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<document id="D2A805B8226C08BA47E9EED31E57BD5D" ID-CLB-Dataset="264461" ID-DOI="10.11646/phytotaxa.609.4.4" ID-GBIF-Dataset="0795a680-d29e-476e-b22d-ab08745c201f" ID-ISSN="1179-3163" ID-Zenodo-Dep="8297126" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1693305759606" checkinUser="plazi" docAuthor="Hoang, Thanh Son &amp; Lin, Che-Wei" docDate="2023" docId="0442EA277E19FF9EFF60F8D9FD19FD33" docLanguage="en" docName="phytotaxa.609.4.4.pdf" docOrigin="Phytotaxa 609 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.609.4.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.14:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="14" docTitle="Begonia datii T. S. Hoang &amp; C. W. Lin. 2023, sp. nov." docType="treatment" docVersion="9" lastPageNumber="287" masterDocId="F87B925F7E19FF9BFFE8FFEDF956FF83" masterDocTitle="Begonia datii (sect. Coelocentrum Begoniaceae) a new species from Quang Binh Province, Central Vietnam" masterLastPageNumber="288" masterPageNumber="282" pageNumber="282" updateTime="1732208532709" updateUser="felipe" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8BAEF3BA7DF7C105220A1AE2A01320E6">Begonia datii (sect. Coelocentrum Begoniaceae) a new species from Quang Binh Province, Central Vietnam</mods:title>
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<mods:namePart id="7E9334AD21BD237A60B73C327A1932A7">Hoang, Thanh Son</mods:namePart>
<mods:nameIdentifier id="2059FA72BFDBBC041BD064D43BE0AC00" type="ORCID">0000-0003-0351-929X</mods:nameIdentifier>
<mods:affiliation id="751F7FEE262363B9805B0D19D87B7D0D">Silviculture Research Institute (SRI) - Vietnamese Academy of Forest Sciences, 46 Duc Thang, Bac Tu Liem, Ha Noi 100000, Vietnam Herbarium of Taiwan Forestry Research Institute, Taiwan Forestry Research Institute, No. 53, Nan-Hai Road, Taipei 100, Taiwan &amp; hoangsonfsiv @ gmail. com; https: // orcid. org / 0000 - 0003 - 0351 - 929 X</mods:affiliation>
<mods:nameIdentifier id="1C893447C4D495AB7BC5D7E07B4F1E28" type="email">hoangsonfsiv@gmail.com</mods:nameIdentifier>
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<mods:namePart id="581080659AB91EDA636E9F8278CE1709">Lin, Che-Wei</mods:namePart>
<mods:nameIdentifier id="8657DCF4B5E98DA8E2DF80B678F7D90B" type="ORCID">0000-0002-8564-5793</mods:nameIdentifier>
<mods:affiliation id="512A4E4943A05C5306D80CDA5AE81F4E">varalba @ gmail. com; https: // orcid. org / 0000 - 0002 - 8564 - 5793</mods:affiliation>
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<mods:title id="A659F387427D53FFC30BD4723BBA4799">Phytotaxa</mods:title>
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<mods:date id="A0EF45C07E8596257F4BF5A59E2095F7">2023</mods:date>
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<mods:number id="1BBCE450811E6CCC17FC7D917C9436B6">2023-08-29</mods:number>
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<mods:number id="68D5BCEBA0E81D747C38371EFD19D95B">609</mods:number>
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<mods:number id="024F94D21D96BEEE4CBC781ABA107FF2">4</mods:number>
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<treatment id="0442EA277E19FF9EFF60F8D9FD19FD33" ID-DOI="http://doi.org/10.5281/zenodo.8301461" ID-GBIF-Taxon="212650356" ID-Zenodo-Dep="8301461" LSID="urn:lsid:plazi:treatment:0442EA277E19FF9EFF60F8D9FD19FD33" httpUri="http://treatment.plazi.org/id/0442EA277E19FF9EFF60F8D9FD19FD33" lastPageId="5" lastPageNumber="287" pageId="0" pageNumber="282">
<subSubSection id="C4F108BA7E19FF9BFF60F8D9FBD1F8CD" box="[136,647,1844,1871]" pageId="0" pageNumber="282" type="nomenclature">
<paragraph id="8C545B317E19FF9BFF60F8D9FBD1F8CD" blockId="0.[136,1114,1844,1871]" box="[136,647,1844,1871]" pageId="0" pageNumber="282">
<heading id="D71CEC5D7E19FF9BFF60F8D9FBD1F8CD" box="[136,647,1844,1871]" fontSize="11" level="2" pageId="0" pageNumber="282" reason="3">
<taxonomicName id="4BEB20B27E19FF9BFF60F8D9FB71F8CD" ID-CoL="d9d99201-d8c6-4723-ae9a-a318377e62ad" authority="T.S.Hoang &amp; C.W.Lin, 2023" authorityName="T. S. Hoang &amp; C. W. Lin." authorityYear="2023" box="[136,551,1844,1871]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="0" pageNumber="282" phylum="Tracheophyta" rank="species" species="datii" status="sp. nov.">
<emphasis id="BE9F87237E19FF9BFF60F8D9F875F8CC" bold="true" box="[136,291,1844,1871]" italics="true" pageId="0" pageNumber="282">Begonia datii</emphasis>
T.S.Hoang &amp; C.W.Lin
</taxonomicName>
,
<emphasis id="BE9F87237E19FF9BFDDAF8D8FBD1F8CD" box="[562,647,1845,1870]" italics="true" pageId="0" pageNumber="282">
<taxonomicNameLabel id="A5AC3A587E19FF9BFDDAF8D8FBD1F8CD" box="[562,647,1845,1870]" pageId="0" pageNumber="282" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C4F108BA7E19FF9BFD66F8D9FD0CF8CD" box="[654,1114,1844,1870]" pageId="0" pageNumber="282" type="multiple">
<paragraph id="8C545B317E19FF9BFD66F8D9FD0CF8CD" blockId="0.[136,1114,1844,1871]" box="[654,1114,1844,1870]" pageId="0" pageNumber="282">
§
<taxonomicName id="4BEB20B27E19FF9BFD4BF8D9FA14F8CD" authorityName="Crosse &amp; P. Fischer" authorityYear="1872" box="[675,834,1844,1870]" class="Gastropoda" family="Eucalodiidae" genus="Coelocentrum" kingdom="Animalia" order="Stylommatophora" pageId="0" pageNumber="282" phylum="Mollusca" rank="genus">
<emphasis id="BE9F87237E19FF9BFD4BF8D9FA14F8CD" box="[675,834,1844,1870]" italics="true" pageId="0" pageNumber="282">Coelocentrum</emphasis>
</taxonomicName>
(
<figureCitation id="14D047B47E19FF9BFCB9F8D9FACFF8CD" box="[849,921,1844,1870]" captionStart="FIGURE 1" captionStartId="1.[136,234,984,1006]" captionTargetBox="[268,1320,190,960]" captionTargetId="figure-20@1.[268,1320,190,960]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of Begonia datii (red star), B. bamaensis (orange hexagon), B. filiformis (blue circles), B. longistyla (pink diamond), B. montaniformis (yellow square), B. morsei Irmsch (green triangle) and B. morsei var. myriotricha Y.M.Shui &amp; W.H.Chen (purple inverted triangle) in China and Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8297128" httpUri="https://zenodo.org/record/8297128/files/figure.png" pageId="0" pageNumber="282">Figs.1</figureCitation>
,
<figureCitation id="14D047B47E19FF9BFC4DF8D9FAE3F8CD" box="[933,949,1844,1870]" captionStart="FIGURE 2" captionStartId="2.[136,234,1942,1964]" captionTargetBox="[246,1352,365,1906]" captionTargetId="figure-95@2.[235,1352,353,1917]" captionTargetPageId="2" captionText="FIGURE 2. Begonia datii T.S.Hoang &amp; C.W.Lin. A. Habit; B, B’. Portion of leaf adaxial and abaxial surfaces; C. Stipule; D–D’’’. Bracts; E, E’. Staminate flower, face and side views; F, F’. Stamens, dorsal and ventral views; G, G’. Pistillate flower, face and side views; H, H’. Style and stigmatic band, dorsal and ventral views; I, I’. Capsules; J. Cross section of ovary." figureDoi="http://doi.org/10.5281/zenodo.8297130" httpUri="https://zenodo.org/record/8297130/files/figure.png" pageId="0" pageNumber="282">2</figureCitation>
,
<figureCitation id="14D047B47E19FF9BFC29F8D9FA99F8CD" box="[961,975,1844,1870]" captionStart="FIGURE 3" captionStartId="3.[136,234,1935,1957]" captionTargetBox="[151,1436,190,1910]" captionTargetId="figure-21@3.[151,1436,190,1910]" captionTargetPageId="3" captionText="FIGURE 3. Begonia datii T.S.Hoang &amp; C.W.Lin. A, B, C. Habit and habitats; D. Rhizome and petioles; E. Stipules; F. Portion of leaf adaxial surface, G. same, abaxial surfaces; H. Inflorescence, also showing bracts; I. Staminate flowers, face and back views; J. Pistillate flowers, face and side views; K. Capsule; L. Cross section of ovary.All from Hoang Son 4435 by T.S. Hoang." figureDoi="http://doi.org/10.5281/zenodo.8297132" httpUri="https://zenodo.org/record/8297132/files/figure.png" pageId="0" pageNumber="282">3</figureCitation>
&amp;
<figureCitation id="14D047B47E19FF9BFC1BF8D9FD4AF8CD" box="[1011,1052,1844,1870]" captionStart="FIGURE 4" captionStartId="4.[136,234,1574,1596]" captionTargetBox="[151,1435,190,1550]" captionTargetId="figure-17@4.[151,1435,190,1550]" captionTargetPageId="4" captionText="FIGURE 4. Morphological comparison of Begonia datii and allied taxa, showing habits and morphological characters: B. datii (A–A’’’), B. bamaensis (B–B’’’), B. filiformis (C–C’’’), B. longistyla (D–D’’’), B. scabrifolia (E–E’’’). A, B, C, D, E. Habit; A’, B’, C’, D’, E’. Leaf, adaxial surfaces; A’’, B’’, C’’, D’’, E’’. Portion of leaf adaxial surfaces; A’’’, B’’’, C’’’, D’’’, E’’’. Flowers. A–A’’’ from Hoang Son 4435 and C–C’’’ from Hoang Son 6433 by T.S. Hoang; B–B’’’ from Peng 18752, D–D’’’ from Peng 20435 and E–E’’’ from Peng 22197 by C.-I Peng." figureDoi="http://doi.org/10.5281/zenodo.8297137" httpUri="https://zenodo.org/record/8297137/files/figure.png" pageId="0" pageNumber="282">4 A</figureCitation>
–A’’’)
</paragraph>
</subSubSection>
<subSubSection id="C4F108BA7E19FF9BFF60F86DFAA2F83A" pageId="0" pageNumber="282" type="materials_examined">
<paragraph id="8C545B317E19FF9BFF60F86DFAA2F83A" blockId="0.[136,1452,1920,1978]" pageId="0" pageNumber="282">
<emphasis id="BE9F87237E19FF9BFF60F86DF994F816" bold="true" box="[136,194,1920,1941]" pageId="0" pageNumber="282">Type:</emphasis>
—
<materialsCitation id="3C83516C7E19FF9BFF33F86DFA1AF83A" ID-GBIF-Occurrence="4402250302" collectingDate="2022-04-20" collectionCode="VAFS" collectorName="Hoang Son" country="Vietnam" elevation="495" location="Thuong Hoa" municipality="Minh Hoa district" pageId="0" pageNumber="282" specimenCount="1" stateProvince="Quang Binh Province" typeStatus="holotype">
<collectingCountry id="F4FC1BA17E19FF9BFF33F86DF81AF816" box="[219,332,1920,1941]" name="Vietnam" pageId="0" pageNumber="282">VIETNAM</collectingCountry>
.
<collectingRegion id="4E2F95D37E19FF9BFEBDF86DFB71F815" box="[341,551,1920,1942]" country="Vietnam" name="Quang Binh" pageId="0" pageNumber="282">Quang Binh Province</collectingRegion>
,
<collectingMunicipality id="6C30C14B7E19FF9BFDD8F86DFB8BF815" box="[560,733,1920,1942]" pageId="0" pageNumber="282">Minh Hoa district</collectingMunicipality>
,
<location id="89340DEA7E19FF9BFD0EF86DFA08F816" LSID="urn:lsid:plazi:treatment:0442EA277E19FF9EFF60F8D9FD19FD33:89340DEA7E19FF9BFD0EF86DFA08F816" box="[742,862,1920,1942]" country="Vietnam" municipality="Minh Hoa district" name="Thuong Hoa" pageId="0" pageNumber="282" stateProvince="Quang Binh Province">Thuong Hoa</location>
commune,
<quantity id="4B13F6D47E19FF9BFC27F86DFD5AF816" box="[975,1036,1920,1942]" metricMagnitude="2" metricUnit="m" metricValue="4.95" pageId="0" pageNumber="282" unit="m" value="495.0">
<elevation id="07C6BC027E19FF9BFC27F86DFD5AF816" box="[975,1036,1920,1942]" metricMagnitude="2" metricUnit="m" metricValue="4.95" pageId="0" pageNumber="282" unit="m" value="495.0">495 m</elevation>
</quantity>
elev., grows on limestone mountains under secondary forests,
<date id="F8557DF17E19FF9BFE99F849F8AEF83A" box="[369,504,1956,1978]" pageId="0" pageNumber="282" value="2022-04-20">
<collectingDate id="E81184197E19FF9BFE99F849F8AEF83A" box="[369,504,1956,1978]" pageId="0" pageNumber="282" value="2022-04-20">20 April 2022</collectingDate>
</date>
,
<emphasis id="BE9F87237E19FF9BFDEAF849FBF5F83A" box="[514,675,1956,1977]" italics="true" pageId="0" pageNumber="282">
<collectorName id="211E3EE77E19FF9BFDEAF849FB3BF83A" box="[514,621,1956,1977]" pageId="0" pageNumber="282">Hoang Son</collectorName>
4435
</emphasis>
(
<typeStatus id="5350E5937E19FF9BFD58F849FA50F839" box="[688,774,1956,1978]" pageId="0" pageNumber="282" type="holotype">holotype</typeStatus>
<collectionCode id="EAFAC3F47E19FF9BFCE3F849FA1EF83A" box="[779,840,1956,1977]" pageId="0" pageNumber="282">VAFS</collectionCode>
,
</materialsCitation>
<materialsCitation id="3C83516C7E19FF9BFCB9F849FAA7F83A" ID-GBIF-Occurrence="4402250301" box="[849,1009,1956,1978]" collectingDate="2022-04-20" collectionCode="VNMN" country="Vietnam" elevation="495" location="Thuong Hoa" municipality="Minh Hoa district" pageId="0" pageNumber="282" specimenCount="1" stateProvince="Quang Binh" typeStatus="isotype">
<typeStatus id="5350E5937E19FF9BFCB9F849FAC0F839" box="[849,918,1956,1978]" pageId="0" pageNumber="282" type="isotype">isotype</typeStatus>
<collectionCode id="EAFAC3F47E19FF9BFC74F849FABCF83A" box="[924,1002,1956,1977]" collectionName="VNMN" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="0" pageNumber="282">VNMN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="D8940BB97E18FF9AFF60FC35FB0EFBB5" ID-DOI="http://doi.org/10.5281/zenodo.8297128" ID-Zenodo-Dep="8297128" httpUri="https://zenodo.org/record/8297128/files/figure.png" pageId="1" pageNumber="283" startId="1.[136,234,984,1006]" targetBox="[268,1320,190,960]" targetPageId="1" targetType="figure">
<paragraph id="8C545B317E18FF9AFF60FC35FB0EFBB5" blockId="1.[136,1451,984,1078]" pageId="1" pageNumber="283">
<emphasis id="BE9F87237E18FF9AFF60FC35F854FC6E" bold="true" box="[136,258,984,1006]" pageId="1" pageNumber="283">FIGURE 1.</emphasis>
Distribution map of
<taxonomicName id="4BEB20B27E18FF9AFE21FC35FB1BFC6D" authorityName="T. S. Hoang &amp; C. W. Lin. A. Habit" authorityYear="2023" box="[457,589,984,1006]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="283" phylum="Tracheophyta" rank="species" species="datii">
<emphasis id="BE9F87237E18FF9AFE21FC35FB1BFC6D" bold="true" box="[457,589,984,1006]" italics="true" pageId="1" pageNumber="283">Begonia datii</emphasis>
</taxonomicName>
(red star),
<taxonomicName id="4BEB20B27E18FF9AFD5DFC34FA6FFC6D" box="[693,825,984,1006]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="283" phylum="Tracheophyta" rank="species" species="bamaensis">
<emphasis id="BE9F87237E18FF9AFD5DFC34FA6FFC6D" bold="true" box="[693,825,984,1006]" italics="true" pageId="1" pageNumber="283">B. bamaensis</emphasis>
</taxonomicName>
(orange hexagon),
<taxonomicName id="4BEB20B27E18FF9AFC1AFC34FD3FFC6D" baseAuthorityName="Irmscher" baseAuthorityYear="1939" box="[1010,1129,984,1006]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="283" phylum="Tracheophyta" rank="species" species="filiformis">
<emphasis id="BE9F87237E18FF9AFC1AFC34FD3FFC6D" bold="true" box="[1010,1129,984,1006]" italics="true" pageId="1" pageNumber="283">B. filiformis</emphasis>
</taxonomicName>
(blue circles),
<taxonomicName id="4BEB20B27E18FF9AFB10FC34FC25FC6D" baseAuthorityName="Shui &amp; Chen" baseAuthorityYear="2005" box="[1272,1395,984,1006]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="283" phylum="Tracheophyta" rank="species" species="longistyla">
<emphasis id="BE9F87237E18FF9AFB10FC34FC25FC6D" bold="true" box="[1272,1395,984,1006]" italics="true" pageId="1" pageNumber="283">B. longistyla</emphasis>
</taxonomicName>
(pink diamond),
<taxonomicName id="4BEB20B27E18FF9AFF19FC10F8F6FB91" box="[241,416,1020,1042]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="283" phylum="Tracheophyta" rank="species" species="montaniformis">
<emphasis id="BE9F87237E18FF9AFF19FC10F8F6FB91" bold="true" box="[241,416,1020,1042]" italics="true" pageId="1" pageNumber="283">B. montaniformis</emphasis>
</taxonomicName>
(yellow square),
<taxonomicName id="4BEB20B27E18FF9AFDA2FC10FBA5FB91" authority="Irmsch" authorityName="Irmsch" box="[586,755,1020,1042]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="283" phylum="Tracheophyta" rank="species" species="morsei">
<emphasis id="BE9F87237E18FF9AFDA2FC10FBFFFB92" bold="true" box="[586,681,1020,1042]" italics="true" pageId="1" pageNumber="283">B. morsei</emphasis>
Irmsch
</taxonomicName>
(green triangle) and
<taxonomicName id="4BEB20B27E18FF9AFC56FC10FCFDFB91" authority="Y. M. Shui &amp; W. H. Chen" authorityName="Y. M. Shui &amp; W. H. Chen" box="[958,1451,1020,1042]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="283" phylum="Tracheophyta" rank="variety" species="morsei" variety="myriotricha">
<emphasis id="BE9F87237E18FF9AFC56FC10FD93FB91" bold="true" box="[958,1221,1020,1042]" pageId="1" pageNumber="283">
<emphasis id="BE9F87237E18FF9AFC56FC10FD48FB92" bold="true" box="[958,1054,1020,1042]" italics="true" pageId="1" pageNumber="283">B. morsei</emphasis>
var.
<emphasis id="BE9F87237E18FF9AFBB9FC11FD93FB91" bold="true" box="[1105,1221,1020,1042]" italics="true" pageId="1" pageNumber="283">myriotricha</emphasis>
</emphasis>
Y.M.Shui &amp; W.H.Chen
</taxonomicName>
(purple inverted triangle) in China and Vietnam.
</paragraph>
</caption>
<subSubSection id="C4F108BA7E18FF9AFF60FB89FAB2FB68" pageId="1" pageNumber="283" type="diagnosis">
<paragraph id="8C545B317E18FF9AFF60FB89FAB2FB68" blockId="1.[136,1452,1124,2087]" pageId="1" pageNumber="283">
<emphasis id="BE9F87237E18FF9AFF60FB89F853FBFD" bold="true" box="[136,261,1124,1150]" pageId="1" pageNumber="283">Diagnosis:</emphasis>
—It resembles
<taxonomicName id="4BEB20B27E18FF9AFE5AFB8BFB6FFBFC" baseAuthorityName="Irmscher" baseAuthorityYear="1939" box="[434,569,1125,1151]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="1" pageNumber="283" phylum="Tracheophyta" rank="species" species="filiformis">
<emphasis id="BE9F87237E18FF9AFE5AFB8BFB6FFBFC" box="[434,569,1125,1151]" italics="true" pageId="1" pageNumber="283">B. filiformis</emphasis>
</taxonomicName>
in its habit, leaf shape and texture, but it can be easily distinguished from the latter by its peduncle and inflorescence covered with villous hairs (vs. glandular-hispid), bracts caducous (vs. persistent or semi-persistent), tepals pinkish-white to pink (vs. yellowish-green), glabrous outer tepals of pistillate flowers (vs. glandular-hispid abaxially), ovary and wings glabrous (vs. glandular-hispid).
</paragraph>
</subSubSection>
<subSubSection id="C4F108BA7E18FF99FF55FB18FA59FECA" lastPageId="2" lastPageNumber="284" pageId="1" pageNumber="283" type="description">
<paragraph id="8C545B317E18FF99FF55FB18FA59FECA" blockId="1.[136,1452,1124,2087]" lastBlockId="2.[136,1452,158,329]" lastPageId="2" lastPageNumber="284" pageId="1" pageNumber="283">
Monoecious rhizomatous herb.
<emphasis id="BE9F87237E18FF9AFDF7FB19FBDFFA8D" bold="true" box="[543,649,1268,1294]" pageId="1" pageNumber="283">Rhizome</emphasis>
stout, crimson to reddish-brown,
<quantity id="4B13F6D47E18FF9AFBEAFB18FD3BFA8C" box="[1026,1133,1269,1295]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="2.0" metricValueMin="0.8" pageId="1" pageNumber="283" unit="mm" value="14.0" valueMax="20.0" valueMin="8.0">8–20 mm</quantity>
thick, internodes congested, villous.
<emphasis id="BE9F87237E18FF9AFF0FFAF5F810FAB1" bold="true" box="[231,326,1304,1330]" pageId="1" pageNumber="283">Stipules</emphasis>
persistent, yellowish-red to reddish-brown, narrowly triangular, 6–14 ×
<quantity id="4B13F6D47E18FF9AFB75FAF4FC40FAB0" box="[1181,1302,1305,1332]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="8.0" metricValueMin="3.5" pageId="1" pageNumber="283" unit="mm" value="5.75" valueMax="8.0" valueMin="3.5">3.5–8 mm</quantity>
, herbaceous, strongly keeled, midrib sparsely hirsute, margin entire, apex aristate, arista
<quantity id="4B13F6D47E18FF9AFC33FAD0FD18FAD4" box="[987,1102,1341,1368]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="1" pageNumber="283" unit="mm" value="3.75" valueMax="5.0" valueMin="2.5">2.5–5 mm</quantity>
long.
<emphasis id="BE9F87237E18FF9AFB7EFAD0FDBEFAD4" bold="true" box="[1174,1256,1341,1367]" pageId="1" pageNumber="283">Leaves</emphasis>
alternate, petiole terete, maroon to red,
<quantity id="4B13F6D47E18FF9AFE9EFA8CF882FAF8" box="[374,468,1377,1403]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" metricValueMax="3.1" metricValueMin="0.9" pageId="1" pageNumber="283" unit="cm" value="20.0" valueMax="31.0" valueMin="9.0">9–31 cm</quantity>
long,
<quantity id="4B13F6D47E18FF9AFDFFFA8CFB29FAF8" box="[535,639,1377,1403]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="1" pageNumber="283" unit="mm" value="7.0" valueMax="10.0" valueMin="4.0">4–10 mm</quantity>
diameter, densely white villous; leaf blade asymmetric, thick chartaceous, succulent, widely ovate, 13–22.5 ×
<quantity id="4B13F6D47E18FF9AFDCEFA68FBF4FA1C" box="[550,674,1413,1440]" metricMagnitude="-1" metricUnit="m" metricValue="1.225" metricValueMax="1.6" metricValueMin="0.85" pageId="1" pageNumber="283" unit="cm" value="12.25" valueMax="16.0" valueMin="8.5">8.5–16 cm</quantity>
, broad side
<quantity id="4B13F6D47E18FF9AFCD9FA68FAFCFA1C" box="[817,938,1413,1440]" metricMagnitude="-2" metricUnit="m" metricValue="7.65" metricValueMax="10.0" metricValueMin="5.3" pageId="1" pageNumber="283" unit="cm" value="7.65" valueMax="10.0" valueMin="5.3">5.3–10 cm</quantity>
wide, basal lobes cordate,
<quantity id="4B13F6D47E18FF9AFB00FA68FC3EFA1C" box="[1256,1384,1413,1439]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="6.7" metricValueMin="3.8" pageId="1" pageNumber="283" unit="cm" value="5.25" valueMax="6.7" valueMin="3.8">3.8–6.7 cm</quantity>
long, margin denticulate with a line of pinkish-white puberulous hairs, apex acuminate to caudate; adaxial surface emerald green, dark olive green to blackish-green, embellished with silvery white patches or semicontinuous stripes of variable sizes between major veins, rarely completely green, densely covered by small raised cones each topped by a pinkish-white to magenta puberulous hairs, giving a wrinkled texture; abaxially purplish-red to magenta; venation basally
<emphasis id="BE9F87237E18FF9AFA62F9FBFCF2F9AC" box="[1418,1444,1558,1583]" italics="true" pageId="1" pageNumber="283">ca</emphasis>
. 7 palmate, midrib distinct,
<emphasis id="BE9F87237E18FF9AFE5CF9D7F898F9D0" box="[436,462,1594,1619]" italics="true" pageId="1" pageNumber="283">ca</emphasis>
. 2 secondary veins on each side, other primary veins branching dichotomously or nearly so, tertiary veins reticulate; major veins densely pinkish-white to magenta puberulous on adaxial surface, densely pink to magenta puberulous on all veins abaxilly.
<emphasis id="BE9F87237E18FF9AFD60F96DFA67F919" bold="true" box="[648,817,1664,1690]" pageId="1" pageNumber="283">Inflorescences</emphasis>
axillary, bisexual, cymose panicle arising directly from rhizome,
<emphasis id="BE9F87237E18FF9AFF19F94BF85DF93C" box="[241,267,1702,1727]" italics="true" pageId="1" pageNumber="283">ca</emphasis>
. 4 orders of branching; peduncle red,
<quantity id="4B13F6D47E18FF9AFD58F948FA1EF93C" box="[688,840,1701,1728]" metricMagnitude="-1" metricUnit="m" metricValue="1.935" metricValueMax="2.55" metricValueMin="1.3199999999999998" pageId="1" pageNumber="283" unit="cm" value="19.35" valueMax="25.5" valueMin="13.2">13.2–25.5 cm</quantity>
long, villous.
<emphasis id="BE9F87237E18FF9AFC0FF948FD63F93C" bold="true" box="[999,1077,1701,1727]" pageId="1" pageNumber="283">Bracts</emphasis>
caducous, yellowish-red, at basal node of inflorescence ovate, 10–17 ×
<quantity id="4B13F6D47E18FF9AFDD8F924FBCBF960" box="[560,669,1737,1763]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="1" pageNumber="283" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">7–10 mm</quantity>
, abaxially villous or glabrous, margin with a line of villous; bracts on upper nodes similar but gradually smaller.
<emphasis id="BE9F87237E18FF9AFD9DF901FA18F885" bold="true" box="[629,846,1772,1798]" pageId="1" pageNumber="283">Staminate flower:</emphasis>
pedicel pinkish-yellow to rosy pink,
<quantity id="4B13F6D47E18FF9AFB16F900FC3FF884" box="[1278,1385,1773,1800]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.5" metricValueMin="1.0" pageId="1" pageNumber="283" unit="cm" value="2.25" valueMax="3.5" valueMin="1.0">1–3.5 cm</quantity>
long, villous or glabrous, tepals 4, pinkish-white to pink, outer 2 ovate to widely ovate, 8–15 ×
<quantity id="4B13F6D47E18FF9AFBB3F8FCFD83F8A8" box="[1115,1237,1809,1835]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="1" pageNumber="283" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">10–13 mm</quantity>
, abaxially glabrous or villous, margin entire, apex rounded to obtuse, inner 2 narrowly obovate, 7–10 ×
<quantity id="4B13F6D47E18FF9AFBD0F8D8FDFAF8CC" box="[1080,1196,1845,1872]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="1" pageNumber="283" unit="mm" value="3.75" valueMax="4.5" valueMin="3.0">3–4.5 mm</quantity>
, glabrous, apex obtuse or rounded; androecium nearly actinomorphic,
<emphasis id="BE9F87237E18FF9AFD4DF8B7FBE9F8F0" box="[677,703,1882,1907]" italics="true" pageId="1" pageNumber="283">ca</emphasis>
.
<quantity id="4B13F6D47E18FF9AFD27F8B4FA45F8F0" box="[719,787,1881,1908]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="283" unit="mm" value="5.0">5 mm</quantity>
across; stamens golden yellow, 65–100; filaments
<emphasis id="BE9F87237E18FF9AFAB6F8B7FC29F8F0" box="[1374,1407,1882,1907]" italics="true" pageId="1" pageNumber="283">ca.</emphasis>
<quantity id="4B13F6D47E18FF9AFA60F8B4F9E2F814" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="283" unit="mm" value="1.5">1.5 mm</quantity>
long, fused at base; anthers obovate,
<emphasis id="BE9F87237E18FF9AFDB3F893FB23F814" box="[603,629,1918,1943]" italics="true" pageId="1" pageNumber="283">ca</emphasis>
.
<quantity id="4B13F6D47E18FF9AFD6CF890FB93F814" box="[644,709,1917,1943]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="283" unit="mm" value="1.0">1 mm</quantity>
long, 2-locular, apex truncate to slightly retuse.
<emphasis id="BE9F87237E18FF9AFB0EF891FCFDF815" bold="true" box="[1254,1451,1916,1942]" pageId="1" pageNumber="283">Pistillate flower:</emphasis>
pedicel pinkish-yellow to rosy pink,
<quantity id="4B13F6D47E18FF9AFDCBF84CFBCAF838" box="[547,668,1953,1979]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.3" metricValueMin="1.2" pageId="1" pageNumber="283" unit="mm" value="17.5" valueMax="23.0" valueMin="12.0">12–23 mm</quantity>
long, glabrous or very sparsely villous, tepals 3, pink, glabrous; outer 2 suborbicular,
<quantity id="4B13F6D47E18FF9AFEDDF828F8F6F85C" box="[309,416,1989,2015]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.2" metricValueMin="0.8" pageId="1" pageNumber="283" unit="mm" value="10.0" valueMax="12.0" valueMin="8.0">8–12 mm</quantity>
across, margin entire, apex rounded, inner 1 oblanceolate to narrowly elliptic,
<emphasis id="BE9F87237E18FF9AFAF8F82BFC7CF85C" box="[1296,1322,1990,2015]" italics="true" pageId="1" pageNumber="283">ca</emphasis>
. 5 ×
<quantity id="4B13F6D47E18FF9AFA8CF828FCF1F85C" box="[1380,1447,1989,2015]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="283" unit="mm" value="3.0">3 mm</quantity>
, apex obtuse; ovary trigonous-ellipsoid,
<quantity id="4B13F6D47E18FF9AFDABF804FBF8F780" box="[579,686,2025,2051]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="1" pageNumber="283" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">7–10 mm</quantity>
long,
<emphasis id="BE9F87237E18FF9AFD1CF807FA58F780" box="[756,782,2026,2051]" italics="true" pageId="1" pageNumber="283">ca</emphasis>
.
<quantity id="4B13F6D47E18FF9AFCF5F804FA0BF780" box="[797,861,2025,2051]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="283" unit="mm" value="4.0">4 mm</quantity>
thick (wings excluded), rosy pink to pinkish-yellow, glabrous; 3-winged, wings unequal, rosy pink to yellowish-pink, glabrous, margin entire; wings crescent-shaped,
<quantity id="4B13F6D47E18FF99FA9BF7E0F9E2FF3A" lastPageId="2" lastPageNumber="284" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="1" pageNumber="283" unit="mm" value="10.5" valueMax="12.0" valueMin="9.0">9–12 mm</quantity>
long, lateral wings narrower,
<quantity id="4B13F6D47E1BFF99FE17FF73FB26FF3A" box="[511,624,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="4.5" metricValueMin="2.0" pageId="2" pageNumber="284" unit="mm" value="3.25" valueMax="4.5" valueMin="2.0">2–4.5 mm</quantity>
wide, abaxial wing
<quantity id="4B13F6D47E1BFF99FCA7FF73FA96FF3A" box="[847,960,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="6.5" metricValueMin="3.0" pageId="2" pageNumber="284" unit="mm" value="4.75" valueMax="6.5" valueMin="3.0">3–6.5 mm</quantity>
wide; ovary 1-locular with intruded parietal placentation, placentae 3, bilamellate; styles 3, fused at base, golden yellow,
<emphasis id="BE9F87237E1BFF99FC0BFF2EFD52FF5F" box="[995,1028,195,220]" italics="true" pageId="2" pageNumber="284">ca.</emphasis>
<quantity id="4B13F6D47E1BFF99FBE3FF2FFD37FF5E" box="[1035,1121,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="284" unit="mm" value="3.5">3.5 mm</quantity>
long, stigma spirally twisted.
<emphasis id="BE9F87237E1BFF99FF60FF0BF9BEFE83" bold="true" box="[136,232,230,256]" pageId="2" pageNumber="284">Capsule</emphasis>
capsule trigonous-ellipsoid,
<quantity id="4B13F6D47E1BFF99FDDEFF0BFBE4FE82" box="[566,690,230,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="2" pageNumber="284" unit="mm" value="15.5" valueMax="18.0" valueMin="13.0">13–18 mm</quantity>
long,
<quantity id="4B13F6D47E1BFF99FD17FF0BFA09FE82" box="[767,863,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="2" pageNumber="284" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
thick (wings excluded), reddish-green when fresh; wings unequal,
<quantity id="4B13F6D47E1BFF99FEDFFEE7F8F9FEA6" box="[311,431,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="2" pageNumber="284" unit="mm" value="17.5" valueMax="20.0" valueMin="15.0">15–20 mm</quantity>
long, lateral wings
<quantity id="4B13F6D47E1BFF99FD61FEE7FBB3FEA6" box="[649,741,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="2" pageNumber="284" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
wide, abaxial wing crescent-shaped to sub-triangular,
<quantity id="4B13F6D47E1BFF99FAAAFEE7FCFAFEA6" box="[1346,1452,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" metricValueMax="1.3" metricValueMin="0.7" pageId="2" pageNumber="284" unit="mm" value="10.0" valueMax="13.0" valueMin="7.0">7–13 mm</quantity>
wide, cuneate at the base, rounded to truncate at the apex.
</paragraph>
</subSubSection>
<caption id="D8940BB97E1BFF99FF60F87BFAABF877" ID-DOI="http://doi.org/10.5281/zenodo.8297130" ID-Zenodo-Dep="8297130" httpUri="https://zenodo.org/record/8297130/files/figure.png" pageId="2" pageNumber="284" startId="2.[136,234,1942,1964]" targetBox="[246,1352,365,1906]" targetPageId="2" targetType="figure">
<paragraph id="8C545B317E1BFF99FF60F87BFAABF877" blockId="2.[136,1452,1942,2036]" pageId="2" pageNumber="284">
<emphasis id="BE9F87237E1BFF99FF60F87BF854F828" bold="true" box="[136,258,1942,1964]" pageId="2" pageNumber="284">FIGURE 2.</emphasis>
<taxonomicName id="4BEB20B27E1BFF99FEEEF87BFB95F82F" authority="T. S. Hoang &amp; C. W. Lin. A. Habit" authorityName="T. S. Hoang &amp; C. W. Lin. A. Habit" authorityYear="2023" box="[262,707,1942,1964]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="2" pageNumber="284" phylum="Tracheophyta" rank="species" species="datii">
<emphasis id="BE9F87237E1BFF99FEEEF87BF8DCF82F" bold="true" box="[262,394,1942,1964]" italics="true" pageId="2" pageNumber="284">Begonia datii</emphasis>
T.S.Hoang &amp; C.W.Lin. A. Habit
</taxonomicName>
; B, B’. Portion of leaf adaxial and abaxial surfaces; C. Stipule; D–D’’’. Bracts; E, E’. Staminate flower, face and side views; F, F’. Stamens, dorsal and ventral views; G, G’. Pistillate flower, face and side views; H, H’. Style and stigmatic band, dorsal and ventral views; I, I’. Capsules; J. Cross section of ovary.
</paragraph>
</caption>
<caption id="D8940BB97E1AFF98FF60F862FDFEF86E" ID-DOI="http://doi.org/10.5281/zenodo.8297132" ID-Zenodo-Dep="8297132" httpUri="https://zenodo.org/record/8297132/files/figure.png" pageId="3" pageNumber="285" startId="3.[136,234,1935,1957]" targetBox="[151,1436,190,1910]" targetPageId="3" targetType="figure">
<paragraph id="8C545B317E1AFF98FF60F862FDFEF86E" blockId="3.[136,1451,1935,2029]" pageId="3" pageNumber="285">
<emphasis id="BE9F87237E1AFF98FF60F862F852F827" bold="true" box="[136,260,1935,1957]" pageId="3" pageNumber="285">FIGURE 3.</emphasis>
<taxonomicName id="4BEB20B27E1AFF98FEE2F862FBD8F827" authority="T. S. Hoang &amp; C. W. Lin. A" authorityName="T. S. Hoang &amp; C. W. Lin. A. Habit" authorityYear="2023" box="[266,654,1935,1957]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="3" pageNumber="285" phylum="Tracheophyta" rank="species" species="datii">
<emphasis id="BE9F87237E1AFF98FEE2F862F8C6F826" bold="true" box="[266,400,1935,1957]" italics="true" pageId="3" pageNumber="285">Begonia datii</emphasis>
T.S.Hoang &amp; C.W.Lin. A
</taxonomicName>
, B, C. Habit and habitats; D. Rhizome and petioles; E. Stipules; F. Portion of leaf adaxial surface, G. same, abaxial surfaces; H. Inflorescence, also showing bracts; I. Staminate flowers, face and back views; J. Pistillate flowers, face and side views; K. Capsule; L. Cross section of ovary. All from
<emphasis id="BE9F87237E1AFF98FC98F835FD47F86F" box="[880,1041,2007,2029]" italics="true" pageId="3" pageNumber="285">Hoang Son 4435</emphasis>
by T.S. Hoang.
</paragraph>
</caption>
<caption id="D8940BB97E1DFF9FFF60F9CBF85FF94F" ID-DOI="http://doi.org/10.5281/zenodo.8297137" ID-Zenodo-Dep="8297137" httpUri="https://zenodo.org/record/8297137/files/figure.png" pageId="4" pageNumber="286" startId="4.[136,234,1574,1596]" targetBox="[151,1435,190,1550]" targetPageId="4" targetType="figure">
<paragraph id="8C545B317E1DFF9FFF60F9CBF85FF94F" blockId="4.[136,1452,1574,1740]" pageId="4" pageNumber="286">
<emphasis id="BE9F87237E1DFF9FFF60F9CBF854F9B8" bold="true" box="[136,258,1574,1596]" pageId="4" pageNumber="286">FIGURE 4.</emphasis>
Morphological comparison of
<taxonomicName id="4BEB20B27E1DFF9FFDC3F9CBFBF9F9BF" authorityName="T. S. Hoang &amp; C. W. Lin. A. Habit" authorityYear="2023" box="[555,687,1574,1596]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="286" phylum="Tracheophyta" rank="species" species="datii">
<emphasis id="BE9F87237E1DFF9FFDC3F9CBFBF9F9BF" bold="true" box="[555,687,1574,1596]" italics="true" pageId="4" pageNumber="286">Begonia datii</emphasis>
</taxonomicName>
and allied taxa, showing habits and morphological characters:
<taxonomicName id="4BEB20B27E1DFF9FFAE0F9CBFC19F9BF" authorityName="T. S. Hoang &amp; C. W. Lin. A. Habit" authorityYear="2023" box="[1288,1359,1574,1596]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="286" phylum="Tracheophyta" rank="species" species="datii">
<emphasis id="BE9F87237E1DFF9FFAE0F9CBFC19F9BF" bold="true" box="[1288,1359,1574,1596]" italics="true" pageId="4" pageNumber="286">B. datii</emphasis>
</taxonomicName>
(A–A’’’),
<taxonomicName id="4BEB20B27E1DFF9FFF60F9A7F858F9E3" box="[136,270,1610,1632]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="286" phylum="Tracheophyta" rank="species" species="bamaensis">
<emphasis id="BE9F87237E1DFF9FFF60F9A7F858F9E3" bold="true" box="[136,270,1610,1632]" italics="true" pageId="4" pageNumber="286">B. bamaensis</emphasis>
</taxonomicName>
(B–B’’’),
<taxonomicName id="4BEB20B27E1DFF9FFE9BF9A7F8BBF9E3" baseAuthorityName="Irmscher" baseAuthorityYear="1939" box="[371,493,1610,1632]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="286" phylum="Tracheophyta" rank="species" species="filiformis">
<emphasis id="BE9F87237E1DFF9FFE9BF9A7F8BBF9E3" bold="true" box="[371,493,1610,1632]" italics="true" pageId="4" pageNumber="286">B. filiformis</emphasis>
</taxonomicName>
(C–C’’’),
<taxonomicName id="4BEB20B27E1DFF9FFDBAF9A7FB86F9E3" baseAuthorityName="Shui &amp; Chen" baseAuthorityYear="2005" box="[594,720,1610,1632]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="286" phylum="Tracheophyta" rank="species" species="longistyla">
<emphasis id="BE9F87237E1DFF9FFDBAF9A7FB86F9E3" bold="true" box="[594,720,1610,1632]" italics="true" pageId="4" pageNumber="286">B. longistyla</emphasis>
</taxonomicName>
(D–D’’’),
<taxonomicName id="4BEB20B27E1DFF9FFCD0F9A7FAE9F9E3" box="[824,959,1610,1632]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="4" pageNumber="286" phylum="Tracheophyta" rank="species" species="scabrifolia">
<emphasis id="BE9F87237E1DFF9FFCD0F9A7FAE9F9E3" bold="true" box="[824,959,1610,1632]" italics="true" pageId="4" pageNumber="286">B. scabrifolia</emphasis>
</taxonomicName>
(E–E’’’). A, B, C, D, E. Habit; A’, B’, C’, D’, E’. Leaf, adaxial surfaces; A’’, B’’, C’’, D’’, E’’. Portion of leaf adaxial surfaces; A’’’, B’’’, C’’’, D’’’, E’’’. Flowers. A–A’’’ from
<emphasis id="BE9F87237E1DFF9FFAD7F982F9EEF924" italics="true" pageId="4" pageNumber="286">Hoang Son 4435</emphasis>
and C–C’’’ from
<emphasis id="BE9F87237E1DFF9FFE8AF97EFB53F924" box="[354,517,1682,1704]" italics="true" pageId="4" pageNumber="286">Hoang Son 6433</emphasis>
by T.S. Hoang; B–B’’’ from
<emphasis id="BE9F87237E1DFF9FFCF6F97EFAC4F924" box="[798,914,1682,1704]" italics="true" pageId="4" pageNumber="286">Peng 18752</emphasis>
, D–D’’’ from
<emphasis id="BE9F87237E1DFF9FFBF4F97EFDC6F924" box="[1052,1168,1682,1704]" italics="true" pageId="4" pageNumber="286">Peng 20435</emphasis>
and E–E’’’ from
<emphasis id="BE9F87237E1DFF9FFADFF97EFCFDF924" box="[1335,1451,1682,1704]" italics="true" pageId="4" pageNumber="286">Peng 22197</emphasis>
by C.-I Peng.
</paragraph>
</caption>
<subSubSection id="C4F108BA7E1DFF9FFF55F917FCD5F8DF" pageId="4" pageNumber="286" type="distribution">
<paragraph id="8C545B317E1DFF9FFF55F917FCD5F8DF" blockId="4.[136,1451,1786,2065]" pageId="4" pageNumber="286">
<emphasis id="BE9F87237E1DFF9FFF55F917F8BDF896" bold="true" box="[189,491,1786,1813]" pageId="4" pageNumber="286">Distribution and ecology:</emphasis>
—Currently only known from the
<typeStatus id="5350E5937E1DFF9FFC81F916FACFF896" box="[873,921,1787,1813]" pageId="4" pageNumber="286">type</typeStatus>
locality in Thuong Hoa commune. The Thuong Hoa commune borders the Nakai-Nam Theun National Park in
<collectingCountry id="F4FC1BA17E1DFF9FFCBFF8F2FAC6F8BA" box="[855,912,1823,1849]" name="Laos" pageId="4" pageNumber="286">Laos</collectingCountry>
, therefore, it is possible that this species occurs there also. The species grows on limestone mountains under secondary forests at 495–510 meters above sea level.
</paragraph>
</subSubSection>
<subSubSection id="C4F108BA7E1DFF9FFF55F88AFC15F803" box="[189,1347,1894,1921]" pageId="4" pageNumber="286" type="etymology">
<paragraph id="8C545B317E1DFF9FFF55F88AFC15F803" blockId="4.[136,1451,1786,2065]" box="[189,1347,1894,1921]" pageId="4" pageNumber="286">
<emphasis id="BE9F87237E1DFF9FFF55F88AF81EF802" bold="true" box="[189,328,1895,1921]" pageId="4" pageNumber="286">Etymology:</emphasis>
—The epithet is in honor of Mr. Bui Tien Dat, who is the first to discover this new species.
</paragraph>
</subSubSection>
<subSubSection id="C4F108BA7E1DFF9FFF55F867F99EF792" pageId="4" pageNumber="286" type="conservation">
<paragraph id="8C545B317E1DFF9FFF55F867F99EF792" blockId="4.[136,1451,1786,2065]" pageId="4" pageNumber="286">
<emphasis id="BE9F87237E1DFF9FFF55F867F831F827" bold="true" box="[189,359,1930,1956]" pageId="4" pageNumber="286">Conservation:</emphasis>
—There were only four populations found at the site of collection with a total of less than 200 mature individuals recorded, while the nearby habitats are remote and virtually uninhabited. For the reason that the populations have restricted distribution, it may be considered as Near Threatened species, according to the IUCN 2022.
</paragraph>
</subSubSection>
<subSubSection id="C4F108BA7E1CFF9EFF55FF2FFD19FD33" pageId="5" pageNumber="287" type="discussion">
<paragraph id="8C545B317E1CFF9EFF55FF2FF9A4FE37" blockId="5.[136,1452,194,688]" pageId="5" pageNumber="287">
<emphasis id="BE9F87237E1CFF9EFF55FF2FF85FFF5F" bold="true" box="[189,265,194,220]" pageId="5" pageNumber="287">Notes:</emphasis>
—Within
<taxonomicName id="4BEB20B27E1CFF9EFE92FF2EFBEEFF5F" box="[378,696,194,220]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="287" phylum="Tracheophyta" rank="section" section="Coelocentrum">
<emphasis id="BE9F87237E1CFF9EFE92FF2EF881FF5F" box="[378,471,195,220]" italics="true" pageId="5" pageNumber="287">Begonia</emphasis>
sect.
<emphasis id="BE9F87237E1CFF9EFDF1FF2FFBEEFF5F" box="[537,696,194,220]" italics="true" pageId="5" pageNumber="287">Coelocentrum</emphasis>
</taxonomicName>
, the number of described species has increased by more than
<quantity id="4B13F6D47E1CFF9EFA9AFF2FFCFAFF5F" box="[1394,1452,194,220]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="287" unit="in" value="20.0">20 in</quantity>
the last decade, mostly from disoveries concentrated in southwestern
<collectingCountry id="F4FC1BA17E1CFF9EFC7FFF0BFA8CFE83" box="[919,986,230,256]" name="China" pageId="5" pageNumber="287">China</collectingCountry>
and northern
<collectingCountry id="F4FC1BA17E1CFF9EFB90FF0BFD8DFE83" box="[1144,1243,230,256]" name="Vietnam" pageId="5" pageNumber="287">Vietnam</collectingCountry>
. There are now 22 species in the section known from
<collectingCountry id="F4FC1BA17E1CFF9EFDF8FEE7FB24FEA7" box="[528,626,266,292]" name="Vietnam" pageId="5" pageNumber="287">Vietnam</collectingCountry>
, however, only two of these occur in the central region of the country:
<taxonomicName id="4BEB20B27E1CFF9EFA7BFEE6F8A8FECA" authority="(Peng et al., 2015)" baseAuthorityName="Peng" baseAuthorityYear="2015" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="287" phylum="Tracheophyta" rank="species" species="montaniformis">
<emphasis id="BE9F87237E1CFF9EFA7BFEE6F87DFECB" italics="true" pageId="5" pageNumber="287">B. montaniformis</emphasis>
(
<bibRefCitation id="E87A26C07E1CFF9EFED1FEC3F8A0FECA" author="Peng, C. - I. &amp; Lin, C. W. &amp; Yang, H. A. &amp; Kono, Y. &amp; Nguyen, H. Q." box="[313,502,302,329]" pageId="5" pageNumber="287" pagination="9" refId="ref3469" refString="Peng, C. - I., Lin, C. W., Yang, H. A., Kono, Y. &amp; Nguyen, H. Q. (2015) Six new species of Begonia (Begoniaceae) from limestone areas in Northern Vietnam. Botanical Studies 56: e 9. https: // doi. org / 10.1186 / s 40529 - 015 - 0089 - 3" type="journal article" year="2015">
Peng
<emphasis id="BE9F87237E1CFF9EFE90FEC2F8E6FECB" box="[376,432,302,328]" italics="true" pageId="5" pageNumber="287">et al.</emphasis>
, 2015
</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="14D047B47E1CFF9EFDE4FEC3FB1DFECB" box="[524,587,302,328]" captionStart="FIGURE 1" captionStartId="1.[136,234,984,1006]" captionTargetBox="[268,1320,190,960]" captionTargetId="figure-20@1.[268,1320,190,960]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of Begonia datii (red star), B. bamaensis (orange hexagon), B. filiformis (blue circles), B. longistyla (pink diamond), B. montaniformis (yellow square), B. morsei Irmsch (green triangle) and B. morsei var. myriotricha Y.M.Shui &amp; W.H.Chen (purple inverted triangle) in China and Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8297128" httpUri="https://zenodo.org/record/8297128/files/figure.png" pageId="5" pageNumber="287">Fig. 1</figureCitation>
) and
<taxonomicName id="4BEB20B27E1CFF9EFD62FEC2FA82FECB" authority="(Leveille, 1909)" baseAuthorityName="Leveille" baseAuthorityYear="1909" box="[650,980,302,328]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="287" phylum="Tracheophyta" rank="species" species="cavaleriei">
<emphasis id="BE9F87237E1CFF9EFD62FEC2FA4EFECB" box="[650,792,302,328]" italics="true" pageId="5" pageNumber="287">B. cavaleriei</emphasis>
(
<bibRefCitation id="E87A26C07E1CFF9EFCCEFEC3FA9AFECB" author="Leveille, H." box="[806,972,302,328]" pageId="5" pageNumber="287" pagination="20 - 23" refId="ref2984" refString="Leveille, H. (1909) Decades plantarum novarum XVI, in Fedde. R epertorium Specierum No-varum Regni Vegetabilis 7 (1 - 3): 20 - 23. https: // doi. org / 10.1002 / fedr. 19090070108" type="journal article" year="1909">Léveillé, 1909</bibRefCitation>
)
</taxonomicName>
. The newly discovered
<taxonomicName id="4BEB20B27E1CFF9EFB31FEC2FC39FECB" authorityName="T. S. Hoang &amp; C. W. Lin. A. Habit" authorityYear="2023" box="[1241,1391,302,328]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="287" phylum="Tracheophyta" rank="species" species="datii">
<emphasis id="BE9F87237E1CFF9EFB31FEC2FC39FECB" box="[1241,1391,302,328]" italics="true" pageId="5" pageNumber="287">Begonia datii</emphasis>
</taxonomicName>
from
<collectingRegion id="4E2F95D37E1CFF9EFF60FEBFF82DFEEF" box="[136,379,338,364]" country="Vietnam" name="Quang Binh" pageId="5" pageNumber="287">Quang Binh Province</collectingRegion>
is the third record of the section from Central
<collectingCountry id="F4FC1BA17E1CFF9EFC68FEBFFAB5FEEF" box="[896,995,338,364]" name="Vietnam" pageId="5" pageNumber="287">Vietnam</collectingCountry>
. Although Central
<collectingCountry id="F4FC1BA17E1CFF9EFB5CFEBFFC45FEEF" box="[1204,1299,338,364]" name="Vietnam" pageId="5" pageNumber="287">Vietnam</collectingCountry>
is at the edge of the range of the section, the low number of species in the area could also be an artifact of it being botanically poorly explored.
</paragraph>
<paragraph id="8C545B317E1CFF9EFF55FE53FD19FD33" blockId="5.[136,1452,194,688]" pageId="5" pageNumber="287">
Morphologically,
<taxonomicName id="4BEB20B27E1CFF9EFE6DFE52F883FE5B" authorityName="T. S. Hoang &amp; C. W. Lin. A. Habit" authorityYear="2023" box="[389,469,446,472]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="287" phylum="Tracheophyta" rank="species" species="datii">
<emphasis id="BE9F87237E1CFF9EFE6DFE52F883FE5B" box="[389,469,446,472]" italics="true" pageId="5" pageNumber="287">B. datii</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4BEB20B27E1CFF9EFD77FE52FAB2FE5B" authority="(Irmscher, 1939)" baseAuthorityName="Irmscher" baseAuthorityYear="1939" box="[671,996,446,472]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="287" phylum="Tracheophyta" rank="species" species="filiformis">
<emphasis id="BE9F87237E1CFF9EFD77FE52FA72FE5B" box="[671,804,446,472]" italics="true" pageId="5" pageNumber="287">B. filiformis</emphasis>
(
<bibRefCitation id="E87A26C07E1CFF9EFCD9FE53FA8DFE5B" author="Irmscher, E." box="[817,987,446,472]" pageId="5" pageNumber="287" pagination="431 - 557" refId="ref2892" refString="Irmscher, E. (1939) Die Begoniaceen Chinas und ihre Bedeutung fur die Frage der Formbildung in polymorphen Sippen. Mitteilungen aus dem Institut fur allgemeine Botanik in Hamburg 10: 431 - 557." type="journal article" year="1939">Irmscher, 1939</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="14D047B47E1CFF9EFC1AFE53FD1AFE5B" box="[1010,1100,446,472]" captionStart="FIGURE 4" captionStartId="4.[136,234,1574,1596]" captionTargetBox="[151,1435,190,1550]" captionTargetId="figure-17@4.[151,1435,190,1550]" captionTargetPageId="4" captionText="FIGURE 4. Morphological comparison of Begonia datii and allied taxa, showing habits and morphological characters: B. datii (A–A’’’), B. bamaensis (B–B’’’), B. filiformis (C–C’’’), B. longistyla (D–D’’’), B. scabrifolia (E–E’’’). A, B, C, D, E. Habit; A’, B’, C’, D’, E’. Leaf, adaxial surfaces; A’’, B’’, C’’, D’’, E’’. Portion of leaf adaxial surfaces; A’’’, B’’’, C’’’, D’’’, E’’’. Flowers. A–A’’’ from Hoang Son 4435 and C–C’’’ from Hoang Son 6433 by T.S. Hoang; B–B’’’ from Peng 18752, D–D’’’ from Peng 20435 and E–E’’’ from Peng 22197 by C.-I Peng." figureDoi="http://doi.org/10.5281/zenodo.8297137" httpUri="https://zenodo.org/record/8297137/files/figure.png" pageId="5" pageNumber="287">Fig. 4 C</figureCitation>
–C’’’) from southwestern
<collectingCountry id="F4FC1BA17E1CFF9EFA80FE53FCFDFE5B" box="[1384,1451,446,472]" name="China" pageId="5" pageNumber="287">China</collectingCountry>
to northern
<collectingCountry id="F4FC1BA17E1CFF9EFEE3FE0FF83CFE7F" box="[267,362,482,508]" name="Vietnam" pageId="5" pageNumber="287">Vietnam</collectingCountry>
(
<figureCitation id="14D047B47E1CFF9EFE92FE0FF8ECFE7F" box="[378,442,482,508]" captionStart="FIGURE 1" captionStartId="1.[136,234,984,1006]" captionTargetBox="[268,1320,190,960]" captionTargetId="figure-20@1.[268,1320,190,960]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of Begonia datii (red star), B. bamaensis (orange hexagon), B. filiformis (blue circles), B. longistyla (pink diamond), B. montaniformis (yellow square), B. morsei Irmsch (green triangle) and B. morsei var. myriotricha Y.M.Shui &amp; W.H.Chen (purple inverted triangle) in China and Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8297128" httpUri="https://zenodo.org/record/8297128/files/figure.png" pageId="5" pageNumber="287">Fig. 1</figureCitation>
) in its robust rhizome with congested internodes, widely ovate leaves that are rugose and variegated; a detailed comparison of characters is presented in
<tableCitation id="C1696E8A7E1CFF9EFCD7FDEBFAC4FDA3" box="[831,914,518,544]" captionStart="TABLE 1" captionStartId="5.[136,224,729,753]" captionTargetPageId="5" captionText="TABLE 1. Comparison of Begonia datii, B. bamaensis, B. filiformis, B. longistyla, B. morsei., B. morsei var. myriotricha and B. scabrifolia." httpUri="http://table.plazi.org/id/D8940BB97E1CFF9EFF60FD34F818FC96" pageId="5" pageNumber="287" tableUuid="D8940BB97E1CFF9EFF60FD34F818FC96">Table 1</tableCitation>
. Moreover,
<taxonomicName id="4BEB20B27E1CFF9EFBF1FDEAFC26FDA3" authority="(Liu et al., 2007)" baseAuthorityName="Liu" baseAuthorityYear="2007" box="[1049,1392,518,544]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="287" phylum="Tracheophyta" rank="species" species="bamaensis">
<emphasis id="BE9F87237E1CFF9EFBF1FDEAFDFBFDA3" box="[1049,1197,518,544]" italics="true" pageId="5" pageNumber="287">B. bamaensis</emphasis>
(
<bibRefCitation id="E87A26C07E1CFF9EFB53FDEBFC3EFDA3" author="Liu, Y. &amp; Ku, S. M. &amp; Peng, C. - I." box="[1211,1384,518,544]" pageId="5" pageNumber="287" pagination="465 - 473" refId="ref3028" refString="Liu, Y., Ku, S. M. &amp; Peng, C. - I. (2007) Begonia bamaensis (sect. Coelocentrum, Begoniaceae), a new species from limestone areas in Guangxi, China. Botanical Studies 48: 465 - 473." type="journal article" year="2007">
Liu
<emphasis id="BE9F87237E1CFF9EFB02FDEAFC77FDA3" box="[1258,1313,518,544]" italics="true" pageId="5" pageNumber="287">et al.</emphasis>
, 2007
</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="14D047B47E1CFF9EFA96FDEBF9E4FDC7" captionStart="FIGURE 4" captionStartId="4.[136,234,1574,1596]" captionTargetBox="[151,1435,190,1550]" captionTargetId="figure-17@4.[151,1435,190,1550]" captionTargetPageId="4" captionText="FIGURE 4. Morphological comparison of Begonia datii and allied taxa, showing habits and morphological characters: B. datii (A–A’’’), B. bamaensis (B–B’’’), B. filiformis (C–C’’’), B. longistyla (D–D’’’), B. scabrifolia (E–E’’’). A, B, C, D, E. Habit; A’, B’, C’, D’, E’. Leaf, adaxial surfaces; A’’, B’’, C’’, D’’, E’’. Portion of leaf adaxial surfaces; A’’’, B’’’, C’’’, D’’’, E’’’. Flowers. A–A’’’ from Hoang Son 4435 and C–C’’’ from Hoang Son 6433 by T.S. Hoang; B–B’’’ from Peng 18752, D–D’’’ from Peng 20435 and E–E’’’ from Peng 22197 by C.-I Peng." figureDoi="http://doi.org/10.5281/zenodo.8297137" httpUri="https://zenodo.org/record/8297137/files/figure.png" pageId="5" pageNumber="287">Fig. 4 B</figureCitation>
–B’’’),
<taxonomicName id="4BEB20B27E1CFF9EFEEAFDC6FBD1FDC6" authority="(Shui &amp; Chen, 2005)" baseAuthorityName="Shui &amp; Chen" baseAuthorityYear="2005" box="[258,647,554,581]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="287" phylum="Tracheophyta" rank="species" species="longistyla">
<emphasis id="BE9F87237E1CFF9EFEEAFDC6F8C6FDC7" box="[258,400,554,580]" italics="true" pageId="5" pageNumber="287">B. longistyla</emphasis>
(
<bibRefCitation id="E87A26C07E1CFF9EFE48FDC7FB29FDC6" author="Shui, Y. M. &amp; Chen, W. H." box="[416,639,554,581]" pageId="5" pageNumber="287" pagination="355 - 374" refId="ref3684" refString="Shui, Y. M. &amp; Chen, W. H. (2005) New Data of Sect. Coelocentrum (Begonia) in Begoniaceae. Acta Botanica Yunnanica 27 (4): 355 - 374." type="journal article" year="2005">Shui &amp; Chen, 2005</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="14D047B47E1CFF9EFD7FFDC7FBAFFDC7" box="[663,761,554,580]" captionStart="FIGURE 4" captionStartId="4.[136,234,1574,1596]" captionTargetBox="[151,1435,190,1550]" captionTargetId="figure-17@4.[151,1435,190,1550]" captionTargetPageId="4" captionText="FIGURE 4. Morphological comparison of Begonia datii and allied taxa, showing habits and morphological characters: B. datii (A–A’’’), B. bamaensis (B–B’’’), B. filiformis (C–C’’’), B. longistyla (D–D’’’), B. scabrifolia (E–E’’’). A, B, C, D, E. Habit; A’, B’, C’, D’, E’. Leaf, adaxial surfaces; A’’, B’’, C’’, D’’, E’’. Portion of leaf adaxial surfaces; A’’’, B’’’, C’’’, D’’’, E’’’. Flowers. A–A’’’ from Hoang Son 4435 and C–C’’’ from Hoang Son 6433 by T.S. Hoang; B–B’’’ from Peng 18752, D–D’’’ from Peng 20435 and E–E’’’ from Peng 22197 by C.-I Peng." figureDoi="http://doi.org/10.5281/zenodo.8297137" httpUri="https://zenodo.org/record/8297137/files/figure.png" pageId="5" pageNumber="287">Fig. 4 D</figureCitation>
–D’’’),
<taxonomicName id="4BEB20B27E1CFF9EFCA1FDC6FC57FDC7" authority="var. morsei (Irmscher, 1939)" authorityName="var. morsei (Irmscher" authorityYear="1939" baseAuthorityName="Irmscher" baseAuthorityYear="1939" box="[841,1281,554,580]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="287" phylum="Tracheophyta" rank="variety" species="morsei" variety="morsei">
<emphasis id="BE9F87237E1CFF9EFCA1FDC6FAE3FDC7" box="[841,949,555,580]" italics="true" pageId="5" pageNumber="287">B. morsei</emphasis>
var.
<emphasis id="BE9F87237E1CFF9EFC06FDC6FD6CFDC7" box="[1006,1082,555,580]" italics="true" pageId="5" pageNumber="287">morsei</emphasis>
(
<bibRefCitation id="E87A26C07E1CFF9EFBA2FDC7FDAFFDC7" author="Irmscher, E." box="[1098,1273,554,580]" pageId="5" pageNumber="287" pagination="431 - 557" refId="ref2892" refString="Irmscher, E. (1939) Die Begoniaceen Chinas und ihre Bedeutung fur die Frage der Formbildung in polymorphen Sippen. Mitteilungen aus dem Institut fur allgemeine Botanik in Hamburg 10: 431 - 557." type="journal article" year="1939">Irmscher, 1939</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4BEB20B27E1CFF9EFAE6FDC6FB53FDEA" authority="(Shui &amp; Chen, 2005)" baseAuthorityName="Shui &amp; Chen" baseAuthorityYear="2005" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="287" phylum="Tracheophyta" rank="variety" species="morsei" variety="myriotricha">
<emphasis id="BE9F87237E1CFF9EFAE6FDC6FC2CFDC7" box="[1294,1402,555,580]" italics="true" pageId="5" pageNumber="287">B. morsei</emphasis>
var.
<emphasis id="BE9F87237E1CFF9EFF60FDA3F85AFDEB" box="[136,268,590,616]" italics="true" pageId="5" pageNumber="287">myriotricha</emphasis>
(
<bibRefCitation id="E87A26C07E1CFF9EFEF5FDA3F8A8FDEA" author="Shui, Y. M. &amp; Chen, W. H." box="[285,510,590,617]" pageId="5" pageNumber="287" pagination="355 - 374" refId="ref3684" refString="Shui, Y. M. &amp; Chen, W. H. (2005) New Data of Sect. Coelocentrum (Begonia) in Begoniaceae. Acta Botanica Yunnanica 27 (4): 355 - 374." type="journal article" year="2005">Shui &amp; Chen, 2005</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4BEB20B27E1CFF9EFDD7FDA2FAF0FDEB" authority="(Liu et al., 2020)" baseAuthorityName="Liu" baseAuthorityYear="2020" box="[575,934,590,616]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="287" phylum="Tracheophyta" rank="species" species="scabrifolia">
<emphasis id="BE9F87237E1CFF9EFDD7FDA2FB8FFDEB" box="[575,729,590,616]" italics="true" pageId="5" pageNumber="287">B. scabrifolia</emphasis>
(
<bibRefCitation id="E87A26C07E1CFF9EFD02FDA3FAC8FDEB" author="Liu, Y. &amp; Tseng, Y. H. &amp; Yang, H. A. &amp; Hu, A. Q. &amp; Xu, W. B. &amp; Lin, C. W. &amp; Kono, Y. &amp; Chang, C. C. &amp; Peng, C. - I. &amp; Chung, K. F." box="[746,926,590,616]" pageId="5" pageNumber="287" pagination="21" refId="ref3079" refString="Liu, Y., Tseng, Y. H., Yang, H. A., Hu, A. Q., Xu, W. B., Lin, C. W., Kono, Y., Chang, C. C., Peng, C. - I. &amp; Chung, K. F. (2020) Six new species of Begonia from Guangxi, China. Botanical Studies 61: 21. https: // doi. org / 10.1186 / s 40529 - 020 - 00298 - y" type="journal article" year="2020">
Liu
<emphasis id="BE9F87237E1CFF9EFCF3FDA2FA03FDEB" box="[795,853,590,616]" italics="true" pageId="5" pageNumber="287">et al.</emphasis>
, 2020
</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="14D047B47E1CFF9EFC5FFDA3FD41FDEB" box="[951,1047,590,616]" captionStart="FIGURE 4" captionStartId="4.[136,234,1574,1596]" captionTargetBox="[151,1435,190,1550]" captionTargetId="figure-17@4.[151,1435,190,1550]" captionTargetPageId="4" captionText="FIGURE 4. Morphological comparison of Begonia datii and allied taxa, showing habits and morphological characters: B. datii (A–A’’’), B. bamaensis (B–B’’’), B. filiformis (C–C’’’), B. longistyla (D–D’’’), B. scabrifolia (E–E’’’). A, B, C, D, E. Habit; A’, B’, C’, D’, E’. Leaf, adaxial surfaces; A’’, B’’, C’’, D’’, E’’. Portion of leaf adaxial surfaces; A’’’, B’’’, C’’’, D’’’, E’’’. Flowers. A–A’’’ from Hoang Son 4435 and C–C’’’ from Hoang Son 6433 by T.S. Hoang; B–B’’’ from Peng 18752, D–D’’’ from Peng 20435 and E–E’’’ from Peng 22197 by C.-I Peng." figureDoi="http://doi.org/10.5281/zenodo.8297137" httpUri="https://zenodo.org/record/8297137/files/figure.png" pageId="5" pageNumber="287">Fig. 4 E</figureCitation>
–E’’’) are similar in maculation and having rugose leaves, however,
<taxonomicName id="4BEB20B27E1CFF9EFE00FD9EFB6EFD0F" authorityName="T. S. Hoang &amp; C. W. Lin. A. Habit" authorityYear="2023" box="[488,568,626,652]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="287" phylum="Tracheophyta" rank="species" species="datii">
<emphasis id="BE9F87237E1CFF9EFE00FD9EFB6EFD0F" box="[488,568,626,652]" italics="true" pageId="5" pageNumber="287">B. datii</emphasis>
</taxonomicName>
can be distinguished from these five species by a combination of morphological characters (
<tableCitation id="C1696E8A7E1CFF9EFEE0FD7BF80AFD33" box="[264,348,662,688]" captionStart="TABLE 1" captionStartId="5.[136,224,729,753]" captionTargetPageId="5" captionText="TABLE 1. Comparison of Begonia datii, B. bamaensis, B. filiformis, B. longistyla, B. morsei., B. morsei var. myriotricha and B. scabrifolia." httpUri="http://table.plazi.org/id/D8940BB97E1CFF9EFF60FD34F818FC96" pageId="5" pageNumber="287" tableUuid="D8940BB97E1CFF9EFF60FD34F818FC96">Table 1</tableCitation>
), and the geographic distributions of those taxa are distant (
<figureCitation id="14D047B47E1CFF9EFC14FD7BFD68FD33" box="[1020,1086,662,688]" captionStart="FIGURE 1" captionStartId="1.[136,234,984,1006]" captionTargetBox="[268,1320,190,960]" captionTargetId="figure-20@1.[268,1320,190,960]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of Begonia datii (red star), B. bamaensis (orange hexagon), B. filiformis (blue circles), B. longistyla (pink diamond), B. montaniformis (yellow square), B. morsei Irmsch (green triangle) and B. morsei var. myriotricha Y.M.Shui &amp; W.H.Chen (purple inverted triangle) in China and Vietnam." figureDoi="http://doi.org/10.5281/zenodo.8297128" httpUri="https://zenodo.org/record/8297128/files/figure.png" pageId="5" pageNumber="287">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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