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<document id="7DAE813084F9601035FEE510FE7A7C17" ID-DOI="10.11646/zootaxa.4059.1.2" ID-GBIF-Dataset="3acbd49b-3995-44f6-8965-c86a456bf91e" ID-ISSN="1175-5326" ID-Zenodo-Dep="236611" ID-ZooBank="C9F9B55A-B48C-4619-AB45-4102FBD26CBE" IM.bibliography_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1461360591258" checkinUser="plazi" docAuthor="Melo, Ivy Frizo De, Araújo, Carolina Rodrigues &amp; Penteado-Dias, Angélica Maria" docDate="2015" docId="472B87FF5C56164AF9FFF89AFB1CFD84" docLanguage="en" docName="zt04059p050.pdf" docOrigin="Zootaxa 4059 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Exochus izbus Gauld &amp; Sithole 2002" docType="treatment" docVersion="12" lastPageNumber="46" masterDocId="BB12FF875C53164CF968FFB3FFD1FF9C" masterDocTitle="New species of Exochus Gravenhorst and Trieces Townes (Hymenoptera, Ichneumonidae, Metopiinae) and first record of seven species from Brazil" masterLastPageNumber="50" masterPageNumber="40" pageNumber="45" updateTime="1736248570979" updateUser="ExternalLinkService">
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<mods:title id="6EAEE2B101CB0DAAA097BBCF5DBE3993">New species of Exochus Gravenhorst and Trieces Townes (Hymenoptera, Ichneumonidae, Metopiinae) and first record of seven species from Brazil</mods:title>
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<mods:namePart id="FDC6A42382684DC9012EEC841B3327A7">Melo, Ivy Frizo De</mods:namePart>
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<mods:namePart id="E3086DC08D7B78F02F9D8C8978A573CB">Araújo, Carolina Rodrigues</mods:namePart>
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<mods:namePart id="C9CDFA4E35D1112613C8F5D366EFAB32">Penteado-Dias, Angélica Maria</mods:namePart>
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<mods:title id="90829C649D576CA9835A241AAE1AC48D">Zootaxa</mods:title>
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<mods:date id="C58E5EAD5A596CA428A737C3DDCFEA5E">2015</mods:date>
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<mods:number id="05D93AAA55E624BF7AC57E7B8D515F72">4059</mods:number>
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<treatment id="472B87FF5C56164AF9FFF89AFB1CFD84" ID-DOI="http://doi.org/10.5281/zenodo.6116285" ID-GBIF-Taxon="119648248" ID-Zenodo-Dep="6116285" LSID="urn:lsid:plazi:treatment:472B87FF5C56164AF9FFF89AFB1CFD84" httpUri="http://treatment.plazi.org/id/472B87FF5C56164AF9FFF89AFB1CFD84" lastPageId="6" lastPageNumber="46" pageId="5" pageNumber="45" scope_family="Ichneumonidae" scope_order="Hymenoptera">
<subSubSection id="879865625C561649F9FFF89AFF3AF8F9" pageId="5" pageNumber="45" type="nomenclature">
<paragraph id="CF3D36E95C561649F9FFF89AFDB3F8DF" blockId="5.[151,610,1833,1893]" box="[151,610,1833,1859]" pageId="5" pageNumber="45">
<heading id="947581855C561649F9FFF89AFDB3F8DF" bold="true" box="[151,610,1833,1859]" fontSize="11" level="1" pageId="5" pageNumber="45" reason="1">
<taxonomicName id="08824D6A5C561649F9FFF89AFDB3F8DF" ID-CoL="3DMYM" authority="Gauld &amp; Sithole, 2002" authorityName="Gauld &amp; Sithole" authorityYear="2002" box="[151,610,1833,1859]" class="Insecta" family="Ichneumonidae" genus="Exochus" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="45" phylum="Arthropoda" rank="species" species="izbus">
<emphasis id="FDF6EAFB5C561649F9FFF89AFDB3F8DF" bold="true" box="[151,610,1833,1859]" pageId="5" pageNumber="45">
<emphasis id="FDF6EAFB5C561649F9FFF89AFE97F8DF" bold="true" box="[151,326,1833,1859]" italics="true" pageId="5" pageNumber="45">Exochus izbus</emphasis>
<bibRefCitation id="AB134B185C561649F825F89AFDB3F8DF" author="Gauld, I. &amp; Sithole, R. &amp; Gomes, J. U. &amp; Godoy, C." box="[333,610,1833,1859]" firstAuthor="Gauld" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="5" pageNumber="45" pagination="1 - 768" part="66" refId="ref6711" refString="Gauld, I., Sithole, R., Gomes, J. U. &amp; Godoy, C. (2002) The Ichneumonidae of Costa Rica, 4. Memoirs of the American Entomological Institute, 66, 1 - 768." title="The Ichneumonidae of Costa Rica, 4" type="journal article" year="2002">Gauld &amp; Sithole, 2002</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="CF3D36E95C561649F9FFF8FFFF3AF8F9" blockId="5.[151,610,1833,1893]" box="[151,235,1868,1893]" pageId="5" pageNumber="45">
(
<figureCitation id="57B92A6C5C561649F9F7F8FFFF33F8F9" box="[159,226,1868,1893]" captionStart="FIGURES 2 5" captionStartId="3.[151,264,1174,1197]" captionTargetBox="[340,1246,193,1152]" captionTargetId="figure@3.[340,1247,193,1153]" captionTargetPageId="3" captionText="FIGURES 2 5. Exochus spp. 2 4, Exochus ablatus, ♀: 2, habitus, lateral; 3, head, dorsolateral; 4, mesoscutum, propodeum and metasoma, dorsal. 5, Exochus izbus, ♀: 5, habitus, lateral." figureDoi="http://doi.org/10.5281/zenodo.236613" httpUri="https://zenodo.org/record/236613/files/figure.png" pageId="5" pageNumber="45">Fig. 5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="879865625C561649F9FFF827FD13F868" pageId="5" pageNumber="45" type="diagnosis">
<paragraph id="CF3D36E95C561649F9FFF827FD13F868" blockId="5.[151,1436,1940,2037]" pageId="5" pageNumber="45">
<emphasis id="FDF6EAFB5C561649F9FFF827FEC0F831" bold="true" box="[151,273,1940,1965]" pageId="5" pageNumber="45">Diagnosis.</emphasis>
This species has an almost completely carinate propodeum with a well-defined area superomedia and very strongly tapered mandibles that are apically twisted, so when they are closed they appear more or less unidentate and needle-sharp (
<bibRefCitation id="AB134B185C561649F888F86FFD66F868" author="Gauld, I. &amp; Sithole, R. &amp; Gomes, J. U. &amp; Godoy, C." box="[480,695,2012,2037]" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="5" pageNumber="45" pagination="1 - 768" part="66" refId="ref6711" refString="Gauld, I., Sithole, R., Gomes, J. U. &amp; Godoy, C. (2002) The Ichneumonidae of Costa Rica, 4. Memoirs of the American Entomological Institute, 66, 1 - 768." title="The Ichneumonidae of Costa Rica, 4" type="journal article" year="2002">
Gauld,
<emphasis id="FDF6EAFB5C561649FB5CF86EFDB4F869" box="[564,613,2012,2037]" italics="true" pageId="5" pageNumber="45">et al</emphasis>
., 2002
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="879865625C55164AF9AFFF24FAADFF48" pageId="6" pageNumber="46" type="discussion">
<paragraph id="CF3D36E95C55164AF9AFFF24FAADFF48" blockId="6.[151,1437,151,536]" pageId="6" pageNumber="46">
<emphasis id="FDF6EAFB5C55164AF9AFFF24FEEAFF2C" bold="true" box="[199,315,151,176]" pageId="6" pageNumber="46">Remarks.</emphasis>
<bibRefCitation id="AB134B185C55164AF82BFF24FDD9FF2C" author="Gauld, I. &amp; Sithole, R. &amp; Gomes, J. U. &amp; Godoy, C." box="[323,520,151,176]" etAl="et al." firstAuthor="Gauld" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="6" pageNumber="46" pagination="1 - 768" part="66" refId="ref6711" refString="Gauld, I., Sithole, R., Gomes, J. U. &amp; Godoy, C. (2002) The Ichneumonidae of Costa Rica, 4. Memoirs of the American Entomological Institute, 66, 1 - 768." title="The Ichneumonidae of Costa Rica, 4" type="journal article" year="2002">
Gauld
<emphasis id="FDF6EAFB5C55164AF8E7FF2AFE18FF2C" box="[399,457,151,176]" italics="true" pageId="6" pageNumber="46">et al.</emphasis>
2002
</bibRefCitation>
reported considerable variation in color in this species, however we did not find such variation in the color pattern of the species when comparing our material with the
<typeStatus id="1039884B5C55164AFD02FF0EFB07FF49" box="[1130,1238,189,213]" pageId="6" pageNumber="46" type="paratype">paratypes</typeStatus>
at the BMNH.
</paragraph>
</subSubSection>
<subSubSection id="879865625C55164AF9AFFF6CFAE7FE81" pageId="6" pageNumber="46" type="discussion">
<paragraph id="CF3D36E95C55164AF9AFFF6CFAE7FE81" blockId="6.[151,1437,151,536]" pageId="6" pageNumber="46">
<emphasis id="FDF6EAFB5C55164AF9AFFF6CFEB1FF64" bold="true" box="[199,352,223,248]" pageId="6" pageNumber="46">Distribution.</emphasis>
<collectingCountry id="B79576795C55164AF803FF53FE3AFF64" box="[363,491,223,248]" name="Costa Rica" pageId="6" pageNumber="46">Costa Rica</collectingCountry>
,
<collectingCountry id="B79576795C55164AF893FF6CFD92FF64" box="[507,579,223,248]" name="Brazil" pageId="6" pageNumber="46">Brazil</collectingCountry>
. The specimens were collected in Brazilian savannah (Itapeva Ecological Station) and semideciduous forest (Mogi Guaçu Ecological Station), in São Paulo State, Southeast
<collectingCountry id="B79576795C55164AFD81FEB7FAE0FE81" box="[1257,1329,260,285]" name="Brazil" pageId="6" pageNumber="46">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="879865625C55164AF9AFFE94FE75FEDC" box="[199,420,295,320]" pageId="6" pageNumber="46" type="discussion">
<paragraph id="CF3D36E95C55164AF9AFFE94FE75FEDC" blockId="6.[151,1437,151,536]" box="[199,420,295,320]" pageId="6" pageNumber="46">
<emphasis id="FDF6EAFB5C55164AF9AFFE94FEF6FEDC" bold="true" box="[199,295,295,320]" pageId="6" pageNumber="46">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="879865625C55164AF9AFFEFFFB1CFD84" pageId="6" pageNumber="46" type="materials_examined">
<paragraph id="CF3D36E95C55164AF9AFFEFFFB1CFD84" blockId="6.[151,1437,151,536]" pageId="6" pageNumber="46">
<emphasis id="FDF6EAFB5C55164AF9AFFEFFFDFDFEF9" bold="true" box="[199,556,332,357]" pageId="6" pageNumber="46">
Material examined. (
<collectionCode id="A993AE2C5C55164AF8A1FEFFFDF2FEF9" box="[457,547,332,357]" pageId="6" pageNumber="46">BMNH</collectionCode>
)
</emphasis>
<typeStatus id="1039884B5C55164AFB5FFEFEFD75FEF9" box="[567,676,333,357]" pageId="6" pageNumber="46" type="paratype">Paratypes</typeStatus>
4 ♀,
<specimenCount id="D984FD605C55164AFB97FEFFFCE1FEF8" box="[767,816,332,356]" pageId="6" pageNumber="46" type="generic">2 ♂</specimenCount>
, Guanacaste National Park, Cerro el Hacha,
<quantity id="087A9B0C5C55164AFC21FEFFFA49FEF8" box="[1353,1432,332,356]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="46" unit="m" value="400.0">400 m</quantity>
,
<date id="BB3C10295C55164AF9FFFEDCFF3DFE14" box="[151,236,367,392]" pageId="6" pageNumber="46" value="1986-11">xi.1986</date>
<date id="BB3C10295C55164AF995FEDCFE92FE14" box="[253,323,367,392]" pageId="6" pageNumber="46" value="1987-01">i.1987</date>
, Gauld &amp; Janzen colls.
<emphasis id="FDF6EAFB5C55164AFB08FEDCFD13FE14" bold="true" box="[608,706,367,392]" pageId="6" pageNumber="46">
(
<collectionCode id="A993AE2C5C55164AFB0FFEDCFD68FE14" box="[615,697,367,392]" httpUri="http://grbio.org/cool/5zzs-n4tg" name="Universidade Federal de Sao Carlos" pageId="6" pageNumber="46">DCBU</collectionCode>
)
</emphasis>
2 ♀,
<collectingCountry id="B79576795C55164AFA64FEDCFC82FE14" box="[780,851,367,392]" name="Brazil" pageId="6" pageNumber="46">Brazil</collectingCountry>
, SP, Itapeva, Itapeva Ecological Station, Brazilian savannah,
<geoCoordinate id="AAB6502E5C55164AF866FE27FE44FE30" box="[270,405,404,428]" direction="south" orientation="latitude" pageId="6" pageNumber="46" precision="1" value="-24.069084">24°48.7” S</geoCoordinate>
<geoCoordinate id="AAB6502E5C55164AF8F5FE27FDEDFE30" box="[413,572,404,429]" direction="west" orientation="longitude" pageId="6" pageNumber="46" precision="1" value="-49.06475">49°353.1” W</geoCoordinate>
, Malaise trap,
<date id="BB3C10295C55164AFB8CFE27FCB2FE30" box="[740,867,404,428]" pageId="6" pageNumber="46" value="2008-04-14">14.IV.2008</date>
, A.M.P Dias coll;
<specimenCount id="D984FD605C55164AFD51FE27FBB7FE30" box="[1081,1126,404,428]" pageId="6" pageNumber="46" type="generic">1 ♂</specimenCount>
, same locality, same date,1 ♀,
<collectingCountry id="B79576795C55164AF9D3FE04FED3FE4C" box="[187,258,439,464]" name="Brazil" pageId="6" pageNumber="46">Brazil</collectingCountry>
, SP, Mogi Guaçu, Mogi Guaçu Ecological Station,
<geoCoordinate id="AAB6502E5C55164AFA24FE04FC09FE4C" box="[844,984,439,464]" direction="south" orientation="latitude" pageId="6" pageNumber="46" precision="15" value="-22.265833">22°1557” S</geoCoordinate>
<geoCoordinate id="AAB6502E5C55164AFA89FE04FBA8FE53" box="[993,1145,439,464]" direction="west" orientation="longitude" pageId="6" pageNumber="46" precision="15" value="-47.17389">47°1026” W</geoCoordinate>
, Malaise trap,
<date id="BB3C10295C55164AFC49FE04FA49FE4C" box="[1313,1432,439,464]" pageId="6" pageNumber="46" value="2007-02-05">05.II.2007</date>
, A.M.P. Dias coll; 1 ♀,
<collectingCountry id="B79576795C55164AF8F0FE6FFE0EFE68" box="[408,479,476,500]" name="Brazil" pageId="6" pageNumber="46">Brazil</collectingCountry>
, SP, Luiz Antônio, Jataí Ecological Station,
<date id="BB3C10295C55164AFAA7FE6FFB80FE68" box="[975,1105,476,500]" pageId="6" pageNumber="46" value="1993-09-28">28.IX.1993</date>
, A.S.Soares coll; 1 ♀,
<collectingCountry id="B79576795C55164AFC39FE6FFA49FE68" box="[1361,1432,476,500]" name="Brazil" pageId="6" pageNumber="46">Brazil</collectingCountry>
, SP, São Carlos, Embrapa, Pecuária Sudeste, Canchim Farm, forest,
<date id="BB3C10295C55164AFAE4FE4CFBD0FD84" box="[908,1025,511,536]" pageId="6" pageNumber="46" value="1985-05-03">03.V.1985</date>
, A.M.P. Dias coll.
</paragraph>
</subSubSection>
</treatment>
</document>