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<document id="27415BE18F6789CD1858B3B7B14D88E3" ID-DOI="10.11646/zootaxa.5484.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13208985" ID-ZooBank="E7473BE4-5C6B-4D08-90CD-DCE99BD35BB9" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1722785721163" checkinUser="plazi" docAuthor="Miranda, Gil F. G., Soares, Matheus M. M. &amp; Thompson, Christian" docDate="2024" docId="F814866ED56FE70A33C24FD0FE74FC20" docLanguage="en" docName="zootaxa.5484.1.1.pdf" docOrigin="Zootaxa 5484 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5484.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Orthonevra cthulhu Miranda, Soares &amp; Thompson 2024, sp. nov" docType="treatment" docVersion="4" lastPageNumber="24" masterDocId="042DFE16D57AE71D33554F47FFD4FF85" masterDocTitle="The Neotropical Orthonevra Macquart, 1829 (Diptera: Syrphidae)" masterLastPageNumber="78" masterPageNumber="1" pageNumber="22" updateTime="1723148386458" updateUser="ExternalLinkService">
<mods:mods id="7BD8E64915B9AA64F235BEC192CCA1E9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B96D8975691AFF8C1A9850FD6A0A4053">
<mods:title id="7133C634ECC4E18574D4CC3B49DD34CF">The Neotropical Orthonevra Macquart, 1829 (Diptera: Syrphidae)</mods:title>
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<mods:name id="45C36C9C8A45D0E53BBCDBBBF82590C3" type="personal">
<mods:role id="C3D7D6C053AA98012386992A77894DF6">
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<mods:namePart id="07A4E8206F4220B3C4B1918886B5EAC5">Miranda, Gil F. G.</mods:namePart>
<mods:nameIdentifier id="3D05788F486D600F41135D60E1BE10C0" type="ORCID">0000-0001-7919-2639</mods:nameIdentifier>
<mods:affiliation id="ABD3CD850FB94A61B6B614B87F2CC8BE">National Collection of Insects, Arachnids and Nematodes, Agriculture and Agri-Food Canada, 960 Carling Ave., Ottawa, ON, K 1 A 0 C 6, Canada / Forrest Lab, Department of Biology, University of Ottawa, Gendron Hall, room 350, 30 Marie Curie, Ottawa, ON, K 1 N 6 N 5, Canada. gilfgm @ gmail. com; https: // orcid. org / 0000 - 0001 - 7919 - 2639</mods:affiliation>
<mods:nameIdentifier id="EB28FDE6D9FC7AD5102333D9E730835F" type="email">gilfgm@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="A78F78101D80B030CAF50FFD3C293583" type="personal">
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<mods:namePart id="E1074D52A7BF96E837B5FE07F8DC9973">Soares, Matheus M. M.</mods:namePart>
<mods:nameIdentifier id="B32656FF60DAE9FA0084853AC814CBED" type="ORCID">0000-0002-2355-1441</mods:nameIdentifier>
<mods:affiliation id="B6BD17EFB32A5E5CA95AC874F42EB0E2">Laboratório de Sistemática de Diptera, Coordenação de Biodiversidade, Instituto Nacional de Pesquisas da Amazônia, Avenida André Araújo, 2936, Bairro Petrópolis, Manaus, AM, 69.067 - 375, Brazil. matheusmmsoares @ gmail. com; https: // orcid. org / 0000 - 0002 - 2355 - 1441 In memoriam * Corresponding author</mods:affiliation>
<mods:nameIdentifier id="09BF46FB6760935E9A2C1BE6BB8E3820" type="email">matheusmmsoares@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="A9977D5C8FBDDE9B793D05406E882EE3" type="personal">
<mods:role id="245B4FD3D62ED92D846E7E61E0B8A98E">
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<mods:namePart id="E186DB94E2FC3012314ED50E4F20A317">Thompson, Christian</mods:namePart>
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<mods:typeOfResource id="F6BB927D40454473A2FC74D2E648725C">text</mods:typeOfResource>
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<mods:title id="3C9C6B1EFEFF8CA9ADEC147E85C433B5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1FF0FCDBB389336F0D2F33E9E1B3C6E5">
<mods:date id="660B5BE2CB87ABAF1517C9C5FBBE737F">2024</mods:date>
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<mods:number id="BBB7549029ADC9B0D19355A070ED1868">2024-07-22</mods:number>
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<mods:number id="B4866344CDFC5715C64F62366CF82513">5484</mods:number>
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<mods:classification id="763A6CD9989B2B59D56FFF8FA450FA64">journal article</mods:classification>
<mods:identifier id="BD2D68731A1E140274B3A70B16862F55" type="DOI">10.11646/zootaxa.5484.1.1</mods:identifier>
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<treatment id="F814866ED56FE70A33C24FD0FE74FC20" ID-DOI="http://doi.org/10.5281/zenodo.13274534" ID-Zenodo-Dep="13274534" LSID="urn:lsid:plazi:treatment:F814866ED56FE70A33C24FD0FE74FC20" httpUri="http://treatment.plazi.org/id/F814866ED56FE70A33C24FD0FE74FC20" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22">
<subSubSection id="38A764F3D56FE70833C24FD0FD92FF71" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="70023778D56FE70833C24FD0FCA6FF34" blockId="21.[151,882,151,178]" box="[151,882,151,178]" pageId="21" pageNumber="22">
<heading id="2B4A8014D56FE70833C24FD0FCA6FF34" bold="true" box="[151,882,151,178]" fontSize="11" level="2" pageId="21" pageNumber="22" reason="4">
<emphasis id="42C9EB6AD56FE70833C24FD0FCA6FF34" bold="true" box="[151,882,151,178]" pageId="21" pageNumber="22">
<taxonomicName id="B7BD4CFBD56FE70833C24FD0FCD9FF34" authority="Miranda, Soares &amp; Thompson" authorityName="Miranda, Soares &amp; Thompson" authorityYear="2024" box="[151,781,151,178]" class="Insecta" family="Syrphidae" genus="Orthonevra" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cthulhu">
<emphasis id="42C9EB6AD56FE70833C24FD0FE5EFF37" bold="true" box="[151,394,151,178]" italics="true" pageId="21" pageNumber="22">Orthonevra cthulhu</emphasis>
Miranda, Soares &amp; Thompson
</taxonomicName>
<taxonomicNameLabel id="59FA5611D56FE70830414FD0FCB9FF34" box="[788,877,151,177]" pageId="21" pageNumber="22" rank="species">sp. nov</taxonomicNameLabel>
.
</emphasis>
</heading>
</paragraph>
<paragraph id="70023778D56FE70833C24F9BFD92FF71" blockId="21.[151,582,219,280]" box="[151,582,219,244]" pageId="21" pageNumber="22">
98-4.
<bibRefCitation id="142C4A89D56FE70833874F9BFE4AFF71" author="Thompson, F. C." box="[210,414,219,244]" pageId="21" pageNumber="22" refId="ref53258" refString="Thompson, F. C. (2006) Primer Taller de Identificacion. Syrphidae de Neotropico. Universidad del Valle, Cali, Valle del Cauca, 618 pp." type="book" year="2006">Thompson 2006: 21</bibRefCitation>
(key reference).
</paragraph>
</subSubSection>
<subSubSection id="38A764F3D56FE70833C24FB9FE1AFEE5" pageId="21" pageNumber="22" type="description">
<paragraph id="70023778D56FE70833C24FB9FE6FFE9D" blockId="21.[151,582,219,280]" box="[151,443,254,280]" pageId="21" pageNumber="22">
<taxonomicName id="B7BD4CFBD56FE70833C24FB9FEC3FE9D" authorityName="Macquart" authorityYear="1829" box="[151,279,254,280]" class="Insecta" family="Syrphidae" genus="Orthonevra" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">Orthonevra</taxonomicName>
3 (FCT notes)
</paragraph>
<paragraph id="70023778D56FE70833C24E01FE1AFEE5" blockId="21.[151,462,326,352]" box="[151,462,326,352]" pageId="21" pageNumber="22">
<figureCitation id="E8862BFDD56FE70833C24E01FF08FEE5" box="[151,220,326,352]" captionStart="FIGURE 9" captionStartId="22.[151,250,1930,1954]" captionTargetBox="[151,1436,633,1907]" captionTargetId="figure-231@22.[151,1436,633,1907]" captionTargetPageId="22" captionText="FIGURE 9. Orthonevra cthulhu sp. nov. a–c. Female: a. Head and thorax, dorsal view. b. Habitus lateral. c. Head, lateral view. d–f. Male: d. Head and thorax, dorsal view. e. Habitus lateral. f. Head, oblique dorsal view. Abbreviations: sc: scale-like pile; scl: scutellum; r1v: sub-apical vitta of cell r1." figureDoi="http://doi.org/10.5281/zenodo.13209005" httpUri="https://zenodo.org/record/13209005/files/figure.png" pageId="21" pageNumber="22">Figs 9</figureCitation>
and
<figureCitation id="E8862BFDD56FE70832464E01FEE6FEE5" box="[275,306,326,352]" captionStart="FIGURE 10" captionStartId="23.[151,250,1900,1924]" captionTargetBox="[199,1375,957,1865]" captionTargetId="figure-295@23.[199,1387,957,1877]" captionTargetPageId="23" captionText="FIGURE 10. Orthonevra cthulhu sp. nov., genitalia. a. Female, dorsal view. b. Female, ventral view. c–e. Male: c. Epandrium, lateral view. d. Hypandrium, lateral view. e. Phallus, lateral view. Abbreviations: btp: basal tubular process; cerc: cercus; hypc: ventral carina of the hypandrium; hypo: hypoproct; epi: epiproct; pg: postgonite; sur: surstylus; s7 &amp; 8: sterna 7 &amp; 8; t7 &amp; 8: terga 7 &amp; 8." figureDoi="http://doi.org/10.5281/zenodo.13209007" httpUri="https://zenodo.org/record/13209007/files/figure.png" pageId="21" pageNumber="22">10</figureCitation>
. Map:
<figureCitation id="E8862BFDD56FE708322A4E01FE1AFEE5" box="[383,462,326,352]" captionStart="FIGURE 39" captionStartId="70.[151,250,1913,1937]" captionTargetBox="[151,1435,669,1889]" captionTargetId="figure-252@70.[151,1435,669,1889]" captionTargetPageId="70" captionText="FIGURE 39. Distribution map for Orthonevra argentina (Brèthes, 1922), O. batman sp. nov., O. chilensis Thompson, 1999, O. cthulhu sp. nov., O. quadristriata (Shannon &amp; Aubertin, 1933), O. roseae sp. nov., O. shannoni (Curran, 1926), and O. thompsoni sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13209085" httpUri="https://zenodo.org/record/13209085/files/figure.png" pageId="21" pageNumber="22">Fig. 39</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="38A764F3D56FE70833C24ECAFB2BFE2D" box="[151,1279,397,424]" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="70023778D56FE70833C24ECAFB2BFE2D" blockId="21.[151,1437,397,1972]" box="[151,1279,397,424]" pageId="21" pageNumber="22">
<emphasis id="42C9EB6AD56FE70833C24ECAFEC6FE22" bold="true" box="[151,274,397,423]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
Similar to
<taxonomicName id="B7BD4CFBD56FE70832DA4EC9FDC9FE2D" box="[399,541,398,424]" class="Insecta" family="Syrphidae" genus="Orthonevra" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="argentina">
<emphasis id="42C9EB6AD56FE70832DA4EC9FDC9FE2D" box="[399,541,398,424]" italics="true" pageId="21" pageNumber="22">O. argentina</emphasis>
</taxonomicName>
and
<taxonomicName id="B7BD4CFBD56FE70831064EC9FD67FE2D" baseAuthorityName="Sedman" baseAuthorityYear="1964" box="[595,691,398,424]" class="Insecta" family="Syrphidae" genus="Orthonevra" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="flukei">
<emphasis id="42C9EB6AD56FE70831064EC9FD67FE2D" box="[595,691,398,424]" italics="true" pageId="21" pageNumber="22">O. flukei</emphasis>
</taxonomicName>
but
<taxonomicName id="B7BD4CFBD56FE70831B04EC9FC8FFE2D" authorityName="Miranda &amp; Soares &amp; Thompson" authorityYear="2024" box="[741,859,398,424]" class="Insecta" family="Syrphidae" genus="Orthonevra" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cthulhu">
<emphasis id="42C9EB6AD56FE70831B04EC9FC8FFE2D" box="[741,859,398,424]" italics="true" pageId="21" pageNumber="22">O. cthulhu</emphasis>
</taxonomicName>
has a short sub-apical vitta in cell r1.
</paragraph>
</subSubSection>
<subSubSection id="38A764F3D56FE70A33924EF6FF09FF50" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22" type="description">
<paragraph id="70023778D56FE70833924EF6FAABFC25" blockId="21.[151,1437,397,1972]" pageId="21" pageNumber="22">
<emphasis id="42C9EB6AD56FE70833924EF6FE85FE4E" bold="true" box="[199,337,433,459]" pageId="21" pageNumber="22">Description</emphasis>
. MALE.
<emphasis id="42C9EB6AD56FE70832EB4EF6FE28FE49" bold="true" box="[446,508,433,460]" italics="true" pageId="21" pageNumber="22">Head</emphasis>
(
<figureCitation id="E8862BFDD56FE708315E4EF5FDADFE49" box="[523,633,434,460]" captionStart="FIGURE 9" captionStartId="22.[151,250,1930,1954]" captionTargetBox="[151,1436,633,1907]" captionTargetId="figure-231@22.[151,1436,633,1907]" captionTargetPageId="22" captionText="FIGURE 9. Orthonevra cthulhu sp. nov. a–c. Female: a. Head and thorax, dorsal view. b. Habitus lateral. c. Head, lateral view. d–f. Male: d. Head and thorax, dorsal view. e. Habitus lateral. f. Head, oblique dorsal view. Abbreviations: sc: scale-like pile; scl: scutellum; r1v: sub-apical vitta of cell r1." figureDoi="http://doi.org/10.5281/zenodo.13209005" httpUri="https://zenodo.org/record/13209005/files/figure.png" pageId="21" pageNumber="22">Fig. 9d–f</figureCitation>
): metallic bluish green, face with convexity on dorsal 1/
<quantity id="B7459A9DD56FE70837BB4EF5FACFFE49" box="[1262,1307,434,460]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="21" pageNumber="22" unit="in" value="2.0">2 in</quantity>
profile and ventral 1/4 slightly produced anteriorly as a convexity, strongly regulose except medially between convexity and anterior oral margin, pile white and scale-like; with white microtrichose subtriangular macula laterally, positioned immediately ventral to level of antennal base but far removed from it. Antenna brownish yellow, post-pedicel darker apically; pedicel twice the length of the scape, postpedicel oval elongated, slightly longer than the length of the pedicel. Mala slightly produced apico-ventrally, smooth. Gena shiny, smooth, and with white pile. Frontal triangle shiny, regulose, with a medial longitudinal sulcus starting on the eye contiguity and ending dorsal to the elevation of the lunule, with sparse white pile, scale-like laterally. Vertical triangle isosceles triangle-shaped, 4 × the length of the eye contiguity, metallic green, bare except for dark erect pile on ocellar triangle; ocellar triangle equilateral, making up 2/3 of the vertical triangle. Occiput not visible laterally on dorsal 1/2, ventral 1/2 homogeneously covered with white microtrichia which gets longer dorsal to the gena, with a row of inconspicuous dark pile that becomes longer and white close to the gena. Eyes holoptic; with a sub-anterior sinuous vitta, a sub-anterior circle dorsally, a strongly sinuous middle vitta, and an uninterrupted medial fascia, eyes darker on anterior margin and on ventral 1/2 of posterior margin, anterior margin connected dorsally and ventrally to sub-anterior vitta (ventral to dorsal circle).
</paragraph>
<paragraph id="70023778D56FE70833924CEEFC36FB45" blockId="21.[151,1437,397,1972]" pageId="21" pageNumber="22">
<emphasis id="42C9EB6AD56FE70833924CEEFEC9FC41" bold="true" box="[199,285,937,964]" italics="true" pageId="21" pageNumber="22">Thorax</emphasis>
(
<figureCitation id="E8862BFDD56FE70832784CEDFE48FC41" box="[301,412,938,964]" captionStart="FIGURE 9" captionStartId="22.[151,250,1930,1954]" captionTargetBox="[151,1436,633,1907]" captionTargetId="figure-231@22.[151,1436,633,1907]" captionTargetPageId="22" captionText="FIGURE 9. Orthonevra cthulhu sp. nov. a–c. Female: a. Head and thorax, dorsal view. b. Habitus lateral. c. Head, lateral view. d–f. Male: d. Head and thorax, dorsal view. e. Habitus lateral. f. Head, oblique dorsal view. Abbreviations: sc: scale-like pile; scl: scutellum; r1v: sub-apical vitta of cell r1." figureDoi="http://doi.org/10.5281/zenodo.13209005" httpUri="https://zenodo.org/record/13209005/files/figure.png" pageId="21" pageNumber="22">Fig. 9d–f</figureCitation>
): metallic green, pile white, short, mostly appressed and sparsely distributed, bare on dark areas; scutum with four complete and separate dark matte vittae, that taper anteriorly, with purple reflections intermixed, sub-medial vittae slightly interrupted at transverse sulcus, and with a short narrow lateral vittate macula posterior to transverse sulcus; notopleuron metallic green with a dark matte area dorsal; scutellum mostly dark metallic green, apical margin of a different lighter hue, with short, appressed, white pile. Pleuron with white pile on antepronotum, proepisternum, posterior anepisternum, on dorsal and ventral patches on katepisternum, anterior anepimeron, and katatergum, mostly with sparse microtrichia. Calypter white with white marginal pile, ventral lobe with longer marginal pile. Plumule white and long. Halter yellowish-white.
</paragraph>
<paragraph id="70023778D56FE70833924B8DFA84FAA9" blockId="21.[151,1437,397,1972]" pageId="21" pageNumber="22">
<emphasis id="42C9EB6AD56FE70833924B8DFF29FB61" bold="true" box="[199,253,1226,1252]" italics="true" pageId="21" pageNumber="22">Legs</emphasis>
(
<figureCitation id="E8862BFDD56FE70832594B8DFEA1FB61" box="[268,373,1226,1252]" captionStart="FIGURE 9" captionStartId="22.[151,250,1930,1954]" captionTargetBox="[151,1436,633,1907]" captionTargetId="figure-231@22.[151,1436,633,1907]" captionTargetPageId="22" captionText="FIGURE 9. Orthonevra cthulhu sp. nov. a–c. Female: a. Head and thorax, dorsal view. b. Habitus lateral. c. Head, lateral view. d–f. Male: d. Head and thorax, dorsal view. e. Habitus lateral. f. Head, oblique dorsal view. Abbreviations: sc: scale-like pile; scl: scutellum; r1v: sub-apical vitta of cell r1." figureDoi="http://doi.org/10.5281/zenodo.13209005" httpUri="https://zenodo.org/record/13209005/files/figure.png" pageId="21" pageNumber="22">Fig. 9e, f</figureCitation>
): metallic green, except brownish tibiae with paler base and apex, and tarsi with first two (three on metatarsus) tarsomeres yellow, remaining brown. Legs covered with short white pile, with black ventral setulae on mesotarsus, metafemur, and apex of mesotibia, shorter, paler and sparser laterally on pro and metatarsus.
</paragraph>
<paragraph id="70023778D56FE70833924A71FD6BFA65" blockId="21.[151,1437,397,1972]" pageId="21" pageNumber="22">
<emphasis id="42C9EB6AD56FE70833924A71FED0FAD5" bold="true" box="[199,260,1334,1360]" italics="true" pageId="21" pageNumber="22">Wing</emphasis>
(
<figureCitation id="E8862BFDD56FE70832414A71FEB2FAD5" box="[276,358,1334,1360]" captionStart="FIGURE 9" captionStartId="22.[151,250,1930,1954]" captionTargetBox="[151,1436,633,1907]" captionTargetId="figure-231@22.[151,1436,633,1907]" captionTargetPageId="22" captionText="FIGURE 9. Orthonevra cthulhu sp. nov. a–c. Female: a. Head and thorax, dorsal view. b. Habitus lateral. c. Head, lateral view. d–f. Male: d. Head and thorax, dorsal view. e. Habitus lateral. f. Head, oblique dorsal view. Abbreviations: sc: scale-like pile; scl: scutellum; r1v: sub-apical vitta of cell r1." figureDoi="http://doi.org/10.5281/zenodo.13209005" httpUri="https://zenodo.org/record/13209005/files/figure.png" pageId="21" pageNumber="22">Fig. 9e</figureCitation>
): hyaline, with dark vitta sub-apically (from end R2+3 until posterior end of M1), and dark apex (but fading posterior to vein R4+5), and dark vittate macula sub-apically on cell r1, a fasciate pair apically on r4+5 and a small fasciate macula antero-apically on dm (
<figureCitation id="E8862BFDD56FE70831BF4A39FCEAFA1D" box="[746,830,1406,1432]" captionStart="FIGURE 9" captionStartId="22.[151,250,1930,1954]" captionTargetBox="[151,1436,633,1907]" captionTargetId="figure-231@22.[151,1436,633,1907]" captionTargetPageId="22" captionText="FIGURE 9. Orthonevra cthulhu sp. nov. a–c. Female: a. Head and thorax, dorsal view. b. Habitus lateral. c. Head, lateral view. d–f. Male: d. Head and thorax, dorsal view. e. Habitus lateral. f. Head, oblique dorsal view. Abbreviations: sc: scale-like pile; scl: scutellum; r1v: sub-apical vitta of cell r1." figureDoi="http://doi.org/10.5281/zenodo.13209005" httpUri="https://zenodo.org/record/13209005/files/figure.png" pageId="21" pageNumber="22">Fig. 9e</figureCitation>
), microtrichose except bare medially on cell bm and immediately posterior to pseudovein; basicosta with dense appressed pale to dark pile, with two more prominent black setae apically. Alula 2 × the width of cell c.
</paragraph>
<paragraph id="70023778D56FE70833924AAEFDB2F9C9" blockId="21.[151,1437,397,1972]" pageId="21" pageNumber="22">
<emphasis id="42C9EB6AD56FE70833924AAEFEE1F981" bold="true" box="[199,309,1513,1540]" italics="true" pageId="21" pageNumber="22">Abdomen</emphasis>
(
<figureCitation id="E8862BFDD56FE70832194AADFE72F981" box="[332,422,1514,1540]" captionStart="FIGURE 9" captionStartId="22.[151,250,1930,1954]" captionTargetBox="[151,1436,633,1907]" captionTargetId="figure-231@22.[151,1436,633,1907]" captionTargetPageId="22" captionText="FIGURE 9. Orthonevra cthulhu sp. nov. a–c. Female: a. Head and thorax, dorsal view. b. Habitus lateral. c. Head, lateral view. d–f. Male: d. Head and thorax, dorsal view. e. Habitus lateral. f. Head, oblique dorsal view. Abbreviations: sc: scale-like pile; scl: scutellum; r1v: sub-apical vitta of cell r1." figureDoi="http://doi.org/10.5281/zenodo.13209005" httpUri="https://zenodo.org/record/13209005/files/figure.png" pageId="21" pageNumber="22">Fig. 9e</figureCitation>
): metallic green, terga matte black microtrichose sub-laterally, pile sockets as small protuberances, pile white, appressed and very short; sterna metallic green, with longer white appressed pile; sternum 4 with right side more extended than left.
</paragraph>
<paragraph id="70023778D56FE70833924912FC58F8E9" blockId="21.[151,1437,397,1972]" pageId="21" pageNumber="22">
<emphasis id="42C9EB6AD56FE70833924912FEE6F9F5" bold="true" box="[199,306,1621,1648]" italics="true" pageId="21" pageNumber="22">Genitalia</emphasis>
(
<figureCitation id="E8862BFDD56FE70832134911FE12F9F5" box="[326,454,1622,1648]" captionStart="FIGURE 10" captionStartId="23.[151,250,1900,1924]" captionTargetBox="[199,1375,957,1865]" captionTargetId="figure-295@23.[199,1387,957,1877]" captionTargetPageId="23" captionText="FIGURE 10. Orthonevra cthulhu sp. nov., genitalia. a. Female, dorsal view. b. Female, ventral view. c–e. Male: c. Epandrium, lateral view. d. Hypandrium, lateral view. e. Phallus, lateral view. Abbreviations: btp: basal tubular process; cerc: cercus; hypc: ventral carina of the hypandrium; hypo: hypoproct; epi: epiproct; pg: postgonite; sur: surstylus; s7 &amp; 8: sterna 7 &amp; 8; t7 &amp; 8: terga 7 &amp; 8." figureDoi="http://doi.org/10.5281/zenodo.13209007" httpUri="https://zenodo.org/record/13209007/files/figure.png" pageId="21" pageNumber="22">Fig. 10c–e</figureCitation>
): surstylus bent in a 90° on the basal 1/3, and directed more dorsally, base slightly expanded and with a shallow concave notch on dorsal surface, with a rounded apex slightly directed ventrally, with very short inconspicuous pile (
<figureCitation id="E8862BFDD56FE70832A749D9FD87F93D" box="[498,595,1694,1720]" captionStart="FIGURE 10" captionStartId="23.[151,250,1900,1924]" captionTargetBox="[199,1375,957,1865]" captionTargetId="figure-295@23.[199,1387,957,1877]" captionTargetPageId="23" captionText="FIGURE 10. Orthonevra cthulhu sp. nov., genitalia. a. Female, dorsal view. b. Female, ventral view. c–e. Male: c. Epandrium, lateral view. d. Hypandrium, lateral view. e. Phallus, lateral view. Abbreviations: btp: basal tubular process; cerc: cercus; hypc: ventral carina of the hypandrium; hypo: hypoproct; epi: epiproct; pg: postgonite; sur: surstylus; s7 &amp; 8: sterna 7 &amp; 8; t7 &amp; 8: terga 7 &amp; 8." figureDoi="http://doi.org/10.5281/zenodo.13209007" httpUri="https://zenodo.org/record/13209007/files/figure.png" pageId="21" pageNumber="22">Fig. 10c</figureCitation>
); subepandrial sclerite arms widened towards apex; cercus triangular, with acute corners, with distinct pile. Hypandrium compressed medially, with anterior ventral notch through 1/2 the length of the hypandrium, notch with a carina on each side (
<figureCitation id="E8862BFDD56FE708301149A1FC7CF885" box="[836,936,1766,1792]" captionStart="FIGURE 10" captionStartId="23.[151,250,1900,1924]" captionTargetBox="[199,1375,957,1865]" captionTargetId="figure-295@23.[199,1387,957,1877]" captionTargetPageId="23" captionText="FIGURE 10. Orthonevra cthulhu sp. nov., genitalia. a. Female, dorsal view. b. Female, ventral view. c–e. Male: c. Epandrium, lateral view. d. Hypandrium, lateral view. e. Phallus, lateral view. Abbreviations: btp: basal tubular process; cerc: cercus; hypc: ventral carina of the hypandrium; hypo: hypoproct; epi: epiproct; pg: postgonite; sur: surstylus; s7 &amp; 8: sterna 7 &amp; 8; t7 &amp; 8: terga 7 &amp; 8." figureDoi="http://doi.org/10.5281/zenodo.13209007" httpUri="https://zenodo.org/record/13209007/files/figure.png" pageId="21" pageNumber="22">Fig. 10d</figureCitation>
); postgonites rectangular but apex produced as a tapering extension and directed ventrally and slightly medially, pile concentrated on dorsal corner before the extension, baso-median surface forms a carina that ends dorsally in a short triangular projection; phallus hooked apically, widened medially, basal tubular process directed ventrally.
</paragraph>
<paragraph id="70023778D56FE70833924832FD70F831" blockId="21.[151,1437,397,1972]" pageId="21" pageNumber="22">
<emphasis id="42C9EB6AD56FE70833924832FEECF80A" bold="true" box="[199,312,1909,1935]" pageId="21" pageNumber="22">Variation</emphasis>
. Some specimens with tibiae mostly metallic brownish-green;
<specimenCount id="66BBFCF1D56FE70830A94831FB42F815" box="[1020,1174,1910,1936]" count="1" pageId="21" pageNumber="22" type="generic">one specimen</specimenCount>
from
<collectingRegion id="B279F99AD56FE708378E4831FAC8F815" box="[1243,1308,1910,1936]" country="Brazil" name="Goias" pageId="21" pageNumber="22">Goiás</collectingRegion>
has a short triangular projection medially on the surstylus.
</paragraph>
<paragraph id="70023778D56CE70B33924FD0FBB1FDE4" blockId="22.[151,1437,151,609]" pageId="22" pageNumber="23">
FEMALE (
<figureCitation id="E8862BFDD56CE70B321F4FD0FE18FF34" box="[330,460,151,177]" captionStart="FIGURE 10" captionStartId="23.[151,250,1900,1924]" captionTargetBox="[199,1375,957,1865]" captionTargetId="figure-295@23.[199,1387,957,1877]" captionTargetPageId="23" captionText="FIGURE 10. Orthonevra cthulhu sp. nov., genitalia. a. Female, dorsal view. b. Female, ventral view. c–e. Male: c. Epandrium, lateral view. d. Hypandrium, lateral view. e. Phallus, lateral view. Abbreviations: btp: basal tubular process; cerc: cercus; hypc: ventral carina of the hypandrium; hypo: hypoproct; epi: epiproct; pg: postgonite; sur: surstylus; s7 &amp; 8: sterna 7 &amp; 8; t7 &amp; 8: terga 7 &amp; 8." figureDoi="http://doi.org/10.5281/zenodo.13209007" httpUri="https://zenodo.org/record/13209007/files/figure.png" pageId="22" pageNumber="23">Fig. 10a, b</figureCitation>
). Like male except: head metallic green, face with slight convexity in profile; frons with medial longitudinal sulcus starting anterior to the ocellar triangle and extending throughout 2/3 of the length of the frons; vertex small, metallic green but with purple reflections lateral to ocellar triangle; ocellar triangle occupying most of the vertex; occiput not visible laterally on dorsal 1/3, ventral 2/3 homogeneously covered in white microtrichia which gets longer dorsal to the gena, with very inconspicuous pile on dorsal 1/3, with a few sparse pile on ventral 1/4; eyes do not form circle antero-dorsally; tergum 5 wholly metallic. Genitalia: Tergum 7 as two rectangular sclerotized areas. Sternum 7 as a sclerotized strip with pile on apical margin. Tergum 8 sub-triangular with two heavier sclerotized fasciate areas. Sternum 8 large, baso-lateral margins concave, apical margin with large medial ‘V-shaped’ indentation and a sub-lateral concave indentation on each side, wholly pilose. Sternum 9 as half-oval, sclerotized and distinct plate, with medial internal long sclerotized tube connected to the genital opening medially on sternum 9. Epiproct flattened, sub-rectangular, convex apical margin, with two short baso-lateral apodemes, sparsely pilose. Cercus oval, pilose on apical 1/2, positioned ventro-laterally to epiproct. Hypoproct mostly membranous, with triangular shaped sclerotized area, wholly pilose.
</paragraph>
<caption id="24C267F0D56CE70B33C248CDFDB7F86E" ID-DOI="http://doi.org/10.5281/zenodo.13209005" ID-Zenodo-Dep="13209005" httpUri="https://zenodo.org/record/13209005/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,250,1930,1954]" targetBox="[151,1436,633,1907]" targetPageId="22" targetType="figure">
<paragraph id="70023778D56CE70B33C248CDFDB7F86E" blockId="22.[151,1437,1930,2027]" pageId="22" pageNumber="23">
<emphasis id="42C9EB6AD56CE70B33C248CDFED8F827" bold="true" box="[151,268,1930,1954]" pageId="22" pageNumber="23">FIGURE 9</emphasis>
.
<taxonomicName id="B7BD4CFBD56CE70B324D48CCFE34F826" authorityName="Miranda &amp; Soares &amp; Thompson" authorityYear="2024" box="[280,480,1931,1955]" class="Insecta" family="Syrphidae" genus="Orthonevra" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="cthulhu" status="sp. nov.">
<emphasis id="42C9EB6AD56CE70B324D48CCFE34F826" box="[280,480,1931,1955]" italics="true" pageId="22" pageNumber="23">Orthonevra cthulhu</emphasis>
</taxonomicName>
<emphasis id="42C9EB6AD56CE70B32B348CCFDB3F826" bold="true" box="[486,615,1931,1955]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="59FA5611D56CE70B32B348CCFDE2F826" box="[486,566,1931,1955]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
a–c.
</emphasis>
Female:
<emphasis id="42C9EB6AD56CE70B319148CCFD03F826" bold="true" box="[708,727,1931,1955]" pageId="22" pageNumber="23">a.</emphasis>
Head and thorax, dorsal view.
<emphasis id="42C9EB6AD56CE70B375B48CCFBF7F826" bold="true" box="[1038,1059,1931,1955]" pageId="22" pageNumber="23">b.</emphasis>
Habitus lateral.
<emphasis id="42C9EB6AD56CE70B379D48CCFB0EF826" bold="true" box="[1224,1242,1931,1955]" pageId="22" pageNumber="23">c.</emphasis>
Head, lateral view.
<emphasis id="42C9EB6AD56CE70B33C248E9FF15F843" bold="true" box="[151,193,1966,1990]" pageId="22" pageNumber="23">d–f.</emphasis>
Male:
<emphasis id="42C9EB6AD56CE70B325F48E8FECAF842" bold="true" box="[266,286,1967,1991]" pageId="22" pageNumber="23">d.</emphasis>
Head and thorax, dorsal view.
<emphasis id="42C9EB6AD56CE70B310B48E8FDBBF842" bold="true" box="[606,623,1967,1991]" pageId="22" pageNumber="23">e.</emphasis>
Habitus lateral.
<emphasis id="42C9EB6AD56CE70B304C48E9FCFDF843" bold="true" box="[793,809,1966,1990]" pageId="22" pageNumber="23">f.</emphasis>
Head, oblique dorsal view. Abbreviations: sc: scale-like pile; scl: scutellum; r1v: sub-apical vitta of cell r1.
</paragraph>
</caption>
<paragraph id="70023778D56DE70A33924FD0FF09FF50" blockId="23.[151,1437,150,933]" pageId="23" pageNumber="24">
<emphasis id="42C9EB6AD56DE70A33924FD0FEC9FF34" bold="true" box="[199,285,151,177]" pageId="23" pageNumber="24">Length</emphasis>
. Body
<quantity id="B7459A9DD56DE70A32394FD1FDD3FF34" box="[364,519,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.675" metricValueMax="5.1" metricValueMin="4.25" pageId="23" pageNumber="24" unit="mm" value="4.675" valueMax="5.1" valueMin="4.25">4.25–5.10mm</quantity>
(n=4), wing 3.60–4.00mm (n=4); female
<quantity id="B7459A9DD56DE70A30834FD1FBA6FF34" box="[982,1138,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="5.115" metricValueMax="5.52" metricValueMin="4.71" pageId="23" pageNumber="24" unit="mm" value="5.115" valueMax="5.52" valueMin="4.71">4.71–5.52mm</quantity>
(n=5), wing
<quantity id="B7459A9DD56DE70A36544FD1FA48FF34" box="[1281,1436,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="4.1" metricValueMin="3.5" pageId="23" pageNumber="24" unit="mm" value="3.8" valueMax="4.1" valueMin="3.5">3.50–4.10mm</quantity>
(n=5).
</paragraph>
</subSubSection>
<subSubSection id="38A764F3D56DE70A33924F99FDF7FE98" pageId="23" pageNumber="24" type="distribution">
<paragraph id="70023778D56DE70A33924F99FA4DFF7C" blockId="23.[151,1437,150,933]" box="[199,1433,222,249]" pageId="23" pageNumber="24">
<emphasis id="42C9EB6AD56DE70A33924F99FE8DFF7D" bold="true" box="[199,345,222,248]" pageId="23" pageNumber="24">Distribution</emphasis>
.
<collectingCountry id="08AA77E8D56DE70A32324F98FE7FFF7C" box="[359,427,223,249]" name="Brazil" pageId="23" pageNumber="24">Brazil</collectingCountry>
(
<collectingRegion id="B279F99AD56DE70A32EE4F98FDD5FF7C" box="[443,513,223,249]" country="Brazil" name="Ceara" pageId="23" pageNumber="24">Ceará</collectingRegion>
,
<collectingRegion id="B279F99AD56DE70A315E4F98FD7AFF7C" box="[523,686,223,249]" country="Brazil" name="Espirito Santo" pageId="23" pageNumber="24">Espírito Santo</collectingRegion>
,
<collectingRegion id="B279F99AD56DE70A31EC4F98FD29FF7C" box="[697,765,223,249]" country="Brazil" name="Goias" pageId="23" pageNumber="24">Goiás</collectingRegion>
,
<collectingRegion id="B279F99AD56DE70A305D4F98FCADFF7C" box="[776,889,223,249]" country="Brazil" name="Sao Paulo" pageId="23" pageNumber="24">São Paulo</collectingRegion>
and
<collectingRegion id="B279F99AD56DE70A30FA4F98FB56FF7C" box="[943,1154,223,249]" country="Brazil" name="Rio Grande do Sul" pageId="23" pageNumber="24">Rio Grande do Sul</collectingRegion>
) and
<collectingCountry id="08AA77E8D56DE70A37954F98FAFCFF7C" box="[1216,1320,223,249]" name="Paraguay" pageId="23" pageNumber="24">Paraguay</collectingCountry>
(
<figureCitation id="E8862BFDD56DE70A366D4F98FA58FF7C" box="[1336,1420,223,249]" captionStart="FIGURE 39" captionStartId="70.[151,250,1913,1937]" captionTargetBox="[151,1435,669,1889]" captionTargetId="figure-252@70.[151,1435,669,1889]" captionTargetPageId="70" captionText="FIGURE 39. Distribution map for Orthonevra argentina (Brèthes, 1922), O. batman sp. nov., O. chilensis Thompson, 1999, O. cthulhu sp. nov., O. quadristriata (Shannon &amp; Aubertin, 1933), O. roseae sp. nov., O. shannoni (Curran, 1926), and O. thompsoni sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13209085" httpUri="https://zenodo.org/record/13209085/files/figure.png" pageId="23" pageNumber="24">Fig. 39</figureCitation>
).
</paragraph>
<paragraph id="70023778D56DE70A33924E45FDF7FE98" blockId="23.[151,1437,150,933]" box="[199,547,258,285]" pageId="23" pageNumber="24">
<emphasis id="42C9EB6AD56DE70A33924E45FE41FE98" bold="true" box="[199,405,258,285]" pageId="23" pageNumber="24">Altitudinal range</emphasis>
.
<quantity id="B7459A9DD56DE70A32F64E44FDCBFE98" box="[419,543,259,285]" metricMagnitude="2" metricUnit="m" metricValue="4.95" metricValueMax="8.3" metricValueMin="1.6" pageId="23" pageNumber="24" unit="m" value="495.0" valueMax="830.0" valueMin="160.0">160–830m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="38A764F3D56DE70A33924E60FB2CFE0C" pageId="23" pageNumber="24" type="etymology">
<paragraph id="70023778D56DE70A33924E60FB2CFE0C" blockId="23.[151,1437,150,933]" pageId="23" pageNumber="24">
<emphasis id="42C9EB6AD56DE70A33924E60FE9CFEC4" bold="true" box="[199,328,295,321]" pageId="23" pageNumber="24">Etymology</emphasis>
. The specific epithet is derived from the name of an ancient cosmic entity from the work of H.P. Lovecraft, and alludes to the exquisite eye marking pattern on the males (when viewed from the front) of this species that resembles the description of the entity in that work. It should be treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="38A764F3D56DE70A33924ED5FA9BFE28" box="[199,1359,402,429]" pageId="23" pageNumber="24" type="discussion">
<paragraph id="70023778D56DE70A33924ED5FA9BFE28" blockId="23.[151,1437,150,933]" box="[199,1359,402,429]" pageId="23" pageNumber="24">
<emphasis id="42C9EB6AD56DE70A33924ED5FE9CFE29" bold="true" box="[199,328,402,428]" pageId="23" pageNumber="24">Comments</emphasis>
. This species is restricted to central
<collectingCountry id="08AA77E8D56DE70A318A4ED4FCF7FE28" box="[735,803,403,429]" name="Brazil" pageId="23" pageNumber="24">Brazil</collectingCountry>
and
<collectingCountry id="08AA77E8D56DE70A300F4ED4FC16FE28" box="[858,962,403,429]" name="Paraguay" pageId="23" pageNumber="24">Paraguay</collectingCountry>
and to altitudes lower than
<quantity id="B7459A9DD56DE70A37AC4ED4FA9FFE28" box="[1273,1355,403,429]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="m" value="1000.0">1000m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="38A764F3D56DE70A33924EF0FE74FC20" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="70023778D56DE70A33924EF0FC59FE70" blockId="23.[151,1437,150,933]" pageId="23" pageNumber="24">
<emphasis id="42C9EB6AD56DE70A33924EF0FE37FE55" bold="true" box="[199,483,438,465]" pageId="23" pageNumber="24">Type material examined</emphasis>
:
<materialsCitation id="C0D53D25D56DE70A32A54EF0FC5CFE70" collectingDate="1970-09" collectorName="P. C. Elias" country="Brazil" latitude="-19.522472" location="Baixo Guandu" longLatPrecision="1" longitude="-41.019722" pageId="23" pageNumber="24" specimenCount="1" stateProvince="Espirito Santo" typeStatus="holotype">
<collectingCountry id="08AA77E8D56DE70A32A54EF0FD8DFE54" box="[496,601,439,465]" name="Brazil" pageId="23" pageNumber="24">BRAZIL</collectingCountry>
.
<collectingRegion id="B279F99AD56DE70A31354EF0FCD5FE54" box="[608,769,439,465]" country="Brazil" name="Espirito Santo" pageId="23" pageNumber="24">Espírito Santo</collectingRegion>
,
<location id="756261A3D56DE70A305C4EF0FC7EFE54" LSID="urn:lsid:plazi:treatment:F814866ED56FE70A33C24FD0FE74FC20:756261A3D56DE70A305C4EF0FC7EFE54" box="[777,938,439,465]" country="Brazil" latitude="-19.522472" longLatPrecision="1" longitude="-41.019722" name="Baixo Guandu" pageId="23" pageNumber="24" stateProvince="Espirito Santo">Baixo Guandu</location>
[ca
<geoCoordinate id="158951BFD56DE70A30804EF0FBBDFE54" box="[981,1129,439,465]" degrees="19" direction="south" minutes="31" orientation="latitude" pageId="23" pageNumber="24" precision="1" seconds="20.9" value="-19.522472">19°31'20.9&quot;S</geoCoordinate>
<geoCoordinate id="158951BFD56DE70A373B4EF0FAC5FE54" box="[1134,1297,439,465]" degrees="41" direction="west" minutes="01" orientation="longitude" pageId="23" pageNumber="24" precision="1" seconds="11.0" value="-41.019722">41°01'11.0&quot;W</geoCoordinate>
], ??.
<date id="040311B8D56DE70A366B4EF0FA4CFE54" box="[1342,1432,439,465]" pageId="23" pageNumber="24" value="1970-09">
<collectingDate id="1447E850D56DE70A366B4EF0FA4CFE54" box="[1342,1432,439,465]" pageId="23" pageNumber="24" value="1970-09">ix.1970</collectingDate>
</date>
,
<collectorName id="DD4852AED56DE70A33C24E9CFEDEFE70" box="[151,266,475,501]" pageId="23" pageNumber="24">P. C. Elias</collectorName>
(male
<typeStatus id="AF0689DAD56DE70A320D4E9CFE6EFE70" box="[344,442,475,501]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
<taxonomicName id="B7BD4CFBD56DE70A32944E9CFCDBFE70" authority="MZUSP, USNMENT" authorityName="MZUSP, USNMENT" box="[449,783,475,501]" class="Insecta" family="Syrphidae" genus="Orthonevra" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="cthulhu">
<emphasis id="42C9EB6AD56DE70A32944E9CFDC1FE70" box="[449,533,475,501]" italics="true" pageId="23" pageNumber="24">cthulhu</emphasis>
MZUSP, USNMENT
</taxonomicName>
00028638)
</materialsCitation>
.
</paragraph>
<paragraph id="70023778D56DE70A33924EB8FE74FC20" blockId="23.[151,1437,150,933]" pageId="23" pageNumber="24">
<materialsCitation id="C0D53D25D56DE70A33924EB8FDE7FD9C" box="[199,563,510,537]" country="Brazil" location="BRAZIL" pageId="23" pageNumber="24" specimenCount="1" typeStatus="paratype">
<emphasis id="42C9EB6AD56DE70A33924EB8FE6EFD9D" bold="true" box="[199,442,510,537]" pageId="23" pageNumber="24">
<typeStatus id="AF0689DAD56DE70A33924EB8FEEBFD9C" box="[199,319,511,537]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
examined
</emphasis>
:
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</materialsCitation>
.
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<collectingRegion id="B279F99AD56DE70A316B4EB8FD56FD9C" box="[574,642,511,537]" country="Brazil" name="Ceara" pageId="23" pageNumber="24">Ceará</collectingRegion>
,
<collectingCounty id="99634FF4D56DE70A31DB4EB8FD3EFD9C" box="[654,746,511,537]" pageId="23" pageNumber="24">Ubajara</collectingCounty>
,
<collectingMunicipality id="9066AD02D56DE70A31A04EB8FBD1FD9C" box="[757,1029,511,537]" pageId="23" pageNumber="24">Parque Nac. de Ubajara</collectingMunicipality>
,
<location id="756261A3D56DE70A37454EB8FAC5FD9C" LSID="urn:lsid:plazi:treatment:F814866ED56FE70A33C24FD0FE74FC20:756261A3D56DE70A37454EB8FAC5FD9C" box="[1040,1297,511,537]" country="Brazil" county="Ubajara" latitude="-3.8369443" longLatPrecision="21" longitude="-40.909725" municipality="Parque Nac. de Ubajara" name="Cachoeira do Cafundo" pageId="23" pageNumber="24" stateProvince="Ceara">Cachoeira do Cafundó</location>
,
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,
<collectorName id="DD4852AED56DE70A32784D64FDDCFDB8" box="[301,520,547,573]" pageId="23" pageNumber="24">Armadilha Malaise</collectorName>
,
<date id="040311B8D56DE70A31404D64FD71FDB8" box="[533,677,547,573]" pageId="23" pageNumber="24" value="2013-02-01" valueMax="2013-02-14" valueMin="2013-02-01">
<collectingDate id="1447E850D56DE70A31404D64FD71FDB8" box="[533,677,547,573]" pageId="23" pageNumber="24" value="2013-02-01" valueMax="2013-02-14" valueMin="2013-02-01">1–14.ii.2013</collectingDate>
</date>
,
<collectorName id="DD4852AED56DE70A31E74D64FCE4FDB8" box="[690,816,547,573]" pageId="23" pageNumber="24">J.A. Rafael</collectorName>
&amp;
<collectorName id="DD4852AED56DE70A30024D64FB81FDB8" box="[855,1109,547,573]" pageId="23" pageNumber="24">F. Limeira-de-Oliveira</collectorName>
(
<specimenCount id="66BBFCF1D56DE70A373C4D64FB1EFDB8" box="[1129,1226,547,573]" count="1" pageId="23" pageNumber="24" type="female">1 female</specimenCount>
<collectionCode id="16ACAFBDD56DE70A37874D64FAFCFDB8" box="[1234,1320,547,573]" pageId="23" pageNumber="24">CZMA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C0D53D25D56DE70A36684D64FBD7FD00" collectingDate="2012-12-16" collectingDateMax="2012-12-31" collectingDateMin="2012-12-16" collectionCode="CZMA" collectorName="F. Limeira-de-Oliveira &amp; J. S. Pinto Junior" country="Brazil" county="Ubajara" latitude="-3.8369443" location="Cachoeira do Cafundo" longLatPrecision="21" longitude="-40.909725" municipality="Parque Nac. de Ubajara" pageId="23" pageNumber="24" specimenCount="4" specimenCount-female="4" stateProvince="Ceara" typeStatus="paratype">
<collectingCounty id="99634FF4D56DE70A36684D64FA4DFDB8" box="[1341,1433,547,573]" pageId="23" pageNumber="24">Ubajara</collectingCounty>
,
<collectingMunicipality id="9066AD02D56DE70A33C24D00FE65FDE4" box="[151,433,583,609]" pageId="23" pageNumber="24">Parque Nac. de Ubajara</collectingMunicipality>
,
<location id="756261A3D56DE70A32954D00FD13FDE4" LSID="urn:lsid:plazi:treatment:F814866ED56FE70A33C24FD0FE74FC20:756261A3D56DE70A32954D00FD13FDE4" box="[448,711,583,609]" country="Brazil" county="Ubajara" latitude="-3.8369443" longLatPrecision="21" longitude="-40.909725" municipality="Parque Nac. de Ubajara" name="Cachoeira do Cafundo" pageId="23" pageNumber="24" stateProvince="Ceara">Cachoeira do Cafundó</location>
,
<geoCoordinate id="158951BFD56DE70A31824D01FC80FDE4" box="[727,852,582,609]" degrees="03" direction="south" minutes="50" orientation="latitude" pageId="23" pageNumber="24" precision="15" seconds="13" value="-3.8369443">03º50'13&quot;S</geoCoordinate>
<geoCoordinate id="158951BFD56DE70A300A4D01FC33FDE4" box="[863,999,582,609]" degrees="40" direction="west" minutes="54" orientation="longitude" pageId="23" pageNumber="24" precision="15" seconds="35" value="-40.909725">40º54'35&quot;W</geoCoordinate>
,
<collectorName id="DD4852AED56DE70A30A04D00FB07FDE4" box="[1013,1235,583,609]" pageId="23" pageNumber="24">Armadilha Malaise</collectorName>
, 18–30.xi &amp;
<date id="040311B8D56DE70A36274D00FECDFD00" pageId="23" pageNumber="24" value="2012-12-16" valueMax="2012-12-31" valueMin="2012-12-16">
<collectingDate id="1447E850D56DE70A36274D00FECDFD00" pageId="23" pageNumber="24" value="2012-12-16" valueMax="2012-12-31" valueMin="2012-12-16">16– 31.xii.2012</collectingDate>
</date>
,
<collectorName id="DD4852AED56DE70A327C4D2CFDFDFD00" box="[297,553,619,645]" pageId="23" pageNumber="24">F. Limeira-de-Oliveira</collectorName>
&amp;
<collectorName id="DD4852AED56DE70A31034D2CFCC2FD00" box="[598,790,619,645]" pageId="23" pageNumber="24">J.S. Pinto Júnior</collectorName>
(
<specimenCount id="66BBFCF1D56DE70A307C4D2CFC4EFD00" box="[809,922,619,645]" count="4" pageId="23" pageNumber="24" type="female">4 females</specimenCount>
<collectionCode id="16ACAFBDD56DE70A30F04D2CFC2FFD00" box="[933,1019,619,645]" pageId="23" pageNumber="24">CZMA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C0D53D25D56DE70A37464D2CFB38FD2C" collectingDate="1970-09" collectionCode="MZUSP" collectorName="P. C. Elias" country="Brazil" latitude="-19.522472" location="Baixo Guandu" longLatPrecision="1" longitude="-41.019722" pageId="23" pageNumber="24" specimenCode="USNMENT01492702-3" specimenCount="2" specimenCount-male="2" stateProvince="Espirito Santo" typeStatus="paratype">
<collectingRegion id="B279F99AD56DE70A37464D2CFB6DFD00" box="[1043,1209,619,645]" country="Brazil" name="Espirito Santo" pageId="23" pageNumber="24">Espírito Santo</collectingRegion>
,
<location id="756261A3D56DE70A379D4D2CFABBFD00" LSID="urn:lsid:plazi:treatment:F814866ED56FE70A33C24FD0FE74FC20:756261A3D56DE70A379D4D2CFABBFD00" box="[1224,1391,619,645]" country="Brazil" latitude="-19.522472" longLatPrecision="1" longitude="-41.019722" name="Baixo Guandu" pageId="23" pageNumber="24" stateProvince="Espirito Santo">Baixo Guandu</location>
[ca
<geoCoordinate id="158951BFD56DE70A33C24DC8FEFFFD2C" box="[151,299,655,681]" degrees="19" direction="south" minutes="31" orientation="latitude" pageId="23" pageNumber="24" precision="1" seconds="20.9" value="-19.522472">19°31'20.9&quot;S</geoCoordinate>
<geoCoordinate id="158951BFD56DE70A32604DC8FE0CFD2C" box="[309,472,655,681]" degrees="41" direction="west" minutes="01" orientation="longitude" pageId="23" pageNumber="24" precision="1" seconds="11.0" value="-41.019722">41°01'11.0&quot;W</geoCoordinate>
], ??.
<date id="040311B8D56DE70A315E4DC8FDB1FD2C" box="[523,613,655,681]" pageId="23" pageNumber="24" value="1970-09">
<collectingDate id="1447E850D56DE70A315E4DC8FDB1FD2C" box="[523,613,655,681]" pageId="23" pageNumber="24" value="1970-09">ix.1970</collectingDate>
</date>
,
<collectorName id="DD4852AED56DE70A31264DC8FD3FFD2C" box="[627,747,655,681]" pageId="23" pageNumber="24">P. C. Elias</collectorName>
(
<specimenCount id="66BBFCF1D56DE70A31AB4DC8FC83FD2C" box="[766,855,655,681]" count="2" pageId="23" pageNumber="24" type="male">2 males</specimenCount>
<collectionCode id="16ACAFBDD56DE70A30344DC8FC6AFD2C" box="[865,958,655,681]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="24">MZUSP</collectionCode>
,
<specimenCode id="201B9F03D56DE70A30994DC8FB31FD2C" box="[972,1253,655,681]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="24">USNMENT01492702–3</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="C0D53D25D56DE70A37AE4DC8FE80FD74" collectingDate="2019-07-29" collectionCode="INPA" collectorName="Armadilha Luminosa &amp; Col. Lopes, W. R." country="Brazil" county="Varjao" latitude="-17.06261" location="Sra. Aparecida" longLatPrecision="1" longitude="-49.644253" municipality="Chacara" pageId="23" pageNumber="24" specimenCount="4" specimenCount-male="4" stateProvince="Goias" typeStatus="paratype">
<collectingRegion id="B279F99AD56DE70A37AE4DC8FAEBFD2C" box="[1275,1343,655,681]" country="Brazil" name="Goias" pageId="23" pageNumber="24">Goiás</collectingRegion>
,
<collectingCounty id="99634FF4D56DE70A36194DC8FA4CFD2C" box="[1356,1432,655,681]" pageId="23" pageNumber="24">Varjão</collectingCounty>
,
<collectingMunicipality id="9066AD02D56DE70A33C24DF4FF27FD48" box="[151,243,691,717]" pageId="23" pageNumber="24">Chácara</collectingMunicipality>
N.
<location id="756261A3D56DE70A324A4DF4FE11FD48" LSID="urn:lsid:plazi:treatment:F814866ED56FE70A33C24FD0FE74FC20:756261A3D56DE70A324A4DF4FE11FD48" box="[287,453,691,717]" country="Brazil" county="Varjao" latitude="-17.06261" longLatPrecision="1" longitude="-49.644253" municipality="Chacara" name="Sra. Aparecida" pageId="23" pageNumber="24" stateProvince="Goias">Sra. Aparecida</location>
[ca
<geoCoordinate id="158951BFD56DE70A32AD4DF5FD58FD48" box="[504,652,690,717]" degrees="17" direction="south" minutes="03" orientation="latitude" pageId="23" pageNumber="24" precision="1" seconds="45.4" value="-17.06261">17°03'45.4&quot;S</geoCoordinate>
<geoCoordinate id="158951BFD56DE70A31C14DF4FCECFD48" box="[660,824,691,717]" degrees="49" direction="west" minutes="38" orientation="longitude" pageId="23" pageNumber="24" precision="1" seconds="39.3" value="-49.644253">49°38'39.3&quot;W</geoCoordinate>
],
<collectorName id="DD4852AED56DE70A301F4DF4FBE9FD48" box="[842,1085,691,717]" pageId="23" pageNumber="24">Armadilha Luminosa</collectorName>
,
<date id="040311B8D56DE70A371F4DF4FB18FD48" box="[1098,1228,691,717]" pageId="23" pageNumber="24" value="2019-07-29">
<collectingDate id="1447E850D56DE70A371F4DF4FB18FD48" box="[1098,1228,691,717]" pageId="23" pageNumber="24" value="2019-07-29">29.vii.2019</collectingDate>
</date>
,
<collectorName id="DD4852AED56DE70A378C4DF4FA49FD48" box="[1241,1437,691,717]" pageId="23" pageNumber="24">Col. Lopes, W.R.</collectorName>
(
<specimenCount id="66BBFCF1D56DE70A33F54D90FF2FFD74" box="[160,251,727,753]" count="4" pageId="23" pageNumber="24" type="male">4 males</specimenCount>
<collectionCode id="16ACAFBDD56DE70A325D4D90FE98FD74" box="[264,332,727,753]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="23" pageNumber="24">INPA</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C0D53D25D56DE70A32314D90FE4FFC90" collectingDate="1961-02-25" collectionCode="MZUSP" collectorName="K. Lenko" country="Brazil" latitude="-23.493084" location="Barueri" longLatPrecision="1" longitude="-46.908" pageId="23" pageNumber="24" specimenCode="USNMENT01492704" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="paratype">
<collectingRegion id="B279F99AD56DE70A32314D90FE09FD74" box="[356,477,727,753]" country="Brazil" name="Sao Paulo" pageId="23" pageNumber="24">São Paulo</collectingRegion>
,
<location id="756261A3D56DE70A32B94D90FD94FD74" LSID="urn:lsid:plazi:treatment:F814866ED56FE70A33C24FD0FE74FC20:756261A3D56DE70A32B94D90FD94FD74" box="[492,576,727,753]" country="Brazil" latitude="-23.493084" longLatPrecision="1" longitude="-46.908" name="Barueri" pageId="23" pageNumber="24" stateProvince="Sao Paulo">Barueri</location>
[ca
<geoCoordinate id="158951BFD56DE70A312F4D91FCDBFD74" box="[634,783,726,753]" degrees="23" direction="south" minutes="29" orientation="latitude" pageId="23" pageNumber="24" precision="1" seconds="35.1" value="-23.493084">23°29'35.1&quot;S</geoCoordinate>
<geoCoordinate id="158951BFD56DE70A304E4D91FC6BFD74" box="[795,959,726,753]" degrees="46" direction="west" minutes="54" orientation="longitude" pageId="23" pageNumber="24" precision="1" seconds="28.8" value="-46.908">46°54'28.8&quot;W</geoCoordinate>
],
<date id="040311B8D56DE70A30834D91FB9EFD74" box="[982,1098,726,753]" pageId="23" pageNumber="24" value="1961-02-25">
<collectingDate id="1447E850D56DE70A30834D91FB9EFD74" box="[982,1098,726,753]" pageId="23" pageNumber="24" value="1961-02-25">25.ii.1961</collectingDate>
</date>
,
<collectorName id="DD4852AED56DE70A370E4D90FB1EFD74" box="[1115,1226,727,753]" pageId="23" pageNumber="24">K. Lenko</collectorName>
(
<specimenCount id="66BBFCF1D56DE70A37B54D90FAFBFD74" box="[1248,1327,727,753]" count="1" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
<collectionCode id="16ACAFBDD56DE70A366E4D90FA4CFD74" box="[1339,1432,727,753]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="24">MZUSP</collectionCode>
,
<specimenCode id="201B9F03D56DE70A33C24DBCFE41FC90" box="[151,405,763,789]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="24">USNMENT01492704</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="C0D53D25D56DE70A32FD4DBCFDA1FCBC" collectingDate="1946-01" collectionCode="MZUSP" collectorName="Faz. Sao Joao" country="Brazil" latitude="-20.594667" location="Orida Verda" longLatPrecision="1" longitude="-49.289555" pageId="23" pageNumber="24" specimenCode="USNMENT01492711" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo" typeStatus="paratype">
<location id="756261A3D56DE70A32FD4DBCFDE6FC90" LSID="urn:lsid:plazi:treatment:F814866ED56FE70A33C24FD0FE74FC20:756261A3D56DE70A32FD4DBCFDE6FC90" box="[424,562,763,789]" country="Brazil" latitude="-20.594667" longLatPrecision="1" longitude="-49.289555" name="Orida Verda" pageId="23" pageNumber="24" stateProvince="Sao Paulo">Orida Verda</location>
[
<emphasis id="42C9EB6AD56DE70A31114DBCFDB7FC91" box="[580,611,763,788]" italics="true" pageId="23" pageNumber="24">sic</emphasis>
. Onda Verde, ca
<geoCoordinate id="158951BFD56DE70A307E4DBDFC6BFC90" box="[811,959,762,789]" degrees="20" direction="south" minutes="35" orientation="latitude" pageId="23" pageNumber="24" precision="1" seconds="40.8" value="-20.594667">20°35'40.8&quot;S</geoCoordinate>
<geoCoordinate id="158951BFD56DE70A309D4DBCFBB8FC90" box="[968,1132,763,789]" degrees="49" direction="west" minutes="17" orientation="longitude" pageId="23" pageNumber="24" precision="1" seconds="22.4" value="-49.289555">49°17'22.4&quot;W</geoCoordinate>
],
<collectorName id="DD4852AED56DE70A37D54DBCFAF5FC90" box="[1152,1313,763,789]" pageId="23" pageNumber="24">Faz. Sao Joao</collectorName>
, ??.
<date id="040311B8D56DE70A36184DBCFA4CFC90" box="[1357,1432,763,789]" pageId="23" pageNumber="24" value="1946-01">
<collectingDate id="1447E850D56DE70A36184DBCFA4CFC90" box="[1357,1432,763,789]" pageId="23" pageNumber="24" value="1946-01">i.1946</collectingDate>
</date>
, (
<specimenCount id="66BBFCF1D56DE70A33F74C58FED6FCBC" box="[162,258,799,825]" count="1" pageId="23" pageNumber="24" type="female">1 female</specimenCount>
<collectionCode id="16ACAFBDD56DE70A325C4C58FEB2FCBC" box="[265,358,799,825]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="24">MZUSP</collectionCode>
,
<specimenCode id="201B9F03D56DE70A32274C58FDBAFCBC" box="[370,622,799,825]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="23" pageNumber="24">USNMENT01492711</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="C0D53D25D56DE70A31D44C58FF3FFC04" collectingDate="2021-09" collectorName="Distrito Capital" country="Paraguay" latitude="-27.919498" location="Esmeralda" longLatPrecision="1" longitude="-51.34275" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Rio Grande do Sul" typeStatus="paratype">
<collectingRegion id="B279F99AD56DE70A31D44C58FC82FCBC" box="[641,854,799,825]" country="Brazil" name="Rio Grande do Sul" pageId="23" pageNumber="24">Rio Grande do Sul</collectingRegion>
,
<location id="756261A3D56DE70A30344C58FC0EFCBC" LSID="urn:lsid:plazi:treatment:F814866ED56FE70A33C24FD0FE74FC20:756261A3D56DE70A30344C58FC0EFCBC" box="[865,986,799,825]" country="Paraguay" latitude="-27.919498" longLatPrecision="1" longitude="-51.34275" name="Esmeralda" pageId="23" pageNumber="24" stateProvince="Rio Grande do Sul">Esmeralda</location>
,
<geoCoordinate id="158951BFD56DE70A30B34C59FBAEFCBC" box="[998,1146,798,825]" degrees="27" direction="south" minutes="55" orientation="latitude" pageId="23" pageNumber="24" precision="1" seconds="10.2" value="-27.919498">27°55'10.2&quot;S</geoCoordinate>
<geoCoordinate id="158951BFD56DE70A37D74C59FAF4FCBC" box="[1154,1312,798,825]" degrees="51" direction="west" minutes="20" orientation="longitude" pageId="23" pageNumber="24" precision="1" seconds="33.9" value="-51.34275">51°20'33.9&quot;W</geoCoordinate>
, iv.
<date id="040311B8D56DE70A36124C58FA48FCBC" box="[1351,1436,799,825]" pageId="23" pageNumber="24" value="2021-09">
<collectingDate id="1447E850D56DE70A36124C58FA48FCBC" box="[1351,1436,799,825]" pageId="23" pageNumber="24" value="2021-09">ix.2021</collectingDate>
</date>
(
<specimenCount id="66BBFCF1D56DE70A33F74C04FF2BFCD8" box="[162,255,835,861]" count="1" pageId="23" pageNumber="24" type="male">1 male</specimenCount>
, photographic record) [https://www.inaturalist.org/observations/93639307].
<collectingCountry id="08AA77E8D56DE70A37C34C04FAF9FCD8" box="[1174,1325,835,861]" name="Paraguay" pageId="23" pageNumber="24">PARAGUAY</collectingCountry>
.
<collectorName id="DD4852AED56DE70A36124C04FF3FFC04" pageId="23" pageNumber="24">Distrito Capital</collectorName>
</materialsCitation>
,
<materialsCitation id="C0D53D25D56DE70A33AF4C20FE4FFC20" collectionCode="ZMHU" collectorName="J. D. Anisits" country="Paraguay" latitude="-25.244833" location="Assuncion" longLatPrecision="1" longitude="-57.571472" pageId="23" pageNumber="24" specimenCode="USNMENT01492690" specimenCount="1" specimenCount-male="1" stateProvince="Asuncion" typeStatus="paratype">
<collectingRegion id="B279F99AD56DE70A33AF4C20FEA4FC04" box="[250,368,871,897]" country="Paraguay" name="Asuncion" pageId="23" pageNumber="24">Assuncion</collectingRegion>
[ca
<geoCoordinate id="158951BFD56DE70A32FE4C21FDEBFC04" box="[427,575,870,897]" degrees="25" direction="south" minutes="14" orientation="latitude" pageId="23" pageNumber="24" precision="1" seconds="41.4" value="-25.244833">25°14'41.4&quot;S</geoCoordinate>
<geoCoordinate id="158951BFD56DE70A311E4C21FD3BFC04" box="[587,751,870,897]" degrees="57" direction="west" minutes="34" orientation="longitude" pageId="23" pageNumber="24" precision="1" seconds="17.3" value="-57.571472">57°34'17.3&quot;W</geoCoordinate>
],
<collectorName id="DD4852AED56DE70A30534C20FCE8FC04" box="[774,828,871,897]" pageId="23" pageNumber="24">Villa</collectorName>
[
<collectorName id="DD4852AED56DE70A30054C20FC40FC04" box="[848,916,871,897]" pageId="23" pageNumber="24">Nova</collectorName>
], 25.xi.[19]05,
<collectorName id="DD4852AED56DE70A371C4C20FB04FC04" box="[1097,1232,871,897]" pageId="23" pageNumber="24">J.D. Anisits</collectorName>
(
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<collectionCode id="16ACAFBDD56DE70A36164C20FA4CFC04" box="[1347,1432,871,897]" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="23" pageNumber="24">ZMHU</collectionCode>
,
<specimenCode id="201B9F03D56DE70A33C24CCCFE41FC20" box="[151,405,907,933]" collectionCode="ZMHU" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="23" pageNumber="24">USNMENT01492690</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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