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<document id="58E6F4B7FA2C4D941F202E5D1058D301" ID-CLB-Dataset="46567" ID-DOI="10.5281/zenodo.204537" ID-GBIF-Dataset="29ee5ed1-fdc0-4cff-9166-d63f024a2e2d" ID-ISSN="1175-5326" ID-Zenodo-Dep="204537" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460375661013" checkinUser="plazi" docAuthor="Kolibáč, Jiří" docDate="2011" docId="152787A5CF48FFE27CA7FDC1FE657DDD" docLanguage="en" docName="zt02928p063.pdf" docOrigin="Zootaxa 2928" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Promanodes alleni Kolibáč, 2011, sp. nov." docType="treatment" docVersion="9" lastPageNumber="62" masterDocId="E91EFFDDCF4AFFE77C30FFADFF9E7C34" masterDocTitle="Promanodes alleni sp. nov., the second species of the Tertiary genus Promanodes Kolibáč, Schmied, Wappler et Kubisz, 2010, with improved diagnosis of the genus and remarks on its phylogeny (Coleoptera: Trogossitidae)" masterLastPageNumber="63" masterPageNumber="57" pageNumber="59" updateTime="1698296240622" updateUser="ExternalLinkService">
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<mods:title id="0D088B9045962BA4C6FB76C411A795B3">Promanodes alleni sp. nov., the second species of the Tertiary genus Promanodes Kolibáč, Schmied, Wappler et Kubisz, 2010, with improved diagnosis of the genus and remarks on its phylogeny (Coleoptera: Trogossitidae)</mods:title>
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<mods:namePart id="C3722535C6431A0747DB4D2F8C32C16D">Kolibáč, Jiří</mods:namePart>
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<mods:date id="7F0B49D436947D7D70BF8536858CBB77">2011</mods:date>
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<treatment id="152787A5CF48FFE27CA7FDC1FE657DDD" ID-DOI="http://doi.org/10.5281/zenodo.6191274" ID-GBIF-Taxon="119413056" ID-Zenodo-Dep="6191274" LSID="urn:lsid:zoobank.org:act:3E7882A0-1341-4843-9BCE-7C3531B23B58" httpUri="http://treatment.plazi.org/id/152787A5CF48FFE27CA7FDC1FE657DDD" lastPageId="5" lastPageNumber="62" pageId="2" pageNumber="59">
<subSubSection id="D5946538CF48FFE57CA7FDC1FE957E93" pageId="2" pageNumber="59" type="nomenclature">
<paragraph id="9D3136B3CF48FFE57CA7FDC1FE417EB2" blockId="2.[151,479,620,679]" box="[151,479,620,646]" pageId="2" pageNumber="59">
<heading id="C67981DFCF48FFE57CA7FDC1FE417EB2" bold="true" box="[151,479,620,646]" fontSize="11" level="1" pageId="2" pageNumber="59" reason="1">
<emphasis id="AFFAEAA1CF48FFE57CA7FDC1FE417EB2" bold="true" box="[151,479,620,646]" pageId="2" pageNumber="59">
<taxonomicName id="5A8E4D30CF48FFE57CA7FDC1FEE77EB2" box="[151,377,620,646]" class="Insecta" family="Trogossitidae" genus="Promanodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="alleni" status="sp. nov.">
<emphasis id="AFFAEAA1CF48FFE57CA7FDC1FEE77EB2" bold="true" box="[151,377,620,646]" italics="true" pageId="2" pageNumber="59">Promanodes alleni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4C957DACF48FFE57DB1FDC1FE417EB2" box="[385,479,620,646]" pageId="2" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9D3136B3CF48FFE57CA7FD22FE957E93" blockId="2.[151,479,620,679]" box="[151,267,655,679]" pageId="2" pageNumber="59">
(
<figureCitation id="05B52A36CF48FFE57CAFFD22FE9C7E93" box="[159,258,655,679]" captionStart-0="FIGURES 1 4" captionStart-1="FIGURES 5 7" captionStartId-0="3.[151,264,1976,1999]" captionStartId-1="4.[151,264,1966,1989]" captionTargetBox-0="[189,1378,206,1942]" captionTargetBox-1="[207,1378,200,1935]" captionTargetId-0="figure@3.[189,1379,193,1943]" captionTargetId-1="figure@4.[196,1383,193,1938]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 1 4. Promanodes alleni sp. nov.: (1) body dorsally; (2) body ventrally; (3) head and pronotum dorsally; (4) hind legs." captionText-1="FIGURES 5 7. Promanodes alleni sp. nov.: (5) body ventrally, partial reconstruction; (6) hind legs (excl. coxae); (7) head ventrally." httpUri-0="https://zenodo.org/record/204538/files/figure.png" httpUri-1="https://zenodo.org/record/204539/files/figure.png" pageId="2" pageNumber="59">Figs 17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D5946538CF48FFE57CA7FD7AFDDB7FF3" pageId="2" pageNumber="59" type="materials_examined">
<paragraph id="9D3136B3CF48FFE57CA7FD7AFDA17F03" blockId="2.[151,1437,727,2011]" pageId="2" pageNumber="59">
<emphasis id="AFFAEAA1CF48FFE57CA7FD7AFE957EDB" bold="true" box="[151,267,727,751]" pageId="2" pageNumber="59">
<typeStatus id="42358811CF48FFE57CA7FD7AFE987EDB" box="[151,262,727,751]" pageId="2" pageNumber="59" type="holotype">Holotype</typeStatus>
.
</emphasis>
Sex unknown, Baltic amber. Exact place of collecting unknown (East Baltic coast?), probably Eocene. Amber piece yellowish-orange, elongate, approximate size 44×9×
<quantity id="5A769B56CF48FFE57FA7FD56FC7E7F27" box="[919,992,763,787]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="59" unit="mm" value="4.0">4 mm</quantity>
. Deposited in the Moravian museum, Brno,
<collectingCountry id="E5997623CF48FFE57CEDFCB2FEA97F03" box="[221,311,799,823]" name="Czech Republic" pageId="2" pageNumber="59">Czechia</collectingCountry>
(J. Kolibáč collection).
</paragraph>
<paragraph id="9D3136B3CF48FFE57CF5FCEEFDDB7FF3" blockId="2.[151,1437,727,2011]" pageId="2" pageNumber="59">
<emphasis id="AFFAEAA1CF48FFE57CF5FCEEFE557F68" bold="true" box="[197,459,835,860]" pageId="2" pageNumber="59">Differential diagnosis.</emphasis>
Body shape as same as in
<taxonomicName id="5A8E4D30CF48FFE57EC4FCE9FCFB7F6F" box="[756,869,835,859]" class="Insecta" family="Trogossitidae" genus="Promanodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="serafini">
<emphasis id="AFFAEAA1CF48FFE57EC4FCE9FCFB7F6F" box="[756,869,835,859]" italics="true" pageId="2" pageNumber="59">P. serafini</emphasis>
</taxonomicName>
but the new species is distinctly smaller (ca.
<quantity id="5A769B56CF48FFE5796CFCEEFA057F6F" box="[1372,1435,835,859]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="59" unit="mm" value="3.0">3 mm</quantity>
vs.
<quantity id="5A769B56CF48FFE57CF2FCCAFE997F4B" box="[194,263,871,895]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="59" unit="mm" value="5.0">5 mm</quantity>
in
<taxonomicName id="5A8E4D30CF48FFE57D02FCC5FE397F4B" box="[306,423,871,895]" class="Insecta" family="Trogossitidae" genus="Promanodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="serafini">
<emphasis id="AFFAEAA1CF48FFE57D02FCC5FE397F4B" box="[306,423,871,895]" italics="true" pageId="2" pageNumber="59">P. serafini</emphasis>
</taxonomicName>
), head, pronotum and elytra with distinct pubescence (head bare, pronotum and elytra pubescent along margins in
<taxonomicName id="5A8E4D30CF48FFE57DE1FC21FDDC7F97" box="[465,578,907,931]" class="Insecta" family="Trogossitidae" genus="Promanodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="serafini">
<emphasis id="AFFAEAA1CF48FFE57DE1FC21FDDC7F97" box="[465,578,907,931]" italics="true" pageId="2" pageNumber="59">P. serafini</emphasis>
</taxonomicName>
), pronotum with much finer sculpture than that in
<taxonomicName id="5A8E4D30CF48FFE5784CFC21FB727F97" box="[1148,1260,907,931]" class="Insecta" family="Trogossitidae" genus="Promanodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="serafini">
<emphasis id="AFFAEAA1CF48FFE5784CFC21FB727F97" box="[1148,1260,907,931]" italics="true" pageId="2" pageNumber="59">P. serafini</emphasis>
</taxonomicName>
, procoxal cavities maybe perfectly closed externally.
</paragraph>
</subSubSection>
<subSubSection id="D5946538CF48FFE27CF5FC7EFD947D95" lastPageId="5" lastPageNumber="62" pageId="2" pageNumber="59" type="description">
<paragraph id="9D3136B3CF48FFE57CF5FC7EFCF478F7" blockId="2.[151,1437,727,2011]" pageId="2" pageNumber="59">
<emphasis id="AFFAEAA1CF48FFE57CF5FC7EFEC97FD8" bold="true" box="[197,343,979,1004]" pageId="2" pageNumber="59">Description.</emphasis>
Body flat, lateral margins of pronotum and elytra distinctly explanate; dorsal and ventral surface probably unicolorous, dark. Head with coarse but sparse punctation (similar to that in
<taxonomicName id="5A8E4D30CF48FFE5786DFC55FB50783B" box="[1117,1230,1015,1039]" class="Insecta" family="Trogossitidae" genus="Promanodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="serafini">
<emphasis id="AFFAEAA1CF48FFE5786DFC55FB50783B" box="[1117,1230,1015,1039]" italics="true" pageId="2" pageNumber="59">P. serafini</emphasis>
</taxonomicName>
) formed by round and relatively deep punctures (interspaces among punctures as large as or larger than their diameter); sculpture of pronotum fine and sparse; each elytron with 7 carinae, 7th carina (running along elytral margin) coalescent with 6th in basal quarter of elytron; two regular rows of punctures situated between adjacent carinae; head with thick, relatively long, erect, sparse pubesce; pronotum and elytra with fine, short and sparse pubescence. Ventral side without pubescence, smooth or with very fine and sparse punctures only.
</paragraph>
<paragraph id="9D3136B3CF48FFE57CF5FB62FA13798B" blockId="2.[151,1437,727,2011]" pageId="2" pageNumber="59">
Head prognathous, approximately as wide as long; gular sutures not observed; eyes relatively large and distinctly elevated, coarsely faceted; space between eyes about four times wider than eye diameter; frontoclypeal suture deep, incurvate backwards, but not so distinct as in
<taxonomicName id="5A8E4D30CF48FFE57F17FAB5FC06791B" box="[807,920,1303,1327]" class="Insecta" family="Trogossitidae" genus="Promanodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="serafini">
<emphasis id="AFFAEAA1CF48FFE57F17FAB5FC06791B" box="[807,920,1303,1327]" italics="true" pageId="2" pageNumber="59">P. serafini</emphasis>
</taxonomicName>
; distinct subantennal grooves present. Antennae probably 10-segmented: scapus robust, pedicellus short but thick, segments 3 to 7 short (not clearly visible), antennal club 3-segmented, large, weakly asymmetrical. Mandibles robust, distinctly bidentate at apex. Maxilla with 4-segmented palpi; the last segment coniform and elongate (not securiform) little longer than two previous segments together. Labial palpi distinctly shorter than maxillary ones; the last segment widened at apex, truncate.
</paragraph>
<paragraph id="9D3136B3CF48FFE57CF5FA66FE2B7A47" blockId="2.[151,1437,727,2011]" pageId="2" pageNumber="59">Prothorax distinctly transverse; lateral margins of pronotum broadly explanate, denticulate and distinctly pubescent along side margins; anterior corners not projecting. Prosternal intercoxal process long and slender, not dilated at apex; prosternal process area distinctly elevated above prosternum. Coxal cavities widely transverse, narrowly separated, surely externally closed in more than half of coxal width or maybe perfectly closed by postcoxal projection of hypomeron.</paragraph>
<paragraph id="9D3136B3CF48FFE57CF5F9D2FAF07B37" blockId="2.[151,1437,727,2011]" pageId="2" pageNumber="59">
Mesothorax wide and flat; mesepimeron and mesepisternum not distinctly visible; mesocoxal cavities weakly transverse, narrowly separated; mesosternal process slender, narrowed towards apex (as in
<taxonomicName id="5A8E4D30CF48FFE578A4F909FA9B7A8F" box="[1172,1285,1699,1723]" class="Insecta" family="Trogossitidae" genus="Promanodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="serafini">
<emphasis id="AFFAEAA1CF48FFE578A4F909FA9B7A8F" box="[1172,1285,1699,1723]" italics="true" pageId="2" pageNumber="59">P. serafini</emphasis>
</taxonomicName>
). Elytral epipleure somewhat narrower than in
<taxonomicName id="5A8E4D30CF48FFE57E10F965FD0F7AEB" box="[544,657,1735,1759]" class="Insecta" family="Trogossitidae" genus="Promanodes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="serafini">
<emphasis id="AFFAEAA1CF48FFE57E10F965FD0F7AEB" box="[544,657,1735,1759]" italics="true" pageId="2" pageNumber="59">P. serafini</emphasis>
</taxonomicName>
, the widest at humeral portion, than fluently narrowed; elytra widest at 2/3 of length, their humeral portion as wide as pronotal base. Elytra with distinct 7 carinae, rounded at apex.
</paragraph>
<paragraph id="9D3136B3CF48FFE57CF5F8A2FD557B5B" blockId="2.[151,1437,727,2011]" pageId="2" pageNumber="59">Metathorax wide, flat, very sparsely and finely punctured; metepisterna relatively wide and nearly parallelsided. Discriminal line (discrimen) inconspicuous, probably absent; paracoxal sutures distinct. Winged (apices of wings visible), wing venation hidden under elytra.</paragraph>
<paragraph id="9D3136B3CF48FFE27CF5F8D1FD8C7D20" blockId="2.[151,1437,727,2011]" lastBlockId="5.[151,1436,151,489]" lastPageId="5" lastPageNumber="62" pageId="2" pageNumber="59">Legs. All pairs of coxae narrowly separated; procoxae relatively small, transverse; mesocoxae somewhat projecting, nearly rounded; metacoxae elongate, flat, reaching elytral epipleure; trochanters very small, triangular; all pairs of femora conspicuously clavate; tibiae slender, with at least one long and straight apical spine, without hooked apical spine and without conspicuous lateral spines along sides of tibiae and apical spines at tibial apex. Tarsi elongate, extremely long as long as respective tibiae; tarsomeres without lobes; the last tarsomere as long as tarsomeres 14 together; basal tarsomeres shorter than each of others; claws long as long as each of tarsomeres 24; empodium distictly bisetose.</paragraph>
<caption id="C9F1663BCF49FFE47CA7F815FF597BD9" httpUri="https://zenodo.org/record/204538/files/figure.png" pageId="3" pageNumber="60" targetBox="[189,1378,206,1942]" targetPageId="3">
<paragraph id="9D3136B3CF49FFE47CA7F815FF597BD9" blockId="3.[151,1436,1976,2029]" pageId="3" pageNumber="60">
<emphasis id="AFFAEAA1CF49FFE47CA7F815FEA27BFA" bold="true" box="[151,316,1976,1999]" pageId="3" pageNumber="60">FIGURES 14.</emphasis>
<taxonomicName id="5A8E4D30CF49FFE47D73F815FD9A7BFA" box="[323,516,1976,1998]" class="Insecta" family="Trogossitidae" genus="Promanodes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="alleni" status="sp. nov.">
<emphasis id="AFFAEAA1CF49FFE47D73F815FD9A7BFA" box="[323,516,1976,1998]" italics="true" pageId="3" pageNumber="60">Promanodes alleni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4C957DACF49FFE47E3CF814FDC37BFA" box="[524,605,1977,1998]" pageId="3" pageNumber="60" rank="species">
<emphasis id="AFFAEAA1CF49FFE47E3CF814FDC37BFA" bold="true" box="[524,605,1977,1998]" pageId="3" pageNumber="60">sp. nov.</emphasis>
</taxonomicNameLabel>
: (1) body dorsally; (2) body ventrally; (3) head and pronotum dorsally; (4) hind legs.
</paragraph>
</caption>
<caption id="C9F1663BCF4EFFE37CA7F803FF687BD7" httpUri="https://zenodo.org/record/204539/files/figure.png" pageId="4" pageNumber="61" targetBox="[207,1378,200,1935]" targetPageId="4">
<paragraph id="9D3136B3CF4EFFE37CA7F803FF687BD7" blockId="4.[151,1436,1966,2019]" pageId="4" pageNumber="61">
<emphasis id="AFFAEAA1CF4EFFE37CA7F803FEA27BF0" bold="true" box="[151,316,1966,1989]" pageId="4" pageNumber="61">FIGURES 57.</emphasis>
<taxonomicName id="5A8E4D30CF4EFFE37D75F802FD987BF1" box="[325,518,1967,1989]" class="Insecta" family="Trogossitidae" genus="Promanodes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="alleni" status="sp. nov.">
<emphasis id="AFFAEAA1CF4EFFE37D75F802FD987BF1" box="[325,518,1967,1989]" italics="true" pageId="4" pageNumber="61">Promanodes alleni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B4C957DACF4EFFE37E3FF802FDFF7BF0" box="[527,609,1967,1988]" pageId="4" pageNumber="61" rank="species">
<emphasis id="AFFAEAA1CF4EFFE37E3FF802FDFF7BF0" bold="true" box="[527,609,1967,1988]" pageId="4" pageNumber="61">sp. nov.</emphasis>
</taxonomicNameLabel>
: (5) body ventrally, partial reconstruction; (6) hind legs (excl. coxae); (7) head ventrally.
</paragraph>
</caption>
<paragraph id="9D3136B3CF4FFFE27CF5FEB0FCE87D01" blockId="5.[151,1436,151,489]" box="[197,886,285,309]" pageId="5" pageNumber="62">
Abdomen distinctly with six visible ventrites as in
<taxonomicName id="5A8E4D30CF4FFFE27ECEFEB3FCF17D01" box="[766,879,285,309]" class="Insecta" family="Trogossitidae" genus="Promanodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="serafini">
<emphasis id="AFFAEAA1CF4FFFE27ECEFEB3FCF17D01" box="[766,879,285,309]" italics="true" pageId="5" pageNumber="62">P. serafini</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9D3136B3CF4FFFE27CF5FEECFD947D95" blockId="5.[151,1436,151,489]" pageId="5" pageNumber="62">
<emphasis id="AFFAEAA1CF4FFFE27CF5FEECFEE57D6D" bold="true" box="[197,379,321,345]" pageId="5" pageNumber="62">Measurements.</emphasis>
Body length (without mandibles) 3.15; width of head incl. eyes 0.77; pronotum length in middle 0.53; pronotum maximal width 1.30; elytral length 2.33; elytron width at base 0.53; elytron maximal width 0.70. (All measurements in mm.)
</paragraph>
</subSubSection>
<subSubSection id="D5946538CF4FFFE27CF5FE00FE657DDD" pageId="5" pageNumber="62" type="etymology">
<paragraph id="9D3136B3CF4FFFE27CF5FE00FE657DDD" blockId="5.[151,1436,151,489]" pageId="5" pageNumber="62">
<emphasis id="AFFAEAA1CF4FFFE27CF5FE00FED27DF1" bold="true" box="[197,332,429,453]" pageId="5" pageNumber="62">Etymology.</emphasis>
The new species is named in honour of Mr. Albert Allen (Boise, Idaho,
<collectingCountry id="E5997623CF4FFFE278A6FE00FB4C7DF1" box="[1174,1234,429,453]" name="United States of America" pageId="5" pageNumber="62">USA</collectingCountry>
), who generously donated the specimen for study.
</paragraph>
</subSubSection>
</treatment>
</document>