550 lines
70 KiB
XML
550 lines
70 KiB
XML
<document id="E20D947D433A35398960467077E090CD" ID-DOI="10.37520/aemnp.2023.014" ID-ISSN="1804-6487" ID-Zenodo-Dep="10134093" ID-ZooBank="D982BDAC-CB44-41BE-A35C-92D630F96BF7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1699897763845" checkinUser="felipe" docAuthor="Bilton, David T." docDate="2023" docId="03F6746EFFDDB624FC6CFC8BFB2AB150" docLanguage="en" docName="ActaEntMusNatPra.63.2.249-264.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 63 (2)" docSource="http://dx.doi.org/10.37520/aemnp.2023.014" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Helophorus (Ropalohelophorus) aethiops J. Balfour-Browne 1954" docType="treatment" docVersion="2" lastPageNumber="255" masterDocId="FFCF0C16FFD9B622FFE4FFB6FFE2B204" masterDocTitle="Out of the Palaearctic: the Helophorus water beetles of the Afrotropical Region (Coleoptera: Helophoridae)" masterLastPageNumber="264" masterPageNumber="249" pageNumber="253" updateTime="1700044289635" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="50291928E9BC23A72B89F5155E28933B">Out of the Palaearctic: the Helophorus water beetles of the Afrotropical Region (Coleoptera: Helophoridae)</mods:title>
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<mods:name id="E32ADF657EDDAD2FD61277EB0E5CDAAD" type="personal">
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<mods:namePart id="5E815C6F09D6073FBD9DADEDA74DC889">Bilton, David T.</mods:namePart>
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<mods:affiliation id="144A6A52C94B112C051896B0438DDA87">Marine Biology and Ecology Research Centre, School of Biological and Marine Sciences, University of Plymouth, Drake Circus Plymouth PL & Department of Zoology, University of Johannesburg, PO Box 524, Auckland Park, 2006 Johannesburg, South Africa</mods:affiliation>
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<mods:title id="71724DFAAF780B281B96E616C4E39128">Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part id="8AF8D2C2D14B59201FF75208156BF8C3">
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<mods:date id="51278963AD533DEDADB0AEEA71A983DE">2023</mods:date>
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<mods:detail id="7E3A44CDE4A2AC5091D0F65A7D5C3E4A" type="series">
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<mods:title id="359165DC1F9093FCB93B35D88D12D0F3">Acta. Ent. Mus. Natl. Pragae</mods:title>
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<mods:number id="9B64F6628E0E32CC25044B5B75241850">2023-09-10</mods:number>
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<mods:number id="0A7F8D1C2DCC6CC80663D288E407A3EE">63</mods:number>
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<mods:location id="64E974F3AB0A39F7B6927B92680629E4">
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<mods:url id="8D630EF7B2720C35278FF969071F7D94">http://dx.doi.org/10.37520/aemnp.2023.014</mods:url>
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<mods:classification id="77CB949A07B33FB41FD81B1886B4C0DD">journal article</mods:classification>
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<mods:identifier id="2431C6B9675EA16CA341AA64CB3E6FD5" type="DOI">10.37520/aemnp.2023.014</mods:identifier>
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<mods:identifier id="C459DD7248C8223E054B3C9F1A8ACF0E" type="ISSN">1804-6487</mods:identifier>
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<mods:identifier id="6EF8C176CEAF179393B79A362E425FCC" type="Zenodo-Dep">10134093</mods:identifier>
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</mods:mods>
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<treatment id="03F6746EFFDDB624FC6CFC8BFB2AB150" LSID="urn:lsid:plazi:treatment:03F6746EFFDDB624FC6CFC8BFB2AB150" httpUri="http://treatment.plazi.org/id/03F6746EFFDDB624FC6CFC8BFB2AB150" lastPageId="6" lastPageNumber="255" pageId="4" pageNumber="253">
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<subSubSection id="C34596F3FFDDB626FC6CFC8BFB14B171" pageId="4" pageNumber="253" type="nomenclature">
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<paragraph id="8BE0C578FFDDB626FC6CFC8BFB14B171" blockId="4.[904,1360,829,885]" pageId="4" pageNumber="253">
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<heading id="D0A87214FFDDB626FC6CFC8BFB14B171" centered="true" fontSize="10" level="2" pageId="4" pageNumber="253" reason="3">
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<taxonomicName id="4C5FBEFBFFDDB626FC6CFC8BFB14B171" ID-CoL="3KHLV" authority="J. Balfour-Browne, 1954" authorityName="J. Balfour-Browne" authorityYear="1954" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="aethiops" subGenus="Ropalohelophorus">
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<emphasis id="B92B196AFFDDB626FC6CFC8BFB14B171" bold="true" pageId="4" pageNumber="253">
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<emphasis id="B92B196AFFDDB626FC6CFC8BFBE8B151" bold="true" box="[904,1034,829,853]" italics="true" pageId="4" pageNumber="253">Helophorus</emphasis>
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(
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<emphasis id="B92B196AFFDDB626FBFEFC8BFB07B151" bold="true" box="[1050,1253,829,853]" italics="true" pageId="4" pageNumber="253">Ropalohelophorus</emphasis>
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)
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<emphasis id="B92B196AFFDDB626FB10FC8BFAB2B151" bold="true" box="[1268,1360,829,853]" italics="true" pageId="4" pageNumber="253">aethiops</emphasis>
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J.
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<bibRefCitation id="EFCEB889FFDDB626FC18FCE8FB14B171" author="BALFOUR-BROWNE J." box="[1020,1270,862,885]" pageId="4" pageNumber="253" pagination="96 - 114" refId="ref16244" refString="BALFOUR-BROWNE J. 1954: A preliminary contribution to the knowledge of the SouthAfrican Palpicornia (Coleoptera). (Professor J. Omer-Cooper's investigation of the inland waters of Africa.) Journal of the Entomolological Society of South Africa 17: 96 - 114." type="journal article" year="1954">Balfour-Browne, 1954</bibRefCitation>
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34596F3FFDDB626FC2DFCCAFAF6B18B" box="[969,1300,892,911]" pageId="4" pageNumber="253" type="description">
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||
<paragraph id="8BE0C578FFDDB626FC2DFCCAFAF6B18B" blockId="4.[969,1300,892,911]" box="[969,1300,892,911]" pageId="4" pageNumber="253">
|
||
(
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<figureCitation id="1364D9FDFFDDB626FC2BFCCAFBCCB18B" box="[975,1070,892,911]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="4" pageNumber="253">Figs 1B–C</figureCitation>
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,
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<figureCitation id="1364D9FDFFDDB626FBD3FCCAFB85B18B" box="[1079,1127,892,911]" captionStart="Fig" captionStartId="3.[153,184,1541,1560]" captionTargetBox="[172,1396,197,1494]" captionTargetId="figure-186@3.[157,1442,186,1522]" captionTargetPageId="3" captionText="Fig. 2. Afrotropical Helophorus, aedeagi. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiopicus, paralectotype; C – H. aethiops J. Balfour-Browne, 1954, holotype; D–F – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; G – H. cooperi Orchymont, 1947, holotype; H – H. brumopluvialis sp. nov., holotype; I–J – H. brumopluvialis sp. nov., Hopefield, South Africa; K – H. nyandaruaensis sp. nov., holotype; L – H. simiensis sp. nov., holotype. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.10134097" httpUri="https://zenodo.org/record/10134097/files/figure.png" pageId="4" pageNumber="253">2C–F</figureCitation>
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,
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<figureCitation id="1364D9FDFFDDB626FB8BFCCAFB68B18B" box="[1135,1162,892,911]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="4" pageNumber="253">3B</figureCitation>
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,
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<figureCitation id="1364D9FDFFDDB626FB77FCCAFB1BB18B" box="[1171,1273,892,911]" captionStart="Fig" captionStartId="8.[151,182,2027,2046]" captionTargetBox="[203,1385,189,1962]" captionTargetId="figure-13@8.[203,1385,189,1962]" captionTargetPageId="8" captionText="Fig. 4. SouthAfrican Helophorus, SEMs.A, C, E & G – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; B, D, F & H – H. brumopluvialis sp. nov., Hopefield, South Africa.A–B – head detail; C–D – pronota; E–F – pronotal detail, median groove to far left; G–H – left elytral striation close to apex." figureDoi="http://doi.org/10.5281/zenodo.10134101" httpUri="https://zenodo.org/record/10134101/files/figure.png" pageId="4" pageNumber="253">4A, C, E, G</figureCitation>
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,
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<figureCitation id="1364D9FDFFDDB626FAE6FCCAFAEFB18B" box="[1282,1293,892,911]" captionStart="Fig" captionStartId="14.[1132,1165,1128,1147]" captionTargetBox="[152,1098,191,1200]" captionTargetId="figure-437@14.[152,1098,191,1200]" captionTargetPageId="14" captionText="Fig. 5. Afrotropical Helophorus, known distributions.Map by https:// mapswire.com." figureDoi="http://doi.org/10.5281/zenodo.10134103" httpUri="https://zenodo.org/record/10134103/files/figure.png" pageId="4" pageNumber="253">5</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34596F3FFDDB626FCD8FC0FFB1BB1C8" box="[828,1273,953,972]" pageId="4" pageNumber="253" type="reference_group">
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<paragraph id="8BE0C578FFDDB626FCD8FC0FFB1BB1C8" blockId="4.[828,1273,953,972]" box="[828,1273,953,972]" pageId="4" pageNumber="253">
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||
<treatmentCitationGroup id="AB4FE256FFDDB626FCD8FC0FFB1BB1C8" box="[828,1273,953,972]" pageId="4" pageNumber="253">
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<treatmentCitation id="0AFEE369FFDDB626FCD8FC0FFB16B1C8" author="BALFOUR-BROWNE J." box="[828,1268,953,972]" page="105" pageId="4" pageNumber="253" year="1954">
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<taxonomicName id="4C5FBEFBFFDDB626FCD8FC0FFB16B1C8" ID-CoL="3KHLV" authority="J. Balfour-Browne, 1954: 105" authorityName="J. Balfour-Browne" authorityPageNumber="105" authorityYear="1954" box="[828,1268,953,972]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="aethiops">
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||
<emphasis id="B92B196AFFDDB626FCD8FC0FFC0CB1C8" box="[828,1006,953,972]" italics="true" pageId="4" pageNumber="253">Helophorus aethiops</emphasis>
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||
J.
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||
<bibRefCitation id="EFCEB889FFDDB626FBE2FC0FFB16B1C8" author="BALFOUR-BROWNE J." box="[1030,1268,953,972]" pageId="4" pageNumber="253" pagination="96 - 114" refId="ref16244" refString="BALFOUR-BROWNE J. 1954: A preliminary contribution to the knowledge of the SouthAfrican Palpicornia (Coleoptera). (Professor J. Omer-Cooper's investigation of the inland waters of Africa.) Journal of the Entomolological Society of South Africa 17: 96 - 114." type="journal article" year="1954">Balfour-Browne, 1954: 105</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C34596F3FFDDB627FCD8FC55FD67B585" lastPageId="5" lastPageNumber="254" pageId="4" pageNumber="253" type="materials_examined">
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||
<paragraph id="8BE0C578FFDDB626FCD8FC55FBE3B62F" blockId="4.[828,1437,995,2080]" pageId="4" pageNumber="253">
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||
<emphasis id="B92B196AFFDDB626FCD8FC55FC55B1F2" bold="true" box="[828,951,995,1014]" pageId="4" pageNumber="253">
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||
<typeStatus id="54E47BDAFFDDB626FCD8FC55FC8BB1F2" box="[828,873,995,1014]" pageId="4" pageNumber="253">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F34885E8FFDDB626FC5BFC55FBD3B1F2" box="[959,1073,995,1014]" name="South Africa" pageId="4" pageNumber="253">South Africa</collectingCountry>
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,
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<collectingRegion id="499B0B9AFFDDB626FBD8FC55FAE4B1F2" box="[1084,1286,995,1014]" country="South Africa" name="Eastern Cape" pageId="4" pageNumber="253">Eastern Cape Province</collectingRegion>
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, Albany District, Gowie’s Kloof, Makhanda (formerly Grahamstown),
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<geoCoordinate id="EE6BA3BFFFDDB626FAF9FC4BFA7EB614" box="[1309,1436,1021,1040]" degrees="33" direction="south" minutes="18" orientation="latitude" pageId="4" pageNumber="253" precision="1" seconds="58.71" value="-33.316307">33°18′58.71″S</geoCoordinate>
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<geoCoordinate id="EE6BA3BFFFDDB626FCD8FBAEFC59B62F" box="[828,955,1048,1067]" degrees="26" direction="east" minutes="31" orientation="longitude" pageId="4" pageNumber="253" precision="1" seconds="29.68" value="26.52491">26°31′29.68″E</geoCoordinate>
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,
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<specimenCount id="9D590EF1FFDDB626FC20FBAEFBE3B62F" box="[964,1025,1048,1067]" count="570" pageId="4" pageNumber="253" type="generic">
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||
<quantity id="4CA7689DFFDDB626FC20FBAEFC1FB62F" box="[964,1021,1048,1067]" metricMagnitude="2" metricUnit="m" metricValue="5.7" pageId="4" pageNumber="253" unit="m" value="570.0">570 m</quantity>
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||
.
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</specimenCount>
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</paragraph>
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<paragraph id="8BE0C578FFDDB626FCD8FB85FBF9B7F4" blockId="4.[828,1437,995,2080]" pageId="4" pageNumber="253">
|
||
<emphasis id="B92B196AFFDDB626FCD8FB85FC25B642" bold="true" box="[828,967,1075,1094]" pageId="4" pageNumber="253">Type material.</emphasis>
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||
<materialsCitation id="3B37CF25FFDDB626FC2BFB85FC6FB6B4" collectionCode="BMNH" collectorName="J. Omer Cooper" country="South Africa" determinerName="J. Balfour-Browne" location="Albany Dist." pageId="4" pageNumber="253" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54E47BDAFFDDB626FC2BFB85FBC9B642" box="[975,1067,1075,1094]" pageId="4" pageNumber="253" type="holotype">HOLOTYPE</typeStatus>
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||
: J, labelled: “J” “Type” [circular label with red margin] “
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<collectingCountry id="F34885E8FFDDB626FC3BFBFBFB9EB664" box="[991,1148,1101,1120]" name="South Africa" pageId="4" pageNumber="253">SOUTH AFRICA</collectingCountry>
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||
//
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||
<location id="8E8093A3FFDDB626FB72FBFBFAE5B664" LSID="urn:lsid:plazi:treatment:03F6746EFFDDB624FC6CFC8BFB2AB150:8E8093A3FFDDB626FB72FBFBFAE5B664" box="[1174,1287,1101,1120]" country="South Africa" name="Albany Dist." pageId="4" pageNumber="253">Albany Dist.</location>
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// Gowies Kloof // 6.viii.39.” “
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<collectorName id="26AAA0AEFFDDB626FC52FBDEFBA3B67F" box="[950,1089,1128,1147]" pageId="4" pageNumber="253">J. Omer Cooper</collectorName>
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||
// coll.” “
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<taxonomicName id="4C5FBEFBFFDDB626FB7DFBDEFB1DB67F" authorityName="Fabricius" authorityYear="1775" box="[1177,1279,1128,1147]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="genus">Helophorus</taxonomicName>
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(s.str.) // aethiops Type!/
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||
<determinerName id="7E3C29BCFFDDB626FC93FB35FBF6B692" box="[887,1044,1155,1174]" pageId="4" pageNumber="253">J. Balfour-Browne</determinerName>
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||
det.” [genus and species name and Type!Hw.] (
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||
<collectionCode id="ED4E5DBDFFDDB626FCA6FB2BFC65B6B4" box="[834,903,1181,1200]" pageId="4" pageNumber="253">BMNH</collectionCode>
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||
)
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||
</materialsCitation>
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||
.
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||
<typeStatus id="54E47BDAFFDDB626FC73FB2BFC12B6B4" box="[919,1008,1181,1200]" pageId="4" pageNumber="253" type="paratype">PARATYPES</typeStatus>
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||
:
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<specimenCount id="9D590EF1FFDDB626FC1AFB2BFBCDB6B4" box="[1022,1071,1175,1200]" count="2" pageId="4" pageNumber="253" type="female">2 ♀♀</specimenCount>
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||
, same data as
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<typeStatus id="54E47BDAFFDDB626FB51FB2BFAE0B6B4" box="[1205,1282,1181,1200]" pageId="4" pageNumber="253" type="holotype">holotype</typeStatus>
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||
; 2 JJ, “
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||
<collectingCountry id="F34885E8FFDDB626FAB7FB2BFC6BB6CF" name="South Africa" pageId="4" pageNumber="253">SOUTH AFRICA</collectingCountry>
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||
// Albany Dist. // Grahamstown. //
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<date id="FFE1E3B8FFDDB626FB53FB0EFB1BB6CF" box="[1207,1273,1208,1227]" pageId="4" pageNumber="253" value="1939-05">v. 1939</date>
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.” “J. Omer Cooper // coll.” “Brit. Mus. // 1939–625.”; 2 JJ, “
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<collectingCountry id="F34885E8FFDDB626FB59FB65FAB8B6E2" box="[1213,1370,1235,1254]" name="South Africa" pageId="4" pageNumber="253">SOUTH AFRICA</collectingCountry>
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||
// Grahamstown. // Teafontein Farm //
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<date id="FFE1E3B8FFDDB626FBB6FB5BFB55B704" box="[1106,1207,1261,1280]" pageId="4" pageNumber="253" value="1939-07-29">29.vii.1939</date>
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.” “J. Omer Cooper // coll.” “Brit. Mus. // 1939–625.”; 1 J, “
|
||
<collectingCountry id="F34885E8FFDDB626FB86FABEFB1DB71F" box="[1122,1279,1288,1307]" name="South Africa" pageId="4" pageNumber="253">SOUTH AFRICA</collectingCountry>
|
||
// Albany Dist. // Piggot Bridge // Rd. 10.ii.39.” “J. Omer Cooper // coll.” “Brit. Mus. // 1939–625.”; 1 J, “
|
||
<collectingCountry id="F34885E8FFDDB626FC3AFA8BFB9BB754" box="[990,1145,1341,1360]" name="South Africa" pageId="4" pageNumber="253">SOUTH AFRICA</collectingCountry>
|
||
// Albany Dist. // Piggot Bridge // Rd. 20.iii.39.” “J. Omer Cooper // coll.” “Brit. Mus. // 1939–625.” “
|
||
<taxonomicName id="4C5FBEFBFFDDB626FA9FFAEEFC64B782" authorityName="Fabricius" authorityYear="1775" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="genus">Helophorus</taxonomicName>
|
||
(s.str.) // aethiops
|
||
<typeStatus id="54E47BDAFFDDB626FBF8FAC5FB88B782" box="[1052,1130,1395,1414]" pageId="4" pageNumber="253" type="paratype">Paratype</typeStatus>
|
||
!/ J. Balfour-Browne det.” [genus and species name and
|
||
<typeStatus id="54E47BDAFFDDB626FC3CFA3BFBC5B7A4" box="[984,1063,1421,1440]" pageId="4" pageNumber="253" type="paratype">Paratype</typeStatus>
|
||
! Hw.]; 2 JJ, “
|
||
<collectingCountry id="F34885E8FFDDB626FB4CFA3BFA6AB7A4" box="[1192,1416,1421,1440]" name="Namibia" pageId="4" pageNumber="253">SOUTH-WEST AFRICA</collectingCountry>
|
||
. // Windhoek, Avis Dam. // 7.vii.37. // J. Omer-Cooper.” “J. Omer Cooper // coll.” “Brit. Mus. // 1939–625.” All BMNH and with circular
|
||
<typeStatus id="54E47BDAFFDDB626FAB6FA75FA7FB7D2" box="[1362,1437,1475,1494]" pageId="4" pageNumber="253" type="paratype">Paratype</typeStatus>
|
||
labels with yellow border.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDDB627FCD8FA4EFD67B585" blockId="4.[828,1437,995,2080]" lastBlockId="5.[151,761,189,1921]" lastPageId="5" lastPageNumber="254" pageId="4" pageNumber="253">
|
||
<emphasis id="B92B196AFFDDB626FCD8FA4EFBC8B40F" bold="true" box="[828,1066,1528,1547]" pageId="4" pageNumber="253">Other material examined.</emphasis>
|
||
24 JJ
|
||
<specimenCount id="9D590EF1FFDDB626FB97FA4EFB4DB40F" box="[1139,1199,1522,1547]" count="21" pageId="4" pageNumber="253" type="female">21 ♀♀</specimenCount>
|
||
, “
|
||
<date id="FFE1E3B8FFDDB626FB27FA4EFAC5B40F" box="[1219,1319,1528,1547]" pageId="4" pageNumber="253" value="2013-12-29">29/xii/2013</date>
|
||
<collectingCountry id="F34885E8FFDDB626FACAFA4EFA7EB40F" box="[1326,1436,1528,1547]" name="South Africa" pageId="4" pageNumber="253">South Africa</collectingCountry>
|
||
KZN // Temp. pond in grassland beside // R56 road ca.
|
||
<quantity id="4CA7689DFFDDB626FAF4F9A5FAA4B422" box="[1296,1350,1555,1574]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="253" unit="km" value="25.0">25 km</quantity>
|
||
ENE of // Kokstad D T Bilton leg.” (
|
||
<collectionCode id="ED4E5DBDFFDDB626FBFEF99BFBADB444" box="[1050,1103,1581,1600]" country="South Africa" name="Albany Museum" pageId="4" pageNumber="253">AMG</collectionCode>
|
||
,
|
||
<collectionCode id="ED4E5DBDFFDDB626FBB3F99BFB9DB444" box="[1111,1151,1581,1600]" pageId="4" pageNumber="253">CBP</collectionCode>
|
||
,
|
||
<collectionCode id="ED4E5DBDFFDDB626FB63F99BFB5CB444" box="[1159,1214,1581,1600]" pageId="4" pageNumber="253">ISAM</collectionCode>
|
||
,
|
||
<collectionCode id="ED4E5DBDFFDDB626FB23F99BFAE8B444" box="[1223,1290,1581,1600]" country="France" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="253">MNHN</collectionCode>
|
||
,
|
||
<collectionCode id="ED4E5DBDFFDDB626FAF6F99BFAB3B444" box="[1298,1361,1581,1600]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="253">NMPC</collectionCode>
|
||
,
|
||
<collectionCode id="ED4E5DBDFFDDB626FABDF99BFA71B444" box="[1369,1427,1581,1600]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="4" pageNumber="253">NMW</collectionCode>
|
||
); 1 J, “
|
||
<date id="FFE1E3B8FFDDB626FC8FF9FEFC32B45F" box="[875,976,1608,1627]" pageId="4" pageNumber="253" value="2013-12-29">29/xii/2013</date>
|
||
<collectingCountry id="F34885E8FFDDB626FC30F9FEFBDDB45F" box="[980,1087,1608,1627]" name="South Africa" pageId="4" pageNumber="253">South Africa</collectingCountry>
|
||
KZN // Temp. pond in grassland beside // R56 road ca.
|
||
<quantity id="4CA7689DFFDDB626FC4EF9D5FC3DB472" box="[938,991,1635,1654]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="253" unit="km" value="25.0">25 km</quantity>
|
||
ENE of // Kokstad D T Bilton leg.” “DNA voucher // IBE-DV277” (
|
||
<collectionCode id="ED4E5DBDFFDDB626FC5CF9CBFC06B494" box="[952,996,1661,1680]" pageId="4" pageNumber="253">CBP</collectionCode>
|
||
);
|
||
<specimenCount id="9D590EF1FFDDB626FC16F9CBFBEDB494" box="[1010,1039,1655,1680]" count="1" pageId="4" pageNumber="253" type="female">1♀</specimenCount>
|
||
, “
|
||
<date id="FFE1E3B8FFDDB626FBC5F9CBFB67B494" box="[1057,1157,1661,1680]" pageId="4" pageNumber="253" value="2013-12-29">29/xii/2013</date>
|
||
<collectingCountry id="F34885E8FFDDB626FB6DF9CBFB16B494" box="[1161,1268,1661,1680]" name="South Africa" pageId="4" pageNumber="253">South Africa</collectingCountry>
|
||
KZN // Temp. pond in grassland beside // R56 road ca.
|
||
<quantity id="4CA7689DFFDDB626FB80F92EFB78B4AF" box="[1124,1178,1688,1707]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="253" unit="km" value="25.0">25 km</quantity>
|
||
ENE of // Kokstad D T Bilton leg.” “DNA voucher // IBE-DV278” (
|
||
<collectionCode id="ED4E5DBDFFDDB626FB66F905FB4CB4C2" box="[1154,1198,1715,1734]" pageId="4" pageNumber="253">CBP</collectionCode>
|
||
); 7 JJ
|
||
<specimenCount id="9D590EF1FFDDB626FB10F905FACCB4C1" box="[1268,1326,1708,1734]" count="14" pageId="4" pageNumber="253" type="female">14 ♀♀</specimenCount>
|
||
, “Stn, No. // 111” “S. AFRICA: // Matatiele //
|
||
<quantity id="4CA7689DFFDDB626FBB3F97BFB75B4E4" box="[1111,1175,1741,1760]" metricMagnitude="3" metricUnit="m" metricValue="1.46304" pageId="4" pageNumber="253" unit="ft" value="4800.0">4800ft.</quantity>
|
||
,
|
||
<date id="FFE1E3B8FFDDB626FB7AF97BFB1EB4E4" box="[1182,1276,1741,1760]" pageId="4" pageNumber="253" value="1954-03-28">28.iii.1954</date>
|
||
” “Shallow, muddy // roadside ditch // very weedy, // grasses and Juncus” “J. Balfour-Browne // Brit. Mus. 1954–797” (
|
||
<collectionCode id="ED4E5DBDFFDDB626FBA7F8B5FB6BB512" box="[1091,1161,1795,1814]" pageId="4" pageNumber="253">BMNH</collectionCode>
|
||
);
|
||
<specimenCount id="9D590EF1FFDDB626FB7FF8B5FB2FB511" box="[1179,1229,1788,1814]" count="2" pageId="4" pageNumber="253" type="female">2 ♀♀</specimenCount>
|
||
, “Stn. No. // 134” “S. AFRICA: // Farm Dam,
|
||
<quantity id="4CA7689DFFDDB626FBF6F8ABFBB0B534" box="[1042,1106,1821,1840]" metricMagnitude="3" metricUnit="m" metricValue="1.2954" pageId="4" pageNumber="253" unit="ft" value="4250.0">4250ft.</quantity>
|
||
, // nr. Mooi River, //
|
||
<date id="FFE1E3B8FFDDB626FAEFF8ABFABEB534" box="[1291,1372,1821,1840]" pageId="4" pageNumber="253" value="1954-04-02">2.iv.1954</date>
|
||
” “Mud with Juncus // and Nitella” “J. Balfour-Browne // Brit. Mus. 1954–797” (
|
||
<collectionCode id="ED4E5DBDFFDDB626FCA6F8E5FC65B562" box="[834,903,1875,1894]" pageId="4" pageNumber="253">BMNH</collectionCode>
|
||
); 1 J, “Stn. No. // 84” “S. AFRICA: // E.
|
||
<collectingRegion id="499B0B9AFFDDB626FB0EF8E5FA85B562" box="[1258,1383,1875,1894]" country="South Africa" name="Western Cape" pageId="4" pageNumber="253">Cape Province</collectingRegion>
|
||
// van Staaden’s Pass, //
|
||
<date id="FFE1E3B8FFDDB626FC3EF8DBFBDAB584" box="[986,1080,1901,1920]" pageId="4" pageNumber="253" value="1954-03-21">21.iii.1954</date>
|
||
” “In gravel at edge // of fast stream” “J. Balfour-Browne // Brit. Mus. 1954–797” “
|
||
<taxonomicName id="4C5FBEFBFFDDB626FB4CF83EFA7EB59F" authorityName="J. Balfour-Browne" authorityYear="1954" box="[1192,1436,1928,1947]" class="Insecta" family="Helophoridae" genus="Helophorus" isSensuStricto="true" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="253" phylum="Arthropoda" rank="species" species="aethiops">Helophorus (s. str.) aethiops</taxonomicName>
|
||
J. B-B // J. Balfour-Browne det. //
|
||
<date id="FFE1E3B8FFDDB626FB82F815FB45B5B2" box="[1126,1191,1955,1974]" pageId="4" pageNumber="253" value="1975-05">v. 1975</date>
|
||
” (
|
||
<collectionCode id="ED4E5DBDFFDDB626FB5FF815FAE2B5B2" box="[1211,1280,1955,1974]" pageId="4" pageNumber="253">BMNH</collectionCode>
|
||
);
|
||
<specimenCount id="9D590EF1FFDDB626FAEBF815FACDB5B1" box="[1295,1327,1948,1974]" count="1" pageId="4" pageNumber="253" type="female">1 ♀</specimenCount>
|
||
, “Stn. No. // 160” “S. SAFRICA: // O.F.S., nr. // Harrismith, //
|
||
<quantity id="4CA7689DFFDDB626FB1EF80BFAD8B5D4" box="[1274,1338,1981,2000]" metricMagnitude="3" metricUnit="m" metricValue="1.6764" pageId="4" pageNumber="253" unit="ft" value="5500.0">5500ft.</quantity>
|
||
,
|
||
<date id="FFE1E3B8FFDDB626FAA0F80BFA76B5D4" box="[1348,1428,1981,2000]" pageId="4" pageNumber="253" value="1954-04-07">7.iv.1954</date>
|
||
” “Farm dam, // very grassy.” “J. Balfour-Browne // Brit. Mus. 1954–797” (
|
||
<collectionCode id="ED4E5DBDFFDDB626FCA6F845FC65BA02" box="[834,903,2035,2054]" pageId="4" pageNumber="253">BMNH</collectionCode>
|
||
); 2 JJ, 8 exx., “S. AFRICA:// Riversdale Dist. // nr.Albertinia //
|
||
<date id="FFE1E3B8FFDDB626FCAAF7BBFC47BA24" box="[846,933,2061,2080]" pageId="4" pageNumber="253" value="1947-02-16">16.ii/1947</date>
|
||
// J. Omer-Cooper” “Brit. Mus. // 1978-162” (
|
||
<collectionCode id="ED4E5DBDFFDDB626FAD9F7BBFA60BA24" box="[1341,1410,2061,2080]" pageId="4" pageNumber="253">BMNH</collectionCode>
|
||
); 1 J
|
||
<specimenCount id="9D590EF1FFDCB627FF49FF75FF2FB2D2" box="[173,205,189,214]" count="1" pageId="5" pageNumber="254" type="female">1 ♀</specimenCount>
|
||
, “S. AFRICA: // ca.
|
||
<quantity id="4CA7689DFFDCB627FE65FF75FE5BB2D2" box="[385,441,195,214]" metricMagnitude="4" metricUnit="m" metricValue="1.9312128" pageId="5" pageNumber="254" unit="mi" value="12.0">12 mi.</quantity>
|
||
from Queenstown // on road to Lady Frere, // ca.
|
||
<quantity id="4CA7689DFFDCB627FEE6FF68FEA0B2F5" box="[258,322,222,241]" metricMagnitude="3" metricUnit="m" metricValue="1.15824" pageId="5" pageNumber="254" unit="ft" value="3800.0">3800ft.</quantity>
|
||
,
|
||
<date id="FFE1E3B8FFDCB627FEAFFF68FE4BB2F5" box="[331,425,222,241]" pageId="5" pageNumber="254" value="1954-03-26">26.iii.1954</date>
|
||
” “in small // grassy dam” “J. Balfour- -Browne // Brit. Mus. 1954–797” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FE5BFF4FFDE6B308" box="[447,516,249,268]" pageId="5" pageNumber="254">BMNH</collectionCode>
|
||
); 1 J
|
||
<specimenCount id="9D590EF1FFDCB627FDD3FF4FFD84B30F" box="[567,614,242,268]" count="4" pageId="5" pageNumber="254" type="female">4 ♀♀</specimenCount>
|
||
, “TRANSVAAL // Middleburg Dist. // Middleburg //
|
||
<date id="FFE1E3B8FFDCB627FE32FEA5FDD1B322" box="[470,563,275,294]" pageId="5" pageNumber="254" value="1948-11-29">29.xi.1948</date>
|
||
” “
|
||
<collectingCountry id="F34885E8FFDCB627FDADFEA5FD07B322" box="[585,741,275,294]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
// J. Omer-Cooper” “J. Omer-Cooper // B.M. 1981–272” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FD95FE98FD54B345" box="[625,694,302,321]" pageId="5" pageNumber="254">BMNH</collectionCode>
|
||
); 2 JJ, 5 exx., “Dry stream // with pools” “TRANSVAAL // Waterburg Dist. // Potgietersrust. //? Matini R. //
|
||
<date id="FFE1E3B8FFDCB627FE5BFED5FDFEB372" box="[447,540,355,374]" pageId="5" pageNumber="254" value="1948-11-25">25.xi.1948</date>
|
||
” “
|
||
<collectingCountry id="F34885E8FFDCB627FDD7FED5FD2DB372" box="[563,719,355,374]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
// J. Omer-Cooper” “J. Omer-Cooper // B.M. 1981–272” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FD88FEC8FD53B395" box="[620,689,382,401]" pageId="5" pageNumber="254">BMNH</collectionCode>
|
||
);
|
||
<specimenCount id="9D590EF1FFDCB627FD25FEC8FD10B395" box="[705,754,376,401]" count="2" pageId="5" pageNumber="254" type="female">2 ♀♀</specimenCount>
|
||
, “TRANSVAAL // Pietersburg Dist. // Bandolier Kop //
|
||
<date id="FFE1E3B8FFDCB627FD75FE2FFD0DB3A8" box="[657,751,409,428]" pageId="5" pageNumber="254" value="1948-11-23">23.xi.1948</date>
|
||
” “
|
||
<collectingCountry id="F34885E8FFDCB627FF7BFE05FEA6B3C2" box="[159,324,435,454]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
// J. Omer-Cooper” “J. Omer-Cooper // B.M. 1981–272” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FEE4FE78FEA7B3E5" box="[256,325,462,481]" pageId="5" pageNumber="254">BMNH</collectionCode>
|
||
);
|
||
<specimenCount id="9D590EF1FFDCB627FEB7FE78FE62B3E5" box="[339,384,456,481]" count="3" pageId="5" pageNumber="254" type="female">3 ♀♀</specimenCount>
|
||
, “TRANSVAAL // Pietersburg Dist. // Dam, 18 mls South // of Pietersburg //
|
||
<date id="FFE1E3B8FFDCB627FE5AFE5FFDF8B3F8" box="[446,538,489,508]" pageId="5" pageNumber="254" value="1948-11-25">25.xi.1948</date>
|
||
” “
|
||
<collectingCountry id="F34885E8FFDCB627FDD6FE5FFD2DB3F8" box="[562,719,489,508]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
// J. Omer-Cooper” “J. Omer-Cooper // B.M. 1981–272” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FD88FDB5FD53B012" box="[620,689,515,534]" pageId="5" pageNumber="254">BMNH</collectionCode>
|
||
); 2 JJ, 18 exx., “
|
||
<collectingCountry id="F34885E8FFDCB627FF0CFDA8FE22B035" box="[232,448,542,561]" name="South Africa" pageId="5" pageNumber="254">ORANGE FREE STATE</collectingCountry>
|
||
// Kroonstad Dist. // Vredefort // Nr. Honing Spruit //
|
||
<date id="FFE1E3B8FFDCB627FEC9FD8FFE74B048" box="[301,406,569,588]" pageId="5" pageNumber="254" value="1947-08-29">29.viii.1947</date>
|
||
” “
|
||
<collectingCountry id="F34885E8FFDCB627FE48FD8FFDABB048" box="[428,585,569,588]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
// J. Omer-Cooper” “J. Omer-Cooper // B.M. 1981–272” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FE08FDE5FDD3B062" box="[492,561,595,614]" pageId="5" pageNumber="254">BMNH</collectionCode>
|
||
); 1 J, 7 exx., “
|
||
<collectingRegion id="499B0B9AFFDCB627FD5BFDE5FF36B085" country="South Africa" name="Western Cape" pageId="5" pageNumber="254">CAPE PROV.</collectingRegion>
|
||
// Cathcart Dist. // Cathcart //
|
||
<date id="FFE1E3B8FFDCB627FE34FDD8FDC5B085" box="[464,551,622,641]" pageId="5" pageNumber="254" value="1950-02-20">20.ii.1950</date>
|
||
” “
|
||
<collectingCountry id="F34885E8FFDCB627FDD8FDD8FD37B085" box="[572,725,622,641]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
// J. Omer-Cooper” “J. Omer-Cooper // B.M. 1981–272” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FD8CFD3FFD4FB098" box="[616,685,649,668]" pageId="5" pageNumber="254">BMNH</collectionCode>
|
||
); 1 exx., “
|
||
<collectingRegion id="499B0B9AFFDCB627FF7BFD15FEF8B0B2" box="[159,282,675,694]" country="South Africa" name="Western Cape" pageId="5" pageNumber="254">CAPE PROV.</collectingRegion>
|
||
// Colesburg Dist. // Colesburg. Dam // outside town. //
|
||
<date id="FFE1E3B8FFDCB627FF73FD08FF0DB0D5" box="[151,239,702,721]" pageId="5" pageNumber="254" value="1947-02-22">22.ii.1947</date>
|
||
” “
|
||
<collectingCountry id="F34885E8FFDCB627FEE1FD08FE42B0D5" box="[261,416,702,721]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
// J. Omer-Cooper” “J. Omer-Cooper // B.M. 1981–272” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FECBFD6FFE91B0E8" box="[303,371,729,748]" pageId="5" pageNumber="254">BMNH</collectionCode>
|
||
); 1 exx., “
|
||
<collectingRegion id="499B0B9AFFDCB627FE27FD6FFDDEB0E8" box="[451,572,729,748]" country="South Africa" name="Western Cape" pageId="5" pageNumber="254">CAPE PROV.</collectingRegion>
|
||
// Aliwal North Dist.// Dam in mountains //
|
||
<date id="FFE1E3B8FFDCB627FEA3FD45FE47B102" box="[327,421,755,774]" pageId="5" pageNumber="254" value="1948-03-13">13.iii.1948</date>
|
||
” “
|
||
<collectingCountry id="F34885E8FFDCB627FE5DFD45FDB0B102" box="[441,594,755,774]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
// J. Omer-Cooper” “J. Omer-Cooper // B.M. 1981–272” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FE3DFCB8FDFCB125" box="[473,542,782,801]" pageId="5" pageNumber="254">BMNH</collectionCode>
|
||
); 1 exx., “TRANSVAAL. // Lydenburg Dist. // Belfast. //
|
||
<date id="FFE1E3B8FFDCB627FE53FC9FFDF5B138" box="[439,535,809,828]" pageId="5" pageNumber="254" value="1948-11-30">30/xi/1948</date>
|
||
” “
|
||
<collectingCountry id="F34885E8FFDCB627FDCAFC9FFD2FB138" box="[558,717,809,828]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
// J. Omer-Cooper” “J. Omer-Cooper // B.M. 1981–272” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FD8CFCF5FD4FB152" box="[616,685,835,854]" pageId="5" pageNumber="254">BMNH</collectionCode>
|
||
); 2 exx., “
|
||
<collectingRegion id="499B0B9AFFDCB627FF7BFCE8FEF8B175" box="[159,282,862,881]" country="South Africa" name="Western Cape" pageId="5" pageNumber="254">CAPE PROV.</collectingRegion>
|
||
// Cathcart Dist. // Cathcart //
|
||
<date id="FFE1E3B8FFDCB627FDFFFCE8FD91B175" box="[539,627,862,881]" pageId="5" pageNumber="254" value="1950-02-20">20.ii.1950</date>
|
||
” “
|
||
<collectingCountry id="F34885E8FFDCB627FD6CFCE8FF2BB188" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
// E. M. Rodgers” “J. Omer-Cooper // B.M. 1981–272” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FD4EFCCFFD0DB188" box="[682,751,889,908]" pageId="5" pageNumber="254">BMNH</collectionCode>
|
||
); 2 exx., “
|
||
<collectingRegion id="499B0B9AFFDCB627FF06FC25FEBDB1A2" box="[226,351,915,934]" country="South Africa" name="Western Cape" pageId="5" pageNumber="254">CAPE PROV.</collectingRegion>
|
||
// Queenstown Dist. // Dam nr. Sterkstroom // 25 mls. Queenstown - // Aliwal.
|
||
<date id="FFE1E3B8FFDCB627FE58FC18FDF4B1C5" box="[444,534,942,961]" pageId="5" pageNumber="254" value="1948-02-13">13.ii.1948</date>
|
||
.” “
|
||
<collectingCountry id="F34885E8FFDCB627FDD5FC18FD2CB1C5" box="[561,718,942,961]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
// J. Omer-Cooper” “J. Omer-Cooper // B.M. 1981–272” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FD8CFC7FFD4FB1D8" box="[616,685,969,988]" pageId="5" pageNumber="254">BMNH</collectionCode>
|
||
); 1 exx., “S. W. AFRICA // Okarupa, ca. 17 ml. // E. of Okahandja, //
|
||
<quantity id="4CA7689DFFDCB627FD5FFC55FD1BB1F2" box="[699,761,995,1014]" metricMagnitude="3" metricUnit="m" metricValue="1.49352" pageId="5" pageNumber="254" unit="ft" value="4900.0">4900ft.</quantity>
|
||
<date id="FFE1E3B8FFDCB627FF73FC48FF0DB615" box="[151,239,1022,1041]" pageId="5" pageNumber="254" value="1954-05-22">22.v.1954</date>
|
||
.” “Pools in overflow // stream from dam, // much weed and algae.” “J. Balfour-Browne // Brit. Mus. 1954–797” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FD8CFBAFFD4FB628" box="[616,685,1049,1068]" pageId="5" pageNumber="254">BMNH</collectionCode>
|
||
); 1 exx., “
|
||
<collectingRegion id="499B0B9AFFDCB627FF7BFB85FEFEB642" box="[159,284,1075,1094]" country="South Africa" name="Western Cape" pageId="5" pageNumber="254">CAPE PROV.</collectingRegion>
|
||
// Uniondale Dist. // Lange Kloof Mts. //
|
||
<date id="FFE1E3B8FFDCB627FD77FB85FD12B642" box="[659,752,1075,1094]" pageId="5" pageNumber="254" value="1947-11-22">22.xi.1947</date>
|
||
” [all except
|
||
<collectingRegion id="499B0B9AFFDCB627FF1DFBF8FE96B665" box="[249,372,1102,1121]" country="South Africa" name="Western Cape" pageId="5" pageNumber="254">CAPE PROV.</collectingRegion>
|
||
Hw] “
|
||
<collectingCountry id="F34885E8FFDCB627FE4AFBF8FDA8B665" box="[430,586,1102,1121]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
// J. Omer-Cooper” “J. Omer-Cooper // B.M. 1981–272” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FE03FBDFFDCEB678" box="[487,556,1129,1148]" pageId="5" pageNumber="254">BMNH</collectionCode>
|
||
); 1 J, “Stn. No. // 78” [78 Hw.] “S. AFRICA // Witte Elbosch, // Groote River, //
|
||
<date id="FFE1E3B8FFDCB627FD76FB35FD12B692" box="[658,752,1155,1174]" pageId="5" pageNumber="254" value="1954-03-19">19.iii.1954</date>
|
||
” “Rapid stream, // weeds and moss, // dead edges” “
|
||
<collectingCountry id="F34885E8FFDCB627FDA8FB28FD04B6B5" box="[588,742,1182,1201]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
// J. Omer-Cooper” “J. Omer-Cooper // B.M. 1981–272” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FD99FB0FFD20B6C8" box="[637,706,1209,1228]" pageId="5" pageNumber="254">BMNH</collectionCode>
|
||
); 1 J, “
|
||
<collectingCountry id="F34885E8FFDCB627FF7BFB65FF0CB6E2" box="[159,238,1235,1254]" name="Namibia" pageId="5" pageNumber="254">Namibia</collectingCountry>
|
||
: WINDHOEK // DIST. Krumneck 20 // Farm dam on Oanob R. // SE 2216 Dd //
|
||
<date id="FFE1E3B8FFDCB627FEA1FB58FE43B705" box="[325,417,1262,1281]" pageId="5" pageNumber="254" value="1987-03-25">25.iii.1987</date>
|
||
// B.A. Curtis // SMI 3210” “Namibian National // Insect Collection, // National Museum, // OP.O. Box 1203, // Windhoek,
|
||
<collectingCountry id="F34885E8FFDCB627FEEAFA95FEBEB732" box="[270,348,1315,1334]" name="Namibia" pageId="5" pageNumber="254">Namibia</collectingCountry>
|
||
.” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FE97FA95FE5CB732" box="[371,446,1315,1334]" country="Namibia" name="National Museum of Namibia" pageId="5" pageNumber="254">NMNW</collectionCode>
|
||
); 1 J, “
|
||
<collectingCountry id="F34885E8FFDCB627FDE6FA95FD41B732" box="[514,675,1315,1334]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
: // Aliwal North, //
|
||
<date id="FFE1E3B8FFDCB627FF08FA88FE94B755" box="[236,374,1342,1361]" pageId="5" pageNumber="254" value="1954-08-18" valueMax="1954-08-25" valueMin="1954-08-18">18-25.viii.1954</date>
|
||
.” “‘in tank’ // H. Andreae” “
|
||
<taxonomicName id="4C5FBEFBFFDCB627FD98FA88FD06B755" authorityName="Fabricius" authorityYear="1775" box="[636,740,1342,1361]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="254" phylum="Arthropoda" rank="genus">Helophorus</taxonomicName>
|
||
// aethiops J. B-B. // J. Balfour-Browne det. // 1975 [name and date Hw] “SAM-COL/ AO 01926” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FE9EFAC5FE56B782" box="[378,436,1395,1414]" pageId="5" pageNumber="254">ISAM</collectionCode>
|
||
); 2 JJ, 3 exx., “
|
||
<collectingCountry id="F34885E8FFDCB627FDA0FAC5FD06B782" box="[580,740,1395,1414]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
: // Aliwal North, //
|
||
<date id="FFE1E3B8FFDCB627FEC5FA38FE4AB7A5" box="[289,424,1422,1441]" pageId="5" pageNumber="254" value="1954-08-18" valueMax="1954-08-25" valueMin="1954-08-18">18-25.viii.1954</date>
|
||
.” “‘in tank’ // H. Andreae” “
|
||
<taxonomicName id="4C5FBEFBFFDCB627FD76FA38FD1AB7A5" authorityName="Fabricius" authorityYear="1775" box="[658,760,1422,1441]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="254" phylum="Arthropoda" rank="genus">Helophorus</taxonomicName>
|
||
// aethiops // Balf.- Browne” [Hw.] “SAM-COL/ AO 01926” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FD50FA1FFD0CB7B8" box="[692,750,1449,1468]" pageId="5" pageNumber="254">ISAM</collectionCode>
|
||
); 1 J
|
||
<specimenCount id="9D590EF1FFDCB627FF24FA75FF13B7D2" box="[192,241,1469,1494]" count="3" pageId="5" pageNumber="254" type="female">3 ♀♀</specimenCount>
|
||
, “
|
||
<collectingCountry id="F34885E8FFDCB627FEE1FA75FE61B7D2" box="[261,387,1475,1494]" name="South Africa" pageId="5" pageNumber="254">SUDAFRIKA</collectingCountry>
|
||
, Natal //
|
||
<quantity id="4CA7689DFFDCB627FE32FA75FDE5B7D2" box="[470,519,1475,1494]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="5" pageNumber="254" unit="km" value="40.0">40km</quantity>
|
||
S Bethlehem //
|
||
<date id="FFE1E3B8FFDCB627FD7CFA75FD15B7D2" box="[664,759,1475,1494]" pageId="5" pageNumber="254" value="1993-12-30">30.12.1993</date>
|
||
// leg. Wewalka (10)” “1997 //
|
||
<taxonomicName id="4C5FBEFBFFDCB627FE43FA68FDE8B7F5" authorityName="J. Balfour-Browne" authorityYear="1954" box="[423,522,1502,1521]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="254" phylum="Arthropoda" rank="species" species="aethiops">H. aethiops</taxonomicName>
|
||
J B-B // R. B. Angus det.” [name and year Hw.] (
|
||
<collectionCode id="ED4E5DBDFFDCB627FE82FA4FFE40B408" box="[358,418,1529,1548]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="254">NMW</collectionCode>
|
||
); 1 J, “
|
||
<date id="FFE1E3B8FFDCB627FE09FA4FFDA6B408" box="[493,580,1529,1548]" pageId="5" pageNumber="254" value="2018-03-09">9/iii/2018</date>
|
||
<collectingCountry id="F34885E8FFDCB627FDA9FA4FFD5DB408" box="[589,703,1529,1548]" name="South Africa" pageId="5" pageNumber="254">South Africa</collectingCountry>
|
||
EC // Winterberge – seepage // below Fenella Falls // D T Bilton leg.” “DNA specimen // E.E. IBE-AN1090” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FE4AF998FE0BB445" box="[430,489,1582,1601]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="254">NMW</collectionCode>
|
||
); 1 J, “
|
||
<date id="FFE1E3B8FFDCB627FDCCF998FD65B445" box="[552,647,1582,1601]" pageId="5" pageNumber="254" value="2018-03-12">12/iii/2018</date>
|
||
<collectingCountry id="F34885E8FFDCB627FD68F998FD15B445" box="[652,759,1582,1601]" name="South Africa" pageId="5" pageNumber="254">South Africa</collectingCountry>
|
||
EC // Compassberg, small stream // above farmhouse // D T Bilton leg.” “DNA specimen // E.E. IBE-AN1092” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FE0DF9D5FDC6B472" box="[489,548,1635,1654]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="254">NMW</collectionCode>
|
||
); 1 J, “
|
||
<date id="FFE1E3B8FFDCB627FD87F9D5FD23B472" box="[611,705,1635,1654]" pageId="5" pageNumber="254" value="2018-03-11">11/iii/2018</date>
|
||
<collectingCountry id="F34885E8FFDCB627FD22F9D5FF2FB495" name="South Africa" pageId="5" pageNumber="254">South Africa</collectingCountry>
|
||
EC // Toorberg – temporary pool //
|
||
<quantity id="4CA7689DFFDCB627FDECF9C8FDD6B495" box="[520,564,1662,1681]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="254" unit="km" value="1.0">1 km</quantity>
|
||
NW of Longfontein // D T Bilton leg.” “DNA specimen // E.E. IBE-AN1095” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FD62F92FFD23B4A8" box="[646,705,1689,1708]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="5" pageNumber="254">NMW</collectionCode>
|
||
); 1 J, “
|
||
<collectingCountry id="F34885E8FFDCB627FF7BF905FED8B4C2" box="[159,314,1715,1734]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
, W.
|
||
<collectingRegion id="499B0B9AFFDCB627FEB8F905FE68B4C2" box="[348,394,1715,1734]" country="South Africa" name="Western Cape" pageId="5" pageNumber="254">Cape</collectingRegion>
|
||
,
|
||
<date id="FFE1E3B8FFDCB627FE75F905FE11B4C2" box="[401,499,1715,1734]" pageId="5" pageNumber="254" value="2003-09-04">4 Sept 2003</date>
|
||
// leg. C.R.Turner,
|
||
<geoCoordinate id="EE6BA3BFFFDCB627FD71F905FD15B4C2" box="[661,759,1715,1734]" degrees="33" direction="south" minutes="39" orientation="latitude" pageId="5" pageNumber="254" precision="15" seconds="19" value="-33.65528">33º39’19”S</geoCoordinate>
|
||
<geoCoordinate id="EE6BA3BFFFDCB627FF73F978FF1FB4E5" box="[151,253,1742,1761]" degrees="21" direction="east" minutes="01" orientation="longitude" pageId="5" pageNumber="254" precision="15" seconds="05" value="21.018055">21º01’05”E</geoCoordinate>
|
||
, alt:
|
||
<quantity id="4CA7689DFFDCB627FECCF978FEB9B4E5" box="[296,347,1742,1761]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="254" unit="m" value="310.0">310m</quantity>
|
||
, // R62 c.
|
||
<quantity id="4CA7689DFFDCB627FE4FF978FE3DB4E5" box="[427,479,1742,1761]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="254" unit="km" value="32.0">32km</quantity>
|
||
east of Ladismith, Large Bridge // with waning river slight flow with silty/fine sand, // rock based pools with sparse vegetation,// some sedges and little reeds.” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FD8DF8B5FD77B512" box="[617,661,1795,1814]" pageId="5" pageNumber="254">CBP</collectionCode>
|
||
); 1J
|
||
<specimenCount id="9D590EF1FFDCB627FD21F8B5FD12B512" box="[709,752,1789,1814]" count="2" pageId="5" pageNumber="254" type="female">2♀♀</specimenCount>
|
||
, “
|
||
<collectingCountry id="F34885E8FFDCB627FF7BF8A8FED8B535" box="[159,314,1822,1841]" name="South Africa" pageId="5" pageNumber="254">SOUTH AFRICA</collectingCountry>
|
||
, W.
|
||
<collectingRegion id="499B0B9AFFDCB627FEB8F8A8FE68B535" box="[348,394,1822,1841]" country="South Africa" name="Western Cape" pageId="5" pageNumber="254">Cape</collectingRegion>
|
||
,
|
||
<date id="FFE1E3B8FFDCB627FE75F8A8FE11B535" box="[401,499,1822,1841]" pageId="5" pageNumber="254" value="2003-09-04">4 Sept 2003</date>
|
||
// leg. C.R.Turner,
|
||
<geoCoordinate id="EE6BA3BFFFDCB627FD71F8A8FD15B535" box="[661,759,1822,1841]" degrees="33" direction="south" minutes="39" orientation="latitude" pageId="5" pageNumber="254" precision="15" seconds="19" value="-33.65528">33º39’19”S</geoCoordinate>
|
||
<geoCoordinate id="EE6BA3BFFFDCB627FF73F88FFF1FB548" box="[151,253,1849,1868]" degrees="21" direction="east" minutes="01" orientation="longitude" pageId="5" pageNumber="254" precision="15" seconds="05" value="21.018055">21º01’05”E</geoCoordinate>
|
||
, alt:
|
||
<quantity id="4CA7689DFFDCB627FEC1F88FFEBAB548" box="[293,344,1849,1868]" metricMagnitude="2" metricUnit="m" metricValue="3.1" pageId="5" pageNumber="254" unit="m" value="310.0">310m</quantity>
|
||
, // R62 c.
|
||
<quantity id="4CA7689DFFDCB627FE47F88FFE35B548" box="[419,471,1849,1868]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="254" unit="km" value="32.0">32km</quantity>
|
||
east of Ladismith, Large Bridge // with waning river slight flow with silty/fine sand, // rock based pools with sparse vegetation, // some sedges and little reeds.” (
|
||
<collectionCode id="ED4E5DBDFFDCB627FDABF8D8FD98B585" box="[591,634,1902,1921]" pageId="5" pageNumber="254">CTP</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34596F3FFDCB624FF73F82CFD5DB5F1" lastPageId="6" lastPageNumber="255" pageId="5" pageNumber="254" type="description">
|
||
<paragraph id="8BE0C578FFDCB627FF73F82CFB1BB091" blockId="5.[151,760,1945,2065]" lastBlockId="5.[828,1437,190,2069]" pageId="5" pageNumber="254">
|
||
<emphasis id="B92B196AFFDCB627FF73F82CFE9EB5B5" bold="true" box="[151,380,1945,1969]" pageId="5" pageNumber="254">
|
||
Redescription.
|
||
<emphasis id="B92B196AFFDCB627FEA0F82FFE9EB5B5" bold="true" box="[324,380,1945,1969]" italics="true" pageId="5" pageNumber="254">Male</emphasis>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1364D9FDFFDCB627FE68F82CFE3DB5B5" box="[396,479,1946,1969]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="5" pageNumber="254">Fig. 1B</figureCitation>
|
||
).
|
||
<emphasis id="B92B196AFFDCB627FE11F82CFDCDB5B5" box="[501,559,1946,1969]" italics="true" pageId="5" pageNumber="254">Head</emphasis>
|
||
black, with strong purple aeneous reflection; strongly shining. Granules effaced, most reduced to a central seta-bearing pore; more evident laterally on clypeus and immediately in front of Y-groove, where some coalesce into irregular, transverse ridges. Y-groove shallow; floor with dense, coarse, rugose punctures, shining as rest of head; stem evenly diverging anteriorly; lateral branches narrow, straight proximally then weakly arcuate laterally in front of eyes. Clypeus slightly elevated in centre; anterior margin weakly arcuate. Labrum with shining, impunctate, basal portion with anterior margin bisinuous around centre; anterior portion with coarse, rugose punctures, each puncture bearing a short, stout, whitish semi-erect seta; anterior margin arcuate with broad, shallow anteromedian emargination.Antennae 8-segmented, yellowish brown, with cupule and club segments darker. Maxillary palpi yellow, apex of terminal segment infuscated; moderately elongate, terminal segment longest, curved and clearly asymmetrical.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDCB627FCBEFD28FAD7B7F1" blockId="5.[828,1437,190,2069]" pageId="5" pageNumber="254">
|
||
<emphasis id="B92B196AFFDCB627FCBEFD28FC26B0B1" box="[858,964,670,693]" italics="true" pageId="5" pageNumber="254">Pronotum</emphasis>
|
||
(
|
||
<figureCitation id="1364D9FDFFDCB627FC31FD28FBCCB0B1" box="[981,1070,670,693]" captionStart="Fig" captionStartId="7.[151,182,1318,1337]" captionTargetBox="[244,1341,192,1289]" captionTargetId="figure-310@7.[244,1341,192,1289]" captionTargetPageId="7" captionText="Fig. 3. Afrotropical Helophorus, pronota.A – H. aethiopicus Régimbart, 1907, lectotype; B – H.aethiops J. Balfour-Browne, 1954, holotype; C – H. cooperi Orchymont, 1947, holotype; D – H. brumopluvialis sp. nov., holotype; E – H. nyandaruaensis sp. nov., holotype; F – H. simiensis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.10134099" httpUri="https://zenodo.org/record/10134099/files/figure.png" pageId="5" pageNumber="254">Figs 3B</figureCitation>
|
||
&
|
||
<figureCitation id="1364D9FDFFDCB627FBB2FD28FB74B0B1" box="[1110,1174,670,693]" captionStart="Fig" captionStartId="8.[151,182,2027,2046]" captionTargetBox="[203,1385,189,1962]" captionTargetId="figure-13@8.[203,1385,189,1962]" captionTargetPageId="8" captionText="Fig. 4. SouthAfrican Helophorus, SEMs.A, C, E & G – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; B, D, F & H – H. brumopluvialis sp. nov., Hopefield, South Africa.A–B – head detail; C–D – pronota; E–F – pronotal detail, median groove to far left; G–H – left elytral striation close to apex." figureDoi="http://doi.org/10.5281/zenodo.10134101" httpUri="https://zenodo.org/record/10134101/files/figure.png" pageId="5" pageNumber="254">4C, E</figureCitation>
|
||
) predominantly reddish brown, with strong purple aeneous reflections, particularly on internal intervals; anterior and lateral borders paler, yellowish brown; shining over entire dorsal surface. Broadest between middle and anterior margins; weakly arched in frontal and dorsal views and somewhat flattened over internal intervals, particularly anteriorly. Anterior margin almost straight over central 0.5, then sinuate to broadly rounded front angles; posterior margin broadly bisinuate around centre. Sides straight over posterior 0.4, then weakly arcuate around broadest point to anterior angles. Lateral margins setose and very weakly crenulated to serrate. Internal and middle intervals flat, very weakly to weakly granulate, strongly shining; most granules reduced to pore and seta; granules more evident on external interval, surface strongly shining and somewhat rugose. Grooves smooth and shining, with sparse, shallow punctures and small raised areas but no clear trace of granules. Median groove narrowed anteriorly and posteriorly, open at both ends more widely so posteriorly. Submedian and submarginal grooves broadly open anteriorly and posteriorly. Submedi- an grooves relatively broad, sinuate. Submarginal grooves straight internally, weakly sinuate externally, broader than submedian. Marginal grooves curved parallel to the side of pronotum, width uniform. All grooves relatively shallow, but with sides abrupt and vertical, particularly evident in submedian, submarginal and marginal grooves.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDCB627FCBEFA48FAD4B4D1" blockId="5.[828,1437,190,2069]" pageId="5" pageNumber="254">
|
||
<emphasis id="B92B196AFFDCB627FCBEFA48FC7FB411" box="[858,925,1534,1557]" italics="true" pageId="5" pageNumber="254">Elytra</emphasis>
|
||
(
|
||
<figureCitation id="1364D9FDFFDCB627FC4BFA48FBE6B411" box="[943,1028,1534,1557]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="5" pageNumber="254">Fig. 1B</figureCitation>
|
||
) brownish-yellow, with diffuse, irregular infuscation; sometimes coalescing into more or less discrete patches around suture and on disc. Shape elongate oval, bluntly rounded at apex. In lateral profile flat over anterior 0.5, then gradually descending to apex. Puncture rows weakly to moderately striate, particularly laterally; intervals weakly arched, more evident laterally.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDCB627FCBEF968FC68B531" blockId="5.[828,1437,190,2069]" pageId="5" pageNumber="254">
|
||
<emphasis id="B92B196AFFDCB627FCBEF968FC6EB4F1" box="[858,908,1758,1781]" italics="true" pageId="5" pageNumber="254">Legs</emphasis>
|
||
yellowish-brown, apex of last tarsal segment and claws infuscated. Swimming hairs distinct and clearly visible.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDCB624FCBEF888FE22B151" blockId="5.[828,1437,190,2069]" lastBlockId="6.[151,761,190,2069]" lastPageId="6" lastPageNumber="255" pageId="5" pageNumber="254">
|
||
<emphasis id="B92B196AFFDCB627FCBEF888FC7FB551" box="[858,925,1854,1877]" italics="true" pageId="5" pageNumber="254">Venter</emphasis>
|
||
black, with pronotal hypomeron, antennal pocket and pseudepipleura and epipleura orange-brown to straw yellow. Mentum shining, centre without microreticulation, with sparse, medium punctures; lateral marginal areas with weak isodiametric microreticulation centrally; margins with fine yellow setae, denser towards posterolateral angles. Submentum shining, without microreticulation; anterior half with sparse, shallow, coarse punctures and fine, hair-like vestiture. Gula and genae rugose, with dense hydrofuge vestiture. Gular sutures well marked as pits anteriorly. Vestiture only absent around margin of compound eyes and their posterolateral corners. Prosternum dull, rugose, with dense hydrofuge vestiture and sparse, long, erect setae approximately 2× length of vestiture. Pronotal hypomeron dull, with dense hydrofuge vestiture. Pronotal epipleura shining, glabrous, with sparse shallow punctures. Anterior antennal pocket well developed, with long, dense setae internally and long, curved setae at posterolateral margins. Elytral epipleura dull, glabrous, narrowing posteriorly at level of metaventrite and continuing until posterior margin of ventrite. Elytral flanks very narrowly visible from below. Ventral surface of mesothorax, metathorax and abdomen dull, with dense hydrofuge vestiture and longer, erect setae; longer setae concentrated on centre of metaventrite and abdominal ventrites 2–5; setae on ventrites arising from longitudinal tubercles, which sometimes appear as shining patches. Hind margin of abdominal ventrite 5 weakly crenulate.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDFB624FF51FCE8FF0AB631" blockId="6.[151,761,190,2069]" pageId="6" pageNumber="255">
|
||
<emphasis id="B92B196AFFDFB624FF51FCE8FEFFB171" box="[181,285,862,885]" italics="true" pageId="6" pageNumber="255">Aedeagus</emphasis>
|
||
(
|
||
<figureCitation id="1364D9FDFFDFB624FEC8FCE8FE62B171" box="[300,384,862,885]" captionStart="Fig" captionStartId="3.[153,184,1541,1560]" captionTargetBox="[172,1396,197,1494]" captionTargetId="figure-186@3.[157,1442,186,1522]" captionTargetPageId="3" captionText="Fig. 2. Afrotropical Helophorus, aedeagi. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiopicus, paralectotype; C – H. aethiops J. Balfour-Browne, 1954, holotype; D–F – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; G – H. cooperi Orchymont, 1947, holotype; H – H. brumopluvialis sp. nov., holotype; I–J – H. brumopluvialis sp. nov., Hopefield, South Africa; K – H. nyandaruaensis sp. nov., holotype; L – H. simiensis sp. nov., holotype. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.10134097" httpUri="https://zenodo.org/record/10134097/files/figure.png" pageId="6" pageNumber="255">Fig. 2C</figureCitation>
|
||
), elongate,
|
||
<quantity id="4CA7689DFFDFB624FE1EFCE8FDA9B171" box="[506,587,862,885]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="255" unit="mm" value="0.5">0.5 mm</quantity>
|
||
in length. Basal piece equal in length to parameres; outer margins of parameres weakly and evenly arcuate to apex; interior margins of parameres very weakly concave apically. Median lobe with apex weakly pointed; sides diverging posteriorly to base of struts, weakly concave; struts slightly longer than to tube.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDFB624FF51FB8BFD1BB651" blockId="6.[151,761,190,2069]" box="[181,761,1085,1109]" pageId="6" pageNumber="255">
|
||
<emphasis id="B92B196AFFDFB624FF51FB8BFEEDB651" bold="true" box="[181,271,1085,1109]" italics="true" pageId="6" pageNumber="255">Female.</emphasis>
|
||
Externally as male, but most somewhat larger.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDFB624FF51FBE8FD1BB6D1" blockId="6.[151,761,190,2069]" pageId="6" pageNumber="255">
|
||
<emphasis id="B92B196AFFDFB624FF51FBE8FEBFB671" bold="true" box="[181,349,1118,1141]" italics="true" pageId="6" pageNumber="255">Measurements.</emphasis>
|
||
Males:
|
||
<typeStatus id="54E47BDAFFDFB624FE52FBE8FDFFB671" box="[438,541,1118,1141]" pageId="6" pageNumber="255" type="holotype">Holotype</typeStatus>
|
||
: BL =
|
||
<quantity id="4CA7689DFFDFB624FD8DFBE8FD2EB671" box="[617,716,1118,1141]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="255" unit="mm" value="3.5">3.50 mm</quantity>
|
||
; EL = 2.40; EW =
|
||
<quantity id="4CA7689DFFDFB624FED6FBC8FE77B691" box="[306,405,1150,1173]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="255" unit="mm" value="1.4">1.40 mm</quantity>
|
||
. Males: BL =
|
||
<quantity id="4CA7689DFFDFB624FDCAFBC8FD2FB691" box="[558,717,1150,1173]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" metricValueMax="3.6" metricValueMin="2.6" pageId="6" pageNumber="255" unit="mm" value="3.1" valueMax="3.6" valueMin="2.6">2.60–3.60 mm</quantity>
|
||
; EL =
|
||
<quantity id="4CA7689DFFDFB624FF4BFB28FEB2B6B1" box="[175,336,1182,1205]" metricMagnitude="-3" metricUnit="m" metricValue="2.175" metricValueMax="2.45" metricValueMin="1.9" pageId="6" pageNumber="255" unit="mm" value="2.175" valueMax="2.45" valueMin="1.9">1.90–2.45 mm</quantity>
|
||
; EW =
|
||
<quantity id="4CA7689DFFDFB624FE4CFB28FDABB6B1" box="[424,585,1182,1205]" metricMagnitude="-3" metricUnit="m" metricValue="1.275" metricValueMax="1.45" metricValueMin="1.1" pageId="6" pageNumber="255" unit="mm" value="1.275" valueMax="1.45" valueMin="1.1">1.10–1.45 mm</quantity>
|
||
. Females: BL =
|
||
<quantity id="4CA7689DFFDFB624FF73FB08FED1B6D1" box="[151,307,1214,1237]" metricMagnitude="-3" metricUnit="m" metricValue="3.825" metricValueMax="4.2" metricValueMin="3.45" pageId="6" pageNumber="255" unit="mm" value="3.825" valueMax="4.2" valueMin="3.45">3.45–4.20 mm</quantity>
|
||
; EL =
|
||
<quantity id="4CA7689DFFDFB624FE97FB08FDEDB6D1" box="[371,527,1214,1237]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="6" pageNumber="255" unit="mm" value="2.5" valueMax="2.7" valueMin="2.3">2.30–2.70 mm</quantity>
|
||
; EW =
|
||
<quantity id="4CA7689DFFDFB624FDBDFB08FD17B6D1" box="[601,757,1214,1237]" metricMagnitude="-3" metricUnit="m" metricValue="1.525" metricValueMax="1.7" metricValueMin="1.35" pageId="6" pageNumber="255" unit="mm" value="1.525" valueMax="1.7" valueMin="1.35">1.35–1.70 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE0C578FFDFB624FF73FB68FD5DB5F1" blockId="6.[151,761,190,2069]" pageId="6" pageNumber="255">
|
||
<emphasis id="B92B196AFFDFB624FF73FB68FEE9B6F1" bold="true" box="[151,267,1246,1269]" pageId="6" pageNumber="255">Variation.</emphasis>
|
||
As with most
|
||
<taxonomicName id="4C5FBEFBFFDFB624FE50FB68FDD1B6F1" authorityName="Fabricius" authorityYear="1775" box="[436,563,1246,1269]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B92B196AFFDFB624FE50FB68FDD1B6F1" box="[436,563,1246,1269]" italics="true" pageId="6" pageNumber="255">Helophorus</emphasis>
|
||
</taxonomicName>
|
||
from which long series are available, there is some variation in many characters, both within and between populations. In particular, the aeneous reflections on the head and pronotum vary from purplish to greenish, with the background colour of the pronotum varying from pinkish brown to pitchy and the elytral colouration varying from relatively pale to relatively dark (see
|
||
<figureCitation id="1364D9FDFFDFB624FE8FFA08FE03B7D1" box="[363,481,1470,1493]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="6" pageNumber="255">Figs 1B–C</figureCitation>
|
||
). The degree of elytral infuscation also varies, from almost absent to dominant over the disc. (
|
||
<figureCitation id="1364D9FDFFDFB624FF19FA48FE96B411" box="[253,372,1534,1557]" captionStart="Fig" captionStartId="2.[151,182,1188,1207]" captionTargetBox="[167,1388,192,1167]" captionTargetId="figure-391@2.[151,1436,186,1175]" captionTargetPageId="2" captionText="Fig. 1. Afrotropical Helophorus, habitus. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiops J. Balfour-Browne, 1954, holotype; C – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; D – H. cooperi Orchymont, 1947, holotype; E – H. brumopluvialis sp. nov., holotype; F – H. nyandaruaensis sp. nov., holotype; G – H. simiensis sp. nov., holotype. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10134095" httpUri="https://zenodo.org/record/10134095/files/figure.png" pageId="6" pageNumber="255">Figs 1B–C</figureCitation>
|
||
). In some specimens the granulation of the pronotal intervals is more evident than others, but granules, if evident, are always weak and flat. The degree of sculpture visible in the pronotal grooves also varies somewhat, this being correlated with the degree of interval granulation evident, but grooves are always as described above in terms of their relative widths, and their abrupt, vertical margins. In the aedeagus, there is variation in the size (
|
||
<quantity id="4CA7689DFFDFB624FF2BF948FE8FB511" box="[207,365,1790,1813]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="6" pageNumber="255" unit="mm" value="0.525" valueMax="0.55" valueMin="0.5">0.50–0.55 mm</quantity>
|
||
) and in the shape of paramere apices between specimens, this variation being seen within populations when longer series are available (
|
||
<figureCitation id="1364D9FDFFDFB624FDB8F888FD33B551" box="[604,721,1854,1877]" captionStart="Fig" captionStartId="3.[153,184,1541,1560]" captionTargetBox="[172,1396,197,1494]" captionTargetId="figure-186@3.[157,1442,186,1522]" captionTargetPageId="3" captionText="Fig. 2. Afrotropical Helophorus, aedeagi. A – H. aethiopicus Régimbart, 1907, lectotype; B – H. aethiopicus, paralectotype; C – H. aethiops J. Balfour-Browne, 1954, holotype; D–F – H. aethiops J. Balfour-Browne, 1954, Kokstad, South Africa; G – H. cooperi Orchymont, 1947, holotype; H – H. brumopluvialis sp. nov., holotype; I–J – H. brumopluvialis sp. nov., Hopefield, South Africa; K – H. nyandaruaensis sp. nov., holotype; L – H. simiensis sp. nov., holotype. Scale bar = 100 μm." figureDoi="http://doi.org/10.5281/zenodo.10134097" httpUri="https://zenodo.org/record/10134097/files/figure.png" pageId="6" pageNumber="255">Figs 2C–F</figureCitation>
|
||
). In some specimens the paramere apices appear pointed, as in the
|
||
<typeStatus id="54E47BDAFFDFB624FF5BF8C8FEC2B591" box="[191,288,1918,1941]" pageId="6" pageNumber="255" type="holotype">holotype</typeStatus>
|
||
; in others they are internally produced. The basal piece varies from being equal to or slightly longer than the parameres and on the median lobe the struts vary from being equal to or slightly longer than the tube.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34596F3FFDFB624FF73F848FB2AB150" pageId="6" pageNumber="255" type="diagnosis">
|
||
<paragraph id="8BE0C578FFDFB624FF73F848FB2AB150" blockId="6.[151,761,190,2069]" lastBlockId="6.[828,1437,189,852]" pageId="6" pageNumber="255">
|
||
<emphasis id="B92B196AFFDFB624FF73F848FE6DBA11" bold="true" box="[151,399,2046,2069]" pageId="6" pageNumber="255">Differential diagnosis.</emphasis>
|
||
This species can be distinguished by its 8-segmented antennae, a pronotum with relatively weak granulation but relatively broad, sharply-defined grooves, relatively large serial punctures on the elytra, with narrow intervals and features of the aedeagus, including the shape of the paramere apices and relatively small size.
|
||
<emphasis id="B92B196AFFDFB624FCD8FEEBFBBBB370" bold="true" box="[828,1113,349,372]" pageId="6" pageNumber="255">Distribution and ecology.</emphasis>
|
||
Apparently widely distributed in summer and year-round rainfall regions of
|
||
<collectingCountry id="F34885E8FFDFB624FAF5FECBFA7AB390" box="[1297,1432,381,404]" name="South Africa" pageId="6" pageNumber="255">South Africa</collectingCountry>
|
||
, extending along the Great Escarpment into
|
||
<collectingCountry id="F34885E8FFDFB624FAE3FE2BFA81B3B0" box="[1287,1379,413,436]" name="Namibia" pageId="6" pageNumber="255">Namibia</collectingCountry>
|
||
(
|
||
<figureCitation id="1364D9FDFFDFB624FA95FE2BFCA9B3D0" captionStart="Fig" captionStartId="14.[1132,1165,1128,1147]" captionTargetBox="[152,1098,191,1200]" captionTargetId="figure-437@14.[152,1098,191,1200]" captionTargetPageId="14" captionText="Fig. 5. Afrotropical Helophorus, known distributions.Map by https:// mapswire.com." figureDoi="http://doi.org/10.5281/zenodo.10134103" httpUri="https://zenodo.org/record/10134103/files/figure.png" pageId="6" pageNumber="255">Fig. 5</figureCitation>
|
||
). Gaps between known South African and Namibian records likely reflect lack of collecting rather than genuine absences. In the southwest extends as far west as the Riversdale district in the Western Cape Province. Not to date known to overlap geographically with
|
||
<taxonomicName id="4C5FBEFBFFDFB624FB3EFD8BFA7EB050" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" box="[1242,1436,573,596]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="brumopluvialis" status="sp. nov.">
|
||
<emphasis id="B92B196AFFDFB624FB3EFD8BFA7EB050" box="[1242,1436,573,596]" italics="true" pageId="6" pageNumber="255">H. brumopluvialis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A218A411FFDFB624FCD8FDEBFC6FB070" box="[828,909,605,628]" pageId="6" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Most collections are from the austral summer, but some are from other months in areas of year-round rainfall. Characteristic of shallow, seasonally flooded waters, typically with aquatic grasses, algae and macrophytes. Cytochrome oxidase-I DNA sequence divergence between this species and
|
||
<taxonomicName id="4C5FBEFBFFDFB624FC02FD4BFB44B110" authority="Bilton, 2023" authorityName="Bilton" authorityYear="2023" box="[998,1190,765,788]" class="Insecta" family="Helophoridae" genus="Helophorus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="255" phylum="Arthropoda" rank="species" species="brumopluvialis" status="sp. nov.">
|
||
<emphasis id="B92B196AFFDFB624FC02FD4BFB44B110" box="[998,1190,765,788]" italics="true" pageId="6" pageNumber="255">H. brumopluvialis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A218A411FFDFB624FB4FFD4BFB1FB110" box="[1195,1277,765,788]" pageId="6" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is 5.5–6.0% (I. Ribera, pers. comm.), suggesting that these two species diverged during the Plio-Pleistocene.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |