214 lines
26 KiB
XML
214 lines
26 KiB
XML
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<mods:title id="95A207C8F601F602B133E0D290FB67FA">The genus Pterostichus in China II: the subgenus Circinatus Sciaky, a species revision and phylogeny (Carabidae, Pterostichini)</mods:title>
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<mods:namePart id="F8B717E55996C3BBE6D4CE13B355A203">Shi, Hongliang</mods:namePart>
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<mods:namePart id="9544F893EF423B3D3A5BC1B3A03A5110">Liang, Hongbin</mods:namePart>
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<treatment id="18AC6C240490228EFBAD7EDE409ADD13" ID-GBIF-Taxon="152063177" LSID="urn:lsid:plazi:treatment:18AC6C240490228EFBAD7EDE409ADD13" httpUri="http://treatment.plazi.org/id/18AC6C240490228EFBAD7EDE409ADD13" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
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<subSubSection id="AEBA17A51453009009025AD032B14F01" pageId="37" pageNumber="38" type="multiple">
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<paragraph id="64C788E78AEC9D457AEC8F7562292B1A" pageId="37" pageNumber="38">Taxon classification Animalia Coleoptera Carabidae</paragraph>
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</subSubSection>
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<subSubSection id="CAB3D85F1D5DDE63F86A0892C8B10628" pageId="37" pageNumber="38" type="nomenclature">
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<paragraph id="5D27E02AED329F1678BBD766F0A5D072" pageId="37" pageNumber="38">
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<taxonomicName id="10F1CA9442D5901F2DDC1AC1A84CFD3A" ID-CoL="4Q3HP" LSID="Chinese common name: 波纳通缘步甲 (Bō Nà Tōng Yuán Bù Jiă)" authority="Jedlicka, 1934" authorityName="Jedlicka" authorityYear="1934" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Circinatus) pohnerti" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pohnerti" subGenus="Circinatus">
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Pterostichus (Circinatus) pohnerti
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<normalizedToken id="EA692E52437AA0918120A916AD785BE8" originalValue="Jedlička">Jedlicka</normalizedToken>
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, 1934
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</taxonomicName>
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Figures 26, 47, 56, 85, 111
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</paragraph>
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</subSubSection>
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<subSubSection id="DFFE1D246428C6504B7DD641947D33BD" pageId="37" pageNumber="38" type="reference_group">
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<paragraph id="1E1B0FFD701360EE7D00F91CDCD0F961" pageId="37" pageNumber="38">
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<taxonomicName id="AA9AC419FBE9A4EF8895B4E07C821CD4" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Circinatus) pohnerti" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pohnerti" subGenus="Circinatus">Pterostichus (Circinatus) pohnerti</taxonomicName>
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<bibRefCitation id="0D5ABD0F431BD733520E61AB8BF6DCFA" author="Jedlicka, A" journalOrPublisher="Casopis Ceskoslovenske spolecnosti entomologicke" pageId="59" pageNumber="60" pagination="13 - 19" title="Carabidi z vychodni Asie. Carabiden aus Ostasien (5. Teil)." volume="30" year="1934">
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<normalizedToken id="EF2063D657DD4B3939E3EC2EC2252FED" originalValue="Jedlička">Jedlicka</normalizedToken>
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1934
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</bibRefCitation>
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: 19 (Original:
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<taxonomicName id="7BD05B7CE454483C1BA7DE4CBFDA3CD8" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">Pterostichus</taxonomicName>
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(incertae sedis); holotype deposited in NMPC);
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<bibRefCitation id="DAD46C574E136219A6571545C5010D4C" author="Jedlicka, A" journalOrPublisher="Abhandlungen und Berichte aus dem Staatlichen Museum fuer Tierkunde in Dresden" pageId="59" pageNumber="60" pagination="177 - 346" title="Monographie des Tribus Pterostichini aus Ostasien (Pterostichi, Trigonotomi, Myadi) (Coleoptera-Carabidae)." volume="26" year="1962">
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<normalizedToken id="8CCDFE467101C237C28F90DADD567952" originalValue="Jedlička">Jedlicka</normalizedToken>
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1962
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</bibRefCitation>
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: 298 (
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<taxonomicName id="296E71EA8913F16A9BE7AD70713F0F9E" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">Pterostichus</taxonomicName>
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(incertae sedis));
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<bibRefCitation id="C522EABC9900187091C816C89BFDFD13" author="Sciaky, R" journalOrPublisher="Annuario del Museo Civico di Storia Naturale di Brescia" pageId="60" pageNumber="61" pagination="217 - 231" title="Circinatus new subgenus and three new species of Pterostichus from China (ColeopteraCarabidae). Natura Bresciana." volume="30" year="1996">Sciaky 1996</bibRefCitation>
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: 221;
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<bibRefCitation id="18E688BBF664CCE574B91B2A97D0CEE2" author="Bousquet, Y" editor="Loebl, I" journalOrPublisher="Apollo Books, Denmark" pageId="59" pageNumber="60" pagination="469 - 521" title="Carabidae: tribe Pterostichini Bonelli, 1810." volumeTitle="Catalogue of Palaearctic Coleoptera, Volume 1, Archostemata- Myxophaga- Adephaga" year="2003">Bousquet 2003</bibRefCitation>
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: 488;
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<bibRefCitation id="61CD09E2AF72528D5BADC98FD9A515D6" author="Lorenz, W" journalOrPublisher="Wolfgang Lorenz, Tutzing, Germany" pageId="59" pageNumber="60" title="Systematic list of extant ground beetles of the world (InsectaColeopteraGeadephaga: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae)" year="2005">Lorenz 2005</bibRefCitation>
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: 280;
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<bibRefCitation id="DB0281D4AE6239BE8EDE61BC1D066138" author="Allegro, G" journalOrPublisher="Bollettino del Museo Civico di Storia Naturale di Verona (Botanica Zoologia)" pageId="59" pageNumber="60" pagination="3 - 18" title="Review of the Chinese subgenus Circinatus of Pterostichus, with description of six new species (Coleoptera, Carabidae)." volume="34" year="2010">Allegro and Sciaky 2010</bibRefCitation>
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: 7.
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</paragraph>
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</subSubSection>
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<subSubSection id="1A3C678E6503CAF336087E79067BD41A" pageId="37" pageNumber="38" type="type locality">
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<paragraph id="172F058FB8DF57369F8EED0C5105C7AE" pageId="37" pageNumber="38">Type locality.</paragraph>
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<paragraph id="2FB497E38358BFA5E6DD99079A13F32F" pageId="37" pageNumber="38">
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The original literature mentioned the locality
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<normalizedToken id="8742107DD531FF02FC9A5D7042AE7D40" originalValue="“Tatsienlu”">"Tatsienlu"</normalizedToken>
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, without further details. But the label of the holotype indicates that the type locality is
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<normalizedToken id="E8A2709A1FE91A583EDDC58A5A6984F5" originalValue="“Kiulung”">"Kiulung"</normalizedToken>
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(= Jiulong), a county west of Gonggashan Mountain.
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</paragraph>
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</subSubSection>
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<subSubSection id="BD9A34F78AAF21082EFA496D50E8A665" pageId="37" pageNumber="38" type="type material">
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<paragraph id="133BDE4B190CD078E64D2C248E79E3B8" pageId="37" pageNumber="38">Type material examined.</paragraph>
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<paragraph id="F3B655EE9670161EC8966CC6DAB766D3" pageId="37" pageNumber="38">
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Holotype of
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<taxonomicName id="DBE595D6CA9C5307942153BB2459E4E9" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus pohnerti" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pohnerti">Pterostichus pohnerti</taxonomicName>
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<normalizedToken id="42CEC8928BBB66C39A35915CB810F1BC" originalValue="Jedlička">Jedlicka</normalizedToken>
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(NMPC), male, body length = 10.9 mm, board mounted, with left foreleg, right hind tarsomeres (apical four segments), left antenna (apical ten antennomeres) and right antenna (apical six antennomeres) missing, genitalia dissected and stored in micro vial pinned under specimen, "Tatsienlu-Kiulung / China Em. Reitter";
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<normalizedToken id="BAA0D74C1BC0CE844948516DCF9B172A" originalValue="”TYPE”">"TYPE"</normalizedToken>
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[red label]; "Mus. Nat. Prague / Inv. 22407" [orange label]; "
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<taxonomicName id="88F3E25D54AE796F2DB4BD8515D2989F" genus="Pohnerti" lsidName="Pohnerti" pageId="37" pageNumber="38" rank="genus">Pohnerti</taxonomicName>
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/ type, sp. n., / DET. ING.
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<normalizedToken id="8F7CB76F2808698B3BA59CD46A6D9E1D" originalValue="JEDLIČKA”">JEDLICKA"</normalizedToken>
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[pink label].
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</paragraph>
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</subSubSection>
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<subSubSection id="217D2473AD1848A0ADC0648D30C9EA0C" pageId="37" pageNumber="38" type="type material">
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<paragraph id="D64CAFD10A8CF381D4E389549C9BFC9A" pageId="37" pageNumber="38">Non-type material examined</paragraph>
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<paragraph id="D71CB4093B1D6AF6FA271706FF558DC1" pageId="37" pageNumber="38">
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(66 specimens). 3 males, 4 females (IZAS), "China, Sichuan, Luding county, Moxi town, way btw. Moxi and Yajiageng pass, mixed forest, 3204 m,
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<geoCoordinate id="9137BC8C0AE8E70005C43BBBB8C0EBDA" direction="north" orientation="latitude" precision="1" value="29.8411">N 29.84110</geoCoordinate>
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,
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<geoCoordinate id="838F535093209B0F2A89501B3795CB44" direction="east" orientation="longitude" precision="1" value="102.04802">E 102.04802</geoCoordinate>
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, by pit fall trap, 2012.VII.11, SHI Hongliang & WANG
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<normalizedToken id="1956995DFD80BFB28A8CF8CFDB1D2480" originalValue="Ya’nan">Ya'nan</normalizedToken>
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lgt."; 1 male, 3 females (IZAS), "Sichuan, Luding, Moxi, 1990.VII.6, T. Deuve & XIE Weiping lgt."; 1 male (IZAS), "Sichuan, Luding, Hailuogou, 3000 m, 2004.VIII.8, BAI Ming lgt."; 4 males, 5 females (IZAS), "China, Sichuan, Luding, Hailuogou, 3200, 2011.7.16, Huang Hao"; 13 males, 5 females (IZAS), "China, Sichuan Prov., Luding, Hailuogou, Caohaizi,
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<geoCoordinate id="19819620D311FE5F3D41AA79614F7E9D" direction="north" orientation="latitude" precision="1" value="29.58228">N29.58228</geoCoordinate>
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,
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<geoCoordinate id="86F2488A110121473F8EF672D52D467C" direction="east" orientation="longitude" precision="1" value="102.02171">E102.02171</geoCoordinate>
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, 2775 m, 2009.V.18, Liang H.B. lgt."; 1 male, 8 females (IZAS), "China, Sichuan Prov., Luding, Hailuogou, Guanjingtai,
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<geoCoordinate id="5D686DC70F5362D29817F6F149637CC2" direction="north" orientation="latitude" precision="1" value="29.56721">N29.56721</geoCoordinate>
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,
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<geoCoordinate id="765486D5298D8D13B886078A25749ED4" direction="east" orientation="longitude" precision="1" value="101.97994">E101.97994</geoCoordinate>
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, 3155 m, 2009.V.16, Yang G.Y. lgt."; 2 males (IZAS), "China, Sichuan, Tianquan, Erlangshan, 2500 m, 2011.VIII.5, Huang Hao lgt."; 9 males, 7 females (IZAS), "China, Sichuan, Baoxing county, Jiajinshan mt., Mahuanggou, mixed forest,
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<geoCoordinate id="5219B657FD53E0829AD1EA2AB30C9568" direction="north" orientation="latitude" precision="1" value="30.85669">N30.85669</geoCoordinate>
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,
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<geoCoordinate id="80C5E74E8451FB724AA69E166A3FC6DF" direction="east" orientation="longitude" precision="1" value="102.76615">E102.76615</geoCoordinate>
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, 2012.VI.29 D 2681 m, Liu Ye, Shi Hongliang leg.".
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</paragraph>
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</subSubSection>
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<subSubSection id="49161743B85B99FD1724937BBFE562CA" pageId="37" pageNumber="38" type="diagnosis">
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<paragraph id="4F2BB9D9D2847998F73BE08DF6E7A159" pageId="37" pageNumber="38">Diagnosis.</paragraph>
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<paragraph id="16AADDDECDBE24C8679AB336C0D000D6" pageId="37" pageNumber="38">Pronotum with single mid-lateral seta; posterior seta distant from hind angle, hind angle completely rounded; outer basal foveal groove distinct, approx half length of inner one, forming a short ridge close to hind angle; basal fovea area finely punctate; elytron with sharp humeral tooth; elytral with faint linear microsculpture; fifth tarsomeres glabrous beneath; terminal sternum of male not modified.</paragraph>
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<paragraph id="930D32A157B07D53C06ADD9CFD6E4643" pageId="37" pageNumber="38">
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This species is very similar to
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<taxonomicName id="D0707D44C5FA2DA0D2E11D442D243A7F" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus xilingensis" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="xilingensis">Pterostichus xilingensis</taxonomicName>
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. They are difficult to separate by external appearance (the only difference is the punctures in the pronotal basal fovea (see the key), but this does not always distinguish them), but they can be easily determined by their allopatric
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</paragraph>
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</subSubSection>
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<subSubSection id="F10F9A77CDB26CF80ED70E4367A1423E" pageId="37" pageNumber="38" type="distribution">
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<paragraph id="9E97171806EFA07BFF9174A15657E83A" pageId="37" pageNumber="38">Distribution.</paragraph>
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<paragraph id="FD6F98EC8B7F2DF50D1F4C2D7A2CDBE3" pageId="37" pageNumber="38">
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In male and female genitalia, these two species are different. In
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<taxonomicName id="DB1FCC9AB492325B8177B2BEE9BADADF" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus pohnerti" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="pohnerti">Pterostichus pohnerti</taxonomicName>
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: median lobe of aedeagus stouter; apical lamella triangular, strongly narrowed to apex; endophallus extending in apical direction of aedeagus; female transparent region on sternum VIII V-shaped, with very long and narrow extensions. In
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<taxonomicName id="7B755F941A2F496B31CE3D5D93879825" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus xilingensis" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="xilingensis">Pterostichus xilingensis</taxonomicName>
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: median lobe of aedeagus much slenderer; apical lamella wide, slightly narrowed to apex; endophallus extending in apical-dorsal direction of aedeagus; female transparent region on sternum VIII nearly quadrate, with very short extensions.
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</paragraph>
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</subSubSection>
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<subSubSection id="5052ABF23A4AED7D75D7BD60A3C82139" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="supplemental description">
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<paragraph id="C48666056D49D5910D173543F8B80594" pageId="37" pageNumber="38">Supplemental description.</paragraph>
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<paragraph id="50E82DBFAAB0FD2006727CF4A22E8E78" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
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Male sternum. Penultimate and terminal sterna without special structures. Endophallus (Fig. 47D, E, G) short and straight, extending in apical direction of aedeagus, major portion of endophallus located beyond apical lamella; gonopore (gp) located at level near aedeagal apex, pointing to ventral face of aedeagus, gonopore lobe (gpl, fully everted in Fig. 47) bent in basal direction of
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<pageBreakToken id="8DA0747F4F1C315C04C8ABF61B214606" pageId="38" pageNumber="39" start="start">aedeagus</pageBreakToken>
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. Four distinct parts recognized: basal lobe (bl) close to base of apical lamella, very small, approx triangular; main piece (mp) large and galericulate, with three branches, covering gpl, apical opening of gpl pointing out from ventral notch between mp branches; apical band (ab) close to gp, also covered by mp, half chitinized, and the rest half scaled and extended to inner margin of pa; pre-apical lobe (pa) small and compressed, ear-like, with fine and dense scales. Female genitalia. Spermatheca (Fig. 56) with seminal canal approx 4.5 times as long as receptaculum; receptaculum capitate, club approx one third as long as receptaculum; seminal canal inserted at base of common oviduct, base of seminal canal with sclerotized region long. Stylomere II with two ensiform setae at basal half of outer margin, and a small one near middle of inner margin; two short nematiform setae located in a furrow near apex. Female sternum VIII (Fig. 85B) with fine and dense setae on posterior margin; posterior margin almost straight, strongly notched in middle; anterior region semi-chitinized, deeply notched in middle; middle transparent region V-shaped, adjacent to anterior and posterior notches in middle. Female tergum VIII (Fig. 85A) short, major portion chitinized, apical fourth semi-chitinized, with very sparse spots; anterior margin weakly notched in middle, posterior margin evenly arcuate.
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="3E5A73629B5BCBD5CE89E7B82CF5589E" pageId="38" pageNumber="39" type="distribution">
|
||
<paragraph id="07720BEF71C002DA419C4284DD313E2C" pageId="38" pageNumber="39">Distribution.</paragraph>
|
||
<paragraph id="FDCB6491657D5EF944747CC777AC970D" pageId="38" pageNumber="39">
|
||
<taxonomicName id="2664DDEB80E1C2886958E5AEBA2B4985" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus pohnerti" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pohnerti">Pterostichus pohnerti</taxonomicName>
|
||
is recorded from four counties of central Sichuan province: Luding (Gonggashan range), Jiulong (type locality), Baoxing (south slope of Jiajinshan), and Tianquan (east slope of Erlangshan). This species is very common and the only known
|
||
<taxonomicName id="F3B6689ADD7060CD03785AF2B93420BB" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus (Circinatus)" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="subGenus" subGenus="Circinatus">Circinatus</taxonomicName>
|
||
species in the Gonggashan range. A large number of specimens was collected from several localities around this mountain. Outside the Gonggashan range, it is also distributed in Baoxing and Tianquan, where it overlaps the range of
|
||
<taxonomicName id="75DE061EF4EC70ECBC3229D613B79AD6" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus beneshi" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="beneshi">Pterostichus beneshi</taxonomicName>
|
||
. (Maps 2, 3)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="06A27F94DF8955A368873F8A3B649644" pageId="38" pageNumber="39" type="affinities">
|
||
<paragraph id="6639F8ECA3BB00EDCEA1511A1ECDD94A" pageId="38" pageNumber="39">Affinities.</paragraph>
|
||
<paragraph id="556ECE8807C9FB0FBDAE2287D41F13BD" pageId="38" pageNumber="39">
|
||
<taxonomicName id="B5AE020F63E01BEED66951E9EA0A8015" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus pohnerti" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pohnerti">Pterostichus pohnerti</taxonomicName>
|
||
could be closest to
|
||
<taxonomicName id="DC8E5CC8F3E0FD7B5B399B47AC0600E7" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus xilingensis" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="xilingensis">Pterostichus xilingensis</taxonomicName>
|
||
based on the similarity in external characters. However, in the stout aedeagus and endophallus extending to the aedeagal apex,
|
||
<taxonomicName id="306A341AAB46C85CABDF3EFEB7758C61" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus pohnerti" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pohnerti">Pterostichus pohnerti</taxonomicName>
|
||
could be allied with some species in the
|
||
<taxonomicName id="0EE348B8C4932A6F8BC35460160E2E2D" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus agilis" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="agilis">agilis</taxonomicName>
|
||
-group. Moreover,
|
||
<taxonomicName id="C8E3876F2F25BB57B2137F0622A63968" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus pohnerti" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pohnerti">Pterostichus pohnerti</taxonomicName>
|
||
has the southernmost distribution in the
|
||
<taxonomicName id="72DDDA698A83A8535C6DAA4D2D442B0F" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus pohnerti" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pohnerti">pohnerti</taxonomicName>
|
||
-group, which is the closest distribution to the range of the
|
||
<taxonomicName id="9FF56831488E70E2FA52BBE2131C5E67" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus agilis" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="agilis">agilis</taxonomicName>
|
||
-group. Based on these distributions, we infer that the systematic position of
|
||
<taxonomicName id="F816EA1794FEE8AF69BF0B741C70E439" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus pohnerti" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pohnerti">Pterostichus pohnerti</taxonomicName>
|
||
could be between
|
||
<taxonomicName id="19EEBA388D08DD442511B78A009DA8BF" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus xilingensis" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="xilingensis">Pterostichus xilingensis</taxonomicName>
|
||
and the
|
||
<taxonomicName id="64D27A864D6B1DF003CF8DA706D770DD" class="Insecta" family="Carabidae" genus="Pterostichus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterostichus agilis" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="agilis">agilis</taxonomicName>
|
||
-group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2BC59E49B89881F64206A4734952D724" pageId="38" pageNumber="39" type="geographical variation">
|
||
<paragraph id="A2048EE8D1299D87902330A8B9FB0CA6" pageId="38" pageNumber="39">Geographical variation.</paragraph>
|
||
<paragraph id="23A42175AE5DC59E5D8D2030AB2EF3AF" pageId="38" pageNumber="39">
|
||
We studied males from three localities: Hailuogou (
|
||
<geoCoordinate id="E97C53A9C422CA9EF0113D481E4CC13F" direction="north" orientation="latitude" precision="555" value="29.58">N29.58°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="059707A17920EFC4105A1D140FC688B8" direction="east" orientation="longitude" precision="555" value="102.02">E102.02°</geoCoordinate>
|
||
), Erlangshan (
|
||
<geoCoordinate id="647C3EF3612D4B59FA13AF2B293ED7BF" direction="north" orientation="latitude" precision="555" value="29.88">N29.88°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="0D24B8E30B76964A9156DFECEB52DDBF" direction="east" orientation="longitude" precision="555" value="102.3">E102.30°</geoCoordinate>
|
||
), and Baoxing (
|
||
<geoCoordinate id="9256601F521F38DEF140EF08CD40495E" direction="north" orientation="latitude" precision="555" value="30.86">N30.86°</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE99E36AE2EBCB9B9799FF8416E3AB53" direction="east" orientation="longitude" precision="555" value="102.77">E102.77°</geoCoordinate>
|
||
), and found no differences in external, aedeagal or endophallic characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |