276 lines
36 KiB
XML
276 lines
36 KiB
XML
<document id="13B449031DDF511DA3B712BB7E80C73B" ID-DOI="10.5281/zenodo.181670" ID-GBIF-Dataset="d8f568a5-6853-437d-8dd5-165215bb2cab" ID-ISSN="1175-5326" ID-Zenodo-Dep="181670" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459837607491" checkinUser="plazi" docAuthor="Dankittipakul, Pakawin & Singtripop, Tippawan" docDate="2008" docId="692187BDFFCAFFDBFF0DFD634059768F" docLanguage="en" docName="zt01747p060.pdf" docOrigin="Zootaxa 1747" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Clubiona abnormis Dankittipakul, sp. n." docType="treatment" docVersion="10" lastPageNumber="45" masterDocId="9518FFC5FFC0FFD0FF9AFFDE431A7045" masterDocTitle="Five new species of the spider genus Clubiona Latreille (Araneae: Clubionidae) from Thailand" masterLastPageNumber="60" masterPageNumber="34" pageNumber="44" updateTime="1698223866526" updateUser="plazi">
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<mods:mods id="013CA8C4C1873C4A3831D7D75629D479" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CC21CF4121F1ACD1DBC2FBE7A03FF848">
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<mods:title id="08652CEA65134FFCA899C7D3F16FA66C">Five new species of the spider genus Clubiona Latreille (Araneae: Clubionidae) from Thailand</mods:title>
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</mods:titleInfo>
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<mods:name id="BF6A62201CA86C86D0ACEA870AB472AA" type="personal">
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<mods:role id="F8BDADE5BEC2C9AB7FB83EB0CACB26B7">
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<mods:roleTerm id="5EDC6A1C9326FB0617DD8849B831DB90">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A29D2CF9CABBF00ABB1B68B9C16FC148">Dankittipakul, Pakawin</mods:namePart>
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</mods:name>
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<mods:name id="3904C75F3156E04A612D2C6100BADE7B" type="personal">
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<mods:role id="F0A8C056CDA8EFE76E6FC0E027403702">
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<mods:roleTerm id="A4FD1DCDCE1E31CCDDD1176AB396778E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0721436054FBFF92951572FC3E247A42">Singtripop, Tippawan</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="0D066FCB10F8F93C67CAA648F592B684">text</mods:typeOfResource>
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<mods:relatedItem id="190EEA3E132A611EA2C21195DC7BF485" type="host">
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<mods:titleInfo id="557740D0FE0AF29AB6F9451078F96B4C">
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<mods:title id="103211A97CD7C490C03288266DD2C637">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1CFBB8B6EDAA42AF30B72AF5E4446D46">
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<mods:date id="64D5E3FD4BF7CAEEE0F772F0C4CFD1A0">2008</mods:date>
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<mods:detail id="D33080D2B45B06F31F0E28367C756E24" type="volume">
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<mods:number id="C26DAC994DE4AB2E1F5145487344A1F9">1747</mods:number>
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</mods:detail>
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<mods:extent id="E28953CD003CDFD25C7E0F11682F8970" unit="page">
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<mods:start id="560BF27421663A140B25E7719D827FA6">34</mods:start>
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<mods:end id="6A95794330EBD64EC15E80BD207C01BA">60</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="E3A9B305C9B2D5E5D8FEB4F07DD9428F">journal article</mods:classification>
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<mods:identifier id="478CA6262EDD3E16DFC22764E44DDC60" type="DOI">10.5281/zenodo.181670</mods:identifier>
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<mods:identifier id="FF4556603B5DF70E9E2B046E0D50B3A4" type="GBIF-Dataset">d8f568a5-6853-437d-8dd5-165215bb2cab</mods:identifier>
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<mods:identifier id="6438FAE6072974F0DB09A62243A74D06" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="06D135A4483C66A206ECA218868F8E7B" type="Zenodo-Dep">181670</mods:identifier>
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</mods:mods>
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<treatment id="692187BDFFCAFFDBFF0DFD634059768F" ID-DOI="http://doi.org/10.5281/zenodo.6228356" ID-GBIF-Taxon="119359362" ID-Zenodo-Dep="6228356" LSID="urn:lsid:plazi:treatment:692187BDFFCAFFDBFF0DFD634059768F" httpUri="http://treatment.plazi.org/id/692187BDFFCAFFDBFF0DFD634059768F" lastPageId="11" lastPageNumber="45" pageId="10" pageNumber="44">
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<subSubSection id="A9926520FFCAFFDAFF0DFD63429F72BF" pageId="10" pageNumber="44" type="nomenclature">
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<paragraph id="E13736ABFFCAFFDAFF0DFD63419B7292" blockId="10.[151,641,701,762]" box="[151,641,701,727]" pageId="10" pageNumber="44">
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<heading id="BA7F81C7FFCAFFDAFF0DFD63419B7292" bold="true" box="[151,641,701,727]" fontSize="11" level="1" pageId="10" pageNumber="44" reason="1">
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<emphasis id="D3FCEAB9FFCAFFDAFF0DFD63419B7292" bold="true" box="[151,641,701,727]" pageId="10" pageNumber="44">
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<taxonomicName id="26884D28FFCAFFDAFF0DFD6341227292" ID-CoL="WC3G" authority="Dankittipakul" authorityName="Dankittipakul" box="[151,568,701,727]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="44" phylum="Arthropoda" rank="species" species="abnormis" status="sp. nov.">
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<emphasis id="D3FCEAB9FFCAFFDAFF0DFD63429A7292" bold="true" box="[151,384,701,727]" italics="true" pageId="10" pageNumber="44">Clubiona abnormis</emphasis>
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Dankittipakul
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</taxonomicName>
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<taxonomicNameLabel id="C8CF57C2FFCAFFDAFDA5FD63419B7292" box="[575,641,701,727]" pageId="10" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="E13736ABFFCAFFDAFF0DFD3E429F72BF" blockId="10.[151,641,701,762]" box="[151,389,736,762]" pageId="10" pageNumber="44">
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||
(
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<figureCitation id="79B32A2EFFCAFFDAFF05FD3E423C72BF" box="[159,294,736,762]" captionStart="FIGURES 24 – 29" captionStartId="18.[151,269,1945,1969]" captionTargetBox="[346,1241,167,1937]" captionTargetId="figure@18.[346,1242,167,1940]" captionTargetPageId="18" captionText="FIGURES 24 – 29. Clubiona suthepica sp. n., female paratype (24 – 27). 24 epigyne, ventral; 25 vulva, dorsal; 26 spermathecal base; 27 dilated terminal apparatus of spermathecal head. Clubiona abnormis sp. n., male holotype (28, 29). 28 male palp, dorsal; 29 ditto, retrolateral. Scale bars = 0.1 mm (24 – 25, 28 – 29)." httpUri="https://zenodo.org/record/181676/files/figure.png" pageId="10" pageNumber="44">Figs 28–29</figureCitation>
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,
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<figureCitation id="79B32A2EFFCAFFDAFEA8FD3E426772BF" box="[306,381,736,762]" captionStart="FIGURES 61 – 68" captionStartId="25.[151,269,1520,1544]" captionTargetBox="[163,1415,203,1485]" captionTargetId="figure@25.[151,1436,194,1496]" captionTargetPageId="25" captionText="FIGURES 61 – 68. Clubiona abnormis sp. n., male holotype (61 – 64) and female paratype (65 – 68). 61 habitus, dorsal; 62 male palp, ventral; 63 ditto, prolateral; 64 ditto, retrolateral; 65 epigyne, ventral; 66 left part of vulva in lactic acid, dorsal; 67 schematic diagram of vulva, dorsal; 68 vulva, caudal. Scale bars = 1.0 mm (61); 0.1 mm (62 – 68)." httpUri="https://zenodo.org/record/181683/files/figure.png" pageId="10" pageNumber="44">61–68</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="A9926520FFCAFFDAFF0DFCF4407A73A1" pageId="10" pageNumber="44" type="materials_examined">
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<paragraph id="E13736ABFFCAFFDAFF0DFCF4427E73D1" blockId="10.[151,1437,810,2036]" pageId="10" pageNumber="44">
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<emphasis id="D3FCEAB9FFCAFFDAFF0DFCF442C97301" bold="true" box="[151,467,810,836]" pageId="10" pageNumber="44">
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<typeStatus id="3E338809FFCAFFDAFF0DFCF443CE7301" box="[151,212,810,836]" pageId="10" pageNumber="44">Type</typeStatus>
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material.
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<typeStatus id="3E338809FFCAFFDAFEC2FCF442D57301" box="[344,463,810,836]" pageId="10" pageNumber="44" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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ɗ, Khao Yai National Park, Khao Yai [Pak Chong District, Nakhorn Ratchasima Province, Northeastern
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<collectingCountry id="999F763BFFCAFFDAFE36FC8C41007329" box="[428,538,850,876]" name="Thailand" pageId="10" pageNumber="44">Thailand</collectingCountry>
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], forests surrounding park headquarters,
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<quantity id="26709B4EFFCAFFDAFC62FC8C47947329" box="[1016,1166,850,876]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="10" pageNumber="44" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">800–1000 m</quantity>
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,
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<date id="9536106BFFCAFFDAFB00FC8C46007329" box="[1178,1306,850,876]" pageId="10" pageNumber="44" value="2006-11-14">14.xi.2006</date>
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, P. Dankittipakul (
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<collectionCode id="8799AE6EFFCAFFDAFF6DFCA4424273D1" box="[247,344,890,916]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="10" pageNumber="44">MHNG</collectionCode>
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).
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</paragraph>
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<paragraph id="E13736ABFFCAFFDAFF5CFC7C407A73A1" blockId="10.[151,1437,810,2036]" pageId="10" pageNumber="44">
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<emphasis id="D3FCEAB9FFCAFFDAFF5CFC7C425773F9" bold="true" box="[198,333,930,956]" pageId="10" pageNumber="44">
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<typeStatus id="3E338809FFCAFFDAFF5CFC7C425273F9" box="[198,328,930,956]" pageId="10" pageNumber="44" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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1Ψ, same data as
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<typeStatus id="3E338809FFCAFFDAFDB5FC7C418C73F9" box="[559,662,930,956]" pageId="10" pageNumber="44" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="8799AE6EFFCAFFDAFD33FC7C401373F9" box="[681,777,930,956]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="10" pageNumber="44">MHNG</collectionCode>
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); 1ɗ, 1Ψ, Loei Province, Phu Ruea District, Phu Ruea National Park,
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<quantity id="26709B4EFFCAFFDAFEDDFC1442B873A1" box="[327,418,970,996]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="44" unit="m" value="1100.0">1100 m</quantity>
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,
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<date id="9536106BFFCAFFDAFE34FC14413873A1" box="[430,546,970,996]" pageId="10" pageNumber="44" value="2006-10-16">16.x.2006</date>
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P. Dankittipakul (
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<collectionCode id="8799AE6EFFCAFFDAFD6CFC14404973A1" box="[758,851,970,996]" httpUri="http://grbio.org/cool/5hig-c780" name="University of Texas" pageId="10" pageNumber="44">TNHM</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="A9926520FFCAFFDAFF5CFC2C468F7449" box="[198,1429,1010,1036]" pageId="10" pageNumber="44" type="etymology">
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<paragraph id="E13736ABFFCAFFDAFF5CFC2C468F7449" blockId="10.[151,1437,810,2036]" box="[198,1429,1010,1036]" pageId="10" pageNumber="44">
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<emphasis id="D3FCEAB9FFCAFFDAFF5CFC2C42497449" bold="true" box="[198,339,1010,1036]" pageId="10" pageNumber="44">Etymology.</emphasis>
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The specific epithet is derived from Latin noun
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<emphasis id="D3FCEAB9FFCAFFDAFC13FC2C40EB744E" box="[905,1009,1010,1035]" italics="true" pageId="10" pageNumber="44">abnorme</emphasis>
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(= deviating from a rule; irregular).
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</paragraph>
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</subSubSection>
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<subSubSection id="A9926520FFCAFFDAFF5CFBC443C476C9" pageId="10" pageNumber="44" type="diagnosis">
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<paragraph id="E13736ABFFCAFFDAFF5CFBC443C476C9" blockId="10.[151,1437,810,2036]" pageId="10" pageNumber="44">
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<emphasis id="D3FCEAB9FFCAFFDAFF5CFBC4425C7471" bold="true" box="[198,326,1050,1076]" pageId="10" pageNumber="44">Diagnosis.</emphasis>
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The males of the
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<taxonomicName id="26884D28FFCAFFDAFDB9FBC441937476" box="[547,649,1050,1075]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="44" phylum="Arthropoda" rank="species" species="japonica">
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<emphasis id="D3FCEAB9FFCAFFDAFDB9FBC441937476" box="[547,649,1050,1075]" italics="true" pageId="10" pageNumber="44">japonica</emphasis>
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</taxonomicName>
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-group (known for
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<taxonomicName id="26884D28FFCAFFDAFCEAFBC447337476" box="[880,1065,1050,1075]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="44" phylum="Arthropoda" rank="species" species="melanosticta">
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<emphasis id="D3FCEAB9FFCAFFDAFCEAFBC447337476" box="[880,1065,1050,1075]" italics="true" pageId="10" pageNumber="44">C. melanosticta</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="26884D28FFCAFFDAFBA1FBC447D07476" box="[1083,1226,1050,1075]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="44" phylum="Arthropoda" rank="species" species="scandens">
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<emphasis id="D3FCEAB9FFCAFFDAFBA1FBC447D07476" box="[1083,1226,1050,1075]" italics="true" pageId="10" pageNumber="44">C. scandens</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="26884D28FFCAFFDAFA9DFBC446867476" box="[1287,1436,1050,1075]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="44" phylum="Arthropoda" rank="species" species="suthepica" status="sp. nov.">
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<emphasis id="D3FCEAB9FFCAFFDAFA9DFBC446867476" box="[1287,1436,1050,1075]" italics="true" pageId="10" pageNumber="44">C. suthepica</emphasis>
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</taxonomicName>
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<emphasis id="D3FCEAB9FFCAFFDAFF0DFB9C43C37419" bold="true" box="[151,217,1090,1116]" pageId="10" pageNumber="44">
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<taxonomicNameLabel id="C8CF57C2FFCAFFDAFF0DFB9C43C37419" box="[151,217,1090,1116]" pageId="10" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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) share the following distinctive suite of characters, here contrasted with the corresponding condition in this new species: (1) beak-shaped conductor distinctly sclerotized (
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<taxonomicName id="26884D28FFCAFFDAFC5EFBB5474374C1" box="[964,1113,1131,1156]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="44" phylum="Arthropoda" rank="species" species="abnormis" status="sp. nov.">
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<emphasis id="D3FCEAB9FFCAFFDAFC5EFBB5474374C1" box="[964,1113,1131,1156]" italics="true" pageId="10" pageNumber="44">C. abnormis</emphasis>
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</taxonomicName>
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<emphasis id="D3FCEAB9FFCAFFDAFBFEFBB447B374C1" bold="true" box="[1124,1193,1130,1156]" pageId="10" pageNumber="44">
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<taxonomicNameLabel id="C8CF57C2FFCAFFDAFBFEFBB447B374C1" box="[1124,1193,1130,1156]" pageId="10" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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: heavily sclerotized hook-shaped distal apex of conductor expanding retrolaterally but its basal portion being membranous,
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<figureCitation id="79B32A2EFFCAFFDAFAF3FB4C43AD7491" captionStart="FIGURES 24 – 29" captionStartId="18.[151,269,1945,1969]" captionTargetBox="[346,1241,167,1937]" captionTargetId="figure@18.[346,1242,167,1940]" captionTargetPageId="18" captionText="FIGURES 24 – 29. Clubiona suthepica sp. n., female paratype (24 – 27). 24 epigyne, ventral; 25 vulva, dorsal; 26 spermathecal base; 27 dilated terminal apparatus of spermathecal head. Clubiona abnormis sp. n., male holotype (28, 29). 28 male palp, dorsal; 29 ditto, retrolateral. Scale bars = 0.1 mm (24 – 25, 28 – 29)." httpUri="https://zenodo.org/record/181676/files/figure.png" pageId="10" pageNumber="44">Figs 28</figureCitation>
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,
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<figureCitation id="79B32A2EFFCAFFDAFF58FB6443F97491" box="[194,227,1210,1236]" captionStart="FIGURES 61 – 68" captionStartId="25.[151,269,1520,1544]" captionTargetBox="[163,1415,203,1485]" captionTargetId="figure@25.[151,1436,194,1496]" captionTargetPageId="25" captionText="FIGURES 61 – 68. Clubiona abnormis sp. n., male holotype (61 – 64) and female paratype (65 – 68). 61 habitus, dorsal; 62 male palp, ventral; 63 ditto, prolateral; 64 ditto, retrolateral; 65 epigyne, ventral; 66 left part of vulva in lactic acid, dorsal; 67 schematic diagram of vulva, dorsal; 68 vulva, caudal. Scale bars = 1.0 mm (61); 0.1 mm (62 – 68)." httpUri="https://zenodo.org/record/181683/files/figure.png" pageId="10" pageNumber="44">62</figureCitation>
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); (2) tegulum with sinuate sperm duct, generally forming a double loop (
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<taxonomicName id="26884D28FFCAFFDAFBAEFB6547DE7491" box="[1076,1220,1211,1236]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="44" phylum="Arthropoda" rank="species" species="abnormis" status="sp. nov.">
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<emphasis id="D3FCEAB9FFCAFFDAFBAEFB6547DE7491" box="[1076,1220,1211,1236]" italics="true" pageId="10" pageNumber="44">C. abnormis</emphasis>
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||
</taxonomicName>
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<emphasis id="D3FCEAB9FFCAFFDAFB51FB6446167491" bold="true" box="[1227,1292,1210,1236]" pageId="10" pageNumber="44">
|
||
<taxonomicNameLabel id="C8CF57C2FFCAFFDAFB51FB6446167491" box="[1227,1292,1210,1236]" pageId="10" pageNumber="44" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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: sperm duct U-shaped,
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<figureCitation id="79B32A2EFFCAFFDAFE80FB3C426F74B9" box="[282,373,1250,1276]" captionStart="FIGURES 61 – 68" captionStartId="25.[151,269,1520,1544]" captionTargetBox="[163,1415,203,1485]" captionTargetId="figure@25.[151,1436,194,1496]" captionTargetPageId="25" captionText="FIGURES 61 – 68. Clubiona abnormis sp. n., male holotype (61 – 64) and female paratype (65 – 68). 61 habitus, dorsal; 62 male palp, ventral; 63 ditto, prolateral; 64 ditto, retrolateral; 65 epigyne, ventral; 66 left part of vulva in lactic acid, dorsal; 67 schematic diagram of vulva, dorsal; 68 vulva, caudal. Scale bars = 1.0 mm (61); 0.1 mm (62 – 68)." httpUri="https://zenodo.org/record/181683/files/figure.png" pageId="10" pageNumber="44">Fig. 62</figureCitation>
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); (3) dorsum of opisthosoma with a pattern consisting of dark blotches and stripes (
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<taxonomicName id="26884D28FFCAFFDAFA1BFB3D421C7561" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="44" phylum="Arthropoda" rank="species" species="abnormis" status="sp. nov.">
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<emphasis id="D3FCEAB9FFCAFFDAFA1BFB3D421C7561" italics="true" pageId="10" pageNumber="44">C. abnormis</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="C8CF57C2FFCAFFDAFE96FAD442547561" box="[268,334,1290,1316]" pageId="10" pageNumber="44" rank="species">
|
||
<emphasis id="D3FCEAB9FFCAFFDAFE96FAD442547561" bold="true" box="[268,334,1290,1316]" pageId="10" pageNumber="44">sp. n.</emphasis>
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</taxonomicNameLabel>
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: with a lightly sclerotized dorsal scutum and a reduced opisthosomal pattern,
|
||
<figureCitation id="79B32A2EFFCAFFDAFB40FAD4462B7561" box="[1242,1329,1290,1316]" captionStart="FIGURES 61 – 68" captionStartId="25.[151,269,1520,1544]" captionTargetBox="[163,1415,203,1485]" captionTargetId="figure@25.[151,1436,194,1496]" captionTargetPageId="25" captionText="FIGURES 61 – 68. Clubiona abnormis sp. n., male holotype (61 – 64) and female paratype (65 – 68). 61 habitus, dorsal; 62 male palp, ventral; 63 ditto, prolateral; 64 ditto, retrolateral; 65 epigyne, ventral; 66 left part of vulva in lactic acid, dorsal; 67 schematic diagram of vulva, dorsal; 68 vulva, caudal. Scale bars = 1.0 mm (61); 0.1 mm (62 – 68)." httpUri="https://zenodo.org/record/181683/files/figure.png" pageId="10" pageNumber="44">Fig. 61</figureCitation>
|
||
). Female genital structure is remarkable in which the epigyne represented by a large depression, and by a small posterior U-shaped epigynal atrium with indistinct genital orifices (
|
||
<figureCitation id="79B32A2EFFCAFFDAFCE2FA8440C87531" box="[888,978,1370,1396]" captionStart="FIGURES 61 – 68" captionStartId="25.[151,269,1520,1544]" captionTargetBox="[163,1415,203,1485]" captionTargetId="figure@25.[151,1436,194,1496]" captionTargetPageId="25" captionText="FIGURES 61 – 68. Clubiona abnormis sp. n., male holotype (61 – 64) and female paratype (65 – 68). 61 habitus, dorsal; 62 male palp, ventral; 63 ditto, prolateral; 64 ditto, retrolateral; 65 epigyne, ventral; 66 left part of vulva in lactic acid, dorsal; 67 schematic diagram of vulva, dorsal; 68 vulva, caudal. Scale bars = 1.0 mm (61); 0.1 mm (62 – 68)." httpUri="https://zenodo.org/record/181683/files/figure.png" pageId="10" pageNumber="44">Fig. 65</figureCitation>
|
||
). Furthermore, spermathecal heads are differently formed (
|
||
<figureCitation id="79B32A2EFFCAFFDAFEE6FA5C411C75D9" box="[380,518,1410,1436]" captionStart="FIGURES 61 – 68" captionStartId="25.[151,269,1520,1544]" captionTargetBox="[163,1415,203,1485]" captionTargetId="figure@25.[151,1436,194,1496]" captionTargetPageId="25" captionText="FIGURES 61 – 68. Clubiona abnormis sp. n., male holotype (61 – 64) and female paratype (65 – 68). 61 habitus, dorsal; 62 male palp, ventral; 63 ditto, prolateral; 64 ditto, retrolateral; 65 epigyne, ventral; 66 left part of vulva in lactic acid, dorsal; 67 schematic diagram of vulva, dorsal; 68 vulva, caudal. Scale bars = 1.0 mm (61); 0.1 mm (62 – 68)." httpUri="https://zenodo.org/record/181683/files/figure.png" pageId="10" pageNumber="44">Figs 66–68</figureCitation>
|
||
): the terminal part carrying fertilization ducts but in other females belonging to the
|
||
<taxonomicName id="26884D28FFCAFFDAFF5FFA7542317581" box="[197,299,1451,1476]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="44" phylum="Arthropoda" rank="species" species="japonica">
|
||
<emphasis id="D3FCEAB9FFCAFFDAFF5FFA7542317581" box="[197,299,1451,1476]" italics="true" pageId="10" pageNumber="44">japonica</emphasis>
|
||
</taxonomicName>
|
||
-group the fertilization ducts are surrounded by the curved spermathecal bases. There remains considerable uncertainty about placing this new species in the
|
||
<taxonomicName id="26884D28FFCAFFDAFCE9FA0D40C375A9" box="[883,985,1491,1516]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="44" phylum="Arthropoda" rank="species" species="japonica">
|
||
<emphasis id="D3FCEAB9FFCAFFDAFCE9FA0D40C375A9" box="[883,985,1491,1516]" italics="true" pageId="10" pageNumber="44">japonica</emphasis>
|
||
</taxonomicName>
|
||
-group. The general shape and appearance of this new species is fairly typical of a patterned, dark-colored member of the
|
||
<taxonomicName id="26884D28FFCAFFDAFBE5FA2547FF7651" box="[1151,1253,1531,1556]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="44" phylum="Arthropoda" rank="species" species="japonica">
|
||
<emphasis id="D3FCEAB9FFCAFFDAFBE5FA2547FF7651" box="[1151,1253,1531,1556]" italics="true" pageId="10" pageNumber="44">japonica</emphasis>
|
||
</taxonomicName>
|
||
-group. In spite of the fact that has just been mentioned, it is impossible to discern any obvious derived features (i.e., sclerotized conductor) that would indicate a close relationship to the
|
||
<taxonomicName id="26884D28FFCAFFDAFC3CF99547167621" box="[934,1036,1611,1636]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="44" phylum="Arthropoda" rank="species" species="japonica">
|
||
<emphasis id="D3FCEAB9FFCAFFDAFC3CF99547167621" box="[934,1036,1611,1636]" italics="true" pageId="10" pageNumber="44">japonica</emphasis>
|
||
</taxonomicName>
|
||
-group found in Southeast Asian today.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A9926520FFCAFFDBFF5CF94447FA763F" lastPageId="11" lastPageNumber="45" pageId="10" pageNumber="44" type="description">
|
||
<paragraph id="E13736ABFFCAFFDAFF5CF94443C27699" blockId="10.[151,1437,810,2036]" pageId="10" pageNumber="44">
|
||
<emphasis id="D3FCEAB9FFCAFFDAFF5CF944412276F1" bold="true" box="[198,568,1690,1716]" pageId="10" pageNumber="44">
|
||
Description. Male (
|
||
<typeStatus id="3E338809FFCAFFDAFE22F944413176F1" box="[440,555,1690,1716]" pageId="10" pageNumber="44" type="holotype">holotype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 5.1; prosoma 2.2 long, 2.2 wide; opisthosoma 2.9 long, 1.5 wide.
|
||
</paragraph>
|
||
<paragraph id="E13736ABFFCAFFDAFF5CF935411C7739" blockId="10.[151,1437,810,2036]" pageId="10" pageNumber="44">
|
||
<emphasis id="D3FCEAB9FFCAFFDAFF5CF93542357741" box="[198,303,1771,1796]" italics="true" pageId="10" pageNumber="44">Prosoma</emphasis>
|
||
(
|
||
<figureCitation id="79B32A2EFFCAFFDAFEDAF93442807741" box="[320,410,1770,1796]" captionStart="FIGURES 61 – 68" captionStartId="25.[151,269,1520,1544]" captionTargetBox="[163,1415,203,1485]" captionTargetId="figure@25.[151,1436,194,1496]" captionTargetPageId="25" captionText="FIGURES 61 – 68. Clubiona abnormis sp. n., male holotype (61 – 64) and female paratype (65 – 68). 61 habitus, dorsal; 62 male palp, ventral; 63 ditto, prolateral; 64 ditto, retrolateral; 65 epigyne, ventral; 66 left part of vulva in lactic acid, dorsal; 67 schematic diagram of vulva, dorsal; 68 vulva, caudal. Scale bars = 1.0 mm (61); 0.1 mm (62 – 68)." httpUri="https://zenodo.org/record/181683/files/figure.png" pageId="10" pageNumber="44">Fig. 61</figureCitation>
|
||
). Long-oval, pars cephalica distinctly narrowed, in profile strongly raised, highest just behind fovea; integument smooth, covered with numerous fine hairs. Dorsal shield of prosoma bright yellow, pars cephalica dark yellow in ocular area, without markings. Chelicerae dark brown. Labium and endites yellowish brown. Sternum yellow.
|
||
</paragraph>
|
||
<paragraph id="E13736ABFFCAFFDAFF5CF855429177B1" blockId="10.[151,1437,810,2036]" pageId="10" pageNumber="44">
|
||
<emphasis id="D3FCEAB9FFCAFFDAFF5CF855421F77E1" box="[198,261,1931,1956]" italics="true" pageId="10" pageNumber="44">Eyes.</emphasis>
|
||
Eyes with black rings. Eye sizes and interdistances: AME 0.11, ALE 0.13, PME 0.13, PLE 0.11, AME–AME 0.08, AME–ALE 0.03, PME–PME 0.11, PME–PLE 0.13; MOQ: 0.31 long, 0.33 anterior width, 0.46 posterior width.
|
||
</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF5CFF46426E709F" blockId="11.[151,1437,152,1738]" pageId="11" pageNumber="45">
|
||
<emphasis id="D3FCEAB9FFCBFFDBFF5CFF46421F70F4" box="[198,261,152,177]" italics="true" pageId="11" pageNumber="45">Legs.</emphasis>
|
||
Legs yellow; anterior tibiae, metatarsi and tarsi brown; without dark annulations. Leg formula 4213. Leg measurements
|
||
</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFE2BFF36471C7147" blockId="11.[151,1437,152,1738]" box="[433,1030,232,258]" pageId="11" pageNumber="45">I II III IV</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF0DFECE4701716F" blockId="11.[151,1437,152,1738]" box="[151,1051,272,298]" pageId="11" pageNumber="45">Femora 1.85 2.20 1.72 2.30</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF0DFEE647017117" blockId="11.[151,1437,152,1738]" box="[151,1051,312,338]" pageId="11" pageNumber="45">Patellae + tibiae 2.85 3.28 1.95 3.25</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF0DFEBE4701713F" blockId="11.[151,1437,152,1738]" box="[151,1051,352,378]" pageId="11" pageNumber="45">Metatarsi 1.45 1.72 1.72 2.90</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF0DFE56470171E7" blockId="11.[151,1437,152,1738]" box="[151,1051,392,418]" pageId="11" pageNumber="45">Tarsi 0.73 0.81 0.62 0.65</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF0DFE6E4700718F" blockId="11.[151,1437,152,1738]" box="[151,1050,432,458]" pageId="11" pageNumber="45">Total 6.88 8.01 6.01 9.10</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF5CFE0646527207" blockId="11.[151,1437,152,1738]" pageId="11" pageNumber="45">
|
||
<emphasis id="D3FCEAB9FFCBFFDBFF5CFE06427B71B4" box="[198,353,472,497]" italics="true" pageId="11" pageNumber="45">Opisthosoma</emphasis>
|
||
(
|
||
<figureCitation id="79B32A2EFFCBFFDBFEEBFE0642D271B7" box="[369,456,472,498]" captionStart="FIGURES 61 – 68" captionStartId="25.[151,269,1520,1544]" captionTargetBox="[163,1415,203,1485]" captionTargetId="figure@25.[151,1436,194,1496]" captionTargetPageId="25" captionText="FIGURES 61 – 68. Clubiona abnormis sp. n., male holotype (61 – 64) and female paratype (65 – 68). 61 habitus, dorsal; 62 male palp, ventral; 63 ditto, prolateral; 64 ditto, retrolateral; 65 epigyne, ventral; 66 left part of vulva in lactic acid, dorsal; 67 schematic diagram of vulva, dorsal; 68 vulva, caudal. Scale bars = 1.0 mm (61); 0.1 mm (62 – 68)." httpUri="https://zenodo.org/record/181683/files/figure.png" pageId="11" pageNumber="45">Fig. 61</figureCitation>
|
||
). Elongate-oval; dorsal scutum ovoid, lightly sclerotized, furnished with a thick tuft of hairs anteriorly; dorsum pale yellow, with dark purplish coloring which is formed into U-shaped pattern; a butterfly-shaped marking situated underneath the dorsal scutum. Spinnerets white, except ALS brown.
|
||
</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF5CFD8E40D97377" blockId="11.[151,1437,152,1738]" pageId="11" pageNumber="45">
|
||
<emphasis id="D3FCEAB9FFCBFFDBFF5CFD8E4224722C" box="[198,318,592,617]" italics="true" pageId="11" pageNumber="45">Male palp</emphasis>
|
||
(
|
||
<figureCitation id="79B32A2EFFCBFFDBFED7FD8E42CE722F" box="[333,468,592,618]" captionStart="FIGURES 24 – 29" captionStartId="18.[151,269,1945,1969]" captionTargetBox="[346,1241,167,1937]" captionTargetId="figure@18.[346,1242,167,1940]" captionTargetPageId="18" captionText="FIGURES 24 – 29. Clubiona suthepica sp. n., female paratype (24 – 27). 24 epigyne, ventral; 25 vulva, dorsal; 26 spermathecal base; 27 dilated terminal apparatus of spermathecal head. Clubiona abnormis sp. n., male holotype (28, 29). 28 male palp, dorsal; 29 ditto, retrolateral. Scale bars = 0.1 mm (24 – 25, 28 – 29)." httpUri="https://zenodo.org/record/181676/files/figure.png" pageId="11" pageNumber="45">Figs 28–29</figureCitation>
|
||
,
|
||
<figureCitation id="79B32A2EFFCBFFDBFE7AFD8E4134722F" box="[480,558,592,618]" captionStart="FIGURES 61 – 68" captionStartId="25.[151,269,1520,1544]" captionTargetBox="[163,1415,203,1485]" captionTargetId="figure@25.[151,1436,194,1496]" captionTargetPageId="25" captionText="FIGURES 61 – 68. Clubiona abnormis sp. n., male holotype (61 – 64) and female paratype (65 – 68). 61 habitus, dorsal; 62 male palp, ventral; 63 ditto, prolateral; 64 ditto, retrolateral; 65 epigyne, ventral; 66 left part of vulva in lactic acid, dorsal; 67 schematic diagram of vulva, dorsal; 68 vulva, caudal. Scale bars = 1.0 mm (61); 0.1 mm (62 – 68)." httpUri="https://zenodo.org/record/181683/files/figure.png" pageId="11" pageNumber="45">62–64</figureCitation>
|
||
). Retrolateral tibial apophysis reddish, strongly excavated ventrally, gradually tapering towards its sharply pointed apex. Cymbium with basal concavity on retrolateral side accommodating tibial apophysis. Bulb with anterior flange lightly sclerotized; sperm duct U-shaped. Conductor beakshaped, situated on apical part of the tegulum, basal part membranous, terminal part strongly sclerotized, directed retrolaterad then abruptly bending distad. Embolus thin, filiform; embolic base arising mesolaterally. Tegular apophysis strongly sclerotized, situated anterio-retrolaterally.
|
||
</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF5CFCE1461073EF" blockId="11.[151,1437,152,1738]" pageId="11" pageNumber="45">
|
||
<emphasis id="D3FCEAB9FFCBFFDBFF5CFCE142A2731C" bold="true" box="[198,440,831,857]" pageId="11" pageNumber="45">
|
||
Female (
|
||
<typeStatus id="3E338809FFCBFFDBFEA9FCE142B1731C" box="[307,427,831,857]" pageId="11" pageNumber="45" type="paratype">paratype</typeStatus>
|
||
):
|
||
</emphasis>
|
||
Total length 6.5; prosoma 3.0 long, 2.2 wide; opisthosoma 3.5 long, 2.0 wide. Not strikingly different from the males but slightly larger in size; prosoma less convex, broader anteriorly; chelicerae orange-brown; dorsum of opisthosoma without dorsal scutum; spinnerets all white in color.
|
||
</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF5CFC6642917467" blockId="11.[151,1437,152,1738]" pageId="11" pageNumber="45">
|
||
<emphasis id="D3FCEAB9FFCBFFDBFF5CFC66421F7394" box="[198,261,952,977]" italics="true" pageId="11" pageNumber="45">Eyes.</emphasis>
|
||
Eyes with black rings. Eye sizes and interdistances: AME 0.16, ALE 0.16, PME 0.16, PLE 0.13, AME–AME 0.15, AME–ALE 0.08, PME–PME 0.30, PME–PLE 0.21; MOQ: 0.30 long, 0.41 anterior width, 0.61 posterior width.
|
||
</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF5CFBEE42EB740F" blockId="11.[151,1437,152,1738]" box="[198,497,1072,1098]" pageId="11" pageNumber="45">
|
||
<emphasis id="D3FCEAB9FFCBFFDBFF5CFBEE421F740C" box="[198,261,1072,1097]" italics="true" pageId="11" pageNumber="45">Legs.</emphasis>
|
||
Leg measurements:
|
||
</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFE2BFB86471C7437" blockId="11.[151,1437,152,1738]" box="[433,1030,1112,1138]" pageId="11" pageNumber="45">I II III IV</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF0DFB5E470174DF" blockId="11.[151,1437,152,1738]" box="[151,1051,1152,1178]" pageId="11" pageNumber="45">Femora 1.98 2.45 1.86 2.78</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF0DFB7647017487" blockId="11.[151,1437,152,1738]" box="[151,1051,1192,1218]" pageId="11" pageNumber="45">Patellae + tibiae 2.95 3.15 2.45 3.59</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF0DFB0E470174AF" blockId="11.[151,1437,152,1738]" box="[151,1051,1232,1258]" pageId="11" pageNumber="45">Metatarsi 1.45 1.76 1.46 2.95</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF0DFB2647017557" blockId="11.[151,1437,152,1738]" box="[151,1051,1272,1298]" pageId="11" pageNumber="45">Tarsi 0.65 0.85 0.62 0.78</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF0DFAFE4733757F" blockId="11.[151,1437,152,1738]" box="[151,1065,1312,1338]" pageId="11" pageNumber="45">Total 7.03 8.21 6.39 10.10</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF5CFA9643C4766F" blockId="11.[151,1437,152,1738]" pageId="11" pageNumber="45">
|
||
<emphasis id="D3FCEAB9FFCBFFDBFF5CFA9642BF7524" box="[198,421,1352,1377]" italics="true" pageId="11" pageNumber="45">Epigyne and vulva</emphasis>
|
||
(
|
||
<figureCitation id="79B32A2EFFCBFFDBFE2DFA9641597527" box="[439,579,1352,1378]" captionStart="FIGURES 61 – 68" captionStartId="25.[151,269,1520,1544]" captionTargetBox="[163,1415,203,1485]" captionTargetId="figure@25.[151,1436,194,1496]" captionTargetPageId="25" captionText="FIGURES 61 – 68. Clubiona abnormis sp. n., male holotype (61 – 64) and female paratype (65 – 68). 61 habitus, dorsal; 62 male palp, ventral; 63 ditto, prolateral; 64 ditto, retrolateral; 65 epigyne, ventral; 66 left part of vulva in lactic acid, dorsal; 67 schematic diagram of vulva, dorsal; 68 vulva, caudal. Scale bars = 1.0 mm (61); 0.1 mm (62 – 68)." httpUri="https://zenodo.org/record/181683/files/figure.png" pageId="11" pageNumber="45">Figs 65–68</figureCitation>
|
||
). Epigynal plate a very large lightly sclerotized depression, longer than wide. Epigynal atrium U-shaped, situated posteriorly, margin not rebordered. Genital orifices entering vulva via posterior part, above epigastric furrow. Spermathecal stalks short, longitudinal tube, located posteriorly and connected to spherical membranous bursae; spermathecal bases carrying extraordinary large fertilization ducts; spermathecal heads elongated, extending anteriorly, dilated terminal portion ovoid, with numerous pores.
|
||
</paragraph>
|
||
<paragraph id="E13736ABFFCBFFDBFF5CF9E947FA763F" blockId="11.[151,1437,152,1738]" pageId="11" pageNumber="45">
|
||
<emphasis id="D3FCEAB9FFCBFFDBFF5CF9E9428A7614" bold="true" box="[198,400,1591,1617]" pageId="11" pageNumber="45">Natural history.</emphasis>
|
||
<taxonomicName id="26884D28FFCBFFDBFE00F9E6419A7614" box="[410,640,1592,1617]" class="Arachnida" family="Clubionidae" genus="Clubiona" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="45" phylum="Arthropoda" rank="species" species="abnormis" status="sp. nov.">
|
||
<emphasis id="D3FCEAB9FFCBFFDBFE00F9E6419A7614" box="[410,640,1592,1617]" italics="true" pageId="11" pageNumber="45">Clubiona abnormis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="D3FCEAB9FFCBFFDBFD13F9E941D47614" bold="true" box="[649,718,1591,1617]" pageId="11" pageNumber="45">
|
||
<taxonomicNameLabel id="C8CF57C2FFCBFFDBFD13F9E941D47614" box="[649,718,1591,1617]" pageId="11" pageNumber="45" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
inhabits evergreen hill forest with densely covered canopy. The localities situate on high altitude mountains (above
|
||
<quantity id="26709B4EFFCBFFDBFCB2F9BE409B763F" box="[808,897,1632,1658]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="45" unit="m" value="1000.0">1000 m</quantity>
|
||
asl) in northeastern
|
||
<collectingCountry id="999F763BFFCBFFDBFBEAF9BE47C6763F" box="[1136,1244,1632,1658]" name="Thailand" pageId="11" pageNumber="45">Thailand</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A9926520FFCBFFDBFF5CF9594059768F" pageId="11" pageNumber="45" type="distribution">
|
||
<paragraph id="E13736ABFFCBFFDBFF5CF9594059768F" blockId="11.[151,1437,152,1738]" pageId="11" pageNumber="45">
|
||
<emphasis id="D3FCEAB9FFCBFFDBFF5CF959427D76E4" bold="true" box="[198,359,1671,1697]" pageId="11" pageNumber="45">Distribution.</emphasis>
|
||
Northeastern
|
||
<collectingCountry id="999F763BFFCBFFDBFD96F956416E76E7" box="[524,628,1672,1698]" name="Thailand" pageId="11" pageNumber="45">Thailand</collectingCountry>
|
||
(Khao Yai and Phu Reua National Parks,
|
||
<figureCitation id="79B32A2EFFCBFFDBFBFAF95647AC76E7" box="[1120,1206,1672,1698]" captionStart="FIGURE 69" captionStartId="26.[151,255,1325,1349]" captionTargetBox="[227,1360,194,1298]" captionTargetId="figure@26.[227,1361,194,1299]" captionTargetPageId="26" captionText="FIGURE 69. Records of Clubiona species treated in the present study: * C. filicata; C. suthepica sp. n.; C. melanosticta; C. octoginta sp. n.; C. campylacantha sp. n.; Ο C. filifera sp. n.; C. abnormis sp. n." httpUri="https://zenodo.org/record/181684/files/figure.png" pageId="11" pageNumber="45">Fig. 69</figureCitation>
|
||
), but there are indications that this new species may well have a wider range.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |