177 lines
19 KiB
XML
177 lines
19 KiB
XML
<document id="86E27BF8640104B58FCAC0E6E3C5F59C" ID-DOI="http://dx.doi.org/10.3897/AfrInvertebr.57.8696" ID-GBIF-Dataset="dffce400-a1b8-4e08-863a-5082c6745bf2" ID-Pensoft-Pub="2305-2562-1-67" ID-ZooBank="17A94A5084C2469DAE1DA9E2A78C502B" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="2305-2562-1-67" ModsDocOrigin="African Invertebrates 57 (1)" ModsDocTitle="A review of the genus Gibbasilus Londt, 1986 in southern Africa (Diptera, Asilidae)" checkinTime="1468017305404" checkinUser="pensoft" docAuthor="Londt, Jason G. H." docDate="2016" docId="B915557E8632EDD0EE680D6D1B7AA037" docLanguage="en" docName="AfrInvert 57(1): 67-81" docOrigin="African Invertebrates 57 (1)" docSource="http://dx.doi.org/10.3897/AfrInvertebr.57.8696" docTitle="Gibbasilus alboala Londt, 2016, sp. n." docType="treatment" docUuid="7870527B-1D2D-4F8D-9103-65C45BE6626D" docUuidSource="ZooBank" docVersion="7" lastPageNumber="70" masterDocId="2A2BBE684276A033A53B5268FFFF2036" masterDocTitle="A review of the genus Gibbasilus Londt, 1986 in southern Africa (Diptera, Asilidae)" masterLastPageNumber="81" masterPageNumber="67" pageNumber="69" updateTime="1701315758028" updateUser="plazi">
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<mods:title id="7D45141CD22E6F33F3999A7689BF7985">A review of the genus Gibbasilus Londt, 1986 in southern Africa (Diptera, Asilidae)</mods:title>
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<mods:name id="8E4970682553F2686FEAE91C1A88B22A" type="personal">
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<mods:namePart id="8DBD19CA773DC156B4B6F2F2637A5C5D">Londt, Jason G. H.</mods:namePart>
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<mods:title id="942F3341750C2C916B71FB58AAB41944">African Invertebrates</mods:title>
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<mods:part id="6C7BF6F69EB838AA59002ADD755A555C">
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<mods:date id="4274664A50CB8A75AC1A0DBD56A5CC36">2016</mods:date>
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<mods:number id="3614031A76404D639A7D36C1A7384836">57</mods:number>
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<mods:number id="06A8324C67A25A47F36BA801EE5EE72C">1</mods:number>
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<mods:url id="C2286696A6A498EB33E931B6277C2D8A">http://dx.doi.org/10.3897/AfrInvertebr.57.8696</mods:url>
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<mods:classification id="CE5FAFC4A9D9A21885962ED730E3490A">journal article</mods:classification>
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<mods:identifier id="DEF58C6ECF0D097404B5F3C8E0F9B27D" type="DOI">http://dx.doi.org/10.3897/AfrInvertebr.57.8696</mods:identifier>
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<treatment id="B915557E8632EDD0EE680D6D1B7AA037" ID-GBIF-Taxon="127879936" LSID="urn:lsid:zoobank.org:act:7870527B-1D2D-4F8D-9103-65C45BE6626D" httpUri="http://treatment.plazi.org/id/B915557E8632EDD0EE680D6D1B7AA037" lastPageId="3" lastPageNumber="70" pageId="2" pageNumber="69">
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<subSubSection id="934BA58F10C983C3B5179085ED6A8368" pageId="2" pageNumber="69" type="multiple">
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<paragraph id="9382CFC82FFFF447CC8E7615F1530116" pageId="2" pageNumber="69">Taxon classification Animalia Diptera Asilidae</paragraph>
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</subSubSection>
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<subSubSection id="91F47AB53D0A67459123F0241ECDF707" pageId="2" pageNumber="69" type="nomenclature">
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<paragraph id="9BECB7F2DBC6829DED787CCE318D4CA2" pageId="2" pageNumber="69">
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<taxonomicName id="1C201323BCD9902F663F5FB5DB957110" LSID="http://zoobank.org/7870527B-1D2D-4F8D-9103-65C45BE6626D" class="Insecta" family="Asilidae" genus="Gibbasilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbasilus alboala" order="Diptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="alboala">Gibbasilus alboala</taxonomicName>
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<taxonomicNameLabel id="1818332536DE34E8805BC6E9113AF331" pageId="2" pageNumber="69">sp. n.</taxonomicNameLabel>
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Figs 2, 3-6, 20B
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</paragraph>
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</subSubSection>
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<subSubSection id="8D63162959550F620B3A4347599EEE5D" pageId="2" pageNumber="69" type="etymology">
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<paragraph id="2582A5A6358BAAF23D2D4678DE8AA47D" pageId="2" pageNumber="69">Etymology.</paragraph>
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<paragraph id="4D6CBCFEFE79B7D79CB4B0D62A3507C4" pageId="2" pageNumber="69">L. albus - white, ala - f. wing; refers to the milky white, opaque bases to the wings.</paragraph>
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</subSubSection>
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<subSubSection id="5C2BD009A81EF55524EA3605F7A7CBDC" lastPageId="3" lastPageNumber="70" pageId="2" pageNumber="69" type="description">
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<paragraph id="F7D017A7104EAF5D81E60801FF918FAF" pageId="2" pageNumber="69">Description.</paragraph>
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<paragraph id="EC84508C26C73DAC6DB8A6E3DAE8431F" pageId="2" pageNumber="69">Based on 2♂ specimens, ♀ unknown.</paragraph>
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<paragraph id="22AA014091A3C95E7AF16A591866ADB1" pageId="2" pageNumber="69">Head: Black, fine silver pruinose, black, pale yellow and white setose. Antenna: Black, fine silver pruinose, scape and pedicel black setose. Segmental ratios (scape as 1) = 1: 0.6: 1.2: 0.7 (elements of style = 0.1, 0.5, 0.1). Style composed of three elements (small basal segment-like element, long middle rod-like element, terminal seta-like sensory element). Face black, fine dull silver pruinose, profile plane (straight, no gibbosity evident). Mystax long, well-developed, extending from epistomal margin to antennal sockets, macrosetae pale yellow along epistomal margin, otherwise all black. Frons and vertex black, dull silver pruinose, fine, long, black setose. Ocellar tubercle with long, fine, black setae. Occipital region black, silver pruinose, black setose dorsally, pale yellow laterally, fine white setose ventrally. Palpi 2-segmented, segment 1 white setose, 2 black setose. Proboscis shiny black, fine white setose.</paragraph>
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<paragraph id="A1157C4E8B1ADDF9D9C199D1E1D23B35" pageId="2" pageNumber="69">
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Thorax: Dark red-brown to black, fine, dull, silver pruinose, black, pale orange, pale yellow and white setose. Pronotum black, white setose. Mesonotum black, entirely dull silver pruinose. Acrostichals numerous, mane-like, predominantly black except for few white setae anteriorly and a cluster of fine white setae posteriorly. Dorsocentrals well-developed, black, extending both anterior and posterior of transverse suture. Mesonotal macrosetae: Strong, pale orange, 2 npl, 2 spal, 1 pal. Scutellum black, entirely dull silver pruinose; disc fine white setose, 2 long, black apical scutellar macrosetae. Pleura: Dark red-brown to black, entirely dull silver pruinose, pale yellow and white setose. Katatergal setae weak, pale yellowish. Anatergites uniformly dull silver pruinose, asetose. Mediotergite weakly fine white setose medially. Legs: Coxae dark red-brown to black, fine, dull silver pruinose, fine white setose. Trochanters shiny black, apruinose, fine white setose. Femora slightly inflated, dark red-brown to black except for narrow orange distal tip, macrosetae mostly yellowish (except for 3-5 black proximoventral setae on fore femora). Tibiae mostly brown-orange becoming progressively darker distally, macrosetae mostly orange (a few black at distal tip). Tarsi dark red-brown with black and orange macrosetae. Claws dark red-brown to black, pulvilli and empodia well developed, orange. Wings (Fig. 2): Holotype 6.7
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<normalizedToken id="A3DA011D1B09EA2C7E96B79F9E809002" originalValue="×">x</normalizedToken>
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2.5 mm, paratype 6.1
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<normalizedToken id="5975B029E7615ACCBBCA55CBB68C940C" originalValue="×">x</normalizedToken>
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2.2 mm. Veins brown, cells r1 m3 and cua closed and stalked, membrane unstained, entirely transparent except for partly opaque, milky proximal region (including whole of cells bm, cua, and cup and proximal part of br), microtrichia absent. Halter pale yellow to orange.
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</paragraph>
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<caption id="C84F13ADF5988AE3E4B3285CCE247351" pageId="2" pageNumber="69">
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<paragraph id="33FB3258E3291A5681E96941BD77C115" pageId="2" pageNumber="69">
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Figure 2.
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<taxonomicName id="EF44511E0D35A2BFA6025244DBD56F71" class="Insecta" family="Asilidae" genus="Gibbasilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbasilus alboala" order="Diptera" pageId="2" pageNumber="69" phylum="Arthropoda" rank="species" species="alboala">Gibbasilus alboala</taxonomicName>
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sp. n. ♂ wing (note backlighting obliterates opaque, milky basal membranes).
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</paragraph>
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</caption>
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<paragraph id="EEFCF830F2C7AF0A1F1D40B34A13BEB3" lastPageId="3" lastPageNumber="70" pageId="2" pageNumber="69">
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Abdomen: Entirely dark red-brown to black, extensively dull silver pruinose, except lateral margins of T2 and T3 which are shiny apruinose. Terga fine short white setose, T1-4 with 1-2 laterally situated pale yellow macrosetae (progressively diminishing in size towards terminalia). Sterna fine, longish setose. ♂ genitalia (Figs 3-6): Epandrium
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<pageBreakToken id="2AFEB39955BC3DBC8ED208FC2F4F9905" pageId="3" pageNumber="70" start="start">about</pageBreakToken>
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three times longer than deep in lateral view, with widest part proximal to mid-length; moderately broadly rounded distally; lobes gradually diverging distally in dorsal view. Gonocoxite not quite ⅓ as long as epandrium, smoothly rounded distally. Gonostylus elongate, almost half the length of epandrium, of approximately same width for entire length in ventral view, distally with a subtriangular upwardly directed process, best appreciated in lateral view. Hypandrium short, almost three times wider than long in ventral view, distal margin very slightly indented medially. Aedeagus long and thin, slightly dorsoventrally compressed, S-shaped distally, terminating in a 3-pronged tip. Posterior margin of S8 slightly indented medially with a terminal, subrectangular, dorsoventrally compressed process projecting to approximately midlength of hypandrium. ♀ unknown.
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</paragraph>
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<caption id="007D18FC7ED33EF89E1D70BB58129EC0" pageId="3" pageNumber="70">
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<paragraph id="B78C926AFEB33EF192DC01F7B6CF5BC9" pageId="3" pageNumber="70">
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Figures 3-6.
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<taxonomicName id="D116271D20C99877C5979CE3E0919208" class="Insecta" family="Asilidae" genus="Gibbasilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbasilus alboala" order="Diptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="species" species="alboala">Gibbasilus alboala</taxonomicName>
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sp. n. ♂ terminalia, lateral (3), dorsal (4), ventral (5), detail of gonostylus and aedeagus (6).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="463DF7D2F831EC11F346480162E2760D" pageId="3" pageNumber="70" type="holotype">
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<paragraph id="2D972D420EA9871B57C0A34137FF941C" pageId="3" pageNumber="70">Holotype.</paragraph>
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<paragraph id="41AB74989480363C0E1ACDE13CBE7836" pageId="3" pageNumber="70">
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SOUTH AFRICA: 1♂ "S Africa: Cape #69 / 1 km W Nieuwoudtville /
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<geoCoordinate id="A6FBE0F6B57AE38BD1244FD174B0129E" direction="south" orientation="latitude" precision="925" value="-31.383333">31°23'S</geoCoordinate>
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,
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<geoCoordinate id="2A486DD1FC2EE89A2C9834CE39A1ACF9" direction="east" orientation="longitude" precision="925" value="19.1">19°06'E</geoCoordinate>
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800 m / Date: 4.xi.1991 / Coll: J.G.H. Londt / Caravan Park and area" (NMSA).
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</paragraph>
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</subSubSection>
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<subSubSection id="7969DFA591DAE86D186995C1720937FE" pageId="3" pageNumber="70" type="paratype">
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<paragraph id="16CBB3270772648CE165684459E3AE34" pageId="3" pageNumber="70">Paratype.</paragraph>
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<paragraph id="4CA96D0BF23F7DB1FCEE220D993A8213" pageId="3" pageNumber="70">1♂ with identical label (NMSA).</paragraph>
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</subSubSection>
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<subSubSection id="C5F9894F60ECC0452D40AFCC224F740B" pageId="3" pageNumber="70" type="remarks">
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<paragraph id="6F9A7802E1072D5A0861B505B909483D" pageId="3" pageNumber="70">Remarks.</paragraph>
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<paragraph id="BD3A2CEF1300B8B09EF868ADDD802711" pageId="3" pageNumber="70">
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This species bears a strong resemblance to
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<taxonomicName id="455F2E7872748526C8B82386A14C96C4" lsidName="condylus" pageId="3" pageNumber="70" rank="species" species="condylus">condylus</taxonomicName>
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sp. n., described below, but is immediately recognised by the milky white bases to the wings. This condition, although unique within
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<taxonomicName id="BF09C9973853606E115ED4D4B8CF86DA" class="Insecta" family="Asilidae" genus="Gibbasilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbasilus" order="Diptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="genus">Gibbasilus</taxonomicName>
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, is encountered in other asilid genera such as
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<taxonomicName id="3F94D34E2B91BC4BA46C6F65C02FAE07" class="Insecta" family="Asilidae" genus="Hypenetes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypenetes" order="Diptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="genus">Hypenetes</taxonomicName>
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Loew, 1858 (
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<bibRefCitation id="04DCF8E06ADAE8C400E6B2AC09B28A8F" author="Londt, JGH" journalOrPublisher="Annals of the Natal Museum" pageId="13" pageNumber="80" pagination="377 - 405" title="Afrotropical Asilidae (Diptera) 10 - The genus Hypenetes Loew, 1858 (Dasypogoninae)." volume="26" year="1985">Londt 1985</bibRefCitation>
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). While it remains possible that the possession of milky wing bases is a variable character I consider
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<taxonomicName id="969DEE57656D1BCD866DAF8A4EFFB300" lsidName="alboala" pageId="3" pageNumber="70" rank="species" species="alboala">alboala</taxonomicName>
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and
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<taxonomicName id="A111AF7DB857B13E90F99022ECAE8904" lsidName="condylus" pageId="3" pageNumber="70" rank="species" species="condylus">condylus</taxonomicName>
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to represent separate taxa until shown otherwise. In the absence of females it is not known if milky wing bases are confined to males.
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</paragraph>
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</subSubSection>
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<subSubSection id="9B14B39C995C6176DDAFBC22C542DE3A" pageId="3" pageNumber="70" type="distribution">
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<paragraph id="579A9856483C825BCD672B7593BB82E7" pageId="3" pageNumber="70">Distribution (Fig. 20B), phenology (Table 1) and biology.</paragraph>
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<paragraph id="6A7D983FDA64B78E3DFD1E3148DB30F7" pageId="3" pageNumber="70">Known only from the type locality. Collected in November. The general habitat consisted of endemic fynbos plant species and large boulders.</paragraph>
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<caption id="7E4C4A8FA0AA15AE921E42E2E47AF309" pageId="3" pageNumber="70">
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<paragraph id="2666551384119CB351D81E3FB8B67CFD" pageId="3" pageNumber="70">
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Table 1. Phenology of
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<taxonomicName id="FABBF88F28551A5A7C5F024EE5074776" class="Insecta" family="Asilidae" genus="Gibbasilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gibbasilus" order="Diptera" pageId="3" pageNumber="70" phylum="Arthropoda" rank="genus">Gibbasilus</taxonomicName>
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species. Months, starting at July, abbreviated as shown.
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</paragraph>
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</caption>
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<paragraph id="E84A214270BE9D513C38E28A01D411C2" pageId="3" pageNumber="70">
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<table id="5CC06431140C86BD0027A4AE4148D255" pageId="3" pageNumber="70">
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<tr id="A4760F45281A881F3517510F1BFFB0D3" pageId="3" pageNumber="70">
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<th id="4B6C4C348B2487BD4D8FE47D08867236" colspan="1" pageId="3" pageNumber="70" rowspan="1">J</th>
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<th id="ECA824A1FC8340BF205CC230A183CA02" colspan="1" pageId="3" pageNumber="70" rowspan="1">A</th>
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<th id="8DE517FF0BACA83819EBAFB2B046F01D" colspan="1" pageId="3" pageNumber="70" rowspan="1">S</th>
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<th id="89433A191A9BB3E5A21F45819D7747D3" colspan="1" pageId="3" pageNumber="70" rowspan="1">O</th>
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<th id="34551831D9A48963CCCBD4A9E2F0DD2A" colspan="1" pageId="3" pageNumber="70" rowspan="1">N</th>
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<th id="9D39745FA6A432DF4EB43519F77B222D" colspan="1" pageId="3" pageNumber="70" rowspan="1">D</th>
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<th id="0ACECC1A4C7E1E3E042F11B2D154D129" colspan="1" pageId="3" pageNumber="70" rowspan="1">J</th>
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<th id="2CF7DF03DFEAEBAC965186FB5C8718DB" colspan="1" pageId="3" pageNumber="70" rowspan="1">F</th>
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<th id="29FAAAD3B6A250AF0BB429978222779C" colspan="1" pageId="3" pageNumber="70" rowspan="1">M</th>
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<th id="F5E59189B5D614A151CCC9649D3DEA72" colspan="1" pageId="3" pageNumber="70" rowspan="1">A</th>
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<th id="A1199EC775046BA80BD6865A7ACB5525" colspan="1" pageId="3" pageNumber="70" rowspan="1">M</th>
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<th id="8ADB2168F20F351834530C11414E483C" colspan="1" pageId="3" pageNumber="70" rowspan="1">J</th>
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</tr>
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<tr id="A4FC0E41D3C2E2545E230907DD5389F3" pageId="3" pageNumber="70">
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<td id="359CE11D77900B18A226368FCFCB0725" colspan="1" pageId="3" pageNumber="70" rowspan="1">
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<taxonomicName id="C3F9F0EE47BBF440867C867D7B311128" lsidName="alboala" pageId="3" pageNumber="70" rank="species" species="alboala">alboala</taxonomicName>
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</td>
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</tr>
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||
<tr id="9EEC72717D2BAE86514871325D24FD45" pageId="3" pageNumber="70">
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<td id="320C0038769A8E2BF16918E36A637E97" colspan="1" pageId="3" pageNumber="70" rowspan="1">
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<taxonomicName id="A72BA1E6BB43C9F596B4300640113193" lsidName="arenaceus" pageId="3" pageNumber="70" rank="species" species="arenaceus">arenaceus</taxonomicName>
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</td>
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||
</tr>
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||
<tr id="6186AE6C1CC4E63E22E8A20686DD188F" pageId="3" pageNumber="70">
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<td id="76DD15564E4D8DD6F1A12AADE1BB0653" colspan="1" pageId="3" pageNumber="70" rowspan="1">
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<taxonomicName id="AE3B30B41D9AA21943464F392144D752" lsidName="brevicolis" pageId="3" pageNumber="70" rank="species" species="brevicolis">brevicolis</taxonomicName>
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</td>
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</tr>
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<tr id="95F50186FD751076789EC669E7B47DFB" pageId="3" pageNumber="70">
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<td id="A4F86193BDD4D7665C80BD847F42D061" colspan="1" pageId="3" pageNumber="70" rowspan="1">
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<taxonomicName id="1BC21D4E77630AA1CE6EA7EFEC28B51C" lsidName="centrolobus" pageId="3" pageNumber="70" rank="species" species="centrolobus">centrolobus</taxonomicName>
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</td>
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</tr>
|
||
<tr id="940F14A44E69DF6E5718DD7940036820" pageId="3" pageNumber="70">
|
||
<td id="11C111EAFB6CBABD3A9C7386DC619883" colspan="1" pageId="3" pageNumber="70" rowspan="1">
|
||
<taxonomicName id="00A489D92F20DF644D1F7ED8AEE7ECB6" lsidName="condylus" pageId="3" pageNumber="70" rank="species" species="condylus">condylus</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
<tr id="F19A3B7B664B45E6595CE3C9EE91273E" pageId="3" pageNumber="70">
|
||
<td id="A65A507447A2EF8F6AB238959DBAF176" colspan="1" pageId="3" pageNumber="70" rowspan="1">
|
||
<taxonomicName id="6B571BE8EC74FF5E7E3F62BEC3B32C2E" lsidName="crinitus" pageId="3" pageNumber="70" rank="species" species="crinitus">crinitus</taxonomicName>
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |