182 lines
21 KiB
XML
182 lines
21 KiB
XML
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<mods:title id="137409E9E300FB931F1F14CBAD53C1BE">Redescription of the ‘ older Adelopsis’ species (Coleoptera: Leiodidae: Cholevinae: Ptomaphagini) based on the analysis of type specimens</mods:title>
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<mods:namePart id="06A1E6FCC6D1257DB0960DAD5DAE43BA">Peck, Stewart B.</mods:namePart>
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<mods:title id="FF25E01820FDABF88DB06584CAD468A4">Zootaxa</mods:title>
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<treatment id="B20E4654FFA6FF98BAF4294BFDF8C3FD" ID-DOI="http://doi.org/10.5281/zenodo.5923424" ID-GBIF-Taxon="160406299" ID-Zenodo-Dep="5923424" LSID="urn:lsid:plazi:treatment:B20E4654FFA6FF98BAF4294BFDF8C3FD" httpUri="http://treatment.plazi.org/id/B20E4654FFA6FF98BAF4294BFDF8C3FD" lastPageNumber="14" pageId="13" pageNumber="14">
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<subSubSection id="72BDA4C9FFA6FF98BAF4294BFDA5C410" box="[151,554,549,576]" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="3A18F742FFA6FF98BAF4294BFDA5C410" blockId="13.[151,554,549,576]" box="[151,554,549,576]" pageId="13" pageNumber="14">
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<heading id="6150402EFFA6FF98BAF4294BFDA5C410" bold="true" box="[151,554,549,576]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
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<taxonomicName id="FDA78CC1FFA6FF98BAF4294BFDA5C410" ID-CoL="9Y5S" authority="Jeannel, 1936" authorityName="Jeannel" authorityYear="1936" box="[151,554,549,576]" class="Insecta" family="Leiodidae" genus="Adelopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brunnea">
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<emphasis id="08D32B50FFA6FF98BAF4294BFDA5C410" bold="true" box="[151,554,549,576]" pageId="13" pageNumber="14">
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<emphasis id="08D32B50FFA6FF98BAF4294BFEF6C46F" bold="true" box="[151,377,549,576]" italics="true" pageId="13" pageNumber="14">Adelopsis brunnea</emphasis>
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<bibRefCitation id="5E368AB3FFA6FF98BBE3294BFDA5C410" author="Jeannel, R." box="[384,554,549,575]" pageId="13" pageNumber="14" pagination="1 - 433" refId="ref43714" refString="Jeannel, R. (1936) Monographie des Catopidae. Memoires du Museum National d'Histoire Naturelle, New Series, 1, 1 - 433." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="72BDA4C9FFA6FF98BAF42907FE4DC4F2" pageId="13" pageNumber="14" type="reference_group">
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<paragraph id="3A18F742FFA6FF98BAF42907FD58C4AD" blockId="13.[151,1437,617,734]" box="[151,727,617,642]" pageId="13" pageNumber="14">
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<treatmentCitationGroup id="1AB7D06CFFA6FF98BAF42907FD58C4AD" box="[151,727,617,642]" pageId="13" pageNumber="14">
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<taxonomicName id="FDA78CC1FFA6FF98BAF42907FD99C4AD" ID-CoL="9Y5T" authority="Jeannel, 1936: 66" authorityName="Jeannel" authorityPageNumber="66" authorityYear="1936" box="[151,534,617,642]" class="Insecta" family="Leiodidae" genus="Adelopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brunneus">
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<emphasis id="08D32B50FFA6FF98BAF42907FED1C4AE" box="[151,350,617,641]" italics="true" pageId="13" pageNumber="14">Adelopsis brunneus</emphasis>
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<treatmentCitation id="BB06D153FFA6FF98BB062907FD99C4AD" author="Jeannel, R." box="[357,534,617,642]" page="66" pageId="13" pageNumber="14" year="1936">
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<bibRefCitation id="5E368AB3FFA6FF98BB062907FD99C4AD" author="Jeannel, R." box="[357,534,617,642]" pageId="13" pageNumber="14" pagination="1 - 433" refId="ref43714" refString="Jeannel, R. (1936) Monographie des Catopidae. Memoires du Museum National d'Histoire Naturelle, New Series, 1, 1 - 433." type="journal article" year="1936">Jeannel, 1936: 66</bibRefCitation>
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||
</treatmentCitation>
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</taxonomicName>
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[and
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<figureCitation id="A29CEBC7FFA6FF98B8332907FD41C4AD" box="[592,718,617,642]" captionStart="FIGURES 76–88" captionStartId="21.[151,264,1545,1570]" captionTargetBox="[161,1400,176,1516]" captionTargetId="figure@21.[151,1407,166,1525]" captionTargetPageId="21" captionText="FIGURES 76–88. Adelopsis pteromoria Szymczakowski, 1975 new status, (probable) holotype male. 76–80, aedeagus, left lateral, dorsal, right lateral, and ventral views, and frontal view of aedeagus apex; 81, last ventrite; 82, genital segment; 83, head, lateral view; 84, antenna, dorsal view; 85, meso and metathorax, ventro-lateral view (of left side of specimen); 86, protarsus and protibia; 87, mesotarsus and mesotibia; 88, metatarsus and metatibia. Scale lines = 0.5 mm; a for figs. 76–82; b for figs. 83–88." figureDoi="http://doi.org/10.5281/zenodo.3542783" httpUri="https://zenodo.org/record/3542783/files/figure.png" pageId="13" pageNumber="14">Figs. 86–87</figureCitation>
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].
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</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="3A18F742FFA6FF98BAF429E6FC59C48F" blockId="13.[151,1437,617,734]" box="[151,982,647,672]" pageId="13" pageNumber="14">
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<treatmentCitationGroup id="1AB7D06CFFA6FF98BAF429E6FC59C48F" box="[151,982,647,672]" pageId="13" pageNumber="14">
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||
<taxonomicName id="FDA78CC1FFA6FF98BAF429E6FE4CC48F" ID-CoL="9Y5T" authority="brunneus" box="[151,451,648,672]" class="Insecta" family="Leiodidae" genus="Adelopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="brunneus" subSpecies="brunneus">
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<emphasis id="08D32B50FFA6FF98BAF429E6FE4CC48F" box="[151,451,648,672]" italics="true" pageId="13" pageNumber="14">Adelopsis brunneus brunneus</emphasis>
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</taxonomicName>
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;
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<treatmentCitation id="BB06D153FFA6FF98BBB329E9FD54C48F" author="Szymczakowski, W." box="[464,731,647,672]" page="15" pageId="13" pageNumber="14" year="1975">
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<bibRefCitation id="5E368AB3FFA6FF98BBB329E9FD54C48F" author="Szymczakowski, W." box="[464,731,647,672]" pageId="13" pageNumber="14" pagination="13 - 24" refId="ref45031" refString="Szymczakowski, W. (1975) Formes cavernicoles d' Adelopsis brunneus Jeann. du Venezuela et de l'Ile de Trinidad (Coleoptera: Catopidae). Boletin de la Sociedad Venezolana de Espeleologia, 6, 13 - 24." type="journal article" year="1975">Szymczakowski, 1975: 15</bibRefCitation>
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||
</treatmentCitation>
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||
[and
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||
<figureCitation id="A29CEBC7FFA6FF98B97629E9FCDEC48F" box="[789,849,647,672]" captionStart="FIGURES 1–3" captionStartId="6.[151,264,1683,1708]" captionTargetBox="[344,1242,1091,1660]" captionTargetId="figure@6.[340,1247,1086,1664]" captionTargetPageId="6" captionText="FIGURES 1–3. General view of aedeagus and genital segment of Adelopsis sp. 1, aedeagus, left view; 2, aedeagus, dorsal view; 3, genital segment, ventral view; a, apex; b, base." figureDoi="http://doi.org/10.5281/zenodo.3542767" httpUri="https://zenodo.org/record/3542767/files/figure.png" pageId="13" pageNumber="14">Fig. 2</figureCitation>
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] (
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<typeStatus id="E51C49E0FFA6FF98B90529E6FC1CC48F" box="[870,915,648,672]" pageId="13" pageNumber="14">type</typeStatus>
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seen).
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||
</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="3A18F742FFA6FF98BAF429C9FE4DC4F2" blockId="13.[151,1437,617,734]" pageId="13" pageNumber="14">
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<treatmentCitationGroup id="1AB7D06CFFA6FF98BAF429C9FE4DC4F2" pageId="13" pageNumber="14">
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<taxonomicName id="FDA78CC1FFA6FF98BAF429C9FE3EC490" ID-CoL="9Y5S" authority="brunnea" box="[151,433,679,703]" class="Insecta" family="Leiodidae" genus="Adelopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subSpecies" species="brunnea" subSpecies="brunnea">
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<emphasis id="08D32B50FFA6FF98BAF429C9FE3EC490" box="[151,433,679,703]" italics="true" pageId="13" pageNumber="14">Adelopsis brunnea brunnea</emphasis>
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</taxonomicName>
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; Gnaspini, 1996: 539 (spelling corrected to feminine gender);
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<treatmentCitation id="BB06D153FFA6FF98BE5A29C8FAE0C490" author="Gnaspini, P. & Peck, S. B." box="[1081,1391,678,703]" page="429" pageId="13" pageNumber="14" year="2001">
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<bibRefCitation id="5E368AB3FFA6FF98BE5A29C8FAE0C490" author="Gnaspini, P. & Peck, S. B." box="[1081,1391,678,703]" pageId="13" pageNumber="14" pagination="427 - 463" refId="ref43422" refString="Gnaspini, P. & Peck, S. B. (2001) The Adelopsis of Colombia (Coleoptera, Leiodidae, Cholevinae, Ptomaphagini). Papeis Avulsos de Zoologia, 41, 427 - 463." type="journal article" year="2001">Gnaspini and Peck, 2001: 429</bibRefCitation>
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||
</treatmentCitation>
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||
(assignment to group), 430.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="72BDA4C9FFA6FF98BAF42862FDF8C3FD" pageId="13" pageNumber="14" type="materials_examined">
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||
<paragraph id="3A18F742FFA6FF98BAF42862FD59C565" blockId="13.[151,1436,779,950]" pageId="13" pageNumber="14">
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||
<materialsCitation id="8ACFFD1FFFA6FF98BAF42862FD59C565" ID-GBIF-Occurrence="2451575244" collectionCode="BMNH" pageId="13" pageNumber="14" specimenCount="2" typeStatus="holotype">
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<typeStatus id="E51C49E0FFA6FF98BAF42862FF70C509" box="[151,255,780,806]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
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male [a single specimen in original description, assumed as
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<typeStatus id="E51C49E0FFA6FF98B9C12865FB88C509" box="[930,1031,779,806]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
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] in
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<collectionCode id="5CB66F87FFA6FF98BE532862FB0BC509" box="[1072,1156,780,806]" pageId="13" pageNumber="14">BMNH</collectionCode>
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[not presently in
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<collectionCode id="5CB66F87FFA6FF98BF2B2862FA13C509" box="[1352,1436,780,806]" pageId="13" pageNumber="14">BMNH</collectionCode>
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(when we requested it for analysis), not examined].
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</materialsCitation>
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||
</paragraph>
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<paragraph id="3A18F742FFA6FF98BAA4283AFC26C541" blockId="13.[151,1436,779,950]" box="[199,937,851,878]" pageId="13" pageNumber="14">
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Length:
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<quantity id="FD5F5AA7FFA6FF98BB46283AFEF4C541" box="[293,379,852,878]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="14" unit="mm" value="2.2">2.2 mm</quantity>
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(original description and
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<bibRefCitation id="5E368AB3FFA6FF98B8FF283AFC13C541" author="Szymczakowski, W." box="[668,924,851,878]" pageId="13" pageNumber="14" pagination="13 - 24" refId="ref45031" refString="Szymczakowski, W. (1975) Formes cavernicoles d' Adelopsis brunneus Jeann. du Venezuela et de l'Ile de Trinidad (Coleoptera: Catopidae). Boletin de la Sociedad Venezolana de Espeleologia, 6, 13 - 24." type="journal article" year="1975">Szymczakowski, 1975</bibRefCitation>
|
||
).
|
||
</paragraph>
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||
<paragraph id="3A18F742FFA6FF98BAA42816FE78C5BD" blockId="13.[151,1436,779,950]" box="[199,503,888,914]" pageId="13" pageNumber="14">
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||
Type locality: “
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<collectingCountry id="42B0B7D2FFA6FF98BB112816FE66C5BD" box="[370,489,888,914]" name="Colombia" pageId="13" pageNumber="14">Colombie</collectingCountry>
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||
”.
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||
</paragraph>
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||
<paragraph id="3A18F742FFA6FF98BAA428F2FAC0C599" blockId="13.[151,1436,779,950]" box="[199,1359,923,950]" pageId="13" pageNumber="14">
|
||
<emphasis id="08D32B50FFA6FF98BAA428F2FED7C599" box="[199,344,924,950]" italics="true" pageId="13" pageNumber="14">Distribution.</emphasis>
|
||
<collectingCountry id="42B0B7D2FFA6FF98BB3D28F2FE5EC599" box="[350,465,924,950]" name="Colombia" pageId="13" pageNumber="14">Colombia</collectingCountry>
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||
: known only from “
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||
<typeStatus id="E51C49E0FFA6FF98B8CC28F2FD6DC599" box="[687,738,924,950]" pageId="13" pageNumber="14">type</typeStatus>
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||
locality” (original description;
|
||
<bibRefCitation id="5E368AB3FFA6FF98BE2128F2FACDC599" author="Szymczakowski, W." box="[1090,1346,923,950]" pageId="13" pageNumber="14" pagination="13 - 24" refId="ref45031" refString="Szymczakowski, W. (1975) Formes cavernicoles d' Adelopsis brunneus Jeann. du Venezuela et de l'Ile de Trinidad (Coleoptera: Catopidae). Boletin de la Sociedad Venezolana de Espeleologia, 6, 13 - 24." type="journal article" year="1975">Szymczakowski, 1975</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="3A18F742FFA6FF98BAF4288AFE0DC5D1" blockId="13.[151,1437,996,1491]" box="[151,386,996,1022]" pageId="13" pageNumber="14">
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||
<emphasis id="08D32B50FFA6FF98BAF4288AFE0DC5D1" box="[151,386,996,1022]" italics="true" pageId="13" pageNumber="14">Taxonomic Remarks.</emphasis>
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||
</paragraph>
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||
<paragraph id="3A18F742FFA6FF98BAA42F66FC16C20D" blockId="13.[151,1437,996,1491]" box="[199,921,1032,1058]" pageId="13" pageNumber="14">
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||
1) See Taxonomic Remarks under
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<taxonomicName id="FDA78CC1FFA6FF98B8292F66FCCEC20D" baseAuthorityName="Murray" baseAuthorityYear="1856" box="[586,833,1032,1058]" class="Insecta" family="Leiodidae" genus="Adelopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ascutellaris">
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<emphasis id="08D32B50FFA6FF98B8292F66FCCEC20D" box="[586,833,1032,1058]" italics="true" pageId="13" pageNumber="14">Adelopsis ascutellaris</emphasis>
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</taxonomicName>
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, above.
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</paragraph>
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<paragraph id="3A18F742FFA6FF98BAA42F42FE35C2F9" blockId="13.[151,1437,996,1491]" pageId="13" pageNumber="14">
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||
2)
|
||
<bibRefCitation id="5E368AB3FFA6FF98BA852F42FD95C269" author="Szymczakowski, W." box="[230,538,1067,1094]" pageId="13" pageNumber="14" pagination="13 - 24" refId="ref45031" refString="Szymczakowski, W. (1975) Formes cavernicoles d' Adelopsis brunneus Jeann. du Venezuela et de l'Ile de Trinidad (Coleoptera: Catopidae). Boletin de la Sociedad Venezolana de Espeleologia, 6, 13 - 24." type="journal article" year="1975">Szymczakowski (1975: 14)</bibRefCitation>
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||
stated that it is very probable that
|
||
<taxonomicName id="FDA78CC1FFA6FF98B9C22F42FB8CC269" authorityName="Jeannel" authorityYear="1936" box="[929,1027,1068,1094]" class="Insecta" family="Leiodidae" genus="Adelopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ovalis">
|
||
<emphasis id="08D32B50FFA6FF98B9C22F42FB8CC269" box="[929,1027,1068,1094]" italics="true" pageId="13" pageNumber="14">A. ovalis</emphasis>
|
||
</taxonomicName>
|
||
is a synonym of
|
||
<taxonomicName id="FDA78CC1FFA6FF98BEA52F42FACDC269" authorityName="Jeannel" authorityYear="1936" box="[1222,1346,1068,1094]" class="Insecta" family="Leiodidae" genus="Adelopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brunnea">
|
||
<emphasis id="08D32B50FFA6FF98BEA52F42FACDC269" box="[1222,1346,1068,1094]" italics="true" pageId="13" pageNumber="14">A. brunnea</emphasis>
|
||
</taxonomicName>
|
||
; and he proposed to use the latter as the valid name [but the former has page precedence] if this synonymy is proved right. However, this ‘potential synonymy’ has never been used in the literature afterwards. We here presently consider both as valid species, and do not propose a revalidation because we understand that Szymczakowski´s statement is not a valid taxonomic act.
|
||
</paragraph>
|
||
<paragraph id="3A18F742FFA6FF98BAA42F8EFF73C3A5" blockId="13.[151,1437,996,1491]" pageId="13" pageNumber="14">
|
||
3) As proposed in Gnaspini (1996: 539) and
|
||
<bibRefCitation id="5E368AB3FFA6FF98B8DC2F8EFC3AC2D5" author="Peck, S. B. & P. Gnaspini & A. F. Newton" box="[703,949,1247,1274]" pageId="13" pageNumber="14" pagination="37 - 72" refId="ref44075" refString="Peck, S. B., P. Gnaspini, and A. F. Newton. (1998) Catalogue and generic keys for the Leiodidae of Mexico State, West Indies, and Central and South America (Insecta: Coleoptera). Giornale Italiano de Entomologia, 9, 37 - 72." type="journal article" year="1998">
|
||
Peck
|
||
<emphasis id="08D32B50FFA6FF98B89E2F8EFCA0C2D5" box="[765,815,1248,1274]" italics="true" pageId="13" pageNumber="14">et al</emphasis>
|
||
. (1998: 62)
|
||
</bibRefCitation>
|
||
, all subspecies of
|
||
<taxonomicName id="FDA78CC1FFA6FF98BEE72F8EFA13C2D6" authority="Jeannel 1936" authorityName="Jeannel" authorityYear="1936" box="[1156,1436,1247,1274]" class="Insecta" family="Leiodidae" genus="Adelopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brunnea">
|
||
<emphasis id="08D32B50FFA6FF98BEE72F8EFA8EC2D5" box="[1156,1281,1248,1274]" italics="true" pageId="13" pageNumber="14">A. brunnea</emphasis>
|
||
<bibRefCitation id="5E368AB3FFA6FF98BF6A2F8EFA13C2D6" author="Jeannel, R." box="[1289,1436,1247,1274]" pageId="13" pageNumber="14" pagination="1 - 433" refId="ref43714" refString="Jeannel, R. (1936) Monographie des Catopidae. Memoires du Museum National d'Histoire Naturelle, New Series, 1, 1 - 433." type="journal article" year="1936">Jeannel 1936</bibRefCitation>
|
||
</taxonomicName>
|
||
are here raised to species status. The combination of the variation observed on aedeagus, genital segment, eyes, antenna, male mesotibia, and spermatheca allow taxon recognition (see discussion under each of these taxa). As a result, this would be the only subspecies left in the species. Therefore, we here abandon the use of subspecies in
|
||
<taxonomicName id="FDA78CC1FFA6FF98BFE72E22FF7BC3A5" authorityName="Jeannel" authorityYear="1936" class="Insecta" family="Leiodidae" genus="Adelopsis" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="brunnea">
|
||
<emphasis id="08D32B50FFA6FF98BFE72E22FF7BC3A5" italics="true" pageId="13" pageNumber="14">A. brunnea</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="3A18F742FFA6FF98BAA42EFAFDF8C3FD" blockId="13.[151,1437,996,1491]" pageId="13" pageNumber="14">4) This species is tentatively assigned to subgroup, since the genital segment (which is here considered important for subgroup assignment) is unknown.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |