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<document id="A8D9145EA5ABF3869FCDD2583CFA4C0A" ID-CLB-Dataset="4944" ID-DOI="10.11646/zootaxa.4984.1.15" ID-GBIF-Dataset="51b2c494-8a51-4c80-bfdd-98240d6f4c07" ID-ISSN="1175-5326" ID-Zenodo-Dep="4926887" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623426170995" checkinUser="plazi" docAuthor="Mamani, Claudia Vanesa, Porto, Willians, Iglesias, Patricia P. &amp; González, Abel Pérez-" docDate="2021" docId="2D0A576DFF94312FA895B4E4FCD8FCCA" docLanguage="en" docName="zootaxa.4984.1.15.pdf" docOrigin="Zootaxa 4984 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stenostygnus huberi Mamani &amp; Porto &amp; Iglesias &amp; González 2021, spec. nov." docType="treatment" docUuid="061A42CE-09C0-40FE-AF84-A14F1E773AC6" docUuidSource="ZooBank" docVersion="7" lastPageNumber="196" masterDocId="D1332F15FF9F3121A802B132FF8DFFA9" masterDocTitle="Two new species of Stenostygnus from South America (Opiliones: Biantidae Stenostygninae)" masterLastPageNumber="202" masterPageNumber="182" pageNumber="193" updateTime="1699010268631" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A9B074D4D2AFEF8485C81DFC8F01FDCB">Two new species of Stenostygnus from South America (Opiliones: Biantidae Stenostygninae)</mods:title>
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<mods:namePart id="6895FACC1022D5DAA361EBF3BD8F8119">Mamani, Claudia Vanesa</mods:namePart>
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<mods:namePart id="0FADA6112387E76B2A3CAB7BE31155C3">Porto, Willians</mods:namePart>
<mods:nameIdentifier id="002CB5B26C479A258680627005587F8F" type="ORCID">0000-0002-4209-5664</mods:nameIdentifier>
<mods:affiliation id="965CAB0317D5A3B15EB5BFE767FC4515">https: // orcid. org / 0000 - 0002 - 4209 - 5664</mods:affiliation>
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<mods:namePart id="3DAF4BEE3CDADED2C61194E788E8C4FB">Iglesias, Patricia P.</mods:namePart>
<mods:nameIdentifier id="A04935AA39E0401C9F188DBC7AAA4637" type="ORCID">0000-0002-3979-4253</mods:nameIdentifier>
<mods:affiliation id="6AB7D0A88081C234F408FE7FD60B53EA">Current address: Laboratorio de Genética Evolutiva, Universidad Nacional de Misiones - CONICET, Félix de Azara 1552, N 3300 LQH, Misiones, Argentina; https: // orcid. org / 0000 - 0002 - 3979 - 4253</mods:affiliation>
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<mods:namePart id="3BB576FDBCD5212A0218F0A60A12C3FB">González, Abel Pérez-</mods:namePart>
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<mods:nameIdentifier id="25C6F103EB9F93CBDFA3F6FC8FA28660" type="email">abelaracno@gmail.com</mods:nameIdentifier>
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<mods:date id="251B62756E731F5B9E653C9D3C51EA1D">2021</mods:date>
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<mods:number id="3AD307EBD05C47E12866C649C8220107">2021-06-10</mods:number>
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<treatment id="2D0A576DFF94312FA895B4E4FCD8FCCA" ID-DOI="http://doi.org/10.5281/zenodo.5213031" ID-GBIF-Taxon="184570904" ID-Zenodo-Dep="5213031" LSID="urn:lsid:zoobank.org:act:061A42CE-09C0-40FE-AF84-A14F1E773AC6" httpUri="http://treatment.plazi.org/id/2D0A576DFF94312FA895B4E4FCD8FCCA" lastPageId="14" lastPageNumber="196" pageId="11" pageNumber="193">
<subSubSection id="EDB9B5F0FF94312AA895B4E4FD80FA59" box="[151,525,1493,1520]" pageId="11" pageNumber="193" type="nomenclature">
<paragraph id="A51CE67BFF94312AA895B4E4FD80FA59" blockId="11.[151,931,1493,1585]" box="[151,525,1493,1520]" pageId="11" pageNumber="193">
<heading id="FE545117FF94312AA895B4E4FD80FA59" bold="true" box="[151,525,1493,1520]" fontSize="11" level="1" pageId="11" pageNumber="193" reason="1">
<emphasis id="97D73A69FF94312AA895B4E4FD80FA59" bold="true" box="[151,525,1493,1520]" pageId="11" pageNumber="193">
<taxonomicName id="62A39DF8FF94312AA895B4E4FE03FA59" ID-CoL="9X5HM" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[151,398,1493,1520]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="193" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov.">
<emphasis id="97D73A69FF94312AA895B4E4FE03FA59" bold="true" box="[151,398,1493,1520]" italics="true" pageId="11" pageNumber="193">Stenostygnus huberi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CE48712FF94312AA997B4E4FD80FA59" box="[405,525,1494,1520]" pageId="11" pageNumber="193" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="EDB9B5F0FF94312AA895B4C8FC2EF998" pageId="11" pageNumber="193" type="description">
<paragraph id="A51CE67BFF94312AA895B4C8FE8BF9BD" blockId="11.[151,931,1493,1585]" box="[151,262,1530,1556]" pageId="11" pageNumber="193">
<figureCitation id="3D98FAFEFF94312AA895B4C8FE8BF9BD" box="[151,262,1530,1556]" captionStart-0="FIGURES 7" captionStart-1="FIGURES 8" captionStart-2="FIGURES 9" captionStart-3="FIGURES 11" captionStart-4="FIGURES 12" captionStartId-0="10.[151,264,1943,1967]" captionStartId-1="11.[151,264,1351,1375]" captionStartId-2="12.[151,264,1864,1888]" captionStartId-3="16.[151,264,1327,1351]" captionStartId-4="17.[151,264,1807,1831]" captionTargetBox-0="[175,1413,181,1919]" captionTargetBox-1="[151,1436,181,1328]" captionTargetBox-2="[151,1436,957,1841]" captionTargetBox-3="[158,1429,181,1303]" captionTargetBox-4="[151,1436,181,1783]" captionTargetId-0="figure-17@10.[175,1413,181,1919]" captionTargetId-1="figure-19@11.[151,1436,181,1328]" captionTargetId-2="figure-420@12.[151,1436,957,1841]" captionTargetId-3="figure-17@16.[158,1429,181,1304]" captionTargetId-4="figure-19@17.[151,1436,181,1783]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="16" captionTargetPageId-4="17" captionText-0="FIGURES 7A–C. Stenostygnus huberi spec. nov., male (α) (holotype, MIZA 0103461), habitus (A dorsal, B ventral, C lateral). Scale bars: A, B= 1 mm, C= 500 µm." captionText-1="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." captionText-2="FIGURES 9A–E. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429). A–C Left chelicera (A ectal, B frontal, C detail of fingers), D–E Left pedipalp (D ectal, E detail of femur). Scale bars: A–C= 200 µm; D= 1 mm; E= 500 µm." captionText-3="FIGURES 11A–G. Stenostygnus huberi spec. nov., penis (A–B male (ss) paratype, MNRJ 2429; C male (ss), paratype, MACN 41020; D–G holotype male (α), MIZA 0103461). A, D Dorsal view. B, E, G Lateral view (G with detail of stylus and conduc- tors). C, F Ventral view. LA—lamina apicalis. Ti—Titillator. Scale bars: 100 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." captionText-4="FIGURES 12A–F. Stenostygnus huberi spec. nov., sexual dimorphism (A–B female paratype, MNRJ 2429; C–D male (ss) paratype, MACN 41020; E–F (holotype male (α), MIZA 0103461), A, C, E Dorsal view. B, D, F Detail of patella. Scale bars: A, C, E= 2 mm; B, F= 200 µm; D= 500 µm" figureDoi-0="http://doi.org/10.5281/zenodo.4926907" figureDoi-1="http://doi.org/10.5281/zenodo.4926909" figureDoi-2="http://doi.org/10.5281/zenodo.4926911" figureDoi-3="http://doi.org/10.5281/zenodo.4926915" figureDoi-4="http://doi.org/10.5281/zenodo.4926917" httpUri-0="https://zenodo.org/record/4926907/files/figure.png" httpUri-1="https://zenodo.org/record/4926909/files/figure.png" httpUri-2="https://zenodo.org/record/4926911/files/figure.png" httpUri-3="https://zenodo.org/record/4926915/files/figure.png" httpUri-4="https://zenodo.org/record/4926917/files/figure.png" pageId="11" pageNumber="193">Figs 7–12</figureCitation>
</paragraph>
<paragraph id="A51CE67BFF94312AA895B72BFC2EF998" blockId="11.[151,931,1493,1585]" box="[151,931,1561,1585]" pageId="11" pageNumber="193">
<uri id="D132EA79FF94312AA895B72BFC2EF998" box="[151,931,1561,1585]" pageId="11" pageNumber="193">
urn:lsid:zoobank.org:act:
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</paragraph>
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<paragraph id="A51CE67BFF94312AA895B76DFE7BF8A0" blockId="11.[151,1437,1631,2018]" pageId="11" pageNumber="193">
<emphasis id="97D73A69FF94312AA895B76DFEC9F9D0" bold="true" box="[151,324,1631,1657]" pageId="11" pageNumber="193">Type material.</emphasis>
<materialsCitation id="15CBEC26FF94312AA94BB76DFB3DF934" ID-GBIF-Occurrence="3335045303" collectingDate="2002-12-15" collectingDateMax="2002-12-16" collectingDateMin="2002-12-15" collectionCode="MIZA" collectorName="A. Huber, O &amp; Villarreal M. &amp; Striffler" country="Venezuela" location="Parque Nacional Yacambu" pageId="11" pageNumber="193" specimenCode="MIZA 0103461" specimenCount="1" stateProvince="Lara">
<emphasis id="97D73A69FF94312AA94BB76DFDCBF9D0" bold="true" box="[329,582,1631,1657]" pageId="11" pageNumber="193">
<collectingCountry id="DDB4A6EBFF94312AA94BB76DFE74F9D0" box="[329,505,1631,1657]" name="Venezuela" pageId="11" pageNumber="193">VENEZUELA</collectingCountry>
,
<emphasis id="97D73A69FF94312AAA07B76DFDB0F9D0" bold="true" box="[517,573,1631,1657]" italics="true" pageId="11" pageNumber="193">
<collectingRegion id="67672899FF94312AAA07B76DFDB0F9D0" box="[517,573,1631,1657]" country="Venezuela" name="Lara" pageId="11" pageNumber="193">Lara</collectingRegion>
</emphasis>
:
</emphasis>
H
<emphasis id="97D73A69FF94312AAA62B76DFD7AF9D0" bold="true" box="[608,759,1631,1657]" pageId="11" pageNumber="193">olotype male</emphasis>
(α) (
<specimenCode id="F5054E00FF94312AAB29B76DFC53F9D3" box="[811,990,1631,1658]" collectionCode="MIZA" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="11" pageNumber="193" type="Museum">MIZA 0103461</specimenCode>
),
<location id="A07CB0A0FF94312AABF3B752FA97F9D0" LSID="urn:lsid:plazi:treatment:2D0A576DFF94312FA895B4E4FCD8FCCA:A07CB0A0FF94312AABF3B752FA97F9D0" box="[1009,1306,1631,1658]" country="Venezuela" name="Parque Nacional Yacambu" pageId="11" pageNumber="193" stateProvince="Lara">Parque Nacional Yacambu</location>
,
<date id="D11DC0BBFF94312AAD21B76DFF5FF934" pageId="11" pageNumber="193" value="2002-12-15" valueMax="2002-12-16" valueMin="2002-12-15">
<collectingDate id="C1593953FF94312AAD21B76DFF5FF934" pageId="11" pageNumber="193" value="2002-12-15" valueMax="2002-12-16" valueMin="2002-12-15">15–16 Dec 2002</collectingDate>
</date>
,
<collectorName id="085683ADFF94312AA8DFB7B1FE39F934" box="[221,436,1667,1693]" pageId="11" pageNumber="193">A. Pérez-González</collectorName>
,
<collectorName id="085683ADFF94312AA9BDB7B1FDC0F934" box="[447,589,1667,1693]" pageId="11" pageNumber="193">A. Giupponi</collectorName>
&amp;
<collectorName id="085683ADFF94312AAA71B7B1FCFDF934" box="[627,880,1667,1693]" pageId="11" pageNumber="193">Bernhard A. Huber, O</collectorName>
.
<collectorName id="085683ADFF94312AAB7EB7B1FB83F937" box="[892,1038,1667,1694]" pageId="11" pageNumber="193">Villarreal M.</collectorName>
&amp; B.
<collectorName id="085683ADFF94312AAC54B7B1FB3DF934" box="[1110,1200,1667,1693]" pageId="11" pageNumber="193">Striffler</collectorName>
</materialsCitation>
.
<materialsCitation id="15CBEC26FF94312AACB9B7B1FE56F96B" ID-GBIF-Occurrence="3335045301" collectionCode="SEM" country="Venezuela" location="Paratypes" pageId="11" pageNumber="193" specimenCount="3" specimenCount-male="3" stateProvince="Paratypes" typeStatus="paratype">
<emphasis id="97D73A69FF94312AACB9B7B1FAB0F934" bold="true" box="[1211,1341,1667,1693]" pageId="11" pageNumber="193">
<collectingRegion id="67672899FF94312AACB9B7B1FABAF934" box="[1211,1335,1667,1693]" country="Venezuela" name="Paratypes" pageId="11" pageNumber="193">
<typeStatus id="7A1858D9FF94312AACB9B7B1FABAF934" box="[1211,1335,1667,1693]" pageId="11" pageNumber="193" type="paratype">Paratypes</typeStatus>
</collectingRegion>
:
</emphasis>
<specimenCount id="B3A52DF2FF94312AAD47B7B1FA11F934" box="[1349,1436,1667,1693]" pageId="11" pageNumber="193" type="male">2 males</specimenCount>
(ss) (
<specimenCount id="B3A52DF2FF94312AA8C4B795FEA0F968" box="[198,301,1703,1729]" pageId="11" pageNumber="193" type="male">one male</specimenCount>
<collectionCode id="C3B27EBEFF94312AA936B795FEE3F968" box="[308,366,1703,1729]" pageId="11" pageNumber="193">SEM</collectionCode>
voucher)
</materialsCitation>
,
<materialsCitation id="15CBEC26FF94312AA9E7B795FB5AF968" ID-GBIF-Occurrence="3335045304" box="[485,1239,1703,1730]" collectionCode="MNRJ" country="Venezuela" location="Paratypes" pageId="11" pageNumber="193" specimenCode="MNRJ 2429" specimenCount="7" specimenCount-female="6" specimenCount-juvenile="1" stateProvince="Paratypes" typeStatus="paratype">
<specimenCount id="B3A52DF2FF94312AA9E7B795FDD9F968" box="[485,596,1703,1729]" pageId="11" pageNumber="193" type="female">6 females</specimenCount>
,
<specimenCount id="B3A52DF2FF94312AAA5DB795FD40F968" box="[607,717,1703,1729]" pageId="11" pageNumber="193" type="juvenile">1 juvenile</specimenCount>
(
<specimenCode id="F5054E00FF94312AAAD9B795FCE6F968" box="[731,875,1703,1729]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="11" pageNumber="193">MNRJ 2429</specimenCode>
), with same data as for holotype
</materialsCitation>
.
<materialsCitation id="15CBEC26FF94312AACE2B795FCC5F94C" ID-GBIF-Occurrence="3335045302" collectionCode="SEM" country="Venezuela" location="Paratypes" pageId="11" pageNumber="193" specimenCount="1" specimenCount-male="1" stateProvince="Paratypes" typeStatus="paratype">
<specimenCount id="B3A52DF2FF94312AACE2B795FAA6F968" box="[1248,1323,1703,1729]" pageId="11" pageNumber="193" type="male">1 male</specimenCount>
(ss) (
<collectionCode id="C3B27EBEFF94312AAD63B795FA11F968" box="[1377,1436,1703,1729]" pageId="11" pageNumber="193">SEM</collectionCode>
voucher) (MACN-Ar 41020), with same data as for holotype
</materialsCitation>
.
<specimenCount id="B3A52DF2FF94312AAB50B7F9FC10F94C" box="[850,925,1739,1765]" pageId="11" pageNumber="193" type="male">1 male</specimenCount>
(ss),
<materialsCitation id="15CBEC26FF94312AABD6B7F9FE7FF8A0" ID-GBIF-Occurrence="3335045307" collectionCode="SEM" country="Venezuela" location="Paratypes" pageId="11" pageNumber="193" specimenCount="2" specimenCount-female="1" specimenCount-juvenile="1" stateProvince="Paratypes" typeStatus="paratype">
<specimenCount id="B3A52DF2FF94312AABD6B7F9FBB5F94C" box="[980,1080,1739,1765]" pageId="11" pageNumber="193" type="female">1 female</specimenCount>
,
<specimenCount id="B3A52DF2FF94312AAC46B7F9FB3EF94C" box="[1092,1203,1739,1765]" pageId="11" pageNumber="193" type="juvenile">1 juvenile</specimenCount>
(MACN-Ar 41019), with same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EDB9B5F0FF94312AA8C5B621FEBDF8DF" pageId="11" pageNumber="193" type="etymology">
<paragraph id="A51CE67BFF94312AA8C5B621FEBDF8DF" blockId="11.[151,1437,1631,2018]" pageId="11" pageNumber="193">
<emphasis id="97D73A69FF94312AA8C5B621FEC3F884" bold="true" box="[199,334,1811,1837]" pageId="11" pageNumber="193">Etymology.</emphasis>
Patronymic in honor of the Austrian arachnologist and friend Bernhard A. Huber in recognition of his remarkable contribution to arachnology, and also for being one of the collectors of the
<typeStatus id="7A1858D9FF94312AAC90B60AFB4FF8FB" box="[1170,1218,1848,1874]" pageId="11" pageNumber="193">type</typeStatus>
series; name in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="EDB9B5F0FF94312DA8C5B64DFDF2FE5C" lastPageId="12" lastPageNumber="194" pageId="11" pageNumber="193" type="diagnosis">
<paragraph id="A51CE67BFF94312DA8C5B64DFDF2FE5C" blockId="11.[151,1437,1631,2018]" lastBlockId="12.[151,1437,151,933]" lastPageId="12" lastPageNumber="194" pageId="11" pageNumber="193">
<emphasis id="97D73A69FF94312AA8C5B64DFECFF830" bold="true" box="[199,322,1919,1945]" pageId="11" pageNumber="193">Diagnosis.</emphasis>
Differs from all other species of
<taxonomicName id="62A39DF8FF94312AAABAB6B2FCC7F833" authorityName="Simon" authorityYear="1879" box="[696,842,1920,1946]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="193" phylum="Arthropoda" rank="genus">
<emphasis id="97D73A69FF94312AAABAB6B2FCC7F833" box="[696,842,1920,1946]" italics="true" pageId="11" pageNumber="193">Stenostygnus</emphasis>
</taxonomicName>
by the presence of granules in the interocular region (
<figureCitation id="3D98FAFEFF94312AA8A2B691FF70F814" box="[160,253,1955,1981]" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="11" pageNumber="193">Fig. 8 A</figureCitation>
); patella IV of males with strong spiniform apophysis on the distal-retrolateral surface (Figs 10 A–B); ventral plate of the penis with a deep cleft basally fused and apically “V” shaped (
<figureCitation id="3D98FAFEFF94312AABFCB6F5FBC2F848" box="[1022,1103,1991,2017]" captionStart="FIGURES 11" captionStartId="16.[151,264,1327,1351]" captionTargetBox="[158,1429,181,1303]" captionTargetId="figure-17@16.[158,1429,181,1304]" captionTargetPageId="16" captionText="FIGURES 11A–G. Stenostygnus huberi spec. nov., penis (A–B male (ss) paratype, MNRJ 2429; C male (ss), paratype, MACN 41020; D–G holotype male (α), MIZA 0103461). A, D Dorsal view. B, E, G Lateral view (G with detail of stylus and conduc- tors). C, F Ventral view. LA—lamina apicalis. Ti—Titillator. Scale bars: 100 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926915" httpUri="https://zenodo.org/record/4926915/files/figure.png" pageId="11" pageNumber="193">Fig. 11</figureCitation>
). It can be distinguished from
<taxonomicName id="62A39DF8FF93312DA895B1A5FEE2FF19" authorityName="Simon" authorityYear="1879" box="[151,367,151,177]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="194" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="97D73A69FF93312DA895B1A5FEE2FF19" box="[151,367,151,177]" italics="true" pageId="12" pageNumber="194">Stenostygnus pusio</emphasis>
</taxonomicName>
by the lateral borders of the scutum almost parallel and mesotergal areas almost equally wide (
<figureCitation id="3D98FAFEFF93312DA8A2B189FF70FF7C" box="[160,253,187,213]" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="12" pageNumber="194">Fig. 8 A</figureCitation>
)
<emphasis id="97D73A69FF93312DA90EB189FED9FF7D" box="[268,340,187,212]" italics="true" pageId="12" pageNumber="194">versus</emphasis>
scutum lateral borders slightly arched (giving an oval appearance to the habitus) (
<bibRefCitation id="C1329B8AFF93312DACE8B189FF5FFF50" author="Pinto-da-Rocha, R." pageId="12" pageNumber="194" pagination="194 - 198" refId="ref8901" refString="Pinto-da-Rocha, R. (1995) Redescription of Stenostygnus pusio Simon and synonymy of Caribbiantinae with Stenostygninae (Opiliones, Laniatores, Biantidae). The Journal of Arachnology, 23, 194 - 198." type="journal article" year="1995">Pinto-da-Rocha 1995</bibRefCitation>
, fig. 1); presence on the pedipalp of a meso-medial spine on the femur and a meso-distal spine on the patella (
<figureCitation id="3D98FAFEFF93312DA8A2B031FE8CFEB4" box="[160,257,259,285]" captionStart="FIGURES 9" captionStartId="12.[151,264,1864,1888]" captionTargetBox="[151,1436,957,1841]" captionTargetId="figure-420@12.[151,1436,957,1841]" captionTargetPageId="12" captionText="FIGURES 9A–E. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429). A–C Left chelicera (A ectal, B frontal, C detail of fingers), D–E Left pedipalp (D ectal, E detail of femur). Scale bars: A–C= 200 µm; D= 1 mm; E= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926911" httpUri="https://zenodo.org/record/4926911/files/figure.png" pageId="12" pageNumber="194">Fig. 9 D</figureCitation>
)
<emphasis id="97D73A69FF93312DA911B031FED6FEB5" box="[275,347,259,284]" italics="true" pageId="12" pageNumber="194">versus</emphasis>
without spines in
<taxonomicName id="62A39DF8FF93312DAA2DB031FD07FEB5" authorityName="Simon" authorityYear="1879" box="[559,650,259,285]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="194" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="97D73A69FF93312DAA2DB031FD07FEB5" box="[559,650,259,285]" italics="true" pageId="12" pageNumber="194">S. pusio</emphasis>
</taxonomicName>
;
<emphasis id="97D73A69FF93312DAA98B031FCA8FEB4" box="[666,805,259,285]" italics="true" pageId="12" pageNumber="194">pars distalis</emphasis>
ventrally convex
<emphasis id="97D73A69FF93312DABF6B031FBB1FEB5" box="[1012,1084,259,284]" italics="true" pageId="12" pageNumber="194">versus</emphasis>
concave in
<taxonomicName id="62A39DF8FF93312DACC5B031FAAFFEB5" authorityName="Simon" authorityYear="1879" box="[1223,1314,259,285]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="194" phylum="Arthropoda" rank="species" species="pusio">
<emphasis id="97D73A69FF93312DACC5B031FAAFFEB5" box="[1223,1314,259,285]" italics="true" pageId="12" pageNumber="194">S. pusio</emphasis>
</taxonomicName>
(Pinto-da- Rocha 1995, fig. 7). This species can be easily separated from
<taxonomicName id="62A39DF8FF93312DAB5DB015FC53FEE9" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[863,990,295,321]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="194" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
<emphasis id="97D73A69FF93312DAB5DB015FC53FEE9" box="[863,990,295,321]" italics="true" pageId="12" pageNumber="194">S. martensi</emphasis>
</taxonomicName>
<emphasis id="97D73A69FF93312DABE4B015FBD4FEE8" bold="true" box="[998,1113,295,321]" pageId="12" pageNumber="194">
<taxonomicNameLabel id="8CE48712FF93312DABE4B015FBD4FEE8" box="[998,1113,295,321]" pageId="12" pageNumber="194" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
by the presence of scattered granules on the interocular area and mesotergal areas I to IV (
<figureCitation id="3D98FAFEFF93312DAB3CB079FC17FECC" box="[830,922,331,357]" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="12" pageNumber="194">Fig. 8 A</figureCitation>
)
<emphasis id="97D73A69FF93312DABA5B079FC62FECD" box="[935,1007,331,356]" italics="true" pageId="12" pageNumber="194">versus</emphasis>
smooth areas in
<taxonomicName id="62A39DF8FF93312DACAAB079FAA8FECD" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[1192,1317,331,357]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="194" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
<emphasis id="97D73A69FF93312DACAAB079FAA8FECD" box="[1192,1317,331,357]" italics="true" pageId="12" pageNumber="194">S. martensi</emphasis>
</taxonomicName>
<emphasis id="97D73A69FF93312DAD29B079FA11FECC" bold="true" box="[1323,1436,331,357]" pageId="12" pageNumber="194">
<taxonomicNameLabel id="8CE48712FF93312DAD29B079FA11FECC" box="[1323,1436,331,357]" pageId="12" pageNumber="194" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="3D98FAFEFF93312DA8A2B05DFE8CFE20" box="[160,257,367,393]" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="12" pageNumber="194">Fig. 2 A</figureCitation>
); presence of granules on the coxae III and IV (
<figureCitation id="3D98FAFEFF93312DAB19B05DFCF4FE20" box="[795,889,367,393]" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="12" pageNumber="194">Fig. 8 B</figureCitation>
)
<emphasis id="97D73A69FF93312DAB88B05DFC5FFE21" box="[906,978,367,392]" italics="true" pageId="12" pageNumber="194">versus</emphasis>
smooth in
<taxonomicName id="62A39DF8FF93312DAC52B05DFB42FE21" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[1104,1231,367,393]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="194" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
<emphasis id="97D73A69FF93312DAC52B05DFB42FE21" box="[1104,1231,367,393]" italics="true" pageId="12" pageNumber="194">S. martensi</emphasis>
</taxonomicName>
<emphasis id="97D73A69FF93312DACD4B05DFAC4FE20" bold="true" box="[1238,1353,367,393]" pageId="12" pageNumber="194">
<taxonomicNameLabel id="8CE48712FF93312DACD4B05DFAC4FE20" box="[1238,1353,367,393]" pageId="12" pageNumber="194" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="3D98FAFEFF93312DAD5BB05DFF21FE04" captionStart="FIGURES 2" captionStartId="3.[151,264,1399,1423]" captionTargetBox="[151,1436,181,1375]" captionTargetId="figure-19@3.[151,1436,181,1375]" captionTargetPageId="3" captionText="FIGURES 2A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A, B, E= 500µm; C= 200 µm; D= 50 µm; F= 100µm." figureDoi="http://doi.org/10.5281/zenodo.4926893" httpUri="https://zenodo.org/record/4926893/files/figure.png" pageId="12" pageNumber="194">Fig. 2 B</figureCitation>
); femur without ectal spines (
<figureCitation id="3D98FAFEFF93312DA9FEB0A1FDD5FE04" box="[508,600,403,429]" captionStart="FIGURES 9" captionStartId="12.[151,264,1864,1888]" captionTargetBox="[151,1436,957,1841]" captionTargetId="figure-420@12.[151,1436,957,1841]" captionTargetPageId="12" captionText="FIGURES 9A–E. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429). A–C Left chelicera (A ectal, B frontal, C detail of fingers), D–E Left pedipalp (D ectal, E detail of femur). Scale bars: A–C= 200 µm; D= 1 mm; E= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926911" httpUri="https://zenodo.org/record/4926911/files/figure.png" pageId="12" pageNumber="194">Fig. 9 E</figureCitation>
)
<emphasis id="97D73A69FF93312DAA6AB0A1FD3DFE05" box="[616,688,403,428]" italics="true" pageId="12" pageNumber="194">versus</emphasis>
two small ectal spines in
<taxonomicName id="62A39DF8FF93312DABD6B0A1FBDFFE05" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[980,1106,403,429]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="194" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
<emphasis id="97D73A69FF93312DABD6B0A1FBDFFE05" box="[980,1106,403,429]" italics="true" pageId="12" pageNumber="194">S. martensi</emphasis>
</taxonomicName>
<emphasis id="97D73A69FF93312DAC5BB0A1FB41FE04" bold="true" box="[1113,1228,403,429]" pageId="12" pageNumber="194">
<taxonomicNameLabel id="8CE48712FF93312DAC5BB0A1FB41FE04" box="[1113,1228,403,429]" pageId="12" pageNumber="194" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="3D98FAFEFF93312DACDEB0A1FAB4FE04" box="[1244,1337,403,429]" captionStart="FIGURES 3" captionStartId="4.[151,264,1624,1648]" captionTargetBox="[181,1407,561,1601]" captionTargetId="figure-240@4.[181,1407,561,1601]" captionTargetPageId="4" captionText="FIGURES 3A–F. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692). A–C Right chelicera (A ectal, B frontal, C detail of fingers). D–F Right pedipalp (D mesal, E detail of tibia and tarsus, F detail of femur). Scale bars: A, B, E= 200 µm; C, F= 100 µm; D= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926895" httpUri="https://zenodo.org/record/4926895/files/figure.png" pageId="12" pageNumber="194">Fig. 3 F</figureCitation>
);
<emphasis id="97D73A69FF93312DAD4CB0A1FF7DFE78" italics="true" pageId="12" pageNumber="194">lamina apicalis</emphasis>
medially divided by a cleft that creates a “V” shape (
<figureCitation id="3D98FAFEFF93312DAB45B085FC3EFE78" box="[839,947,439,465]" captionStart="FIGURES 11" captionStartId="16.[151,264,1327,1351]" captionTargetBox="[158,1429,181,1303]" captionTargetId="figure-17@16.[158,1429,181,1304]" captionTargetPageId="16" captionText="FIGURES 11A–G. Stenostygnus huberi spec. nov., penis (A–B male (ss) paratype, MNRJ 2429; C male (ss), paratype, MACN 41020; D–G holotype male (α), MIZA 0103461). A, D Dorsal view. B, E, G Lateral view (G with detail of stylus and conduc- tors). C, F Ventral view. LA—lamina apicalis. Ti—Titillator. Scale bars: 100 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926915" httpUri="https://zenodo.org/record/4926915/files/figure.png" pageId="12" pageNumber="194">Fig. 11 A</figureCitation>
)
<emphasis id="97D73A69FF93312DABC3B085FB84FE79" box="[961,1033,439,464]" italics="true" pageId="12" pageNumber="194">versus</emphasis>
divided into two halves medially in contact in
<taxonomicName id="62A39DF8FF93312DA90EB0E9FE07FE5D" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[268,394,475,501]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="194" phylum="Arthropoda" rank="species" species="martensi" status="sp. nov.">
<emphasis id="97D73A69FF93312DA90EB0E9FE07FE5D" box="[268,394,475,501]" italics="true" pageId="12" pageNumber="194">S. martensi</emphasis>
</taxonomicName>
<emphasis id="97D73A69FF93312DA993B0E9FD8EFE5C" bold="true" box="[401,515,475,501]" pageId="12" pageNumber="194">
<taxonomicNameLabel id="8CE48712FF93312DA993B0E9FD8EFE5C" box="[401,515,475,501]" pageId="12" pageNumber="194" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="3D98FAFEFF93312DAA10B0E9FDFFFE5C" box="[530,626,474,501]" captionStart="FIGURES 5" captionStartId="8.[151,264,1283,1307]" captionTargetBox="[148,1440,181,1259]" captionTargetId="figure-17@8.[148,1440,181,1259]" captionTargetPageId="8" captionText="FIGURES 5A–G. Stenostygnus martensi spec. nov., male (paratype, MNRJ 2692), penis (A, E dorsal, B, F lateral, C, G ventral, D detail of conductor and stylus). CO—conductor. LA—lamina apicalis. SO—stylus opening of ductus ejaculatorius. ST—stylus. Ti—Titillator. Scale bars:A= 50 µm; B, C, E-G= 100 µm; D= 20 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926899" httpUri="https://zenodo.org/record/4926899/files/figure.png" pageId="12" pageNumber="194">Fig. 5 C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="EDB9B5F0FF93312FA8C5B0CCFCE8FC96" lastPageId="14" lastPageNumber="196" pageId="12" pageNumber="194" type="description">
<paragraph id="A51CE67BFF93312DA8C5B0CCFCE1FCBC" blockId="12.[151,1437,151,933]" pageId="12" pageNumber="194">
<emphasis id="97D73A69FF93312DA8C5B0CCFE11FDB0" bold="true" box="[199,412,510,537]" pageId="12" pageNumber="194">Description. Male</emphasis>
(
<typeStatus id="7A1858D9FF93312DA9ABB0CDFD95FDB3" box="[425,536,511,538]" pageId="12" pageNumber="194" type="holotype">Holotype</typeStatus>
): measurements in
<tableCitation id="E821D3C0FF93312DAAE9B0CDFCCDFDB0" box="[747,832,511,537]" captionStart="TABLE 2" captionStartId="14.[151,239,913,937]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in mm) of the type series of Stenostygnus huberi spec. nov. Holotype in bold; TBL—total body length, CL—carapace length, MSL—mesotergal scutum length, CMW—carapace maximum width, MSMW—mesotergal scutum maximum width, Tr—Trochanter, Fe—Femur, Pa—Patella, Ti—Tibia, Ta—Tarsus, T—Total." httpUri="http://table.plazi.org/id/F1DCB6F3FF91312FA895B2A3FAA0FC58" pageId="12" pageNumber="194" tableUuid="F1DCB6F3FF91312FA895B2A3FAA0FC58">Table 2</tableCitation>
. Dorsum: Habitus slightly oval (almost rectangular) in shape (
<figureCitation id="3D98FAFEFF93312DA8E4B311FEC6FD94" box="[230,331,547,573]" captionStart="FIGURES 7" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[175,1413,181,1919]" captionTargetId="figure-17@10.[175,1413,181,1919]" captionTargetPageId="10" captionText="FIGURES 7A–C. Stenostygnus huberi spec. nov., male (α) (holotype, MIZA 0103461), habitus (A dorsal, B ventral, C lateral). Scale bars: A, B= 1 mm, C= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926907" httpUri="https://zenodo.org/record/4926907/files/figure.png" pageId="12" pageNumber="194">Figs 7 A</figureCitation>
;
<figureCitation id="3D98FAFEFF93312DA955B311FE09FD94" box="[343,388,547,573]" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="12" pageNumber="194">8 A</figureCitation>
). Carapace wider than long with the anterior border straight and unarmed. Cheliceral sockets not marked. Eyes widely separated; eye mounds very low, near sulcus I; interocular area with scattered granules, with greater concentration on the eye mounds (
<figureCitation id="3D98FAFEFF93312DAAA6B359FC8AFD2C" box="[676,775,619,645]" captionStart="FIGURES 7" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[175,1413,181,1919]" captionTargetId="figure-17@10.[175,1413,181,1919]" captionTargetPageId="10" captionText="FIGURES 7A–C. Stenostygnus huberi spec. nov., male (α) (holotype, MIZA 0103461), habitus (A dorsal, B ventral, C lateral). Scale bars: A, B= 1 mm, C= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926907" httpUri="https://zenodo.org/record/4926907/files/figure.png" pageId="12" pageNumber="194">Figs 7 C</figureCitation>
;
<figureCitation id="3D98FAFEFF93312DAB10B359FCB0FD2C" box="[786,829,619,645]" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="12" pageNumber="194">8 A</figureCitation>
). Carapace in lateral view slightly convex (
<figureCitation id="3D98FAFEFF93312DAD22B359FA0EFD2C" box="[1312,1411,619,645]" captionStart="FIGURES 7" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[175,1413,181,1919]" captionTargetId="figure-17@10.[175,1413,181,1919]" captionTargetPageId="10" captionText="FIGURES 7A–C. Stenostygnus huberi spec. nov., male (α) (holotype, MIZA 0103461), habitus (A dorsal, B ventral, C lateral). Scale bars: A, B= 1 mm, C= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926907" httpUri="https://zenodo.org/record/4926907/files/figure.png" pageId="12" pageNumber="194">Figs 7 C</figureCitation>
;
<figureCitation id="3D98FAFEFF93312DAD8CB359FF21FD00" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="12" pageNumber="194">8 E</figureCitation>
). Mesotergum in lateral view slightly convex (
<figureCitation id="3D98FAFEFF93312DAAB8B3BDFC95FD00" box="[698,792,655,681]" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="12" pageNumber="194">Fig. 8 E</figureCitation>
), divided into five well defined areas (
<emphasis id="97D73A69FF93312DACCFB3BDFB62FD01" box="[1229,1263,655,680]" italics="true" pageId="12" pageNumber="194">i.e.</emphasis>
with sulci well marked). Mesotergal areas I to IV with scattered granules. Lateral margins and posterior border of scutum with a row of small granules (
<figureCitation id="3D98FAFEFF93312DA99DB3E5FD8FFD58" box="[415,514,727,753]" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="12" pageNumber="194">Fig. 8 A</figureCitation>
). Ozopore with an oval and narrow orifice with a descending channel toward the posterior region (
<figureCitation id="3D98FAFEFF93312DA95BB3C9FE38FCBC" box="[345,437,763,789]" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="12" pageNumber="194">Fig. 8 F</figureCitation>
). Free tergites granulated (
<figureCitation id="3D98FAFEFF93312DAAE2B3C9FCD2FCBC" box="[736,863,763,789]" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="12" pageNumber="194">Fig. 8 A, E</figureCitation>
).
</paragraph>
<paragraph id="A51CE67BFF93312DA8C5B22DFB10FCF4" blockId="12.[151,1437,151,933]" pageId="12" pageNumber="194">
Venter: Without any conspicuous armature (
<figureCitation id="3D98FAFEFF93312DAAB0B22DFCBDFC90" box="[690,816,799,825]" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="12" pageNumber="194">Fig. 8 B, C</figureCitation>
). Spiracles visible (
<figureCitation id="3D98FAFEFF93312DAC09B22DFB06FC90" box="[1035,1163,799,825]" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="12" pageNumber="194">Fig. 8 B, D</figureCitation>
). Coxa I to III with minutes granules. Coxa II with tubercles in the medial region. Coxa IV smooth. (
<figureCitation id="3D98FAFEFF93312DABF4B271FBD4FCF4" box="[1014,1113,835,861]" captionStart="FIGURES 7" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[175,1413,181,1919]" captionTargetId="figure-17@10.[175,1413,181,1919]" captionTargetPageId="10" captionText="FIGURES 7A–C. Stenostygnus huberi spec. nov., male (α) (holotype, MIZA 0103461), habitus (A dorsal, B ventral, C lateral). Scale bars: A, B= 1 mm, C= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926907" httpUri="https://zenodo.org/record/4926907/files/figure.png" pageId="12" pageNumber="194">Figs 7 B</figureCitation>
;
<figureCitation id="3D98FAFEFF93312DAC67B271FB02FCF4" box="[1125,1167,835,861]" captionStart="FIGURES 8" captionStartId="11.[151,264,1351,1375]" captionTargetBox="[151,1436,181,1328]" captionTargetId="figure-19@11.[151,1436,181,1328]" captionTargetPageId="11" captionText="FIGURES 8A–F. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429), habitus (A dorsal, B ventral, C detail of sternum, D detail of spiracle, E lateral, F detail of ozopore). Scale bars: A = 1 mm; B, E= 500 µm; C, F= 100 µm; D= 20 µm." figureDoi="http://doi.org/10.5281/zenodo.4926909" httpUri="https://zenodo.org/record/4926909/files/figure.png" pageId="12" pageNumber="194">8 B</figureCitation>
).
</paragraph>
<paragraph id="A51CE67BFF93312DA8C5B255FCD5FC0C" blockId="12.[151,1437,151,933]" pageId="12" pageNumber="194">
Chelicerae: Without remarkable armature. Basichelicerite with a short bulla. Cheliceral hand with sparse setae (
<figureCitation id="3D98FAFEFF93312DA8A2B2B9FF72FC0C" box="[160,255,907,933]" captionStart="FIGURES 9" captionStartId="12.[151,264,1864,1888]" captionTargetBox="[151,1436,957,1841]" captionTargetId="figure-420@12.[151,1436,957,1841]" captionTargetPageId="12" captionText="FIGURES 9A–E. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429). A–C Left chelicera (A ectal, B frontal, C detail of fingers), D–E Left pedipalp (D ectal, E detail of femur). Scale bars: A–C= 200 µm; D= 1 mm; E= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926911" httpUri="https://zenodo.org/record/4926911/files/figure.png" pageId="12" pageNumber="194">Fig. 9 A</figureCitation>
). Fingers thin, with uniform small teeth (
<figureCitation id="3D98FAFEFF93312DAAC9B2B9FCC6FC0C" box="[715,843,907,933]" captionStart="FIGURES 9" captionStartId="12.[151,264,1864,1888]" captionTargetBox="[151,1436,957,1841]" captionTargetId="figure-420@12.[151,1436,957,1841]" captionTargetPageId="12" captionText="FIGURES 9A–E. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429). A–C Left chelicera (A ectal, B frontal, C detail of fingers), D–E Left pedipalp (D ectal, E detail of femur). Scale bars: A–C= 200 µm; D= 1 mm; E= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926911" httpUri="https://zenodo.org/record/4926911/files/figure.png" pageId="12" pageNumber="194">Fig. 9 B, C</figureCitation>
).
</paragraph>
<caption id="F1DCB6F3FF93312DA895B67AFAA6F82C" ID-DOI="http://doi.org/10.5281/zenodo.4926911" ID-Zenodo-Dep="4926911" httpUri="https://zenodo.org/record/4926911/files/figure.png" pageId="12" pageNumber="194" startId="12.[151,264,1864,1888]" targetBox="[151,1436,957,1841]" targetPageId="12">
<paragraph id="A51CE67BFF93312DA895B67AFAA6F82C" blockId="12.[151,1436,1864,1925]" pageId="12" pageNumber="194">
<emphasis id="97D73A69FF93312DA895B67AFEDFF8C9" bold="true" box="[151,338,1864,1888]" pageId="12" pageNumber="194">FIGURES 9A–E.</emphasis>
<taxonomicName id="62A39DF8FF93312DA95AB67AFDA8F8C9" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[344,549,1864,1888]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="194" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov.">
<emphasis id="97D73A69FF93312DA95AB67AFDA8F8C9" box="[344,549,1864,1888]" italics="true" pageId="12" pageNumber="194">Stenostygnus huberi</emphasis>
</taxonomicName>
<emphasis id="97D73A69FF93312DAA2EB67AFD19F8C9" bold="true" box="[556,660,1864,1888]" pageId="12" pageNumber="194">
<taxonomicNameLabel id="8CE48712FF93312DAA2EB67AFD19F8C9" box="[556,660,1864,1888]" pageId="12" pageNumber="194" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, male (ss) (paratype, MNRJ 2429). A–C Left chelicera (A ectal, B frontal, C detail of fingers), D–E Left pedipalp (D ectal, E detail of femur). Scale bars: A–C= 200 µm; D= 1 mm; E= 500 µm.
</paragraph>
</caption>
<paragraph id="A51CE67BFF93312CA8C5B680FA11FF50" blockId="12.[151,1437,1970,2033]" lastBlockId="13.[151,1436,151,465]" lastPageId="13" lastPageNumber="195" pageId="12" pageNumber="194">
Pedipalps: Raptorial and long, with spines concentrated on tibia and tarsus (
<figureCitation id="3D98FAFEFF93312DAC10B680FBFFF864" box="[1042,1138,1970,1997]" captionStart="FIGURES 9" captionStartId="12.[151,264,1864,1888]" captionTargetBox="[151,1436,957,1841]" captionTargetId="figure-420@12.[151,1436,957,1841]" captionTargetPageId="12" captionText="FIGURES 9A–E. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429). A–C Left chelicera (A ectal, B frontal, C detail of fingers), D–E Left pedipalp (D ectal, E detail of femur). Scale bars: A–C= 200 µm; D= 1 mm; E= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926911" httpUri="https://zenodo.org/record/4926911/files/figure.png" pageId="12" pageNumber="194">Fig. 9 D</figureCitation>
). Coxa elongated, surpassing the anterior border of the carapace (
<emphasis id="97D73A69FF93312DAA51B6E5FDF8F859" box="[595,629,2007,2032]" italics="true" pageId="12" pageNumber="194">i.e.</emphasis>
can be seen in dorsal view) (
<figureCitation id="3D98FAFEFF93312DABB9B6E4FB94F859" box="[955,1049,2006,2033]" captionStart="FIGURES 7" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[175,1413,181,1919]" captionTargetId="figure-17@10.[175,1413,181,1919]" captionTargetPageId="10" captionText="FIGURES 7A–C. Stenostygnus huberi spec. nov., male (α) (holotype, MIZA 0103461), habitus (A dorsal, B ventral, C lateral). Scale bars: A, B= 1 mm, C= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926907" httpUri="https://zenodo.org/record/4926907/files/figure.png" pageId="12" pageNumber="194">Fig. 7 A</figureCitation>
) proximally with one small dorsoectal protuberance; trochanter small and smooth; femur thin and cylindrical, with a meso-medial spine (
<figureCitation id="3D98FAFEFF92312CAD2DB1A5FA03FF18" box="[1327,1422,151,177]" captionStart="FIGURES 9" captionStartId="12.[151,264,1864,1888]" captionTargetBox="[151,1436,957,1841]" captionTargetId="figure-420@12.[151,1436,957,1841]" captionTargetPageId="12" captionText="FIGURES 9A–E. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429). A–C Left chelicera (A ectal, B frontal, C detail of fingers), D–E Left pedipalp (D ectal, E detail of femur). Scale bars: A–C= 200 µm; D= 1 mm; E= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926911" httpUri="https://zenodo.org/record/4926911/files/figure.png" pageId="13" pageNumber="195">Fig. 9 E</figureCitation>
); patella clavate, swollen distally, with a meso-distal spine (
<figureCitation id="3D98FAFEFF92312CAB2CB189FC1FFF7C" box="[814,914,187,213]" captionStart="FIGURES 9" captionStartId="12.[151,264,1864,1888]" captionTargetBox="[151,1436,957,1841]" captionTargetId="figure-420@12.[151,1436,957,1841]" captionTargetPageId="12" captionText="FIGURES 9A–E. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429). A–C Left chelicera (A ectal, B frontal, C detail of fingers), D–E Left pedipalp (D ectal, E detail of femur). Scale bars: A–C= 200 µm; D= 1 mm; E= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926911" httpUri="https://zenodo.org/record/4926911/files/figure.png" pageId="13" pageNumber="195">Fig. 9 D</figureCitation>
); tibia with one ventro-basal spine and armed ventrally with three ectal and three mesal spines; tarsus mesal and ectally with three spines on each side (
<figureCitation id="3D98FAFEFF92312CAD33B1EDFA02FF50" box="[1329,1423,223,249]" captionStart="FIGURES 9" captionStartId="12.[151,264,1864,1888]" captionTargetBox="[151,1436,957,1841]" captionTargetId="figure-420@12.[151,1436,957,1841]" captionTargetPageId="12" captionText="FIGURES 9A–E. Stenostygnus huberi spec. nov., male (ss) (paratype, MNRJ 2429). A–C Left chelicera (A ectal, B frontal, C detail of fingers), D–E Left pedipalp (D ectal, E detail of femur). Scale bars: A–C= 200 µm; D= 1 mm; E= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926911" httpUri="https://zenodo.org/record/4926911/files/figure.png" pageId="13" pageNumber="195">Fig. 9 E</figureCitation>
).
</paragraph>
<paragraph id="A51CE67BFF92312CA8C5B031FB23FE23" blockId="13.[151,1436,151,465]" pageId="13" pageNumber="195">Legs: Thin, cylindrical, and elongated (Fig. 10 A). Coxa IV without remarkable armature; femur IV slightly curved; patella I–III short and swollen; patella IV with a prolateral-distal spiniform apophyses and dorsally roughly granulated (some of the granules looks like small pointed tip tubercles) (Fig. 10 B); tibia IV slightly swollen distally (Fig. 10 B); tarsi III and IV with dense scopula (Fig. 10 C, D). Tarsal formula: 6(3):11(3):6:6.</paragraph>
<paragraph id="A51CE67BFF92312CA8C5B0A1FF7FFE78" blockId="13.[151,1436,151,465]" pageId="13" pageNumber="195">
Color (specimen preserved in 80% ethanol): Body yellowish. Mesotergal areas, with slight brown reticulations (
<figureCitation id="3D98FAFEFF92312CA8A2B085FF69FE78" box="[160,228,439,465]" captionStart="FIGURES 7" captionStartId="10.[151,264,1943,1967]" captionTargetBox="[175,1413,181,1919]" captionTargetId="figure-17@10.[175,1413,181,1919]" captionTargetPageId="10" captionText="FIGURES 7A–C. Stenostygnus huberi spec. nov., male (α) (holotype, MIZA 0103461), habitus (A dorsal, B ventral, C lateral). Scale bars: A, B= 1 mm, C= 500 µm." figureDoi="http://doi.org/10.5281/zenodo.4926907" httpUri="https://zenodo.org/record/4926907/files/figure.png" pageId="13" pageNumber="195">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="A51CE67BFF92312CA895B79BFB39F94F" blockId="13.[151,1437,1705,1766]" pageId="13" pageNumber="195">
<emphasis id="97D73A69FF92312CA895B79BFE88F968" bold="true" box="[151,261,1705,1729]" pageId="13" pageNumber="195">Figure 10.</emphasis>
<taxonomicName id="62A39DF8FF92312CA90EB798FE54F968" authority="Mamani &amp; Porto &amp; Iglesias &amp; González, 2021" authorityName="Mamani &amp; Porto &amp; Iglesias &amp; González" authorityYear="2021" box="[268,473,1705,1729]" class="Arachnida" family="Biantidae" genus="Stenostygnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="13" pageNumber="195" phylum="Arthropoda" rank="species" species="huberi" status="sp. nov.">
<emphasis id="97D73A69FF92312CA90EB798FE54F968" box="[268,473,1705,1729]" italics="true" pageId="13" pageNumber="195">Stenostygnus huberi</emphasis>
</taxonomicName>
<emphasis id="97D73A69FF92312CA9DDB798FDCAF96B" bold="true" box="[479,583,1706,1730]" pageId="13" pageNumber="195">
<taxonomicNameLabel id="8CE48712FF92312CA9DDB798FDCAF96B" box="[479,583,1706,1730]" pageId="13" pageNumber="195" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, male (ss) (
<typeStatus id="7A1858D9FF92312CAAB5B798FC9FF96B" box="[695,786,1706,1730]" pageId="13" pageNumber="195" type="paratype">paratype</typeStatus>
, MNRJ 2429), left leg IV (A retrolateral, B detail of patella and tibia, C detail of tarsus, D detail of scopula). Scale bars: A=
<quantity id="625B4B9EFF92312CAAFBB7FCFCBAF94C" box="[761,823,1742,1766]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="195" unit="mm" value="2.0">2 mm</quantity>
; B=
<quantity id="625B4B9EFF92312CAB65B7FFFC35F94F" box="[871,952,1741,1766]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="195" unit="mm" value="500.0">500 µm</quantity>
; C=
<quantity id="625B4B9EFF92312CABEAB7FCFBB7F94F" box="[1000,1082,1742,1766]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="195" unit="mm" value="200.0">200 µm</quantity>
; D=
<quantity id="625B4B9EFF92312CAC69B7FFFB3DF94F" box="[1131,1200,1741,1766]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="195" unit="mm" value="50.0">50 µm</quantity>
.
</paragraph>
<paragraph id="A51CE67BFF92312FA8C5B626FCCCFE27" blockId="13.[151,1437,1812,2018]" lastBlockId="14.[151,1437,156,867]" lastPageId="14" lastPageNumber="196" pageId="13" pageNumber="195">
Genitalia: Penis with
<emphasis id="97D73A69FF92312CA9BEB626FDC9F887" box="[444,580,1812,1838]" italics="true" pageId="13" pageNumber="195">pars basalis</emphasis>
and
<emphasis id="97D73A69FF92312CAA7FB626FD43F887" box="[637,718,1812,1838]" italics="true" pageId="13" pageNumber="195">distalis</emphasis>
well demarcated,
<emphasis id="97D73A69FF92312CAB9CB626FBA5F887" box="[926,1064,1812,1838]" italics="true" pageId="13" pageNumber="195">pars distalis</emphasis>
is markedly rounded contrasting with the thin and tubular
<emphasis id="97D73A69FF92312CA9B1B60AFDB6F8FB" box="[435,571,1848,1874]" italics="true" pageId="13" pageNumber="195">pars basalis</emphasis>
(
<figureCitation id="3D98FAFEFF92312CAA49B60AFD57F8FB" box="[587,730,1848,1874]" captionStart="FIGURES 11" captionStartId="16.[151,264,1327,1351]" captionTargetBox="[158,1429,181,1303]" captionTargetId="figure-17@16.[158,1429,181,1304]" captionTargetPageId="16" captionText="FIGURES 11A–G. Stenostygnus huberi spec. nov., penis (A–B male (ss) paratype, MNRJ 2429; C male (ss), paratype, MACN 41020; D–G holotype male (α), MIZA 0103461). A, D Dorsal view. B, E, G Lateral view (G with detail of stylus and conduc- tors). C, F Ventral view. LA—lamina apicalis. Ti—Titillator. Scale bars: 100 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926915" httpUri="https://zenodo.org/record/4926915/files/figure.png" pageId="13" pageNumber="195">Fig. 11 A, D</figureCitation>
).
<emphasis id="97D73A69FF92312CAAECB60AFCF6F8FB" box="[750,891,1848,1874]" italics="true" pageId="13" pageNumber="195">Pars distalis</emphasis>
ending in a
<emphasis id="97D73A69FF92312CAC05B60AFB38F8FB" box="[1031,1205,1848,1874]" italics="true" pageId="13" pageNumber="195">lamina apicalis</emphasis>
medially divided by a cleft with a “V” shaped (
<figureCitation id="3D98FAFEFF92312CA9C7B66EFDDCF8DF" box="[453,593,1884,1910]" captionStart="FIGURES 11" captionStartId="16.[151,264,1327,1351]" captionTargetBox="[158,1429,181,1303]" captionTargetId="figure-17@16.[158,1429,181,1304]" captionTargetPageId="16" captionText="FIGURES 11A–G. Stenostygnus huberi spec. nov., penis (A–B male (ss) paratype, MNRJ 2429; C male (ss), paratype, MACN 41020; D–G holotype male (α), MIZA 0103461). A, D Dorsal view. B, E, G Lateral view (G with detail of stylus and conduc- tors). C, F Ventral view. LA—lamina apicalis. Ti—Titillator. Scale bars: 100 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926915" httpUri="https://zenodo.org/record/4926915/files/figure.png" pageId="13" pageNumber="195">Fig. 11 C, F</figureCitation>
). Lateral border of
<emphasis id="97D73A69FF92312CAB2FB66EFC3AF8DF" box="[813,951,1884,1910]" italics="true" pageId="13" pageNumber="195">pars distalis</emphasis>
armed with 6 pairs of leaf-like macrosetae and 2 subapical pairs of very short (truncated, not leaf-like) setae (
<figureCitation id="3D98FAFEFF92312CAB89B6B2FB95F833" box="[907,1048,1920,1946]" captionStart="FIGURES 11" captionStartId="16.[151,264,1327,1351]" captionTargetBox="[158,1429,181,1303]" captionTargetId="figure-17@16.[158,1429,181,1304]" captionTargetPageId="16" captionText="FIGURES 11A–G. Stenostygnus huberi spec. nov., penis (A–B male (ss) paratype, MNRJ 2429; C male (ss), paratype, MACN 41020; D–G holotype male (α), MIZA 0103461). A, D Dorsal view. B, E, G Lateral view (G with detail of stylus and conduc- tors). C, F Ventral view. LA—lamina apicalis. Ti—Titillator. Scale bars: 100 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926915" httpUri="https://zenodo.org/record/4926915/files/figure.png" pageId="13" pageNumber="195">Fig. 11 B, F</figureCitation>
). Dorsally the capsula externa has two wide and apically rounded titillators. The apical borders of the titillators are dorsally projected. The internal surface of titillators is covered by small digitiform projections (
<figureCitation id="3D98FAFEFF92312CAB58B6FAFC68F84B" box="[858,997,1992,2018]" captionStart="FIGURES 11" captionStartId="16.[151,264,1327,1351]" captionTargetBox="[158,1429,181,1303]" captionTargetId="figure-17@16.[158,1429,181,1304]" captionTargetPageId="16" captionText="FIGURES 11A–G. Stenostygnus huberi spec. nov., penis (A–B male (ss) paratype, MNRJ 2429; C male (ss), paratype, MACN 41020; D–G holotype male (α), MIZA 0103461). A, D Dorsal view. B, E, G Lateral view (G with detail of stylus and conduc- tors). C, F Ventral view. LA—lamina apicalis. Ti—Titillator. Scale bars: 100 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926915" httpUri="https://zenodo.org/record/4926915/files/figure.png" pageId="13" pageNumber="195">Fig. 11 A, B</figureCitation>
). Capsula interna with two laminar and sharply pointed conductors bearing an also laminar and pointed stylus with a ventral subapical opening of
<emphasis id="97D73A69FF91312FAD51B1AEFEA6FF73" italics="true" pageId="14" pageNumber="196">ductus ejaculatorius</emphasis>
(
<figureCitation id="3D98FAFEFF91312FA938B1F2FE2BFF73" box="[314,422,192,218]" captionStart="FIGURES 11" captionStartId="16.[151,264,1327,1351]" captionTargetBox="[158,1429,181,1303]" captionTargetId="figure-17@16.[158,1429,181,1304]" captionTargetPageId="16" captionText="FIGURES 11A–G. Stenostygnus huberi spec. nov., penis (A–B male (ss) paratype, MNRJ 2429; C male (ss), paratype, MACN 41020; D–G holotype male (α), MIZA 0103461). A, D Dorsal view. B, E, G Lateral view (G with detail of stylus and conduc- tors). C, F Ventral view. LA—lamina apicalis. Ti—Titillator. Scale bars: 100 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926915" httpUri="https://zenodo.org/record/4926915/files/figure.png" pageId="14" pageNumber="196">Fig. 11 G</figureCitation>
). The cuticular area of the lateral macrosetae has a rugose (folding) appearance (
<figureCitation id="3D98FAFEFF91312FAD26B1F2FA03FF73" box="[1316,1422,192,218]" captionStart="FIGURES 11" captionStartId="16.[151,264,1327,1351]" captionTargetBox="[158,1429,181,1303]" captionTargetId="figure-17@16.[158,1429,181,1304]" captionTargetPageId="16" captionText="FIGURES 11A–G. Stenostygnus huberi spec. nov., penis (A–B male (ss) paratype, MNRJ 2429; C male (ss), paratype, MACN 41020; D–G holotype male (α), MIZA 0103461). A, D Dorsal view. B, E, G Lateral view (G with detail of stylus and conduc- tors). C, F Ventral view. LA—lamina apicalis. Ti—Titillator. Scale bars: 100 µm. Stylus in green, conductors in pink and capsula externa in blue (For interpretation of the references to colour the reader is referred to the pdf version of this article)." figureDoi="http://doi.org/10.5281/zenodo.4926915" httpUri="https://zenodo.org/record/4926915/files/figure.png" pageId="14" pageNumber="196">Fig. 11 B</figureCitation>
), this could be an effect of thinner or less chitinous cuticular layer in this region, that could be (hypothetically) related to the region of
<emphasis id="97D73A69FF91312FA947B03AFE40FE8B" box="[325,461,264,290]" italics="true" pageId="14" pageNumber="196">pars distalis</emphasis>
that became swollen during the turgescent stage in the hydraulic penial functioning, this inflation could be responsible for the macrosetae erection necessary for functional interaction with the ovipositor during copulation. These putative expansible areas could functionally correspond to the spongy body (‘Schwellkörper’
<emphasis id="97D73A69FF91312FA8C8B046FF6DFE24" box="[202,224,372,397]" italics="true" pageId="14" pageNumber="196">in</emphasis>
<bibRefCitation id="C1329B8AFF91312FA8E5B046FE0EFE27" author="Martens, J." box="[231,387,372,398]" pageId="14" pageNumber="196" pagination="347 - 414" refId="ref8555" refString="Martens, J. (1978) Opiliones aus dem Nepal-Himalaya. IV. Biantidae (Arachnida). Senckenbergiana biologica, 58 (5 / 6), 347 - 414." type="journal article" year="1978">Martens 1978</bibRefCitation>
) commonly observed in Asian biantids.
</paragraph>
<paragraph id="A51CE67BFF91312FA8C5B0AAFBDEFE53" blockId="14.[151,1437,156,867]" pageId="14" pageNumber="196">
Male (ss)
<typeStatus id="7A1858D9FF91312FA92CB0AAFE03FE1B" box="[302,398,408,434]" pageId="14" pageNumber="196" type="paratype">paratype</typeStatus>
(MNRJ 2429): measurements as in
<tableCitation id="E821D3C0FF91312FAB1FB0AAFCFFFE1B" box="[797,882,408,434]" captionStart="TABLE 2" captionStartId="14.[151,239,913,937]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in mm) of the type series of Stenostygnus huberi spec. nov. Holotype in bold; TBL—total body length, CL—carapace length, MSL—mesotergal scutum length, CMW—carapace maximum width, MSMW—mesotergal scutum maximum width, Tr—Trochanter, Fe—Femur, Pa—Patella, Ti—Tibia, Ta—Tarsus, T—Total." httpUri="http://table.plazi.org/id/F1DCB6F3FF91312FA895B2A3FAA0FC58" pageId="14" pageNumber="196" tableUuid="F1DCB6F3FF91312FA895B2A3FAA0FC58">Table 2</tableCitation>
. Similar to the male (α) in relation to the armature of the dorsal
<emphasis id="97D73A69FF91312FA92FB08EFEF1FE7C" box="[301,380,444,469]" italics="true" pageId="14" pageNumber="196">scutum</emphasis>
(
<figureCitation id="3D98FAFEFF91312FA98FB08EFD93FE7F" box="[397,542,444,470]" captionStart="FIGURES 12" captionStartId="17.[151,264,1807,1831]" captionTargetBox="[151,1436,181,1783]" captionTargetId="figure-19@17.[151,1436,181,1783]" captionTargetPageId="17" captionText="FIGURES 12A–F. Stenostygnus huberi spec. nov., sexual dimorphism (A–B female paratype, MNRJ 2429; C–D male (ss) paratype, MACN 41020; E–F (holotype male (α), MIZA 0103461), A, C, E Dorsal view. B, D, F Detail of patella. Scale bars: A, C, E= 2 mm; B, F= 200 µm; D= 500 µm" figureDoi="http://doi.org/10.5281/zenodo.4926917" httpUri="https://zenodo.org/record/4926917/files/figure.png" pageId="14" pageNumber="196">Fig. 12 B, C</figureCitation>
). Sexual dimorphism evident in patella IV as in the
<typeStatus id="7A1858D9FF91312FAC6CB08EFB5DFE7F" box="[1134,1232,444,470]" pageId="14" pageNumber="196" type="holotype">holotype</typeStatus>
but male (
<emphasis id="97D73A69FF91312FAD49B08EFAD4FE7C" box="[1355,1369,444,469]" italics="true" pageId="14" pageNumber="196">ss</emphasis>
) with prolateral-distal spiniform apophyses and dorsal tubercles smaller in size (
<figureCitation id="3D98FAFEFF91312FABD5B0D2FBCBFE53" box="[983,1094,480,506]" captionStart="FIGURES 12" captionStartId="17.[151,264,1807,1831]" captionTargetBox="[151,1436,181,1783]" captionTargetId="figure-19@17.[151,1436,181,1783]" captionTargetPageId="17" captionText="FIGURES 12A–F. Stenostygnus huberi spec. nov., sexual dimorphism (A–B female paratype, MNRJ 2429; C–D male (ss) paratype, MACN 41020; E–F (holotype male (α), MIZA 0103461), A, C, E Dorsal view. B, D, F Detail of patella. Scale bars: A, C, E= 2 mm; B, F= 200 µm; D= 500 µm" figureDoi="http://doi.org/10.5281/zenodo.4926917" httpUri="https://zenodo.org/record/4926917/files/figure.png" pageId="14" pageNumber="196">Fig. 12 D</figureCitation>
).
</paragraph>
<paragraph id="A51CE67BFF91312FA895B336FE97FDCF" blockId="14.[151,1437,156,867]" pageId="14" pageNumber="196">
<emphasis id="97D73A69FF91312FA895B336FF63FDB7" bold="true" box="[151,238,516,542]" pageId="14" pageNumber="196">Female</emphasis>
(
<typeStatus id="7A1858D9FF91312FA8FDB336FEE9FDB7" box="[255,356,516,542]" pageId="14" pageNumber="196" type="paratype">paratype</typeStatus>
, MNRJ 2429): measurements as in
<tableCitation id="E821D3C0FF91312FAB00B336FCD7FDB7" box="[770,858,516,542]" captionStart="TABLE 2" captionStartId="14.[151,239,913,937]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in mm) of the type series of Stenostygnus huberi spec. nov. Holotype in bold; TBL—total body length, CL—carapace length, MSL—mesotergal scutum length, CMW—carapace maximum width, MSMW—mesotergal scutum maximum width, Tr—Trochanter, Fe—Femur, Pa—Patella, Ti—Tibia, Ta—Tarsus, T—Total." httpUri="http://table.plazi.org/id/F1DCB6F3FF91312FA895B2A3FAA0FC58" pageId="14" pageNumber="196" tableUuid="F1DCB6F3FF91312FA895B2A3FAA0FC58">Table 2</tableCitation>
. Similar to male in relation to the armature of the dorsal scutum (
<figureCitation id="3D98FAFEFF91312FA940B31AFE61FDEB" box="[322,492,552,578]" captionStart="FIGURES 12" captionStartId="17.[151,264,1807,1831]" captionTargetBox="[151,1436,181,1783]" captionTargetId="figure-19@17.[151,1436,181,1783]" captionTargetPageId="17" captionText="FIGURES 12A–F. Stenostygnus huberi spec. nov., sexual dimorphism (A–B female paratype, MNRJ 2429; C–D male (ss) paratype, MACN 41020; E–F (holotype male (α), MIZA 0103461), A, C, E Dorsal view. B, D, F Detail of patella. Scale bars: A, C, E= 2 mm; B, F= 200 µm; D= 500 µm" figureDoi="http://doi.org/10.5281/zenodo.4926917" httpUri="https://zenodo.org/record/4926917/files/figure.png" pageId="14" pageNumber="196">Fig. 12 A, C, E</figureCitation>
), but without prolateral-distal spiniform apophyses and dorsal tubercles in patella IV (
<figureCitation id="3D98FAFEFF91312FA8A2B37EFE81FDCF" box="[160,268,588,614]" captionStart="FIGURES 12" captionStartId="17.[151,264,1807,1831]" captionTargetBox="[151,1436,181,1783]" captionTargetId="figure-19@17.[151,1436,181,1783]" captionTargetPageId="17" captionText="FIGURES 12A–F. Stenostygnus huberi spec. nov., sexual dimorphism (A–B female paratype, MNRJ 2429; C–D male (ss) paratype, MACN 41020; E–F (holotype male (α), MIZA 0103461), A, C, E Dorsal view. B, D, F Detail of patella. Scale bars: A, C, E= 2 mm; B, F= 200 µm; D= 500 µm" figureDoi="http://doi.org/10.5281/zenodo.4926917" httpUri="https://zenodo.org/record/4926917/files/figure.png" pageId="14" pageNumber="196">Fig. 12 B</figureCitation>
).
</paragraph>
<paragraph id="A51CE67BFF91312FA8C5B35DFCE8FC96" blockId="14.[151,1437,156,867]" pageId="14" pageNumber="196">
<emphasis id="97D73A69FF91312FA8C5B35DFEB2FD20" bold="true" box="[199,319,623,649]" pageId="14" pageNumber="196">Variation.</emphasis>
Measurements (
<tableCitation id="E821D3C0FF91312FA9FBB342FDDFFD23" box="[505,594,624,650]" captionStart="TABLE 2" captionStartId="14.[151,239,913,937]" captionTargetPageId="14" captionText="TABLE 2. Measurements (in mm) of the type series of Stenostygnus huberi spec. nov. Holotype in bold; TBL—total body length, CL—carapace length, MSL—mesotergal scutum length, CMW—carapace maximum width, MSMW—mesotergal scutum maximum width, Tr—Trochanter, Fe—Femur, Pa—Patella, Ti—Tibia, Ta—Tarsus, T—Total." httpUri="http://table.plazi.org/id/F1DCB6F3FF91312FA895B2A3FAA0FC58" pageId="14" pageNumber="196" tableUuid="F1DCB6F3FF91312FA895B2A3FAA0FC58">Table 2</tableCitation>
): Males (n=5): total body length (1.92–2.35), carapace length (0.71–0.75), mesotergal length (1.70–1.93), carapace maximum width (1.08–1.18), mesotergal
<emphasis id="97D73A69FF91312FAC38B3A6FB04FD04" box="[1082,1161,660,685]" italics="true" pageId="14" pageNumber="196">scutum</emphasis>
maximum width (1.42– 1.61), leg I (5.63–6.30), leg II (14.07–14.68), leg III (7.99–9.39), leg IV (11.61–13.77); Females (n=7): total body length (1.86–2.40), carapace length (0.64–0.73), mesotergal length (1.59–1.86), carapace maximum width (1.07– 1.10), mesotergal
<emphasis id="97D73A69FF91312FA960B232FE3CFCB0" box="[354,433,768,793]" italics="true" pageId="14" pageNumber="196">scutum</emphasis>
maximum width (1.48–1.62), leg I (4.97–5.39), leg II (11.82–13.14), leg III (6.88–7.55), leg IV (9.37–10.44). Tarsal formula (n=12): 6–7(3):9–11(3):6:6.
</paragraph>
</subSubSection>
<subSubSection id="EDB9B5F0FF91312FA8C5B275FCD8FCCA" box="[199,853,839,867]" pageId="14" pageNumber="196" type="distribution">
<paragraph id="A51CE67BFF91312FA8C5B275FCD8FCCA" blockId="14.[151,1437,156,867]" box="[199,853,839,867]" pageId="14" pageNumber="196">
<emphasis id="97D73A69FF91312FA8C5B275FEECFCC8" bold="true" box="[199,353,839,865]" pageId="14" pageNumber="196">Distribution.</emphasis>
Known only from the
<typeStatus id="7A1858D9FF91312FAA62B27AFD1DFCCB" box="[608,656,840,866]" pageId="14" pageNumber="196">type</typeStatus>
locality (
<figureCitation id="3D98FAFEFF91312FAAF9B27AFCCAFCCA" box="[763,839,840,867]" captionStart="FIGURE 13" captionStartId="18.[151,250,1113,1137]" captionTargetBox="[151,1436,181,1090]" captionTargetId="figure-17@18.[151,1436,181,1090]" captionTargetPageId="18" captionText="FIGURE 13. Geographical distribution of the genus Stenostygnus in South America. Stenostygnus pusio (circles) in Colombia, Ecuador and Brazil, Stenostygnus huberi spec. nov. (square) in Lara State, Venezuela and Stenostygnus martensi spec. nov. (triangle) in Mérida State, Venezuela." figureDoi="http://doi.org/10.5281/zenodo.4926919" httpUri="https://zenodo.org/record/4926919/files/figure.png" pageId="14" pageNumber="196">Fig.13</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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