524 lines
54 KiB
XML
524 lines
54 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1139.96639" ID-Pensoft-Pub="1313-2970-1139-33" ID-Pensoft-UUID="A2C101C84CB5568A92BA5F0DB93564D2" ID-ZooBank="0A828EAE0A7E4EF690869572E8F15E21" ModsDocID="1313-2970-1139-33" checkinTime="1673408730709" checkinUser="pensoft" docAuthor="Gao, Zhizhong, Hou, Yanmeng & Zhang, Feng" docDate="2023" docId="A5514B016A2D5F4CB83DB46941CC8861" docLanguage="en" docName="ZooKeys 1139: 33-69" docOrigin="ZooKeys 1139" docPubDate="2023-01-09" docSource="http://dx.doi.org/10.3897/zookeys.1139.96639" docTitle="Allochthonius pandus Gao & Hou & Zhang 2023, sp. nov." docType="treatment" docUuid="50DA34BD-CAD3-4A28-AFD6-28078FBF21E1" docUuidSource="ZooBank" docVersion="1" id="A2C101C84CB5568A92BA5F0DB93564D2" lastPageNumber="33" masterDocId="A2C101C84CB5568A92BA5F0DB93564D2" masterDocTitle="Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China" masterLastPageNumber="69" masterPageNumber="33" pageNumber="33" updateTime="1673408730709" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gao, Zhizhong</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6666-8746</mods:nameIdentifier>
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<mods:affiliation>Department of Biology, Xinzhou Teachers University, Xinzhou, Shanxi 034000, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hou, Yanmeng</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0059-3419</mods:nameIdentifier>
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<mods:affiliation>The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhang, Feng</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3347-1031</mods:nameIdentifier>
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<mods:affiliation>The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:nameIdentifier type="email">dudu06042001@163.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-01-09</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1139</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>33</mods:start>
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<mods:end>69</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1139.96639</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1139.96639</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1139-33</mods:identifier>
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<mods:identifier type="ZooBank">0A828EAE0A7E4EF690869572E8F15E21</mods:identifier>
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<mods:identifier type="Pensoft-UUID">A2C101C84CB5568A92BA5F0DB93564D2</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:50DA34BD-CAD3-4A28-AFD6-28078FBF21E1" httpUri="http://treatment.plazi.org/id/A5514B016A2D5F4CB83DB46941CC8861" lastPageNumber="33" pageId="0" pageNumber="33">
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<subSubSection pageId="0" pageNumber="33" type="nomenclature">
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<paragraph pageId="0" pageNumber="33">
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<taxonomicName LSID="https://zoobank.org/50DA34BD-CAD3-4A28-AFD6-28078FBF21E1" authority="Gao & Hou & Zhang, 2023" authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Allochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="pandus" status="sp. nov.">Allochthonius pandus</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="33">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="description">
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<paragraph pageId="0" pageNumber="33">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area with indication of cave locations representing the type localities. Each color represents an administrative region (red: Guizhou Province; purple: Weining County; green: Xishui County; blue: Fenggang County) A Wulibei Cave (Spelaeochthonius wulibeiensis sp. nov.) B Liangfeng Cave (Allochthonius bainiensis sp. nov.) C Daozuo Cave (A. pandus sp. nov.) D Sanjie Cave (A. xinqiaoensis sp. nov.)." figureDoi="10.3897/zookeys.1139.96639.figure1" httpUri="https://binary.pensoft.net/fig/793039" pageId="0" pageNumber="33">Figs 1C</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Daozuo Cave, type locality of Allochthonius pandus sp. nov. A entrance B inside the cave entrance C narrow tunnel to the deepest part of the cave (red arrow) D, E areas where A. pandus sp. nov. specimens were collected F live male of A. pandus sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1139.96639.figure7" httpUri="https://binary.pensoft.net/fig/793045" pageId="0" pageNumber="33">, 7</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Allochthonius pandus sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1139.96639.figure8" httpUri="https://binary.pensoft.net/fig/793046" pageId="0" pageNumber="33">, 8</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Allochthonius pandus sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure9" httpUri="https://binary.pensoft.net/fig/793047" pageId="0" pageNumber="33">, 9</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Allochthonius pandus sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure10" httpUri="https://binary.pensoft.net/fig/793048" pageId="0" pageNumber="33">, 10</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Allochthonius pandus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure11" httpUri="https://binary.pensoft.net/fig/793049" pageId="0" pageNumber="33">, 11</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Allochthonius pandus sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure12" httpUri="https://binary.pensoft.net/fig/793050" pageId="0" pageNumber="33">
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, 12 Chinese name
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<normalizedToken originalValue="弯指异伪蝎">弯指异伪蝎</normalizedToken>
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</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="type material">
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<paragraph pageId="0" pageNumber="33">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Holotype</emphasis>
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: China • ♂; Guizhou Province, Xishui County, Xianyuan Town, Jinshan Village, Daozuo Cave;
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<geoCoordinate degrees="28" direction="north" minutes="18.04" orientation="latitude" precision="9" value="28.300667">28°18.04'N</geoCoordinate>
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,
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<geoCoordinate degrees="106" direction="east" minutes="41.70" orientation="longitude" precision="9" value="106.695">106°41.70'E</geoCoordinate>
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; 1606 m a.s.l.; 24 Jul. 2022; Yanmeng Hou, Lu Zhang, Jianzhou Sun and Wenlong Fan leg.; under a stone in the deep zone; Ps.-MHBU-HBUARA#2022-47701 (Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area with indication of cave locations representing the type localities. Each color represents an administrative region (red: Guizhou Province; purple: Weining County; green: Xishui County; blue: Fenggang County) A Wulibei Cave (Spelaeochthonius wulibeiensis sp. nov.) B Liangfeng Cave (Allochthonius bainiensis sp. nov.) C Daozuo Cave (A. pandus sp. nov.) D Sanjie Cave (A. xinqiaoensis sp. nov.)." figureDoi="10.3897/zookeys.1139.96639.figure1" httpUri="https://binary.pensoft.net/fig/793039" pageId="0" pageNumber="33">1C</figureCitation>
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,
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Daozuo Cave, type locality of Allochthonius pandus sp. nov. A entrance B inside the cave entrance C narrow tunnel to the deepest part of the cave (red arrow) D, E areas where A. pandus sp. nov. specimens were collected F live male of A. pandus sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1139.96639.figure7" httpUri="https://binary.pensoft.net/fig/793045" pageId="0" pageNumber="33">7A-E</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Paratypes</emphasis>
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: • 1♂; the same location as the holotype; 28 Aug. 2020; Zegang Feng, Hongru Xu and Yanmeng Hou leg.; Ps.-MHBU-GZXS-20-24 • 2♀; the same data as the holotype; Ps.-MSWU-HBUARA#2022-47702-HBUARA#2022-47703.
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</paragraph>
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<caption doi="10.3897/zookeys.1139.96639.figure7" httpUri="https://binary.pensoft.net/fig/793045" pageId="0" pageNumber="33" start="Figure 7" startId="F7">
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" pageId="0" pageNumber="33">Figure 7.</emphasis>
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Daozuo Cave, type locality of
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<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Allochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="pandus">
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<emphasis italics="true" pageId="0" pageNumber="33">Allochthonius pandus</emphasis>
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</taxonomicName>
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sp. nov.
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<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
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entrance
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<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
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inside the cave entrance
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<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
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narrow tunnel to the deepest part of the cave (red arrow)
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<emphasis bold="true" pageId="0" pageNumber="33">D, E</emphasis>
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areas where
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<taxonomicName lsidName="A. pandus" pageId="0" pageNumber="33" rank="species" species="pandus">
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<emphasis italics="true" pageId="0" pageNumber="33">A. pandus</emphasis>
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</taxonomicName>
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sp. nov. specimens were collected
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<emphasis bold="true" pageId="0" pageNumber="33">F</emphasis>
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live male of
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<taxonomicName lsidName="A. pandus" pageId="0" pageNumber="33" rank="species" species="pandus">
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<emphasis italics="true" pageId="0" pageNumber="33">A. pandus</emphasis>
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</taxonomicName>
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sp. nov. in its natural environment.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="diagnosis">
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<paragraph pageId="0" pageNumber="33">Diagnosis</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" pageId="0" pageNumber="33">(♂♀).</emphasis>
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The new species can be recognized by the following combination of characters: cheliceral palm with five setae; coxa I with four coxal spines (tridentate blades, each blade with a central fan-shaped spine terminally) on a tubercle; pedipalps slender, femur 9.07-10.15 (♂), 8.50-8.60 (♀), chela 7.00-7.52 (♂), 6.64-7.15 (♀)
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than broad, both chelal fingers with a row of teeth (fixed chelal finger with 31 or 33 teeth; movable chelal finger with 26 or 28 teeth), slightly retrorse and pointed; chela fingers markedly curved in dorsal view.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="etymology">
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<paragraph pageId="0" pageNumber="33">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="33">
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The specific name is derived from the Latin word
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<emphasis italics="true" pageId="0" pageNumber="33">pandus</emphasis>
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(curved) and refers to the character of the curved chelal fingers.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="33" type="description">
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<paragraph pageId="0" pageNumber="33">Description.</paragraph>
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<paragraph pageId="0" pageNumber="33">
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<emphasis bold="true" pageId="0" pageNumber="33">Adult males</emphasis>
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(Figs
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Daozuo Cave, type locality of Allochthonius pandus sp. nov. A entrance B inside the cave entrance C narrow tunnel to the deepest part of the cave (red arrow) D, E areas where A. pandus sp. nov. specimens were collected F live male of A. pandus sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1139.96639.figure7" httpUri="https://binary.pensoft.net/fig/793045" pageId="0" pageNumber="33">7F</figureCitation>
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,
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Allochthonius pandus sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1139.96639.figure8" httpUri="https://binary.pensoft.net/fig/793046" pageId="0" pageNumber="33">8A</figureCitation>
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,
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Allochthonius pandus sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure9" httpUri="https://binary.pensoft.net/fig/793047" pageId="0" pageNumber="33">9</figureCitation>
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,
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Allochthonius pandus sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure10" httpUri="https://binary.pensoft.net/fig/793048" pageId="0" pageNumber="33">10A, B</figureCitation>
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,
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Allochthonius pandus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure11" httpUri="https://binary.pensoft.net/fig/793049" pageId="0" pageNumber="33">11</figureCitation>
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,
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Allochthonius pandus sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure12" httpUri="https://binary.pensoft.net/fig/793050" pageId="0" pageNumber="33">12</figureCitation>
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).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Color</emphasis>
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(Figs
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Daozuo Cave, type locality of Allochthonius pandus sp. nov. A entrance B inside the cave entrance C narrow tunnel to the deepest part of the cave (red arrow) D, E areas where A. pandus sp. nov. specimens were collected F live male of A. pandus sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1139.96639.figure7" httpUri="https://binary.pensoft.net/fig/793045" pageId="0" pageNumber="33">7F</figureCitation>
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,
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Allochthonius pandus sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1139.96639.figure8" httpUri="https://binary.pensoft.net/fig/793046" pageId="0" pageNumber="33">8A</figureCitation>
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,
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Allochthonius pandus sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure9" httpUri="https://binary.pensoft.net/fig/793047" pageId="0" pageNumber="33">9</figureCitation>
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,
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Allochthonius pandus sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure10" httpUri="https://binary.pensoft.net/fig/793048" pageId="0" pageNumber="33">10A, B</figureCitation>
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): generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Cephalothorax</emphasis>
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(Figs
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Allochthonius pandus sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure9" httpUri="https://binary.pensoft.net/fig/793047" pageId="0" pageNumber="33">9B</figureCitation>
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,
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Allochthonius pandus sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure10" httpUri="https://binary.pensoft.net/fig/793048" pageId="0" pageNumber="33">10A</figureCitation>
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,
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Allochthonius pandus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure11" httpUri="https://binary.pensoft.net/fig/793049" pageId="0" pageNumber="33">11A, C</figureCitation>
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): carapace inverted trapezoid, 0.91-0.93
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<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than broad, gently narrowed posteriorly; surface smooth, without furrows but with six lyrifissures and the posterior part with squamous sculpturing; no traces of eyes; epistomal process absent, space between median setae slightly recurved; with 14 setae arranged 4: 4: 2: 2: 2, preocular setae absent, most setae heavy, long, and gently curved. Chaetotaxy of coxae: P 3, I 4, II 4-6, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/3 length of medial seta; coxal spines present on coxa I only, consisting of a tubercle expanded terminally into a characteristic
|
||
<normalizedToken originalValue="“spray”">"spray"</normalizedToken>
|
||
or
|
||
<normalizedToken originalValue="“fan”">"fan"</normalizedToken>
|
||
of four elevated processes which extend apically, subequal in length (Figs
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Allochthonius pandus sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure10" httpUri="https://binary.pensoft.net/fig/793048" pageId="0" pageNumber="33">10A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Allochthonius pandus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure11" httpUri="https://binary.pensoft.net/fig/793049" pageId="0" pageNumber="33">11C</figureCitation>
|
||
); bisetose intercoxal tubercle present between coxae III and IV (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Allochthonius pandus sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure10" httpUri="https://binary.pensoft.net/fig/793048" pageId="0" pageNumber="33">10A</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Chelicera</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Allochthonius pandus sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure9" httpUri="https://binary.pensoft.net/fig/793047" pageId="0" pageNumber="33">9C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Allochthonius pandus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure11" httpUri="https://binary.pensoft.net/fig/793049" pageId="0" pageNumber="33">11B, E</figureCitation>
|
||
): large, approximately as long as carapace, 2.56-2.60
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with a medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with ten acute teeth, distal one largest; movable finger with 15 or 16 retrorse contiguous teeth of equal length, plus four or five vestigial, rounded, and contiguous basal teeth, 19-21 in total; galea absent. Serrula exterior with 19 or 20 blades and serrula interior with 10 or 11 blades. Rallum in two rows and composed of nine blades with fine pinnate, of which the basal-most blade shorter than the others (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Allochthonius pandus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure11" httpUri="https://binary.pensoft.net/fig/793049" pageId="0" pageNumber="33">11E</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Pedipalp</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Allochthonius pandus sp. nov., holotype male A left chela (lateral view) B carapace (dorsal view) C left chelicera (dorsal view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure9" httpUri="https://binary.pensoft.net/fig/793047" pageId="0" pageNumber="33">9A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Allochthonius pandus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure11" httpUri="https://binary.pensoft.net/fig/793049" pageId="0" pageNumber="33">11D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Allochthonius pandus sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure12" httpUri="https://binary.pensoft.net/fig/793050" pageId="0" pageNumber="33">12A, B</figureCitation>
|
||
): long and slender, trochanter 1.65-1.67, femur 9.07-10.15, patella 3.47-3.57, chela 7.00-7.52, hand 2.60-2.96
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than broad; femur 2.44-2.64
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than patella; movable chelal finger 1.59-1.72
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than hand and 0.62-0.64
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Allochthonius pandus sp. nov., holotype male A carapace (dorsal view) B left chelicera (dorsal view), with details of teeth C coxal spines on coxae I (ventral view) D left pedipalp (minus chela, dorsal view) E rallum. Scale bars: 0.20 mm (A, B, D); 0.10 mm (C, E)." figureDoi="10.3897/zookeys.1139.96639.figure11" httpUri="https://binary.pensoft.net/fig/793049" pageId="0" pageNumber="33">11D</figureCitation>
|
||
). Chelal palm slightly constricted towards fingers. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria,
|
||
<emphasis italics="true" pageId="0" pageNumber="33">ib</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="33">isb</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="33">eb</emphasis>
|
||
,
|
||
<emphasis italics="true" pageId="0" pageNumber="33">esb</emphasis>
|
||
, and
|
||
<emphasis italics="true" pageId="0" pageNumber="33">ist</emphasis>
|
||
clustered at the base of fixed finger,
|
||
<emphasis italics="true" pageId="0" pageNumber="33">esb</emphasis>
|
||
slightly distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="33">ist</emphasis>
|
||
;
|
||
<emphasis italics="true" pageId="0" pageNumber="33">it</emphasis>
|
||
slightly distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="33">est</emphasis>
|
||
, situated subdistally;
|
||
<emphasis italics="true" pageId="0" pageNumber="33">et</emphasis>
|
||
situated subdistally, very close to chelal teeth;
|
||
<emphasis italics="true" pageId="0" pageNumber="33">dx</emphasis>
|
||
situated distal to
|
||
<emphasis italics="true" pageId="0" pageNumber="33">et</emphasis>
|
||
, near the tip of fixed finger;
|
||
<emphasis italics="true" pageId="0" pageNumber="33">sb</emphasis>
|
||
situated closer to
|
||
<emphasis italics="true" pageId="0" pageNumber="33">b</emphasis>
|
||
than to
|
||
<emphasis italics="true" pageId="0" pageNumber="33">st</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Allochthonius pandus sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure12" httpUri="https://binary.pensoft.net/fig/793050" pageId="0" pageNumber="33">12A</figureCitation>
|
||
). Microsetae (chemosensory setae) absent on hand and both palpal fingers. Sensilla absent. Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 31 or 33 teeth, slightly retrorse and pointed; movable chelal finger with 24 or 25 teeth (slightly smaller than teeth on fixed chelal finger), plus two or three vestigial, rounded and contiguous basal teeth, 26-28 in total; a small tubercle between the fourteenth and fifteenth teeth present (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Allochthonius pandus sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure12" httpUri="https://binary.pensoft.net/fig/793050" pageId="0" pageNumber="33">12A</figureCitation>
|
||
). Chelal fingers markedly curved in dorsal view (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Allochthonius pandus sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure12" httpUri="https://binary.pensoft.net/fig/793050" pageId="0" pageNumber="33">12B</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Opisthosoma</emphasis>
|
||
: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I-XII: 2: 4-5: 4-5: 6: 6: 6: 6: 6: 3: 2: TT: 0, tergite IX with an unpaired median seta. Sternal chaetotaxy III-XII: 6-8: 9-10: 9-11: 9: 9-10: 9-10: 7-8: 5-6: 0: 2. Anterior genital operculum with eight setae, genital opening pit-like, with five or six marginal setae on each side, 18-19 in total, with a pair of lyrifissures present anterolateral and posteriolateral to genital opening, respectively (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Allochthonius pandus sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure10" httpUri="https://binary.pensoft.net/fig/793048" pageId="0" pageNumber="33">10B</figureCitation>
|
||
).
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Legs</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Allochthonius pandus sp. nov., holotype male A left chela (lateral view), with details of teeth and trichobothrial pattern B left chela (dorsal view) C leg I (lateral view) D leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="10.3897/zookeys.1139.96639.figure12" httpUri="https://binary.pensoft.net/fig/793050" pageId="0" pageNumber="33">12C, D</figureCitation>
|
||
): generally typical, long, and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.64-1.79
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than patella and with one lyrifissure at the base of femur; tarsus 2.42-2.60
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than tibia. Femoropatella of leg IV 5.00-5.17
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep and with one lyrifissure at the base of femur; tibia 6.55-7.30
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep; with a long tactile seta on both tarsal segments: basitarsus 4.00-4.86
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep (TS = 0.21-0.25), telotarsus 13.50-14.50
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep and 2.38-2.72
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than basitarsus (TS = 0.22-0.25). Setae of leg I (trochanter to tibia) 2-3: 9-11: 9-10: 12-13, setae of leg IV (trochanter to basitarsus) 2-3: 2: 4-5: 17-19: 10-11. Arolium slightly shorter than the claws, not divided; claws simple.
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="33">Dimensions of adult males</emphasis>
|
||
(length/breadth or, in the case of the legs, length/depth in mm): body length 1.97-2.23. Pedipalps: trochanter 0.28-0.30/0.17-0.18, femur 1.27-1.32/0.13-0.14, patella 0.50-0.52/0.14-0.15, chela 1.73-1.75/0.23-0.25, hand 0.65-0.68/0.23-0.25, movable chelal finger length 1.08-1.12. Chelicera 0.64-0.65/0.25, movable finger length 0.33-0.34. Carapace 0.53-0.54/0.58. Leg I: trochanter 0.20-0.21/0.15, femur 0.68-0.69/0.08-0.09, patella 0.38-0.42/0.08, tibia 0.30-0.31/0.05-0.06, tarsus 0.75-0.78/0.05-0.06. Leg IV: trochanter 0.29-0.30/0.16-0.17, femoropatella 0.93-0.95/0.18-0.19, tibia 0.72-0.73/0.10-0.11, basitarsus 0.32-0.34/0.07-0.08, telotarsus 0.81-0.87/0.06.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1139.96639.figure8" httpUri="https://binary.pensoft.net/fig/793046" pageId="0" pageNumber="33" start="Figure 8" startId="F8">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 8.</emphasis>
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Allochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Allochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
holotype male, habitus (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
paratype female, habitus (dorsal view). Scale bars: 0.50 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1139.96639.figure9" httpUri="https://binary.pensoft.net/fig/793047" pageId="0" pageNumber="33" start="Figure 9" startId="F9">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 9.</emphasis>
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Allochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Allochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
left chela (lateral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
carapace (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
left chelicera (dorsal view). Scale bars: 0.20 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B, C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1139.96639.figure10" httpUri="https://binary.pensoft.net/fig/793048" pageId="0" pageNumber="33" start="Figure 10" startId="F10">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 10.</emphasis>
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Allochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Allochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A, B</emphasis>
|
||
), paratype female (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
coxae (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
male genital area (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
female genital area (ventral view). Scale bars: 0.20 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B, C</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1139.96639.figure11" httpUri="https://binary.pensoft.net/fig/793049" pageId="0" pageNumber="33" start="Figure 11" startId="F11">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 11.</emphasis>
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Allochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Allochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
carapace (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
left chelicera (dorsal view), with details of teeth
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
coxal spines on coxae I (ventral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">D</emphasis>
|
||
left pedipalp (minus chela, dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">E</emphasis>
|
||
rallum. Scale bars: 0.20 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A, B, D</emphasis>
|
||
); 0.10 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C, E</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zookeys.1139.96639.figure12" httpUri="https://binary.pensoft.net/fig/793050" pageId="0" pageNumber="33" start="Figure 12" startId="F12">
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Figure 12.</emphasis>
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Allochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Allochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., holotype male
|
||
<emphasis bold="true" pageId="0" pageNumber="33">A</emphasis>
|
||
left chela (lateral view), with details of teeth and trichobothrial pattern
|
||
<emphasis bold="true" pageId="0" pageNumber="33">B</emphasis>
|
||
left chela (dorsal view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">C</emphasis>
|
||
leg I (lateral view)
|
||
<emphasis bold="true" pageId="0" pageNumber="33">D</emphasis>
|
||
leg IV (lateral view). Scale bars: 0.20 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<emphasis bold="true" pageId="0" pageNumber="33">Adult females</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Allochthonius pandus sp. nov. A holotype male, habitus (dorsal view) B paratype female, habitus (dorsal view). Scale bars: 0.50 mm." figureDoi="10.3897/zookeys.1139.96639.figure8" httpUri="https://binary.pensoft.net/fig/793046" pageId="0" pageNumber="33">8B</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Allochthonius pandus sp. nov., holotype male (A, B), paratype female (C) A coxae (ventral view) B male genital area (ventral view) C female genital area (ventral view). Scale bars: 0.20 mm (A); 0.10 mm (B, C)." figureDoi="10.3897/zookeys.1139.96639.figure10" httpUri="https://binary.pensoft.net/fig/793048" pageId="0" pageNumber="33">10C</figureCitation>
|
||
). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 4, II 5, III 5, IV 5; tergal chaetotaxy I-XII: 2: 4: 4: 4-6: 6: 6: 6: 6: 4: 2: TT: 0; sternal chaetotaxy IV-XII: 9-10: 10-12: 9: 10-11: 10-12: 7-8: 6: 0: 2; anterior genital operculum with eight or nine setae, posterior margin with nine or ten marginal setae, 17-19 in total; leg IV with a long tactile seta on both tarsal segments: basitarsus 3.78
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep (TS = 0.24), telotarsus 12.86-14.50
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than deep and 2.56-2.65
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than basitarsus (TS = 0.20-0.23). Body length 2.10-2.41. Pedipalps: trochanter 0.33/0.19-0.20 (1.65-1.74
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), femur 1.36-1.38/0.16 (8.50-8.63
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), patella 0.54-0.55/0.18-0.19 (2.89-3.00
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), chela 1.86/0.26-0.28 (6.64-7.15
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), hand 0.72-0.75/0.26-0.28 (2.68-2.77
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), movable chelal finger length 1.16. Chelicera 0.73-0.76/0.26-0.28 (2.71-2.81
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), movable finger length 0.39. Carapace 0.55-0.57/0.65 (0.85-0.88
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
). Leg I: trochanter 0.19/0.17 (1.12
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), femur 0.70-0.75/0.10-0.11 (6.82-7.00
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), patella 0.43-0.44/0.09 (4.78-4.89
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), tibia 0.33-0.35/0.06 (5.50-5.83
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), tarsus 0.79-0.84/0.06 (13.17-14.00
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
). Leg IV: trochanter 0.32-0.34/0.16-0.18 (1.89-2.00
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), femoropatella 0.92-1.02/0.19-0.22 (4.64-4.84
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), tibia 0.67-0.73/0.10-0.11 (6.64-6.70
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), basitarsus 0.34/0.09 (3.78
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
), telotarsus 0.87-0.90/0.06-0.07 (12.86-14.50
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="33" type="remarks">
|
||
<paragraph pageId="0" pageNumber="33">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Allochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Allochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is similar to
|
||
<taxonomicName lsidName="A. xinqiaoensis" pageId="0" pageNumber="33" rank="species" species="xinqiaoensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">A. xinqiaoensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. in having a pair of distinctly curved chelal fingers and the same chaetotaxy of the carapace (4: 4: 2: 2: 2), but differs by the presence of lower number of blades of coxal spines (4 vs. 6), more rallum blades (9 vs. 8), more slender chela (chela 6.64-7.15 (♀)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
vs. 5.44 (♀)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than broad), lower number of setae on the coxae (3: 4: 5: 5: 5 vs. 3: 6: 7-9: 5: 5) and more teeth on the chelal fingers (31-33 vs. 23 teeth on the fixed chelal finger and 26-28 vs. 23 teeth on the movable chelal finger);
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Allochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Allochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from
|
||
<taxonomicName lsidName="A. bainiensis" pageId="0" pageNumber="33" rank="species" species="bainiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">A. bainiensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. by the presence of a pair of distinctly curved chelal fingers.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Allochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Allochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. differs from
|
||
<taxonomicName lsidName="A. brevitus" pageId="0" pageNumber="33" rank="species" species="brevitus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">A. brevitus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="A. yoshizawai" pageId="0" pageNumber="33" rank="species" species="yoshizawai">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">A. yoshizawai</emphasis>
|
||
</taxonomicName>
|
||
in the number of setae on the anterior of the carapace (4 vs. 6), the cheliceral hand (5 vs. 6) and tergite II (4-5 vs. 6 or 2), and the number of rallum blades (9 vs. 11) and the presence of more slender pedipalps (e.g., palpal femur 9.07-10.15 (♂)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than broad in
|
||
<taxonomicName lsidName="A. pandus" pageId="0" pageNumber="33" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">A. pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., while 4.33-4.73 (♂) and 6.50 (♂)
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
in
|
||
<taxonomicName lsidName="A. brevitus" pageId="0" pageNumber="33" rank="species" species="brevitus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">A. brevitus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName lsidName="A. yoshizawai" pageId="0" pageNumber="33" rank="species" species="yoshizawai">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">A. yoshizawai</emphasis>
|
||
</taxonomicName>
|
||
, respectively).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Allochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Allochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from
|
||
<taxonomicName lsidName="A. ishikawai" pageId="0" pageNumber="33" rank="species" species="ishikawai">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">A. ishikawai</emphasis>
|
||
</taxonomicName>
|
||
and all
|
||
<taxonomicName lsidName="A. ishikawai" pageId="0" pageNumber="33" rank="species" species="ishikawai">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">A. ishikawai</emphasis>
|
||
</taxonomicName>
|
||
subspecies by the number of setae on the carapace (14 vs. 16 or more), the presence of lower number of rallum blades (9 vs. 10) and more teeth on both chelal fingers (26-28 vs. 11-17 teeth on the movable finger and 31-33 vs. 9-17 teeth on the fixed chelal finger).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
<taxonomicName authorityName="Gao & Hou & Zhang" authorityYear="2023" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Allochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allochthonius pandus" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="species" species="pandus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Allochthonius pandus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished from the other species of
|
||
<taxonomicName authorityName="Chamberlin" authorityYear="1929" class="Arachnida" family="Pseudotyrannochthoniidae" genus="Allochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Allochthonius" order="Pseudoscorpiones" pageId="0" pageNumber="33" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="33">Allochthonius</emphasis>
|
||
</taxonomicName>
|
||
by the absence of any traces of eyes (
|
||
<bibRefCitation author="Morikawa, K" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="33" refId="B24" refString="Morikawa, K, 1954. On some pseudoscorpions in Japanese lime-grottoes. Memoirs of Ehime University 2(2B): 79-87." title="On some pseudoscorpions in Japanese lime-grottoes. Memoirs of Ehime University 2 (2 B): 79 - 87." year="1954">Morikawa 1954</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Morikawa, K" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="33" refId="B25" refString="Morikawa, K, 1956. Cave pseudoscorpions of Japan (I). Memoirs of Ehime University 2(2B): 271-282." title="Cave pseudoscorpions of Japan (I). Memoirs of Ehime University 2 (2 B): 271 - 282." year="1956">1956</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Morikawa, K" journalOrPublisher="Revue Suisse de Zoologie" pageId="0" pageNumber="33" refId="B26" refString="Morikawa, K, 1960. Systematics studies of Japanese pseudoscorpions. Memoirs of Ehime University 2(2B): 85-172." title="Systematics studies of Japanese pseudoscorpions. Memoirs of Ehime University 2 (2 B): 85 - 172." year="1960">1960</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.33338/ef.5003" author="Hu, JF" journalOrPublisher="Entomologica Fennica" pageId="0" pageNumber="33" pagination="243 - 248" refId="B13" refString="Hu, JF, Zhang, F, 2012. Two new species of the genus Allochthonius Chamberlin from China (Pseudoscorpiones: Pseudotyrannochthoniidae). Entomologica Fennica 22 (4): 243 - 248, DOI: https://doi.org/10.33338/ef.5003" title="Two new species of the genus Allochthonius Chamberlin from China (Pseudoscorpiones: Pseudotyrannochthoniidae)." url="https://doi.org/10.33338/ef.5003" volume="22" year="2012">Hu and Zhang 2012</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.3897/subtbiol.37.58580" author="Viana, ACM" journalOrPublisher="Subterranean Biology" pageId="0" pageNumber="33" pagination="47 - 55" refId="B30" refString="Viana, ACM, Ferreira, RL, 2021. A new troglobitic species of Allochthonius (subgenus Urochthonius) (Pseudoscorpiones, Pseudotyrannochthoniidae) from Japan. Subterranean Biology 37: 47 - 55, DOI: https://doi.org/10.3897/subtbiol.37.58580" title="A new troglobitic species of Allochthonius (subgenus Urochthonius) (Pseudoscorpiones, Pseudotyrannochthoniidae) from Japan." url="https://doi.org/10.3897/subtbiol.37.58580" volume="37" year="2021">Viana and Ferreira 2021</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Viana, ACM" journalOrPublisher="Subterranean Biology" pageId="0" pageNumber="33" publicationUrl="https://wac.nmbe.ch/order/pseudoscorpiones/3" refId="B31" refString="2022. . https://wac.nmbe.ch/order/pseudoscorpiones/3" url="https://wac.nmbe.ch/order/pseudoscorpiones/3" year="2022">WPC 2022</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="33" type="distribution">
|
||
<paragraph pageId="0" pageNumber="33">Distribution and habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="33">
|
||
This species is only known from the type locality, Daozuo Cave (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area with indication of cave locations representing the type localities. Each color represents an administrative region (red: Guizhou Province; purple: Weining County; green: Xishui County; blue: Fenggang County) A Wulibei Cave (Spelaeochthonius wulibeiensis sp. nov.) B Liangfeng Cave (Allochthonius bainiensis sp. nov.) C Daozuo Cave (A. pandus sp. nov.) D Sanjie Cave (A. xinqiaoensis sp. nov.)." figureDoi="10.3897/zookeys.1139.96639.figure1" httpUri="https://binary.pensoft.net/fig/793039" pageId="0" pageNumber="33">1C</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Daozuo Cave, type locality of Allochthonius pandus sp. nov. A entrance B inside the cave entrance C narrow tunnel to the deepest part of the cave (red arrow) D, E areas where A. pandus sp. nov. specimens were collected F live male of A. pandus sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1139.96639.figure7" httpUri="https://binary.pensoft.net/fig/793045" pageId="0" pageNumber="33">7</figureCitation>
|
||
), which is located near a road, 1 km southwest of Jinshan Village (Xishui County) and is surrounded by rural and agricultural fields. This limestone cave has a large, rectangular entrance (~ 1 m high and 30 m wide) and a total length of ~ 300 m, only a narrow tunnel leads to the deepest part of the cave, which is a slightly wider, low-temperature, high-humidity, and completely lightless environment (temperature ~ 11 °C, humidity> 90%). All specimens were collected under stones in the deepest part of the cave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |