosensis Bleeker, and descrption of a new species of Engyprosopon based on misidentification of this species (Pleuronectiformes: Bothidae). Copeia, 2003 (4), 833 - 837. https: // doi. org / 10.1643 / IA 02 - 266.1" title="On the status of Rhombus cocosensis Bleeker, and descrption of a new species of Engyprosopon based on misidentification of this species (Pleuronectiformes: Bothidae)" type="journal article" year="2003">Hensley, 2003</bibRefCitation>
: 834
</treatmentCitation>
</taxonomicName>
(
<typeStatus id="E54933CDFFA7FF8932D9F8A9FD6E984B" box="[687,732,1794,1816]" pageId="21" pageNumber="470">type</typeStatus>
locality:
<location id="3F2DDBB4FFA7FF89334DF8A8FC74984B" LSID="urn:lsid:plazi:treatment:B25B3C79FFA7FF8430E1FA55FBBF9855:3F2DDBB4FFA7FF89334DF8A8FC74984B" box="[827,966,1794,1816]" country="Mozambique" latitude="-13.0" longLatPrecision="1291" longitude="40.583332" name="Porto Amelia" pageId="21" pageNumber="470">Porto Amelia</location>
,
<collectingCountry id="42E5CDFFFFA7FF8933A6F8A9FBEA984B" box="[976,1112,1794,1816]" name="Mozambique" pageId="21" pageNumber="470">Mozambique</collectingCountry>
,
<geoCoordinate id="5FC6EBA8FFA7FF893414F8A9FB01984B" box="[1122,1203,1794,1816]" direction="south" orientation="latitude" pageId="21" pageNumber="470" precision="925" value="-13.0">13°00'S</geoCoordinate>
,
<geoCoordinate id="5FC6EBA8FFA7FF8934C8F8A9FAA1984B" box="[1214,1299,1794,1816]" direction="east" orientation="longitude" pageId="21" pageNumber="470" precision="925" value="40.583332">40°35'E</geoCoordinate>
).
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="72E8DEE4FFA7FF8930E1F8EEFD0698D1" pageId="21" pageNumber="470" type="materials_examined">
<paragraph id="3A4D8D6FFFA7FF8930E1F8EEFD0698D1" blockId="21.[151,1436,1861,2030]" pageId="21" pageNumber="470">
<materialsCitation id="8A9A8732FFA7FF8930E1F8EEFD0298D1" ID-GBIF-Occurrence="1838430533" collectingDate="2015-04-10" collectionCode="NMMB-P" country="Taiwan" location="Kaohsiung" pageId="21" pageNumber="470" specimenCode="NMMB-P22218" specimenCount="14" specimenCount-female="3" specimenCount-male="11" stateProvince="Kaohsiung">
<emphasis id="0886517DFFA7FF8930E1F8EEFE31980D" bold="true" box="[151,387,1861,1886]" pageId="21" pageNumber="470">Material examined.</emphasis>
<specimenCode id="6A542514FFA7FF8931F8F8EDFDFA980D" box="[398,584,1862,1886]" collectionCode="NMMB-P" pageId="21" pageNumber="470">NMMB-P22218</specimenCode>
,
<specimenCount id="2CF446E6FFA7FF893221F8EDFD73980D" box="[599,705,1861,1886]" pageId="21" pageNumber="470" type="male">11 males</specimenCount>
, 49.867.0 mm SL,
<specimenCount id="2CF446E6FFA7FF8933C4F8EDFB96980D" box="[946,1060,1861,1886]" pageId="21" pageNumber="470" type="female">3 females</specimenCount>
,
<quantity id="FD0A208AFFA7FF893445F8EDFB6B980E" box="[1075,1241,1862,1886]" metricMagnitude="-2" metricUnit="m" metricValue="4.7700000000000005" metricValueMax="4.8100000000000005" metricValueMin="4.7299999999999995" pageId="21" pageNumber="470" unit="mm" value="47.7" valueMax="48.1" valueMin="47.3">47.348.1 mm</quantity>
SL, Ke-tzu-liao,
<collectingRegion id="F836438DFFA7FF8930E1F8C2FEA198D1" box="[151,275,1897,1922]" country="Taiwan" name="Kaohsiung" pageId="21" pageNumber="470">Kaohsiung</collectingRegion>
, southwestern
<collectingCountry id="42E5CDFFFFA7FF8931CFF8C2FDBC98D1" box="[441,526,1897,1922]" name="Taiwan" pageId="21" pageNumber="470">Taiwan</collectingCountry>
,
<date id="4E4CABAFFFA7FF89326FF8C1FD0298D1" box="[537,688,1897,1922]" pageId="21" pageNumber="470" value="2015-04-10">
<collectingDate id="5E085247FFA7FF89326FF8C1FD0298D1" box="[537,688,1897,1922]" pageId="21" pageNumber="470" value="2015-04-10">10 Apr. 2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="72E8DEE4FFA7FF8A30B1F826FF439F86" lastPageId="22" lastPageNumber="471" pageId="21" pageNumber="470" type="diagnosis">
<paragraph id="3A4D8D6FFFA7FF8A30B1F826FF439F86" blockId="21.[151,1436,1861,2030]" lastBlockId="22.[151,1436,151,573]" lastPageId="22" lastPageNumber="471" pageId="21" pageNumber="470">
<emphasis id="0886517DFFA7FF8930B1F826FEF398F5" bold="true" box="[199,321,1933,1958]" pageId="21" pageNumber="470">Diagnosis.</emphasis>
Gill rakers with spines; large maxilla (
<quantity id="FD0A208AFFA7FF89328AF825FC3998F5" box="[764,907,1933,1958]" metricMagnitude="-2" metricUnit="m" metricValue="5.8928" metricValueMax="6.1722" metricValueMin="5.6134" pageId="21" pageNumber="470" unit="in" value="2.32" valueMax="2.43" valueMin="2.21">2.212.43 in</quantity>
HL) extending to vertical through anterior 1/3 to 1/2 of lower eye; interorbital width narrow in both sexes, that of males wider than width found in females and juveniles; ocular-side pectoral fin filamentous, greatly elongate in males; ocular-side pectoral fin with two dark cross bands; ventral margin of ocular-side lower jaw with 3 distinct black spots, the posteriormost especially distinct.
</paragraph>
</subSubSection>
<subSubSection id="72E8DEE4FFA4FF8430B1FF74FDE39A5D" lastPageId="24" lastPageNumber="473" pageId="22" pageNumber="471" type="description">
<paragraph id="3A4D8D6FFFA4FF8A30B1FF74FC9F9E13" blockId="22.[151,1436,151,573]" pageId="22" pageNumber="471">
<emphasis id="0886517DFFA4FF8A30B1FF74FEEB9FAB" bold="true" box="[199,345,223,248]" pageId="22" pageNumber="471">Description.</emphasis>
Dorsal-fin rays 7984, anal-fin rays 5764, ocular-side pectoral-fin rays 1012, blind-side pectoral-fin rays 89, caudal-fin rays 3 + 11 + 3, ocular-side pelvic-fin rays 6, blind-side pelvic-fin rays 6, scales in lateral line 4652, gill rakers 0 + 67, vertebrae 10+2425.
</paragraph>
<paragraph id="3A4D8D6FFFA4FF8A30B1FEE6FE1D9D6F" blockId="22.[151,1436,151,573]" pageId="22" pageNumber="471">
In SL: HL 3.453.83, body depth 1.922.21. In HL: snout 4.074.68, upper eye diameter 2.963.50, lower eye diameter 2.943.55, interorbital width
<quantity id="FD0A208AFFA4FF8A323CFEDBFD6A9EDB" box="[586,728,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="2.8067" metricValueMax="3.6576" metricValueMin="1.9558" pageId="22" pageNumber="471" unit="in" value="11.05" valueMax="14.4" valueMin="7.7">7.7014.4 in</quantity>
males,
<quantity id="FD0A208AFFA4FF8A3346FEC4FC0F9EDB" box="[816,957,367,392]" metricMagnitude="-1" metricUnit="m" metricValue="6.984999999999999" metricValueMax="7.0104" metricValueMin="6.9596" pageId="22" pageNumber="471" unit="in" value="27.5" valueMax="27.6" valueMin="27.4">27.427.6 in</quantity>
females, ocular-side upper jaw 2.212.43, blind-side upper jaw 2.172.40, ocular-side lower jaw 1.681.90, blind-side lower jaw 1.681.87, caudal peduncle depth 2.152.75, ocular-side pectoral fin
<quantity id="FD0A208AFFA4FF8A3219FE1CFD4C9E83" box="[623,766,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="1.8542" metricValueMax="2.2352" metricValueMin="1.4731999999999998" pageId="22" pageNumber="471" unit="in" value="0.73" valueMax="0.88" valueMin="0.58">0.580.88 in</quantity>
males,
<quantity id="FD0A208AFFA4FF8A332FFE13FC5A9E83" box="[857,1000,439,464]" metricMagnitude="-2" metricUnit="m" metricValue="3.3146999999999998" metricValueMax="3.5052" metricValueMin="3.1242" pageId="22" pageNumber="471" unit="in" value="1.305" valueMax="1.38" valueMin="1.23">1.231.38 in</quantity>
females, blind-side pectoral fin 1.75 2.14, ocular-side pelvic fin 1.902.98, blind-side pelvic fin 1.902.91, base of ocular-side pelvic fin 2.192.87, base of blind-side pelvic fin 5.849.13, longest dorsal fin ray 1.692.29, longest anal fin ray 1.942.23, middle caudal fin ray 1.121.38.
</paragraph>
<caption id="6E8DDDE7FFA4FF8A30E1F89AFE759835" httpUri="https://zenodo.org/record/1227817/files/figure.png" pageId="22" pageNumber="471" startId="22.[151,250,1841,1863]" targetBox="[216,1425,617,1793]" targetPageId="22">
<paragraph id="3A4D8D6FFFA4FF8A30E1F89AFE759835" blockId="22.[151,1436,1841,1894]" pageId="22" pageNumber="471">
<emphasis id="0886517DFFA4FF8A30E1F89AFEA89814" bold="true" box="[151,282,1841,1863]" pageId="22" pageNumber="471">FIGURE 13</emphasis>
.
<taxonomicName id="FDF2F6ECFFA4FF8A315EF899FD469814" authority="Hensley, 2003" authorityName="Hensley" authorityYear="2003" box="[296,756,1841,1863]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="22" pageNumber="471" phylum="Chordata" rank="species" species="mozambiqense">
<emphasis id="0886517DFFA4FF8A315EF899FDE59814" box="[296,599,1841,1863]" italics="true" pageId="22" pageNumber="471">Engyprosopon mozambiqense</emphasis>
Hensley, 2003
</taxonomicName>
. Ocular side, A. NMMB-P22218, male, 58.8 mm SL. B. NMMB- P22218, female, 47.6 mm SL.
</paragraph>
</caption>
<caption id="6E8DDDE7FFA5FF8B30E1FA63FB499A8D" box="[151,1275,1480,1503]" httpUri="https://zenodo.org/record/1227819/files/figure.png" pageId="23" pageNumber="472" startId="23.[151,250,1480,1502]" targetBox="[151,1436,193,1459]" targetPageId="23">
<paragraph id="3A4D8D6FFFA5FF8B30E1FA63FB499A8D" blockId="23.[151,1275,1480,1503]" box="[151,1275,1480,1503]" pageId="23" pageNumber="472">
<emphasis id="0886517DFFA5FF8B30E1FA63FEAB9A8D" bold="true" box="[151,281,1480,1502]" pageId="23" pageNumber="472">FIGURE 14</emphasis>
.
<taxonomicName id="FDF2F6ECFFA5FF8B3153FA61FD489A8D" authority="Hensley, 2003" authorityName="Hensley" authorityYear="2003" box="[293,762,1480,1503]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="23" pageNumber="472" phylum="Chordata" rank="species" species="mozambiquense">
<emphasis id="0886517DFFA5FF8B3153FA61FDD29A8D" box="[293,608,1480,1503]" italics="true" pageId="23" pageNumber="472">Engyprosopon mozambiquense</emphasis>
Hensley, 2003
</taxonomicName>
. Blind side, same specimens as those in Figure 12.
</paragraph>
</caption>
<paragraph id="3A4D8D6FFFA5FF8B30B1F9AEFCBC99A5" blockId="23.[151,1437,1541,1998]" pageId="23" pageNumber="472">
Body ovate, deepest at middle of body, its depth about 1/2 of SL; dorsal and ventral contours gently arching with gradual posterior taper. Caudal peduncle less than 1/4 of body depth. Head large, its length more than 1/4 of SL, upper profile gently arched with shallow notch anterior to interorbital area; upper profile steep in mature males, not so in females and juveniles. Snout round, much shorter than eye diameter. A short, obtuse rostral spine on snout in males; spine absent in females. Eyes small; eye diameter much less than upper jaw length; anterior margins of both eyes on about same vertical line. Interorbital region very narrow, concave, becoming wider with growth, and wider in males than in females and juveniles (
<figureCitation id="A2C991EAFFA5FF8B32EFF976FCB399A5" box="[665,769,1757,1782]" captionStart="FIGURE 19" captionStartId="29.[151,250,1905,1927]" captionTargetBox="[198,1387,223,1845]" captionTargetId="figure@29.[151,1436,193,1884]" captionTargetPageId="29" captionText="FIGURE 19. Relationship between SL and interorbital width (A), and ocular-side pectoral fin length (B) in Engyprosopon mozambiquense (trangles) and Engyprosopon parvipectorale sp. nov. (circles)." httpUri="https://zenodo.org/record/1227829/files/figure.png" pageId="23" pageNumber="472">Fig. 19A</figureCitation>
).
</paragraph>
<paragraph id="3A4D8D6FFFA5FF8B30B1F8AAFE60989D" blockId="23.[151,1437,1541,1998]" pageId="23" pageNumber="472">
Mouth large, oblique, maxilla extending to vertical through anterior 1/3 to 1/2 of lower eye; anterior tips of both jaws almost on same line when mouth closed. Teeth on upper jaw uniserial, some anterior teeth somewhat larger and stronger; lower jaw teeth uniserial, almost all of same size, similar in size to anterior upper teeth. Gill rakers on first arch flattened and slender (
<figureCitation id="A2C991EAFFA5FF8B321AF8C6FD0D98D5" box="[620,703,1901,1926]" captionStart="FIGURE 15" captionStartId="24.[151,250,699,721]" captionTargetBox="[355,1229,200,679]" captionTargetId="figure@24.[346,1246,193,685]" captionTargetPageId="24" captionText="FIGURE 15. Engyprosopon mozambiquense Hensley, 2003. Gill rakers on first arch from NMMB-P22218." httpUri="https://zenodo.org/record/1227821/files/figure.png" pageId="23" pageNumber="472">Fig. 15</figureCitation>
), distal part of inner margin with some small teeth; no gill rakers on upper limb. Ocular-side scales large, ctenoid with short ctenii, highly deciduous; snout and both jaws naked; cycloid scales on blind side.
</paragraph>
<caption id="6E8DDDE7FFAAFF8430E1FD10FB699D82" box="[151,1243,699,721]" httpUri="https://zenodo.org/record/1227821/files/figure.png" pageId="24" pageNumber="473" startId="24.[151,250,699,721]" targetBox="[355,1229,200,679]" targetPageId="24">
<paragraph id="3A4D8D6FFFAAFF8430E1FD10FB699D82" blockId="24.[151,1243,699,721]" box="[151,1243,699,721]" pageId="24" pageNumber="473">
<emphasis id="0886517DFFAAFF8430E1FD10FEAB9D82" bold="true" box="[151,281,699,721]" pageId="24" pageNumber="473">FIGURE 15</emphasis>
.
<taxonomicName id="FDF2F6ECFFAAFF843153FD17FD489D82" authority="Hensley, 2003" authorityName="Hensley" authorityYear="2003" box="[293,762,699,721]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="24" pageNumber="473" phylum="Chordata" rank="species" species="mozambiquense">
<emphasis id="0886517DFFAAFF843153FD17FDD29D82" box="[293,608,699,721]" italics="true" pageId="24" pageNumber="473">Engyprosopon mozambiquense</emphasis>
Hensley, 2003
</taxonomicName>
. Gill rakers on first arch from NMMB-P22218.
</paragraph>
</caption>
<paragraph id="3A4D8D6FFFAAFF8430B1FD56FF7F9C98" blockId="24.[151,1437,765,1798]" pageId="24" pageNumber="473">
Dorsal-fin origin on blind side ventral to concavity on dorsal margin, no elongate rays. Anal-fin origin below pectoral fin base. Ocular-side pectoral fin greatly elongated into filament in males, second ray longest, much longer than head length; ocular-side pectoral fin shorter than head length in females (
<figureCitation id="A2C991EAFFAAFF843455FCEEFB3E9C0D" box="[1059,1164,837,862]" captionStart="FIGURE 19" captionStartId="29.[151,250,1905,1927]" captionTargetBox="[198,1387,223,1845]" captionTargetId="figure@29.[151,1436,193,1884]" captionTargetPageId="29" captionText="FIGURE 19. Relationship between SL and interorbital width (A), and ocular-side pectoral fin length (B) in Engyprosopon mozambiquense (trangles) and Engyprosopon parvipectorale sp. nov. (circles)." httpUri="https://zenodo.org/record/1227829/files/figure.png" pageId="24" pageNumber="473">Fig. 19B</figureCitation>
). Ocular-side pelvic-fin origin at tip of isthmus; ocular-side fourth or fifth ray opposite to first ray of blind-side fin. Tip of isthmus near vertical line through middle of lower eye. Caudal-fin rays branched, expect for three upper- and three lowermost rays.
</paragraph>
<paragraph id="3A4D8D6FFFAAFF8430B1FC7DFD1A9BF1" blockId="24.[151,1437,765,1798]" pageId="24" pageNumber="473">
<emphasis id="0886517DFFAAFF8430B1FC7DFE0D9CBD" box="[199,447,982,1006]" italics="true" pageId="24" pageNumber="473">Coloration in alcohol</emphasis>
. Ground color on ocular side uniformly light brown, scattered with numerous, small, indistinct dark dots on body: two distinct dark spots on lower jaw, anterior one on symphysial portion of lower jaw, posterior one near posterior end of jaw; dark band from middle of lower eye, crossing upper jaw and extending to lower jaw; blind side of body more or less dark in males, uniformly yellowish white in females. Dorsal and anal fins with a series of dark spots; caudal fin with irregularly scattered dark spots; ocular-side pectoral fin with two dark cross bands, lower band wider than upper.
</paragraph>
<paragraph id="3A4D8D6FFFAAFF8430B1FB06FDE39A5D" blockId="24.[151,1437,765,1798]" pageId="24" pageNumber="473">
<emphasis id="0886517DFFAAFF8430B1FB06FE0C9B95" bold="true" box="[199,446,1197,1222]" pageId="24" pageNumber="473">Sexual dimorphism.</emphasis>
This species shows sexual dimorphism in presence or absence of a rostral spine, interorbital width, curve on anterior dorsal profile of the head, the length of the ocular-side pectoral fin, and in coloration of the blind side of the body.
</paragraph>
</subSubSection>
<subSubSection id="72E8DEE4FFAAFF8430B1FAB1FD7B9A29" pageId="24" pageNumber="473" type="distribution">
<paragraph id="3A4D8D6FFFAAFF8430B1FAB1FD7B9A29" blockId="24.[151,1437,765,1798]" pageId="24" pageNumber="473">
<emphasis id="0886517DFFAAFF8430B1FAB1FED29A60" bold="true" box="[199,352,1306,1331]" pageId="24" pageNumber="473">Distribution.</emphasis>
Widespread in the Indo-West Pacific Ocean:
<collectingCountry id="42E5CDFFFFAAFF8433D0FAB1FC499A60" box="[934,1019,1306,1331]" name="Taiwan" pageId="24" pageNumber="473">Taiwan</collectingCountry>
,
<collectingCountry id="42E5CDFFFFAAFF843466FAB1FB549A60" box="[1040,1254,1306,1331]" name="Philippines" pageId="24" pageNumber="473">Philippine Islands</collectingCountry>
, Indian Ocean (
<collectingCountry id="42E5CDFFFFAAFF8430E8FA96FE859A05" box="[158,311,1341,1366]" name="Mozambique" pageId="24" pageNumber="473">Mozambique</collectingCountry>
, northern
<collectingCountry id="42E5CDFFFFAAFF8431DFFA96FD809A05" box="[425,562,1341,1366]" name="Madagascar" pageId="24" pageNumber="473">Madagascar</collectingCountry>
,
<collectingCountry id="42E5CDFFFFAAFF843248FA96FDC89A05" box="[574,634,1341,1366]" name="India" pageId="24" pageNumber="473">India</collectingCountry>
,
<collectingCountry id="42E5CDFFFFAAFF8432F1FA94FD649A05" box="[647,726,1343,1366]" name="Myanmar" pageId="24" pageNumber="473">Burma</collectingCountry>
,
<collectingCountry id="42E5CDFFFFAAFF843297FA96FC209A05" box="[737,914,1341,1366]" name="India" pageId="24" pageNumber="473">Nicobar Islands</collectingCountry>
and Cocos Islands), and Coral Sea (KA, pers. data). Bathymetric range
<quantity id="FD0A208AFFAAFF8431C5FAC9FDBA9A29" box="[435,520,1378,1402]" metricMagnitude="1" metricUnit="m" metricValue="4.9" metricValueMax="9.0" metricValueMin="0.8" pageId="24" pageNumber="473" unit="m" value="49.0" valueMax="90.0" valueMin="8.0">890 m</quantity>
(
<bibRefCitation id="5E63F09EFFAAFF843261FAC9FD0F9A29" author="Hensley, D. A." box="[535,701,1378,1403]" journalOrPublisher="Copeia, 2003" pageId="24" pageNumber="473" pagination="833 - 837" part="4" publicationUrl="https://doi.org/10.1643/IA02-266.1" refId="ref14895" refString="Hensley, D. A. (2003) On the status of Rhombus cocosensis Bleeker, and descrption of a new species of Engyprosopon based on misidentification of this species (Pleuronectiformes: Bothidae). Copeia, 2003 (4), 833 - 837. https: // doi. org / 10.1643 / IA 02 - 266.1" title="On the status of Rhombus cocosensis Bleeker, and descrption of a new species of Engyprosopon based on misidentification of this species (Pleuronectiformes: Bothidae)" type="journal article" year="2003">Hensley, 2003</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="72E8DEE4FFAAFF8430B1FA2EFBBF9855" pageId="24" pageNumber="473" type="discussion">
<paragraph id="3A4D8D6FFFAAFF8430B1FA2EFEF59959" blockId="24.[151,1437,765,1798]" pageId="24" pageNumber="473">
<emphasis id="0886517DFFAAFF8430B1FA2EFE899ACD" bold="true" box="[199,315,1413,1438]" pageId="24" pageNumber="473">Remarks.</emphasis>
The present species, described by
<bibRefCitation id="5E63F09EFFAAFF8432C9FA2EFCDD9ACD" author="Hensley, D. A." box="[703,879,1413,1438]" journalOrPublisher="Copeia, 2003" pageId="24" pageNumber="473" pagination="833 - 837" part="4" publicationUrl="https://doi.org/10.1643/IA02-266.1" refId="ref14895" refString="Hensley, D. A. (2003) On the status of Rhombus cocosensis Bleeker, and descrption of a new species of Engyprosopon based on misidentification of this species (Pleuronectiformes: Bothidae). Copeia, 2003 (4), 833 - 837. https: // doi. org / 10.1643 / IA 02 - 266.1" title="On the status of Rhombus cocosensis Bleeker, and descrption of a new species of Engyprosopon based on misidentification of this species (Pleuronectiformes: Bothidae)" type="journal article" year="2003">Hensley (2003)</bibRefCitation>
, was previously misidentified as
<taxonomicName id="FDF2F6ECFFAAFF843494FA2CFACB9ACD" authorityName=": Weber &amp; de Beaufort" authorityYear="1929" box="[1250,1401,1415,1438]" class="Actinopterygii" family="Bothidae" genus="Bothus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="24" pageNumber="473" phylum="Chordata" rank="species" species="cocosensis">
<emphasis id="0886517DFFAAFF843494FA2CFACB9ACD" box="[1250,1401,1415,1438]" italics="true" pageId="24" pageNumber="473">E. cocosensis</emphasis>
</taxonomicName>
by
<bibRefCitation id="5E63F09EFFAAFF8430E1FA00FE8C9A91" author="Norman, J. R." box="[151,318,1450,1474]" journalOrPublisher="Records of the Indian Museum" pageId="24" pageNumber="473" pagination="7 - 48" part="29" refId="ref15589" refString="Norman, J. R. (1927) The flatfishes (Heterosomata) of India, with a list of the specimens in the Indian Museum. Records of the Indian Museum, 29 (1), 7 - 48." title="The flatfishes (Heterosomata) of India, with a list of the specimens in the Indian Museum" type="journal article" year="1927">Norman (1927</bibRefCitation>
,
<bibRefCitation id="5E63F09EFFAAFF84313EFA01FE339A91" author="Norman, J. R." bookContentInfo="459 pp." box="[328,385,1450,1474]" journalOrPublisher="British Museum, London" pageId="24" pageNumber="473" publicationUrl="https://doi.org/10.5962/bhl.title.8585" refId="ref15632" refString="Norman, J. R. (1934) A systematic monograph of the flatfishes (Heterosomata). 1. Psettodidae, Bothidae, Pleuronectidae. British Museum, London, 459 pp. https: // doi. org / 10.5962 / bhl. title. 8585" title="A systematic monograph of the flatfishes (Heterosomata). 1. Psettodidae, Bothidae, Pleuronectidae" type="book" year="1934">1934</bibRefCitation>
) and
<bibRefCitation id="5E63F09EFFAAFF8431B6FA01FD5D9A91" author="Hensley, D. A. &amp; Amaoka, K." box="[448,751,1450,1475]" editor="Carpenter, K. E." journalOrPublisher="FAO, Rome" pageId="24" pageNumber="473" pagination="3799 - 3841" refId="ref15048" refString="Hensley, D. A. &amp; Amaoka, K. (2001) Family Bothidae. In: Carpenter, K. E. &amp; Niem, V. E. (Eds.), FAO species identification guide for fishery Purposes. The living marine resources of the western central Pacific. Bony fishes part 4 (Labridae to Latimeriidae), estuarine crocodiles, sea turtles, sea snakes and marine mammals. FAO, Rome, pp. 3799 - 3841." title="Family Bothidae" type="book chapter" volumeTitle="FAO species identification guide for fishery Purposes. The living marine resources of the western central Pacific. Bony fishes part 4 (Labridae to Latimeriidae), estuarine crocodiles, sea turtles, sea snakes and marine mammals" year="2001">Hensley &amp; Amaoka (2001)</bibRefCitation>
.
<bibRefCitation id="5E63F09EFFAAFF84328AFA01FC189A91" author="Hensley, D. A." box="[764,938,1450,1475]" journalOrPublisher="Copeia, 2003" pageId="24" pageNumber="473" pagination="833 - 837" part="4" publicationUrl="https://doi.org/10.1643/IA02-266.1" refId="ref14895" refString="Hensley, D. A. (2003) On the status of Rhombus cocosensis Bleeker, and descrption of a new species of Engyprosopon based on misidentification of this species (Pleuronectiformes: Bothidae). Copeia, 2003 (4), 833 - 837. https: // doi. org / 10.1643 / IA 02 - 266.1" title="On the status of Rhombus cocosensis Bleeker, and descrption of a new species of Engyprosopon based on misidentification of this species (Pleuronectiformes: Bothidae)" type="journal article" year="2003">Hensley (2003)</bibRefCitation>
found that the
<typeStatus id="E54933CDFFAAFF843424FA01FB069A90" box="[1106,1204,1450,1475]" pageId="24" pageNumber="473" type="holotype">holotype</typeStatus>
of
<taxonomicName id="FDF2F6ECFFAAFF8434AFFA00FAC29A91" authorityName=": Weber &amp; de Beaufort" authorityYear="1929" box="[1241,1392,1451,1474]" class="Actinopterygii" family="Bothidae" genus="Bothus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="24" pageNumber="473" phylum="Chordata" rank="species" species="cocosensis">
<emphasis id="0886517DFFAAFF8434AFFA00FAC29A91" box="[1241,1392,1451,1474]" italics="true" pageId="24" pageNumber="473">E. cocosensis</emphasis>
</taxonomicName>
has palmate gill rakers and a lure on the first dorsal-fin ray, these are unique characters to members of the genus
<taxonomicName id="FDF2F6ECFFAAFF8430E1FA59FEF29959" baseAuthorityName="Hensley" baseAuthorityYear="2005" box="[151,320,1522,1546]" class="Actinopterygii" family="Bothidae" genus="Asterorhombus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="24" pageNumber="473" phylum="Chordata" rank="genus">
<emphasis id="0886517DFFAAFF8430E1FA59FEF29959" box="[151,320,1522,1546]" italics="true" pageId="24" pageNumber="473">Asterorhombus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="3A4D8D6FFFAAFF8430B1F9BCFBBF9855" blockId="24.[151,1437,765,1798]" pageId="24" pageNumber="473">
<taxonomicName id="FDF2F6ECFFAAFF8430B1F9BCFD91997D" authorityName="Hensley" authorityYear="2003" box="[199,547,1558,1582]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="24" pageNumber="473" phylum="Chordata" rank="species" species="mozambiquense">
<emphasis id="0886517DFFAAFF8430B1F9BCFD91997D" box="[199,547,1558,1582]" italics="true" pageId="24" pageNumber="473">Engyprosopon mozambiquense</emphasis>
</taxonomicName>
belongs to a species group in
<taxonomicName id="FDF2F6ECFFAAFF843300F9BCFBAA997D" authorityName="Gunther" authorityYear="1862" box="[886,1048,1559,1582]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="24" pageNumber="473" phylum="Chordata" rank="genus">
<emphasis id="0886517DFFAAFF843300F9BCFBAA997D" box="[886,1048,1559,1582]" italics="true" pageId="24" pageNumber="473">Engyprosopon</emphasis>
</taxonomicName>
with serrate gill rakers. It is easily discriminated from congeners of the group in having the combination of some dark spots on the caudal fin, 3 black spots on the ventral margin of the ocular-side lower jaw, two dark cross bands on the ocular-side pectoral fin, a large mouth (length of ocular-side maxilla about
<quantity id="FD0A208AFFAAFF8432BCF929FC8F99C8" box="[714,829,1666,1691]" metricMagnitude="-2" metricUnit="m" metricValue="5.842" metricValueMax="6.096" metricValueMin="5.588" pageId="24" pageNumber="473" unit="in" value="2.3" valueMax="2.4" valueMin="2.2">2.22.4 in</quantity>
HL), and narrower interorbital width compared with respective features of other species (when compared between same sexes and same sizes). This species is widely distributed in the Indian Ocean, Coral Sea, and the
<collectingCountry id="42E5CDFFFFAAFF8432A3F961FC1099B0" box="[725,930,1738,1763]" name="Philippines" pageId="24" pageNumber="473">Philippine Islands</collectingCountry>
. This is the first record of
<taxonomicName id="FDF2F6ECFFAAFF8434BCF960FA2E99B1" authorityName="Hensley" authorityYear="2003" box="[1226,1436,1738,1762]" class="Actinopterygii" family="Bothidae" genus="Engyprosopon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pleuronectiformes" pageId="24" pageNumber="473" phylum="Chordata" rank="species" species="mozambiquense">
<emphasis id="0886517DFFAAFF8434BCF960FA2E99B1" box="[1226,1436,1738,1762]" italics="true" pageId="24" pageNumber="473">E. mozambiquense</emphasis>
</taxonomicName>
from the waters of
<collectingCountry id="42E5CDFFFFAAFF84311AF946FE739855" box="[364,449,1773,1798]" name="Taiwan" pageId="24" pageNumber="473">Taiwan</collectingCountry>
, and is also the northernmost record for this species.
</paragraph>
</subSubSection>
</treatment>
</document>