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<mods:title id="59F5F30CF938904571F244CA6836B165">A New Species Of Semi-Aquatic Freshwater Earthworm Of The Genus Glyphidrilus Horst, 1889 From The Mekong River (Oligochaeta: Almidae)</mods:title>
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<paragraph id="8C6536D0242DFFE1B6894D64C23BF8F5" blockId="3.[896,1367,1835,1892]" box="[896,1367,1835,1860]" pageId="3" pageNumber="268">
<heading id="D72D81BC242DFFE1B6894D64C23BF8F5" bold="true" box="[896,1367,1835,1860]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="268" reason="2">
<taxonomicName id="4BDA4D53242DFFE1B6894D64C23BF8F5" authority="Michaelsen, 1902" authorityName="Michaelsen" authorityYear="1902" box="[896,1367,1835,1860]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="268" phylum="Annelida" rank="species" species="malayanus">
<emphasis id="BEAEEAC2242DFFE1B6894D64C23BF8F5" bold="true" box="[896,1367,1835,1860]" pageId="3" pageNumber="268">
<emphasis id="BEAEEAC2242DFFE1B6894D64C3E6F8F5" bold="true" box="[896,1162,1835,1860]" italics="true" pageId="3" pageNumber="268">Glyphidrilus malayanus</emphasis>
<bibRefCitation id="E84B4B21242DFFE1B19B4D64C23BF8F5" author="Michaelsen, W" box="[1170,1367,1835,1860]" pageId="3" pageNumber="268" pagination="1 - 54" refId="ref7301" refString="Michaelsen, W., 1902. Neue oligochaeten und neue Fundorte altbekannter. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 19: 1 - 54." type="journal article" year="1902">Michaelsen, 1902</bibRefCitation>
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<paragraph id="8C6536D0242DFFE1B12E4D03C3DDF8D5" blockId="3.[896,1367,1835,1892]" box="[1063,1201,1868,1892]" pageId="3" pageNumber="268">
(
<figureCitation id="14E12A55242DFFE1B1264D03C3E2F8D5" box="[1071,1166,1868,1892]" captionStart="Fig" captionStartId="5.[112,147,1918,1940]" captionTargetBox="[113,752,1213,1895]" captionTargetId="figure-285@5.[113,754,1213,1895]" captionTargetPageId="5" captionText="Fig. 3. The colouration of newly collected specimens just after the first step preservation in 30% ethanol:A, Glyphidrilus mekongensis, new species (paratype CUMZ 3216); B, Glyphidrilus malayanus from Tok Bok Hotspring, Kelantan, Malaysia (CUMZ 3218); C, Glyphidrilus singaporensis topotype specimen (CUMZ 3217); D, cocoon characters of Glyphidrilus mekongensis, new species (paratype CUMZ 3216)." figureDoi="http://doi.org/10.5281/zenodo.4508504" httpUri="https://zenodo.org/record/4508504/files/figure.png" pageId="3" pageNumber="268">Figs. 3B</figureCitation>
, 
<figureCitation id="14E12A55242DFFE1B1934D03C3C4F8D5" box="[1178,1192,1868,1892]" captionStart="Fig" captionStartId="7.[113,148,2034,2056]" captionText="Fig. 5. External and internal morphology of holotype (ZMH V5875) of Glyphidrilus malayanus: A, external ventral view; B, internal dorsal view; C, specimen collected from Sungei Kelatan, Kelantan, Malaysia (CUMZ 3219); D, from Sungei Berkoh, Johor, Malaysia (CUMZ 3220); E, from Tok Bok Hotspring, Kelantan, Malaysia (CUMZ 3218), Malaysia." pageId="3" pageNumber="268">5</figureCitation>
)
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<treatmentCitation id="0D7B10C1242DFFE1B6254DC4C38CF811" author="Michaelsen, W" box="[812,1248,1930,1952]" page="35" pageId="3" pageNumber="268" year="1902">
<taxonomicName id="4BDA4D53242DFFE1B6254DC4C38CF811" authority="Michaelsen, 1902: 35" authorityName="Michaelsen" authorityPageNumber="35" authorityYear="1902" box="[812,1248,1930,1952]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="3" pageNumber="268" phylum="Annelida" rank="species" species="malayanus">
<emphasis id="BEAEEAC2242DFFE1B6254DC4C37CF82E" box="[812,1040,1931,1951]" italics="true" pageId="3" pageNumber="268">Glyphidrilus malayanus</emphasis>
<bibRefCitation id="E84B4B21242DFFE1B11D4DC5C38CF811" author="Michaelsen, W" box="[1044,1248,1930,1952]" pageId="3" pageNumber="268" pagination="1 - 54" refId="ref7301" refString="Michaelsen, W., 1902. Neue oligochaeten und neue Fundorte altbekannter. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 19: 1 - 54." type="journal article" year="1902">Michaelsen, 1902: 35</bibRefCitation>
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. 
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locality: Malay Peninsula: Lubock Paku, 
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River. 
<treatmentCitation id="0D7B10C1242DFFE1B1C44DE8C2C4F80D" author="Michaelsen, W" box="[1229,1448,1959,1981]" page="104" pageId="3" pageNumber="268" year="1910">
<bibRefCitation id="E84B4B21242DFFE1B1C44DE8C2C4F80D" author="Michaelsen, W" box="[1229,1448,1959,1981]" pageId="3" pageNumber="268" pagination="1 - 108" refId="ref7327" refString="Michaelsen, W., 1910. Die oligochatenfauna der vonderindischceylonischen region. Abhandulngen aus dem Gebiete der Naturwissenschaften, 19: 1 - 108." type="journal article" year="1910">Michaelsen, 1910: 104</bibRefCitation>
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. Brinkhurst &amp; Jamieson, 1971: 762, Fig. 
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, F.
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<emphasis id="BEAEEAC2242DFFE1B6254DB2C499F7A2" bold="true" box="[812,1013,2045,2067]" pageId="3" pageNumber="268">
<emphasis id="BEAEEAC2242DFFE1B6254DB2C483F7A2" bold="true" box="[812,1007,2045,2067]" italics="true" pageId="3" pageNumber="268">Material examined</emphasis>
.
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— 
<typeStatus id="53618872242DFFE1B1294DB2C317F7A2" box="[1056,1147,2045,2067]" pageId="3" pageNumber="268" type="holotype">Holotype</typeStatus>
from 
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annelid reference collections (
<collectionCode id="EACBAE15242DFFE1B6AF4255C4B2F79E" box="[934,990,2074,2095]" country="Germany" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="3" pageNumber="268" type="Museum">ZMH</collectionCode>
V5875, 
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). Additional records: 110
</paragraph>
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<paragraph id="8C6536D0242AFFE6B5794882C23FFCAC" blockId="4.[112,2015,717,797]" pageId="4" pageNumber="269">
Table 2. Morphological characters comparison of 
<taxonomicName id="4BDA4D53242AFFE6B75F4880C43EFD52" authority="Chanabun &amp; Bantaowong &amp; Sutcharit &amp; Tongkerd &amp; James &amp; Panha, 2012" authorityName="Chanabun &amp; Bantaowong &amp; Sutcharit &amp; Tongkerd &amp; James &amp; Panha" authorityYear="2012" box="[598,850,719,739]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="mekongensis" status="sp. nov.">
<emphasis id="BEAEEAC2242AFFE6B75F4880C43EFD52" box="[598,850,719,739]" italics="true" pageId="4" pageNumber="269">Glyphidrilus mekongensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A59D57B9242AFFE6B6564882C4BAFD52" box="[863,982,717,739]" pageId="4" pageNumber="269" rank="species">new species</taxonomicNameLabel>
, 
<taxonomicName id="4BDA4D53242AFFE6B6E94880C389FD52" authority="(Rosa, 1890)" baseAuthorityName="Rosa" baseAuthorityYear="1890" box="[992,1253,717,739]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="papillatus">
<emphasis id="BEAEEAC2242AFFE6B6E94880C333FD52" box="[992,1119,719,739]" italics="true" pageId="4" pageNumber="269">G. papillatus</emphasis>
(
<bibRefCitation id="E84B4B21242AFFE6B1644882C3B2FD52" author="Rosa, D" box="[1133,1246,717,739]" pageId="4" pageNumber="269" pagination="386 - 400" refId="ref7353" refString="Rosa, D., 1890. Viaggio di Leonardo Fea in Birmania e Regioni vicine, xxv, Moniligastridi, Geoscolecidi ed Eudrilidi (I 2). Annali del Museo Civico di Storia Naturale di Genova, 9: 386 - 400." type="journal article" year="1890">Rosa, 1890</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4BDA4D53242AFFE6B1F94880C140FD52" authority="Michaelsen, 1902" authorityName="Michaelsen" authorityYear="1902" box="[1264,1580,717,739]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="malayanus">
<emphasis id="BEAEEAC2242AFFE6B1F94880C21BFD52" box="[1264,1399,719,739]" italics="true" pageId="4" pageNumber="269">G. malayanus</emphasis>
<bibRefCitation id="E84B4B21242AFFE6B0774882C140FD52" author="Michaelsen, W" box="[1406,1580,717,739]" pageId="4" pageNumber="269" pagination="1 - 54" refId="ref7301" refString="Michaelsen, W., 1902. Neue oligochaeten und neue Fundorte altbekannter. Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 19: 1 - 54." type="journal article" year="1902">Michaelsen, 1902</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4BDA4D53242AFFE6B33E4880C053FD52" authority="Gates, 1958" authorityName="Gates" authorityYear="1958" box="[1591,1855,717,739]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="birmanicus">
<emphasis id="BEAEEAC2242AFFE6B33E4880C1AEFD52" box="[1591,1730,719,739]" italics="true" pageId="4" pageNumber="269">G. birmanicus</emphasis>
<bibRefCitation id="E84B4B21242AFFE6B3C04882C053FD52" author="Gates, G" box="[1737,1855,717,739]" pageId="4" pageNumber="269" pagination="53 - 66" refId="ref7122" refString="Gates, G. E., 1958. On Indian and Burmese earthworms of the genus Glyphidrilus. Records of the Indian Musuem, 53: 53 - 66." type="journal article" year="1958">Gates, 1958</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4BDA4D53242AFFE6B2434880C64CFCB1" authority="Chen &amp; Xu, 1977" authorityName="Chen &amp; Xu" authorityYear="1977" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="yunnanensis">
<emphasis id="BEAEEAC2242AFFE6B2434880C0B3FD52" box="[1866,2015,719,739]" italics="true" pageId="4" pageNumber="269">G. yunnanensis</emphasis>
Chen &amp; Xu, 1977
</taxonomicName>
, 
<taxonomicName id="4BDA4D53242AFFE6B42548A3C52CFCB1" authority="Shen &amp; Yeo, 2005" authorityName="Shen &amp; Yeo" authorityYear="2005" box="[300,576,746,768]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="gatesi">
<emphasis id="BEAEEAC2242AFFE6B42548A3C6EFFCB1" box="[300,387,748,768]" italics="true" pageId="4" pageNumber="269">G. gatesi</emphasis>
Shen &amp; Yeo, 2005
</taxonomicName>
, 
<taxonomicName id="4BDA4D53242AFFE6B74248A3C4C8FCB1" authority="Shen &amp; Yeo, 2005" authorityName="Shen &amp; Yeo" authorityYear="2005" box="[587,932,746,768]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="singaporensis">
<emphasis id="BEAEEAC2242AFFE6B74248A3C58BFCB1" box="[587,743,748,768]" italics="true" pageId="4" pageNumber="269">G singaporensis</emphasis>
Shen &amp; Yeo, 2005
</taxonomicName>
, and 
<taxonomicName id="4BDA4D53242AFFE6B6D048A3C2E2FCB1" authority="Panha &amp; Chanabun, 2011" authorityName="Panha &amp; Chanabun" authorityYear="2011" box="[985,1422,746,768]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="vangviengensis">
<emphasis id="BEAEEAC2242AFFE6B6D048A3C3E7FCB1" box="[985,1163,748,768]" italics="true" pageId="4" pageNumber="269">G. vangviengensis</emphasis>
Panha &amp; 
<bibRefCitation id="E84B4B21242AFFE6B1E448A5C2E2FCB1" author="Chanabun, R &amp; Bantaowong, C &amp; Sutcharit, P &amp; Tongkerd, K &amp; Inkavilay, S" box="[1261,1422,746,768]" pageId="4" pageNumber="269" pagination="213 - 222" refId="ref6938" refString="Chanabun, R., U. Bantaowong, C. Sutcharit, P. Tongkerd, K. Inkavilay, S. W. James &amp; S. Panha, 2011. A new species of semi-aquatic freshwater earthworm of the genus Glyphidrilus Horst, 1889 from Laos (Oligochaeta: Almidae). Tropical Natural History, 11: 213 - 222." type="journal article" year="2011">Chanabun, 2011</bibRefCitation>
</taxonomicName>
. The morphological characters are from the type specimens, except for the character with (*) which are from the original descriptions of 
<bibRefCitation id="E84B4B21242AFFE6B6524948C4B8FCAC" author="Rosa, D" box="[859,980,775,797]" pageId="4" pageNumber="269" pagination="386 - 400" refId="ref7353" refString="Rosa, D., 1890. Viaggio di Leonardo Fea in Birmania e Regioni vicine, xxv, Moniligastridi, Geoscolecidi ed Eudrilidi (I 2). Annali del Museo Civico di Storia Naturale di Genova, 9: 386 - 400." type="journal article" year="1890">Rosa (1890)</bibRefCitation>
, 
<bibRefCitation id="E84B4B21242AFFE6B6D64948C333FCAC" author="Gates, G" box="[991,1119,775,797]" pageId="4" pageNumber="269" pagination="53 - 66" refId="ref7122" refString="Gates, G. E., 1958. On Indian and Burmese earthworms of the genus Glyphidrilus. Records of the Indian Musuem, 53: 53 - 66." type="journal article" year="1958">Gates (1958)</bibRefCitation>
, and Chen &amp; Xu (1977).
</paragraph>
</caption>
<paragraph id="8C6536D0242AFFE6B579490FC00EFA55" pageId="4" pageNumber="269">
<table id="FEDAC470242A001DB579490FC09DFA55" box="[112,2033,832,1508]" gridcols="9" gridrows="14" pageId="4" pageNumber="269">
<tr id="32EA3492242A001DB579490FC09DFCE7" box="[112,2033,832,854]" gridrow="0" pageId="4" pageNumber="269">
<th id="713B5DEE242A001DB579490FC67BFCE7" box="[112,279,832,854]" gridcol="0" gridrow="0" pageId="4" pageNumber="269">
<emphasis id="BEAEEAC2242AFFE6B579490FC78FFCE7" bold="true" box="[112,227,832,854]" pageId="4" pageNumber="269">Characters</emphasis>
</th>
<th id="713B5DEE242A001DB43F490FC535FCE7" box="[310,601,832,854]" gridcol="1" gridrow="0" pageId="4" pageNumber="269">
<emphasis id="BEAEEAC2242AFFE6B43F490FC535FCE7" bold="true" box="[310,601,832,854]" pageId="4" pageNumber="269">
<taxonomicName id="4BDA4D53242AFFE6B43F490FC6B8FCE7" authority="Chanabun &amp; Bantaowong &amp; Sutcharit &amp; Tongkerd &amp; James &amp; Panha, 2012" authorityName="Chanabun &amp; Bantaowong &amp; Sutcharit &amp; Tongkerd &amp; James &amp; Panha" authorityYear="2012" box="[310,468,832,854]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="mekongensis" status="sp. nov.">
<emphasis id="BEAEEAC2242AFFE6B43F490FC6B8FCE7" bold="true" box="[310,468,832,854]" italics="true" pageId="4" pageNumber="269">G. mekongensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A59D57B9242AFFE6B4E8490FC535FCE7" box="[481,601,832,854]" pageId="4" pageNumber="269" rank="species">new species</taxonomicNameLabel>
</emphasis>
</th>
<th id="713B5DEE242A001DB789490FC45FFCE7" box="[640,819,832,854]" gridcol="2" gridrow="0" pageId="4" pageNumber="269">
<emphasis id="BEAEEAC2242AFFE6B789490FC460FCE7" bold="true" box="[640,780,832,854]" pageId="4" pageNumber="269">
<taxonomicName id="4BDA4D53242AFFE6B789490FC46CFCE7" baseAuthorityName="Rosa" baseAuthorityYear="1890" box="[640,768,832,854]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="papillatus">
<emphasis id="BEAEEAC2242AFFE6B789490FC46CFCE7" bold="true" box="[640,768,832,854]" italics="true" pageId="4" pageNumber="269">G. papillatus</emphasis>
</taxonomicName>
*
</emphasis>
</th>
<th id="713B5DEE242A001DB65A490FC492FCE7" box="[851,1022,832,854]" gridcol="3" gridrow="0" pageId="4" pageNumber="269">
<taxonomicName id="4BDA4D53242AFFE6B65A490FC4B1FCE7" authorityName="Michaelsen" authorityYear="1902" box="[851,989,832,854]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="malayanus">
<emphasis id="BEAEEAC2242AFFE6B65A490FC4B1FCE7" bold="true" box="[851,989,832,854]" italics="true" pageId="4" pageNumber="269">G. malayanus</emphasis>
</taxonomicName>
</th>
<th id="713B5DEE242A001DB112490FC3A0FCE7" box="[1051,1228,832,854]" gridcol="4" gridrow="0" pageId="4" pageNumber="269">
<emphasis id="BEAEEAC2242AFFE6B112490FC3D9FCE7" bold="true" box="[1051,1205,832,854]" pageId="4" pageNumber="269">
<taxonomicName id="4BDA4D53242AFFE6B112490FC3C5FCE7" authorityName="Gates" authorityYear="1958" box="[1051,1193,832,854]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="birmanicus">
<emphasis id="BEAEEAC2242AFFE6B112490FC3C5FCE7" bold="true" box="[1051,1193,832,854]" italics="true" pageId="4" pageNumber="269">G. birmanicus</emphasis>
</taxonomicName>
*
</emphasis>
</th>
<th id="713B5DEE242A001DB1EF490FC2E1FCE7" box="[1254,1421,832,854]" gridcol="5" gridrow="0" pageId="4" pageNumber="269">
<emphasis id="BEAEEAC2242AFFE6B1EF490FC2E1FCE7" bold="true" box="[1254,1421,832,854]" pageId="4" pageNumber="269">
<taxonomicName id="4BDA4D53242AFFE6B1EF490FC2EDFCE7" authorityName="Chen &amp; Xu" authorityYear="1977" box="[1254,1409,832,854]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="yunnanensis">
<emphasis id="BEAEEAC2242AFFE6B1EF490FC2EDFCE7" bold="true" box="[1254,1409,832,854]" italics="true" pageId="4" pageNumber="269">G. yunnanensis</emphasis>
</taxonomicName>
*
</emphasis>
</th>
<th id="713B5DEE242A001DB0A7490FC15EFCE7" box="[1454,1586,832,854]" gridcol="6" gridrow="0" pageId="4" pageNumber="269">
<taxonomicName id="4BDA4D53242AFFE6B0A7490FC169FCE7" authorityName="Shen &amp; Yeo" authorityYear="2005" box="[1454,1541,832,854]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="gatesi">
<emphasis id="BEAEEAC2242AFFE6B0A7490FC169FCE7" bold="true" box="[1454,1541,832,854]" italics="true" pageId="4" pageNumber="269">G. gatesi</emphasis>
</taxonomicName>
</th>
<th id="713B5DEE242A001DB35A490FC07BFCE7" box="[1619,1815,832,854]" gridcol="7" gridrow="0" pageId="4" pageNumber="269">
<taxonomicName id="4BDA4D53242AFFE6B35A490FC195FCE7" authorityName="Shen &amp; Yeo" authorityYear="2005" box="[1619,1785,832,854]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="singaporensis">
<emphasis id="BEAEEAC2242AFFE6B35A490FC195FCE7" bold="true" box="[1619,1785,832,854]" italics="true" pageId="4" pageNumber="269">G. singaporensis</emphasis>
</taxonomicName>
</th>
<th id="713B5DEE242A001DB23A490FC09DFCE7" box="[1843,2033,832,854]" gridcol="8" gridrow="0" pageId="4" pageNumber="269">
<taxonomicName id="4BDA4D53242AFFE6B23A490FC084FCE7" authorityName="Panha &amp; Chanabun" authorityYear="2011" box="[1843,2024,832,854]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="4" pageNumber="269" phylum="Annelida" rank="species" species="vangviengensis">
<emphasis id="BEAEEAC2242AFFE6B23A490FC084FCE7" bold="true" box="[1843,2024,832,854]" italics="true" pageId="4" pageNumber="269">G. vangviengensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="32EA3492242A001DB5794927C09DFCCF" box="[112,2033,872,894]" gridrow="1" pageId="4" pageNumber="269">
<th id="713B5DEE242A001DB5794927C67BFCCF" box="[112,279,872,894]" gridcol="0" gridrow="1" pageId="4" pageNumber="269">Length</th>
<td id="713B5DEE242A001DB43F4927C535FCCF" box="[310,601,872,894]" gridcol="1" gridrow="1" pageId="4" pageNumber="269">125–224</td>
<td id="713B5DEE242A001DB7894927C45FFCCF" box="[640,819,872,894]" gridcol="2" gridrow="1" pageId="4" pageNumber="269">100</td>
<td id="713B5DEE242A001DB65A4927C492FCCF" box="[851,1022,872,894]" gridcol="3" gridrow="1" pageId="4" pageNumber="269">52–91</td>
<td id="713B5DEE242A001DB1124927C3A0FCCF" box="[1051,1228,872,894]" gridcol="4" gridrow="1" pageId="4" pageNumber="269">95–103</td>
<td id="713B5DEE242A001DB1EF4927C2E1FCCF" box="[1254,1421,872,894]" gridcol="5" gridrow="1" pageId="4" pageNumber="269">123</td>
<td id="713B5DEE242A001DB0A74927C15EFCCF" box="[1454,1586,872,894]" gridcol="6" gridrow="1" pageId="4" pageNumber="269">50+</td>
<td id="713B5DEE242A001DB35A4927C07BFCCF" box="[1619,1815,872,894]" gridcol="7" gridrow="1" pageId="4" pageNumber="269">101–143</td>
<td id="713B5DEE242A001DB23A4927C09DFCCF" box="[1843,2033,872,894]" gridcol="8" gridrow="1" pageId="4" pageNumber="269">104–160</td>
</tr>
<tr id="32EA3492242A001DB57949C7C09DFC2F" box="[112,2033,904,926]" gridrow="2" pageId="4" pageNumber="269">
<th id="713B5DEE242A001DB57949C7C67BFC2F" box="[112,279,904,926]" gridcol="0" gridrow="2" pageId="4" pageNumber="269">Segments</th>
<td id="713B5DEE242A001DB43F49C7C535FC2F" box="[310,601,904,926]" gridcol="1" gridrow="2" pageId="4" pageNumber="269">
223 
<emphasis id="BEAEEAC2242AFFE6B45349C7C60AFC2F" bold="true" box="[346,358,904,926]" pageId="4" pageNumber="269">–</emphasis>
382
</td>
<td id="713B5DEE242A001DB78949C7C45FFC2F" box="[640,819,904,926]" gridcol="2" gridrow="2" pageId="4" pageNumber="269">330</td>
<td id="713B5DEE242A001DB65A49C7C492FC2F" box="[851,1022,904,926]" gridcol="3" gridrow="2" pageId="4" pageNumber="269">192–313</td>
<td id="713B5DEE242A001DB11249C7C3A0FC2F" box="[1051,1228,904,926]" gridcol="4" gridrow="2" pageId="4" pageNumber="269">?</td>
<td id="713B5DEE242A001DB1EF49C7C2E1FC2F" box="[1254,1421,904,926]" gridcol="5" gridrow="2" pageId="4" pageNumber="269">139</td>
<td id="713B5DEE242A001DB0A749C7C15EFC2F" box="[1454,1586,904,926]" gridcol="6" gridrow="2" pageId="4" pageNumber="269">62+</td>
<td id="713B5DEE242A001DB35A49C7C07BFC2F" box="[1619,1815,904,926]" gridcol="7" gridrow="2" pageId="4" pageNumber="269">181–341</td>
<td id="713B5DEE242A001DB23A49C7C09DFC2F" box="[1843,2033,904,926]" gridcol="8" gridrow="2" pageId="4" pageNumber="269">145–229</td>
</tr>
<tr id="32EA3492242A001DB57949E6C09DFC6D" box="[112,2033,937,988]" gridrow="3" pageId="4" pageNumber="269">
<th id="713B5DEE242A001DB57949E6C67BFC6D" box="[112,279,937,988]" gridcol="0" gridrow="3" pageId="4" pageNumber="269">Clitellum</th>
<td id="713B5DEE242A001DB43F49E6C535FC6D" box="[310,601,937,988]" gridcol="1" gridrow="3" pageId="4" pageNumber="269">19–37, 38</td>
<td id="713B5DEE242A001DB78949E6C45FFC6D" box="[640,819,937,988]" gridcol="2" gridrow="3" pageId="4" pageNumber="269">14–40 24, 25</td>
<td id="713B5DEE242A001DB65A49E6C492FC6D" box="[851,1022,937,988]" gridcol="3" gridrow="3" pageId="4" pageNumber="269">15, 16, 17–23,</td>
<td id="713B5DEE242A001DB11249E6C3A0FC6D" box="[1051,1228,937,988]" gridcol="4" gridrow="3" pageId="4" pageNumber="269">12, 13–43, 44</td>
<td id="713B5DEE242A001DB1EF49E6C2E1FC6D" box="[1254,1421,937,988]" gridcol="5" gridrow="3" pageId="4" pageNumber="269">18–38</td>
<td id="713B5DEE242A001DB0A749E6C15EFC6D" box="[1454,1586,937,988]" gridcol="6" gridrow="3" pageId="4" pageNumber="269">17–26 31</td>
<td id="713B5DEE242A001DB35A49E6C07BFC6D" box="[1619,1815,937,988]" gridcol="7" gridrow="3" pageId="4" pageNumber="269">18–27, ½28, 29, 30,</td>
<td id="713B5DEE242A001DB23A49E6C09DFC6D" box="[1843,2033,937,988]" gridcol="8" gridrow="3" pageId="4" pageNumber="269">19, 20–35, 36, 37</td>
</tr>
<tr id="32EA3492242A001DB57949A9C09DFC4D" box="[112,2033,998,1020]" gridrow="4" pageId="4" pageNumber="269">
<th id="713B5DEE242A001DB57949A9C67BFC4D" box="[112,279,998,1020]" gridcol="0" gridrow="4" pageId="4" pageNumber="269">Wings</th>
<td id="713B5DEE242A001DB43F49A9C535FC4D" box="[310,601,998,1020]" gridcol="1" gridrow="4" pageId="4" pageNumber="269">24–½33, 33, 34, ½35</td>
<td id="713B5DEE242A001DB78949A9C45FFC4D" box="[640,819,998,1020]" gridcol="2" gridrow="4" pageId="4" pageNumber="269">18–23, 24</td>
<td id="713B5DEE242A001DB65A49A9C492FC4D" box="[851,1022,998,1020]" gridcol="3" gridrow="4" pageId="4" pageNumber="269">¾18, 18–21, ½22</td>
<td id="713B5DEE242A001DB11249A9C3A0FC4D" box="[1051,1228,998,1020]" gridcol="4" gridrow="4" pageId="4" pageNumber="269">21–29</td>
<td id="713B5DEE242A001DB1EF49A9C2E1FC4D" box="[1254,1421,998,1020]" gridcol="5" gridrow="4" pageId="4" pageNumber="269">22–32</td>
<td id="713B5DEE242A001DB0A749A9C15EFC4D" box="[1454,1586,998,1020]" gridcol="6" gridrow="4" pageId="4" pageNumber="269">19–½24</td>
<td id="713B5DEE242A001DB35A49A9C07BFC4D" box="[1619,1815,998,1020]" gridcol="7" gridrow="4" pageId="4" pageNumber="269">21–½26, 26</td>
<td id="713B5DEE242A001DB23A49A9C09DFC4D" box="[1843,2033,998,1020]" gridcol="8" gridrow="4" pageId="4" pageNumber="269">24, 25–31, 32</td>
</tr>
<tr id="32EA3492242A001DB5794E48C09DFBE6" box="[112,2033,1031,1111]" gridrow="5" pageId="4" pageNumber="269">
<th id="713B5DEE242A001DB5794E48C67BFBE6" box="[112,279,1031,1111]" gridcol="0" gridrow="5" pageId="4" pageNumber="269">Genital markings Paired</th>
<td id="713B5DEE242A001DB43F4E48C535FBE6" box="[310,601,1031,1111]" gridcol="1" gridrow="5" pageId="4" pageNumber="269">23</td>
<td id="713B5DEE242A001DB7894E48C45FFBE6" box="[640,819,1031,1111]" gridcol="2" gridrow="5" pageId="4" pageNumber="269">12, 13, 14, 15, 16, 17, 18–20, 24–29</td>
<td id="713B5DEE242A001DB65A4E48C492FBE6" box="[851,1022,1031,1111]" gridcol="3" gridrow="5" pageId="4" pageNumber="269">13, 14, 15–17, 22, 23</td>
<td id="713B5DEE242A001DB1124E48C3A0FBE6" box="[1051,1228,1031,1111]" gridcol="4" gridrow="5" pageId="4" pageNumber="269">12, 13–21, 22, 23, 26, 29, 30–31, 33, 34</td>
<td id="713B5DEE242A001DB1EF4E48C2E1FBE6" box="[1254,1421,1031,1111]" gridcol="5" gridrow="5" pageId="4" pageNumber="269">18–21, 32–34</td>
<td id="713B5DEE242A001DB0A74E48C15EFBE6" box="[1454,1586,1031,1111]" gridcol="6" gridrow="5" pageId="4" pageNumber="269">15, 16–18, 24, 25, 26</td>
<td id="713B5DEE242A001DB35A4E48C07BFBE6" box="[1619,1815,1031,1111]" gridcol="7" gridrow="5" pageId="4" pageNumber="269">12–15, 20, 26, 27, 28</td>
<td id="713B5DEE242A001DB23A4E48C09DFBE6" box="[1843,2033,1031,1111]" gridcol="8" gridrow="5" pageId="4" pageNumber="269">18, 19–23, 24, 33, 34</td>
</tr>
<tr id="32EA3492242A001DB5794E12C09DFB21" box="[112,2033,1117,1168]" gridrow="6" pageId="4" pageNumber="269">
<th id="713B5DEE242A001DB5794E12C67BFB21" box="[112,279,1117,1168]" gridcol="0" gridrow="6" pageId="4" pageNumber="269">Unpaired</th>
<td id="713B5DEE242A001DB43F4E12C535FB21" box="[310,601,1117,1168]" gridcol="1" gridrow="6" pageId="4" pageNumber="269">—</td>
<td id="713B5DEE242A001DB7894E12C45FFB21" box="[640,819,1117,1168]" gridcol="2" gridrow="6" pageId="4" pageNumber="269">11–21, 23–33</td>
<td id="713B5DEE242A001DB65A4E12C492FB21" box="[851,1022,1117,1168]" gridcol="3" gridrow="6" pageId="4" pageNumber="269">12–16, 22–25</td>
<td id="713B5DEE242A001DB1124E12C3A0FB21" box="[1051,1228,1117,1168]" gridcol="4" gridrow="6" pageId="4" pageNumber="269">—</td>
<td id="713B5DEE242A001DB1EF4E12C2E1FB21" box="[1254,1421,1117,1168]" gridcol="5" gridrow="6" pageId="4" pageNumber="269">—</td>
<td id="713B5DEE242A001DB0A74E12C15EFB21" box="[1454,1586,1117,1168]" gridcol="6" gridrow="6" pageId="4" pageNumber="269">13–19, 24–27</td>
<td id="713B5DEE242A001DB35A4E12C07BFB21" box="[1619,1815,1117,1168]" gridcol="7" gridrow="6" pageId="4" pageNumber="269">17, 18–25, 28–29, 30</td>
<td id="713B5DEE242A001DB23A4E12C09DFB21" box="[1843,2033,1117,1168]" gridcol="8" gridrow="6" pageId="4" pageNumber="269">12, 13–14, 15</td>
</tr>
<tr id="32EA3492242A001DB5794ED4C09DFB00" box="[112,2033,1179,1201]" gridrow="7" pageId="4" pageNumber="269">
<th id="713B5DEE242A001DB5794ED4C67BFB00" box="[112,279,1179,1201]" gridcol="0" gridrow="7" pageId="4" pageNumber="269">Hearts</th>
<td id="713B5DEE242A001DB43F4ED4C535FB00" box="[310,601,1179,1201]" gridcol="1" gridrow="7" pageId="4" pageNumber="269">7–11</td>
<td id="713B5DEE242A001DB7894ED4C45FFB00" box="[640,819,1179,1201]" gridcol="2" gridrow="7" pageId="4" pageNumber="269">7–11</td>
<td id="713B5DEE242A001DB65A4ED4C492FB00" box="[851,1022,1179,1201]" gridcol="3" gridrow="7" pageId="4" pageNumber="269">9–11</td>
<td id="713B5DEE242A001DB1124ED4C3A0FB00" box="[1051,1228,1179,1201]" gridcol="4" gridrow="7" pageId="4" pageNumber="269">7–11</td>
<td id="713B5DEE242A001DB1EF4ED4C2E1FB00" box="[1254,1421,1179,1201]" gridcol="5" gridrow="7" pageId="4" pageNumber="269">7–11</td>
<td id="713B5DEE242A001DB0A74ED4C15EFB00" box="[1454,1586,1179,1201]" gridcol="6" gridrow="7" pageId="4" pageNumber="269">9–11</td>
<td id="713B5DEE242A001DB35A4ED4C07BFB00" box="[1619,1815,1179,1201]" gridcol="7" gridrow="7" pageId="4" pageNumber="269">9–11</td>
<td id="713B5DEE242A001DB23A4ED4C09DFB00" box="[1843,2033,1179,1201]" gridcol="8" gridrow="7" pageId="4" pageNumber="269">7–11</td>
</tr>
<tr id="32EA3492242A001DB5794EF4C09DFB60" box="[112,2033,1211,1233]" gridrow="8" pageId="4" pageNumber="269">
<th id="713B5DEE242A001DB5794EF4C67BFB60" box="[112,279,1211,1233]" gridcol="0" gridrow="8" pageId="4" pageNumber="269">Seminal vesicles</th>
<td id="713B5DEE242A001DB43F4EF4C535FB60" box="[310,601,1211,1233]" gridcol="1" gridrow="8" pageId="4" pageNumber="269">9–12</td>
<td id="713B5DEE242A001DB7894EF4C45FFB60" box="[640,819,1211,1233]" gridcol="2" gridrow="8" pageId="4" pageNumber="269">9–12</td>
<td id="713B5DEE242A001DB65A4EF4C492FB60" box="[851,1022,1211,1233]" gridcol="3" gridrow="8" pageId="4" pageNumber="269">9–12</td>
<td id="713B5DEE242A001DB1124EF4C3A0FB60" box="[1051,1228,1211,1233]" gridcol="4" gridrow="8" pageId="4" pageNumber="269">9–12</td>
<td id="713B5DEE242A001DB1EF4EF4C2E1FB60" box="[1254,1421,1211,1233]" gridcol="5" gridrow="8" pageId="4" pageNumber="269">9–12</td>
<td id="713B5DEE242A001DB0A74EF4C15EFB60" box="[1454,1586,1211,1233]" gridcol="6" gridrow="8" pageId="4" pageNumber="269">9–12</td>
<td id="713B5DEE242A001DB35A4EF4C07BFB60" box="[1619,1815,1211,1233]" gridcol="7" gridrow="8" pageId="4" pageNumber="269">9–12</td>
<td id="713B5DEE242A001DB23A4EF4C09DFB60" box="[1843,2033,1211,1233]" gridcol="8" gridrow="8" pageId="4" pageNumber="269">9–12</td>
</tr>
<tr id="32EA3492242A001DB5794E93C09DFB43" box="[112,2033,1244,1266]" gridrow="9" pageId="4" pageNumber="269">
<th id="713B5DEE242A001DB5794E93C67BFB43" box="[112,279,1244,1266]" gridcol="0" gridrow="9" pageId="4" pageNumber="269">Intestinal origin</th>
<td id="713B5DEE242A001DB43F4E93C535FB43" box="[310,601,1244,1266]" gridcol="1" gridrow="9" pageId="4" pageNumber="269">15</td>
<td id="713B5DEE242A001DB7894E93C45FFB43" box="[640,819,1244,1266]" gridcol="2" gridrow="9" pageId="4" pageNumber="269">15</td>
<td id="713B5DEE242A001DB65A4E93C492FB43" box="[851,1022,1244,1266]" gridcol="3" gridrow="9" pageId="4" pageNumber="269">16</td>
<td id="713B5DEE242A001DB1124E93C3A0FB43" box="[1051,1228,1244,1266]" gridcol="4" gridrow="9" pageId="4" pageNumber="269">15</td>
<td id="713B5DEE242A001DB1EF4E93C2E1FB43" box="[1254,1421,1244,1266]" gridcol="5" gridrow="9" pageId="4" pageNumber="269">16</td>
<td id="713B5DEE242A001DB0A74E93C15EFB43" box="[1454,1586,1244,1266]" gridcol="6" gridrow="9" pageId="4" pageNumber="269">18</td>
<td id="713B5DEE242A001DB35A4E93C07BFB43" box="[1619,1815,1244,1266]" gridcol="7" gridrow="9" pageId="4" pageNumber="269">15 or 16</td>
<td id="713B5DEE242A001DB23A4E93C09DFB43" box="[1843,2033,1244,1266]" gridcol="8" gridrow="9" pageId="4" pageNumber="269">16</td>
</tr>
<tr id="32EA3492242A001DB5794EB3C09DFAA3" box="[112,2033,1276,1298]" gridrow="10" pageId="4" pageNumber="269">
<th id="713B5DEE242A001DB5794EB3C67BFAA3" box="[112,279,1276,1298]" gridcol="0" gridrow="10" pageId="4" pageNumber="269">Gizzard</th>
<td id="713B5DEE242A001DB43F4EB3C535FAA3" box="[310,601,1276,1298]" gridcol="1" gridrow="10" pageId="4" pageNumber="269">8</td>
<td id="713B5DEE242A001DB7894EB3C45FFAA3" box="[640,819,1276,1298]" gridcol="2" gridrow="10" pageId="4" pageNumber="269">7/8</td>
<td id="713B5DEE242A001DB65A4EB3C492FAA3" box="[851,1022,1276,1298]" gridcol="3" gridrow="10" pageId="4" pageNumber="269">8</td>
<td id="713B5DEE242A001DB1124EB3C3A0FAA3" box="[1051,1228,1276,1298]" gridcol="4" gridrow="10" pageId="4" pageNumber="269">7/8</td>
<td id="713B5DEE242A001DB1EF4EB3C2E1FAA3" box="[1254,1421,1276,1298]" gridcol="5" gridrow="10" pageId="4" pageNumber="269">8</td>
<td id="713B5DEE242A001DB0A74EB3C15EFAA3" box="[1454,1586,1276,1298]" gridcol="6" gridrow="10" pageId="4" pageNumber="269">8</td>
<td id="713B5DEE242A001DB35A4EB3C07BFAA3" box="[1619,1815,1276,1298]" gridcol="7" gridrow="10" pageId="4" pageNumber="269">8</td>
<td id="713B5DEE242A001DB23A4EB3C09DFAA3" box="[1843,2033,1276,1298]" gridcol="8" gridrow="10" pageId="4" pageNumber="269">8</td>
</tr>
<tr id="32EA3492242A001DB5794F52C09DFAE1" box="[112,2033,1309,1360]" gridrow="11" pageId="4" pageNumber="269">
<th id="713B5DEE242A001DB5794F52C67BFAE1" box="[112,279,1309,1360]" gridcol="0" gridrow="11" pageId="4" pageNumber="269">Spermathecae</th>
<td id="713B5DEE242A001DB43F4F52C535FAE1" box="[310,601,1309,1360]" gridcol="1" gridrow="11" pageId="4" pageNumber="269">—</td>
<td id="713B5DEE242A001DB7894F52C45FFAE1" box="[640,819,1309,1360]" gridcol="2" gridrow="11" pageId="4" pageNumber="269">14–17</td>
<td id="713B5DEE242A001DB65A4F52C492FAE1" box="[851,1022,1309,1360]" gridcol="3" gridrow="11" pageId="4" pageNumber="269">14/15–16/17</td>
<td id="713B5DEE242A001DB1124F52C3A0FAE1" box="[1051,1228,1309,1360]" gridcol="4" gridrow="11" pageId="4" pageNumber="269">13/14–17/18</td>
<td id="713B5DEE242A001DB1EF4F52C2E1FAE1" box="[1254,1421,1309,1360]" gridcol="5" gridrow="11" pageId="4" pageNumber="269">—</td>
<td id="713B5DEE242A001DB0A74F52C15EFAE1" box="[1454,1586,1309,1360]" gridcol="6" gridrow="11" pageId="4" pageNumber="269">15–17</td>
<td id="713B5DEE242A001DB35A4F52C07BFAE1" box="[1619,1815,1309,1360]" gridcol="7" gridrow="11" pageId="4" pageNumber="269">14–17 or 13/14–16/17</td>
<td id="713B5DEE242A001DB23A4F52C09DFAE1" box="[1843,2033,1309,1360]" gridcol="8" gridrow="11" pageId="4" pageNumber="269">—</td>
</tr>
<tr id="32EA3492242A001DB5794F18C09DFADC" box="[112,2033,1367,1389]" gridrow="12" pageId="4" pageNumber="269">
<th id="713B5DEE242A001DB5794F18C67BFADC" box="[112,279,1367,1389]" gridcol="0" gridrow="12" pageId="4" pageNumber="269">First nephridia</th>
<td id="713B5DEE242A001DB43F4F18C535FADC" box="[310,601,1367,1389]" gridcol="1" gridrow="12" pageId="4" pageNumber="269">13</td>
<td id="713B5DEE242A001DB7894F18C45FFADC" box="[640,819,1367,1389]" gridcol="2" gridrow="12" pageId="4" pageNumber="269">17</td>
<td id="713B5DEE242A001DB65A4F18C492FADC" box="[851,1022,1367,1389]" gridcol="3" gridrow="12" pageId="4" pageNumber="269">14</td>
<td id="713B5DEE242A001DB1124F18C3A0FADC" box="[1051,1228,1367,1389]" gridcol="4" gridrow="12" pageId="4" pageNumber="269">13</td>
<td id="713B5DEE242A001DB1EF4F18C2E1FADC" box="[1254,1421,1367,1389]" gridcol="5" gridrow="12" pageId="4" pageNumber="269">14</td>
<td id="713B5DEE242A001DB0A74F18C15EFADC" box="[1454,1586,1367,1389]" gridcol="6" gridrow="12" pageId="4" pageNumber="269">14</td>
<td id="713B5DEE242A001DB35A4F18C07BFADC" box="[1619,1815,1367,1389]" gridcol="7" gridrow="12" pageId="4" pageNumber="269">12</td>
<td id="713B5DEE242A001DB23A4F18C09DFADC" box="[1843,2033,1367,1389]" gridcol="8" gridrow="12" pageId="4" pageNumber="269">14</td>
</tr>
<tr id="32EA3492242A001DB5794F38C09DFA55" box="[112,2033,1399,1508]" gridrow="13" pageId="4" pageNumber="269">
<th id="713B5DEE242A001DB5794F38C67BFA55" box="[112,279,1399,1508]" gridcol="0" gridrow="13" pageId="4" pageNumber="269">Type locality</th>
<td id="713B5DEE242A001DB43F4F38C535FA55" box="[310,601,1399,1508]" gridcol="1" gridrow="13" pageId="4" pageNumber="269">Khong Chiam, Ubon Ratchathani, Thailand</td>
<td id="713B5DEE242A001DB7894F38C45FFA55" box="[640,819,1399,1508]" gridcol="2" gridrow="13" pageId="4" pageNumber="269">Burma</td>
<td id="713B5DEE242A001DB65A4F38C492FA55" box="[851,1022,1399,1508]" gridcol="3" gridrow="13" pageId="4" pageNumber="269">Lubock Paku, Pahang River, Malaysia</td>
<td id="713B5DEE242A001DB1124F38C3A0FA55" box="[1051,1228,1399,1508]" gridcol="4" gridrow="13" pageId="4" pageNumber="269">Burma</td>
<td id="713B5DEE242A001DB1EF4F38C2E1FA55" box="[1254,1421,1399,1508]" gridcol="5" gridrow="13" pageId="4" pageNumber="269">Menglun, Loso River, Yunnan, China</td>
<td id="713B5DEE242A001DB0A74F38C15EFA55" box="[1454,1586,1399,1508]" gridcol="6" gridrow="13" pageId="4" pageNumber="269">Sungei Kayu, River Sedili, Johor, Malaysia</td>
<td id="713B5DEE242A001DB35A4F38C07BFA55" box="[1619,1815,1399,1508]" gridcol="7" gridrow="13" pageId="4" pageNumber="269">Jungle Fall Valley, Bukit Timah, Singapore</td>
<td id="713B5DEE242A001DB23A4F38C09DFA55" box="[1843,2033,1399,1508]" gridcol="8" gridrow="13" pageId="4" pageNumber="269">Bank of Song River at Veintiane Province, Laos</td>
</tr>
</table>
</paragraph>
<tableNote id="713C375E242AFFE6B5794FB5C52BF99D" pageId="4" pageNumber="269" targetBox="[112,2033,832,1508]" targetPageId="4">
<paragraph id="8C6536D0242AFFE6B5794FB5C6E5F9A1" blockId="4.[112,583,1530,1580]" box="[112,393,1530,1552]" pageId="4" pageNumber="269">— Indicates character absent</paragraph>
<paragraph id="8C6536D0242AFFE6B5794C59C52BF99D" blockId="4.[112,583,1530,1580]" box="[112,583,1558,1580]" pageId="4" pageNumber="269">? Indicates not shown in the original description</paragraph>
</tableNote>
<paragraph id="8C6536D0242BFFE7B5784AF1C539FE30" blockId="5.[113,752,190,385]" pageId="5" pageNumber="270">
adults and 
<materialsCitation id="3CB23C8D242BFFE7B5D04AF1C66FFEBF" collectingDate="2011-05-27" collectionCode="CUMZ" country="Malaysia" elevation="24" latitude="6.01225" location="Sungei" longLatPrecision="1" longitude="102.18283" pageId="5" pageNumber="270" specimenCode="CUMZ 3219" specimenCount="257" specimenCount-juvenile="257" stateProvince="Kelantan">
<specimenCount id="9ADCFD59242BFFE7B5D04AF1C636FF65" box="[217,346,190,212]" pageId="5" pageNumber="270" type="juvenile">257 juveniles</specimenCount>
(
<specimenCode id="DC7C9EAB242BFFE7B46F4AF1C689FF62" box="[358,485,190,211]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="270">CUMZ 3219</specimenCode>
, 
<figureCitation id="14E12A55242BFFE7B4E64AF1C555FF65" box="[495,569,190,212]" captionStart="Fig" captionStartId="7.[113,148,2034,2056]" captionText="Fig. 5. External and internal morphology of holotype (ZMH V5875) of Glyphidrilus malayanus: A, external ventral view; B, internal dorsal view; C, specimen collected from Sungei Kelatan, Kelantan, Malaysia (CUMZ 3219); D, from Sungei Berkoh, Johor, Malaysia (CUMZ 3220); E, from Tok Bok Hotspring, Kelantan, Malaysia (CUMZ 3218), Malaysia." pageId="5" pageNumber="270">Fig. 5C</figureCitation>
), 
<location id="8905600B242BFFE7B7424AF1C5E2FF65" LSID="urn:lsid:plazi:treatment:047387C6242DFFE7B6894D64C3A5FC07:8905600B242BFFE7B7424AF1C5E2FF65" box="[587,654,190,212]" country="Malaysia" latitude="6.01225" longLatPrecision="1" longitude="102.18283" name="Sungei" pageId="5" pageNumber="270" stateProvince="Kelantan">Sungei</location>
<collectingRegion id="4E1EF832242BFFE7B79A4AF1C580FF65" box="[659,748,190,212]" country="Malaysia" name="Kelantan" pageId="5" pageNumber="270">Kelantan</collectingRegion>
, 
<collectingRegion id="4E1EF832242BFFE7B5784A94C7A6FF40" box="[113,202,219,241]" country="Malaysia" name="Kelantan" pageId="5" pageNumber="270">Kelantan</collectingRegion>
, 
<collectingCountry id="F4CD7640242BFFE7B5DA4A94C641FF40" box="[211,301,219,241]" name="Malaysia" pageId="5" pageNumber="270">Malaysia</collectingCountry>
, 
<geoCoordinate id="E9EE5017242BFFE7B43F4A94C6CFFF40" box="[310,419,219,241]" degrees="6" direction="north" minutes="0" orientation="latitude" pageId="5" pageNumber="270" precision="1" seconds="44.1" value="6.01225">6°0'44.1&quot;N</geoCoordinate>
, 
<geoCoordinate id="E9EE5017242BFFE7B4A44A94C52DFF40" box="[429,577,219,241]" degrees="102" direction="east" minutes="10" orientation="longitude" pageId="5" pageNumber="270" precision="1" seconds="58.2" value="102.18283">102° 10'58.2&quot;E</geoCoordinate>
, 
<quantity id="4B229B35242BFFE7B7434A94C517FF41" box="[586,635,219,240]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="270" unit="m" value="24.0">
<elevation id="07F7D1E3242BFFE7B7434A94C517FF41" box="[586,635,219,240]" metricMagnitude="1" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="270" unit="m" value="24.0">24 m</elevation>
</quantity>
in elevation (
<date id="F8641010242BFFE7B5714AB7C797FEBF" box="[120,251,248,270]" pageId="5" pageNumber="270" value="2011-05-27">
<collectingDate id="E820E9F8242BFFE7B5714AB7C797FEBF" box="[120,251,248,270]" pageId="5" pageNumber="270" value="2011-05-27">27 May 2011</collectingDate>
</date>
)
</materialsCitation>
. 
<materialsCitation id="3CB23C8D242BFFE7B4024AB7C60CFEF6" collectingDate="2011-05-21" collectionCode="CUMZ" country="Malaysia" elevation="35" latitude="2.1197498" location="Sungei Berkoh" longLatPrecision="1" longitude="103.04025" pageId="5" pageNumber="270" specimenCode="CUMZ 3220" specimenCount="28" specimenCount-adult="17" specimenCount-juvenile="11" stateProvince="Johor">
<specimenCount id="9ADCFD59242BFFE7B4024AB7C60EFEBC" box="[267,354,247,269]" pageId="5" pageNumber="270" type="adult">17 adults</specimenCount>
and 
<specimenCount id="9ADCFD59242BFFE7B4984AB7C568FEBC" box="[401,516,247,269]" pageId="5" pageNumber="270" type="juvenile">11 juveniles</specimenCount>
(
<specimenCode id="DC7C9EAB242BFFE7B7194AB7C5E3FEBC" box="[528,655,248,269]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="270">CUMZ 3220</specimenCode>
, 
<figureCitation id="14E12A55242BFFE7B7904AB8C588FEBC" box="[665,740,247,269]" captionStart="Fig" captionStartId="7.[113,148,2034,2056]" captionText="Fig. 5. External and internal morphology of holotype (ZMH V5875) of Glyphidrilus malayanus: A, external ventral view; B, internal dorsal view; C, specimen collected from Sungei Kelatan, Kelantan, Malaysia (CUMZ 3219); D, from Sungei Berkoh, Johor, Malaysia (CUMZ 3220); E, from Tok Bok Hotspring, Kelantan, Malaysia (CUMZ 3218), Malaysia." pageId="5" pageNumber="270">Fig. 5D</figureCitation>
), 
<location id="8905600B242BFFE7B5784B5BC66DFE9B" LSID="urn:lsid:plazi:treatment:047387C6242DFFE7B6894D64C3A5FC07:8905600B242BFFE7B5784B5BC66DFE9B" box="[113,257,276,298]" country="Malaysia" latitude="2.1197498" longLatPrecision="1" longitude="103.04025" name="Sungei Berkoh" pageId="5" pageNumber="270" stateProvince="Johor">Sungei Berkoh</location>
, 
<collectingRegion id="4E1EF832242BFFE7B4004B5BC62CFE9B" box="[265,320,276,298]" country="Malaysia" name="Johor" pageId="5" pageNumber="270">Johor</collectingRegion>
, 
<collectingCountry id="F4CD7640242BFFE7B4414B5BC6CEFE9B" box="[328,418,276,298]" name="Malaysia" pageId="5" pageNumber="270">Malaysia</collectingCountry>
, 
<geoCoordinate id="E9EE5017242BFFE7B4A34B5BC57AFE98" box="[426,534,276,297]" degrees="2" direction="north" minutes="7" orientation="latitude" pageId="5" pageNumber="270" precision="1" seconds="11.1" value="2.1197498">2°7'11.1&quot;N</geoCoordinate>
, 
<geoCoordinate id="E9EE5017242BFFE7B7164B5BC5CCFE98" box="[543,672,276,297]" degrees="103" direction="east" minutes="2" orientation="longitude" pageId="5" pageNumber="270" precision="1" seconds="24.9" value="103.04025">103°2'24.9&quot;E</geoCoordinate>
, 
<quantity id="4B229B35242BFFE7B7A04B5BC5B5FE9B" box="[681,729,276,298]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="270" unit="m" value="35.0">
<elevation id="07F7D1E3242BFFE7B7A04B5BC5B5FE9B" box="[681,729,276,298]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="270" unit="m" value="35.0">35 m</elevation>
</quantity>
in elevation (
<date id="F8641010242BFFE7B5DF4B7EC635FEF6" box="[214,345,305,327]" pageId="5" pageNumber="270" value="2011-05-21">
<collectingDate id="E820E9F8242BFFE7B5DF4B7EC635FEF6" box="[214,345,305,327]" pageId="5" pageNumber="270" value="2011-05-21">21 May 2011</collectingDate>
</date>
)
</materialsCitation>
. 
<materialsCitation id="3CB23C8D242BFFE7B4614B7EC53DFE30" collectingDate="2011-05-26" collectionCode="CUMZ" country="Malaysia" elevation="36" latitude="5.838722" location="Tok Bok Hotspring" longLatPrecision="1" longitude="102.23064" pageId="5" pageNumber="270" specimenCode="CUMZ 3218" specimenCount="106" specimenCount-adult="65" specimenCount-juvenile="41" stateProvince="Kelantan">
<specimenCount id="9ADCFD59242BFFE7B4614B7EC6D3FEF6" box="[360,447,305,327]" pageId="5" pageNumber="270" type="adult">65 adults</specimenCount>
and 
<specimenCount id="9ADCFD59242BFFE7B4E44B7EC50EFEF6" box="[493,610,305,327]" pageId="5" pageNumber="270" type="juvenile">41 juveniles</specimenCount>
(
<specimenCode id="DC7C9EAB242BFFE7B7644B7EC580FEF7" box="[621,748,305,326]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="5" pageNumber="270">CUMZ 3218</specimenCode>
, 
<figureCitation id="14E12A55242BFFE7B5784B01C7AEFED2" box="[113,194,334,356]" captionStart="Fig" captionStartId="5.[112,147,1918,1940]" captionTargetBox="[113,752,1213,1895]" captionTargetId="figure-285@5.[113,754,1213,1895]" captionTargetPageId="5" captionText="Fig. 3. The colouration of newly collected specimens just after the first step preservation in 30% ethanol:A, Glyphidrilus mekongensis, new species (paratype CUMZ 3216); B, Glyphidrilus malayanus from Tok Bok Hotspring, Kelantan, Malaysia (CUMZ 3218); C, Glyphidrilus singaporensis topotype specimen (CUMZ 3217); D, cocoon characters of Glyphidrilus mekongensis, new species (paratype CUMZ 3216)." figureDoi="http://doi.org/10.5281/zenodo.4508504" httpUri="https://zenodo.org/record/4508504/files/figure.png" pageId="5" pageNumber="270">Figs. 3B</figureCitation>
, 
<figureCitation id="14E12A55242BFFE7B5C34B01C78BFED5" box="[202,231,334,356]" captionStart="Fig" captionStartId="7.[113,148,2034,2056]" captionText="Fig. 5. External and internal morphology of holotype (ZMH V5875) of Glyphidrilus malayanus: A, external ventral view; B, internal dorsal view; C, specimen collected from Sungei Kelatan, Kelantan, Malaysia (CUMZ 3219); D, from Sungei Berkoh, Johor, Malaysia (CUMZ 3220); E, from Tok Bok Hotspring, Kelantan, Malaysia (CUMZ 3218), Malaysia." pageId="5" pageNumber="270">5E</figureCitation>
), 
<location id="8905600B242BFFE7B5FF4B01C6C0FED5" LSID="urn:lsid:plazi:treatment:047387C6242DFFE7B6894D64C3A5FC07:8905600B242BFFE7B5FF4B01C6C0FED5" box="[246,428,334,356]" country="Malaysia" latitude="5.838722" longLatPrecision="1" longitude="102.23064" name="Tok Bok Hotspring" pageId="5" pageNumber="270" stateProvince="Kelantan">Tok Bok Hotspring</location>
, 
<collectingRegion id="4E1EF832242BFFE7B4BD4B01C560FED5" box="[436,524,334,356]" country="Malaysia" name="Kelantan" pageId="5" pageNumber="270">Kelantan</collectingRegion>
, 
<collectingCountry id="F4CD7640242BFFE7B71D4B01C501FED5" box="[532,621,334,356]" name="Malaysia" pageId="5" pageNumber="270">Malaysia</collectingCountry>
, 
<geoCoordinate id="E9EE5017242BFFE7B77C4B01C580FED5" box="[629,748,334,356]" degrees="5" direction="north" minutes="50" orientation="latitude" pageId="5" pageNumber="270" precision="1" seconds="19.4" value="5.838722">5°50'19.4&quot;N</geoCoordinate>
, 
<geoCoordinate id="E9EE5017242BFFE7B5784B24C793FE30" box="[113,255,363,385]" degrees="102" direction="east" minutes="13" orientation="longitude" pageId="5" pageNumber="270" precision="1" seconds="50.3" value="102.23064">102°13'50.3&quot;E</geoCoordinate>
, 
<quantity id="4B229B35242BFFE7B4024B24C651FE31" box="[267,317,363,385]" metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="270" unit="m" value="36.0">
<elevation id="07F7D1E3242BFFE7B4024B24C651FE31" box="[267,317,363,385]" metricMagnitude="1" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="270" unit="m" value="36.0">36 m</elevation>
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in elevation (
<date id="F8641010242BFFE7B4CD4B24C526FE30" box="[452,586,363,385]" pageId="5" pageNumber="270" value="2011-05-26">
<collectingDate id="E820E9F8242BFFE7B4CD4B24C526FE30" box="[452,586,363,385]" pageId="5" pageNumber="270" value="2011-05-26">26 May 2011</collectingDate>
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)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C4C0655B242BFFE7B5784BE6C22FFE07" pageId="5" pageNumber="270" type="description">
<paragraph id="8C6536D0242BFFE7B5784BE6C625FC30" blockId="5.[113,753,425,897]" pageId="5" pageNumber="270">
<emphasis id="BEAEEAC2242BFFE7B5784BE6C790FE70" bold="true" box="[113,252,425,449]" pageId="5" pageNumber="270">
<emphasis id="BEAEEAC2242BFFE7B5784BE6C798FE70" bold="true" box="[113,244,425,449]" italics="true" pageId="5" pageNumber="270">Description</emphasis>
.
</emphasis>
— Dimensions: 
<quantity id="4B229B35242BFFE7B4C84BE6C556FE71" box="[449,570,425,449]" metricMagnitude="-2" metricUnit="m" metricValue="7.15" metricValueMax="9.1" metricValueMin="5.2" pageId="5" pageNumber="270" unit="mm" value="71.5" valueMax="91.0" valueMin="52.0">52–91 mm</quantity>
body length by 1.4–2.5 mm in the anterior body region, body cylindrical in the anterior part but quadrangular in transverse section after clitellum, with 192–313 segments. The body colour is pale brown. Dorsal surface considerably broader than ventral at the posterior end. The clitellar wings on ventro-lateral part of the clitellum in ¾18, 18–21, ½22, with 2.1–2.6 mm long, and 0.5–1.2 mm wide on both sides. Prostomium zygolobous. Dorsal pores absent. Clitellum annular shape from 15, 16, 17–23, 24, 25. Four pairs of setae per segment from 2. Female pores, male pores, and spermathecal pores not visible. Genital markings: lateral series paired or asymmetrical, between line b and c in 13–15; lateral series paired or asymmetrical, lateral to line b in 16, 17, 22, 23; unpaired median series between aa in 12–16, 22–25.
</paragraph>
<paragraph id="8C6536D0242BFFE7B57849E6C3EEFF47" blockId="5.[113,752,937,1153]" lastBlockId="5.[812,1451,190,246]" pageId="5" pageNumber="270">Septa 4/5–8/9 thick, 9/10–14/15 moderately thick, and extremely thin from 15/16 to the end. Gizzard globular in 8. Intestine enlarged from 16. Hearts from 9–11. No nephridia distinguishable in the first 13 segments. Four pairs of seminal vesicles in 9–12, with the pair in 12 longest, parallel with intestine and extending to 15, 16 or 17. Ovaries in 13. Prostate and accessory glands absent. Spermathecae sessile, globular, 0.2–0.3 mm in diameter, on segments 14/15–16/17, two to three on each side per segment.</paragraph>
<paragraph id="8C6536D0242BFFE7B6254B52C22FFE07" blockId="5.[812,1452,285,438]" pageId="5" pageNumber="270">
<emphasis id="BEAEEAC2242BFFE7B6254B52C4EAFE87" bold="true" box="[812,902,285,310]" pageId="5" pageNumber="270">
<emphasis id="BEAEEAC2242BFFE7B6254B52C413FE87" bold="true" box="[812,895,285,310]" italics="true" pageId="5" pageNumber="270">Habitat</emphasis>
.
</emphasis>
— Found on the shore but in proximity to the river water, in the sandy mud topsoil and also under the water at a depth of about 
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. The river bank soil surface was covered with worm casts at Sungei Berkoh and Tok Bok Hotspring. Also found in the sandy clay in Sungei 
<collectingRegion id="4E1EF832242BFFE7B1D54BD1C22CFE07" box="[1244,1344,414,438]" country="Malaysia" name="Kelantan" pageId="5" pageNumber="270">Kelantan</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C4C0655B242BFFE7B6254B92C3A5FC07" pageId="5" pageNumber="270" type="discussion">
<paragraph id="8C6536D0242BFFE7B6254B92C3A5FC07" blockId="5.[812,1452,477,950]" pageId="5" pageNumber="270">
<emphasis id="BEAEEAC2242BFFE7B6254B92C4F8FE47" bold="true" box="[812,916,477,502]" pageId="5" pageNumber="270">
<emphasis id="BEAEEAC2242BFFE7B6254B92C4E1FE47" bold="true" box="[812,909,477,502]" italics="true" pageId="5" pageNumber="270">Remarks</emphasis>
.
</emphasis>
— 
<taxonomicName id="4BDA4D53242BFFE7B6B04B90C3DBFE47" authorityName="Michaelsen" authorityYear="1902" box="[953,1207,479,502]" class="Clitellata" family="Glossoscolecidae" genus="Glyphidrilus" kingdom="Animalia" order="Haplotaxida" pageId="5" pageNumber="270" phylum="Annelida" rank="species" species="malayanus">
<emphasis id="BEAEEAC2242BFFE7B6B04B90C3DBFE47" box="[953,1207,479,502]" italics="true" pageId="5" pageNumber="270">Glyphidrilus malayanus</emphasis>
</taxonomicName>
from Malay Peninsula: Lubock Paku, 
<collectingRegion id="4E1EF832242BFFE7B6CB4BB1C37CFDA7" box="[962,1040,510,534]" country="Malaysia" name="Pahang" pageId="5" pageNumber="270">Pahang</collectingRegion>
River, 
<collectingCountry id="F4CD7640242BFFE7B1504BB1C3D5FDA7" box="[1113,1209,510,534]" name="Malaysia" pageId="5" pageNumber="270">Malaysia</collectingCountry>
has clitellum in 15–25, wings in ¾18–½22, and the absence of median genital markings. The specimens collected from Sungei Berkoh, Tok Bok Hotspring, and Sungei 
<collectingRegion id="4E1EF832242BFFE7B18F4811C38AFDC7" box="[1158,1254,606,630]" country="Malaysia" name="Kelantan" pageId="5" pageNumber="270">Kelantan</collectingRegion>
in this study were slightly different from the type specimen. The positions of the clitellum and wings of the additional specimens are slightly different from the wings location of the type specimen by one or two segments, and median genital markings were present in these specimens. Most of the morphological characteristics are quite similar to the 
<typeStatus id="53618872242BFFE7B12C4951C3EEFC87" box="[1061,1154,798,822]" pageId="5" pageNumber="270" type="holotype">holotype</typeStatus>
specimen. The smaller size is the only distinct character noticed. We added up our own drawings of the 
<typeStatus id="53618872242BFFE7B6D24911C354FCC7" box="[987,1080,862,886]" pageId="5" pageNumber="270" type="holotype">holotype</typeStatus>
shown in 
<figureCitation id="14E12A55242BFFE7B1A04911C271FCC4" box="[1193,1309,862,886]" captionStart="Fig" captionStartId="7.[113,148,2034,2056]" captionText="Fig. 5. External and internal morphology of holotype (ZMH V5875) of Glyphidrilus malayanus: A, external ventral view; B, internal dorsal view; C, specimen collected from Sungei Kelatan, Kelantan, Malaysia (CUMZ 3219); D, from Sungei Berkoh, Johor, Malaysia (CUMZ 3220); E, from Tok Bok Hotspring, Kelantan, Malaysia (CUMZ 3218), Malaysia." pageId="5" pageNumber="270">Fig. 5A, B</figureCitation>
, and of other specimens collected from different localities in 
<figureCitation id="14E12A55242BFFE7B0344931C21FFC27" box="[1341,1395,894,918]" captionStart="Fig" captionStartId="7.[113,148,2034,2056]" captionText="Fig. 5. External and internal morphology of holotype (ZMH V5875) of Glyphidrilus malayanus: A, external ventral view; B, internal dorsal view; C, specimen collected from Sungei Kelatan, Kelantan, Malaysia (CUMZ 3219); D, from Sungei Berkoh, Johor, Malaysia (CUMZ 3220); E, from Tok Bok Hotspring, Kelantan, Malaysia (CUMZ 3218), Malaysia." pageId="5" pageNumber="270">Fig.5</figureCitation>
C–E, and also the colour picture in 
<figureCitation id="14E12A55242BFFE7B17849D1C3A9FC07" box="[1137,1221,926,950]" captionStart="Fig" captionStartId="5.[112,147,1918,1940]" captionTargetBox="[113,752,1213,1895]" captionTargetId="figure-285@5.[113,754,1213,1895]" captionTargetPageId="5" captionText="Fig. 3. The colouration of newly collected specimens just after the first step preservation in 30% ethanol:A, Glyphidrilus mekongensis, new species (paratype CUMZ 3216); B, Glyphidrilus malayanus from Tok Bok Hotspring, Kelantan, Malaysia (CUMZ 3218); C, Glyphidrilus singaporensis topotype specimen (CUMZ 3217); D, cocoon characters of Glyphidrilus mekongensis, new species (paratype CUMZ 3216)." figureDoi="http://doi.org/10.5281/zenodo.4508504" httpUri="https://zenodo.org/record/4508504/files/figure.png" pageId="5" pageNumber="270">Fig. 3B</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
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