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<document id="2B1479993B1D1911889E4C78B92845F3" ID-DOI="10.11646/zootaxa.5433.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10953646" ID-ZooBank="06E5A735-A276-41D7-A9EE-B09642D953B6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712750820505" checkinUser="plazi" docAuthor="Sak, Serdar, Karaytuğ, Süphan &amp; Huys, Rony" docDate="2024" docId="03EB13395D6FFFB6C9CC11469CFAFAB3" docLanguage="en" docName="zootaxa.5433.1.1.pdf" docOrigin="Zootaxa 5433 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5433.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Arenopontia problematica Masry 1970" docType="treatment" docVersion="2" lastPageNumber="9" masterDocId="FFD26B415D69FFBEC95B1029983EFFA9" masterDocTitle="A revision of the genus Arenopontia Kunz, 1937 (Copepoda, Harpacticoida, Arenopontiidae), including the description of five new species" masterLastPageNumber="50" masterPageNumber="1" pageNumber="7" updateTime="1712751723636" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="42790ACFF5E04F2D8F5EE6969C1344E6">A revision of the genus Arenopontia Kunz, 1937 (Copepoda, Harpacticoida, Arenopontiidae), including the description of five new species</mods:title>
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<mods:namePart id="C1A7C995553A119EBF648B5662EDEF82">Sak, Serdar</mods:namePart>
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<mods:namePart id="EC70A15837F1CF78A224F36DB2C29B66">Karaytuğ, Süphan</mods:namePart>
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<mods:namePart id="E1FA8AF903949D7E2E0CC70C2343F454">Huys, Rony</mods:namePart>
<mods:affiliation id="7895DB16D81699BC3103EF6565D5CB5F">Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK.</mods:affiliation>
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<treatment id="03EB13395D6FFFB6C9CC11469CFAFAB3" LSID="urn:lsid:plazi:treatment:03EB13395D6FFFB6C9CC11469CFAFAB3" httpUri="http://treatment.plazi.org/id/03EB13395D6FFFB6C9CC11469CFAFAB3" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="C358F1A45D6FFFB8C9CC11469A4BFE21" box="[151,629,366,393]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BFDA22F5D6FFFB8C9CC11469A4BFE21" blockId="6.[151,629,366,429]" box="[151,629,366,393]" pageId="6" pageNumber="7">
<heading id="D0B515435D6FFFB8C9CC11469A4BFE21" bold="true" box="[151,629,366,393]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<taxonomicName id="4C42D9AC5D6FFFB8C9CC11469A4BFE21" ID-CoL="GFQP" authority="Masry, 1970" authorityName="Masry" authorityYear="1970" box="[151,629,366,393]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="problematica">
<emphasis id="B9367E3D5D6FFFB8C9CC11469A4BFE21" bold="true" box="[151,629,366,393]" pageId="6" pageNumber="7">
<emphasis id="B9367E3D5D6FFFB8C9CC114699EAFE20" bold="true" box="[151,468,366,393]" italics="true" pageId="6" pageNumber="7">Arenopontia problematica</emphasis>
<bibRefCitation id="EFD3DFDE5D6FFFB8C88011469A4BFE21" author="Masry, D." box="[475,629,366,393]" pageId="6" pageNumber="7" pagination="229 - 258" refId="ref28101" refString="Masry, D. (1970) Ecological study of some sandy beaches along the Israeli Mediterranean coast, with a description of the interstitial harpacticoids (Crustacea, Copepoda). Cahiers de Biologie marine, 11, 229 - 258. https: // doi. org / 10.21411 / CBM. A. 4943 D 377" type="journal article" year="1970">Masry, 1970</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C358F1A45D6FFFB8C9CC11BA98D5FE04" box="[151,235,403,429]" pageId="6" pageNumber="7" type="description">
<paragraph id="8BFDA22F5D6FFFB8C9CC11BA98D5FE04" blockId="6.[151,629,366,429]" box="[151,235,403,429]" pageId="6" pageNumber="7">
(
<figureCitation id="1379BEAA5D6FFFB8C9C411BA98DEFE04" box="[159,224,403,429]" captionStart="FIGURE 1" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[196,1390,196,1899]" captionTargetId="figure-17@7.[184,1404,179,1908]" captionTargetPageId="7" captionText="FIGURE 1. Arenopontia problematica Masry, 1970 (♀): (A) P1, anterior [coxa omitted]; (B) P2, anterior [protopod omitted]; (C) P3, anterior [protopod omitted]; (D) P4, anterior [protopod largely omitted]; (E) antennule, dorsal; (F) right caudal ramus, lateral; (G) antenna [coxa omitted]." figureDoi="http://doi.org/10.5281/zenodo.10953648" httpUri="https://zenodo.org/record/10953648/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C358F1A45D6FFFB8C9CC11FE9B38FD9E" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="8BFDA22F5D6FFFB8C9CC11FE9BBDFE46" blockId="6.[151,899,471,496]" box="[151,899,471,496]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="AB5285015D6FFFB8C9CC11FE9BBDFE46" box="[151,899,471,496]" pageId="6" pageNumber="7">
<taxonomicName id="4C42D9AC5D6FFFB8C9CC11FE9A80FE46" ID-CoL="GFQP" authority="Masry, 1970" authorityName="Masry" authorityYear="1970" box="[151,702,471,496]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="problematica" subGenus="Arenopontia">
<emphasis id="B9367E3D5D6FFFB8C9CC11FE992BFE47" box="[151,277,471,494]" italics="true" pageId="6" pageNumber="7">Arenopontia</emphasis>
(
<emphasis id="B9367E3D5D6FFFB8C87F11FE999CFE47" box="[292,418,471,494]" italics="true" pageId="6" pageNumber="7">Arenopontia</emphasis>
)
<emphasis id="B9367E3D5D6FFFB8C8EB11FE9A08FE46" box="[432,566,471,495]" italics="true" pageId="6" pageNumber="7">problematica</emphasis>
<bibRefCitation id="EFD3DFDE5D6FFFB8CB6611F19A80FE46" author="Masry, D." box="[573,702,471,496]" pageId="6" pageNumber="7" pagination="229 - 258" refId="ref28101" refString="Masry, D. (1970) Ecological study of some sandy beaches along the Israeli Mediterranean coast, with a description of the interstitial harpacticoids (Crustacea, Copepoda). Cahiers de Biologie marine, 11, 229 - 258. https: // doi. org / 10.21411 / CBM. A. 4943 D 377" type="journal article" year="1970">Masry, 1970</bibRefCitation>
</taxonomicName>
:
<treatmentCitation id="0AE3843E5D6FFFB8CB9311FE9BBDFE46" author="Bodin, Ph." box="[712,899,471,495]" page="124" pageId="6" pageNumber="7" year="1979">
<bibRefCitation id="EFD3DFDE5D6FFFB8CB9311FE9BBDFE46" author="Bodin, Ph." box="[712,899,471,495]" pageId="6" pageNumber="7" refId="ref24769" refString="Bodin, Ph. (1979) Catalogue des nouveaux Copepodes Harpacticoides marins. (Nouvelle edition). Universite de Bretagne Occidentale, Brest, 228 pp." type="book" year="1979">Bodin (1979: 124)</bibRefCitation>
</treatmentCitation>
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</paragraph>
<paragraph id="8BFDA22F5D6FFFB8C9CC12349B38FD9E" blockId="6.[151,1437,541,2008]" box="[151,774,541,568]" pageId="6" pageNumber="7">
<emphasis id="B9367E3D5D6FFFB8C9CC123499AEFD9E" bold="true" box="[151,400,541,567]" pageId="6" pageNumber="7">Original description.</emphasis>
<bibRefCitation id="EFD3DFDE5D6FFFB8C8CC12379A0AFD9E" author="Masry, D." box="[407,564,541,568]" pageId="6" pageNumber="7" pagination="229 - 258" refId="ref28101" refString="Masry, D. (1970) Ecological study of some sandy beaches along the Israeli Mediterranean coast, with a description of the interstitial harpacticoids (Crustacea, Copepoda). Cahiers de Biologie marine, 11, 229 - 258. https: // doi. org / 10.21411 / CBM. A. 4943 D 377" type="journal article" year="1970">Masry (1970)</bibRefCitation>
: 249–251;
<figureCitation id="1379BEAA5D6FFFB8CBEB12349B3CFD9E" box="[688,770,541,567]" captionStart="FIGURE 12" captionStartId="24.[151,250,1943,1967]" captionTargetBox="[175,1411,192,1909]" captionTargetId="figure-17@24.[161,1426,181,1919]" captionTargetPageId="24" captionText="FIGURE 12. Arenopontia adriatica sp. nov. (♀): (A) P1, anterior; (B) P2, anterior; (C) P3, anterior; (D) P4, anterior." figureDoi="http://doi.org/10.5281/zenodo.10953676" httpUri="https://zenodo.org/record/10953676/files/figure.png" pageId="6" pageNumber="7">Fig. 12</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C358F1A45D6FFFB6C99C12689CCCFE77" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BFDA22F5D6FFFB8C99C12689962FD6E" blockId="6.[151,1437,541,2008]" pageId="6" pageNumber="7">
<emphasis id="B9367E3D5D6FFFB8C99C12689958FDF2" bold="true" box="[199,358,577,603]" pageId="6" pageNumber="7">Type locality.</emphasis>
<materialsCitation id="3B2AA8725D6FFFB8C83612689966FD6E" collectionCode="ICZN" country="Israel" county="Yel Yavne" location="Levantine" municipality="Palmahim" pageId="6" pageNumber="7" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F355E2BF5D6FFFB8C83612689990FDF2" box="[365,430,577,603]" name="Israel" pageId="6" pageNumber="7">Israel</collectingCountry>
,
<location id="8E9DF4F45D6FFFB8C8E112689A14FDF2" LSID="urn:lsid:plazi:treatment:03EB13395D6FFFB6C9CC11469CFAFAB3:8E9DF4F45D6FFFB8C8E112689A14FDF2" box="[442,554,577,603]" country="Israel" county="Yel Yavne" municipality="Palmahim" name="Levantine" pageId="6" pageNumber="7">Levantine</location>
coast.
<bibRefCitation id="EFD3DFDE5D6FFFB8CB22126B9B2DFDF2" author="Masry, D." box="[633,787,577,604]" pageId="6" pageNumber="7" pagination="229 - 258" refId="ref28101" refString="Masry, D. (1970) Ecological study of some sandy beaches along the Israeli Mediterranean coast, with a description of the interstitial harpacticoids (Crustacea, Copepoda). Cahiers de Biologie marine, 11, 229 - 258. https: // doi. org / 10.21411 / CBM. A. 4943 D 377" type="journal article" year="1970">Masry (1970)</bibRefCitation>
did not explicitly designate a type locality but mentioned that he collected material in five different sites:
<location id="8E9DF4F45D6FFFB8CB91124C9B0BFDD6" LSID="urn:lsid:plazi:treatment:03EB13395D6FFFB6C9CC11469CFAFAB3:8E9DF4F45D6FFFB8CB91124C9B0BFDD6" box="[714,821,613,639]" country="Israel" county="Yel Yavne" municipality="Palmahim" name="Nitzanim" pageId="6" pageNumber="7">Nitzanim</location>
,
<location id="8E9DF4F45D6FFFB8CA1F124C9BFBFDD6" LSID="urn:lsid:plazi:treatment:03EB13395D6FFFB6C9CC11469CFAFAB3:8E9DF4F45D6FFFB8CA1F124C9BFBFDD6" box="[836,965,613,639]" country="Israel" county="Yel Yavne" municipality="Palmahim" name="Mikhmoret" pageId="6" pageNumber="7">Mikhmoret</location>
,
<collectingMunicipality id="6B9938555D6FFFB8CA88124C9C76FDD6" box="[979,1096,613,639]" pageId="6" pageNumber="7">Palmahim</collectingMunicipality>
,
<collectingCounty id="629CDAA35D6FFFB8CD0E124C9CF4FD29" box="[1109,1226,613,640]" pageId="6" pageNumber="7">Yel Yavne</collectingCounty>
and south of Tel- Shiqmona. The latter site was labelled on the
<typeStatus id="54F91C8D5D6FFFB8CBF112A09B32FD0A" box="[682,780,649,675]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
slide and is consequently designated as the type locality (
<collectionCode id="ED533AEA5D6FFFB8C9C4128498DCFD6E" box="[159,226,685,711]" pageId="6" pageNumber="7">ICZN</collectionCode>
Art. 76.1)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFDA22F5D6FFFB8C99C12F89D1FFD42" blockId="6.[151,1437,541,2008]" box="[199,1313,721,748]" pageId="6" pageNumber="7">
<emphasis id="B9367E3D5D6FFFB8C99C12F8998FFD42" bold="true" box="[199,433,721,747]" pageId="6" pageNumber="7">Material examined.</emphasis>
<materialsCitation id="3B2AA8725D6FFFB8C8EC12F89D23FD42" box="[439,1309,721,748]" location="Hebrew University" pageId="6" pageNumber="7" specimenCount="1" specimenCount-female="1" stateProvince="Jerusalem" typeStatus="holotype">
<location id="8E9DF4F45D6FFFB8C8EC12F89AB0FD42" LSID="urn:lsid:plazi:treatment:03EB13395D6FFFB6C9CC11469CFAFAB3:8E9DF4F45D6FFFB8C8EC12F89AB0FD42" box="[439,654,721,747]" name="Hebrew University" pageId="6" pageNumber="7" stateProvince="Jerusalem">Hebrew University</location>
,
<collectingRegion id="49866CCD5D6FFFB8CBC212F89B33FD42" box="[665,781,721,747]" country="Israel" name="Jerusalem" pageId="6" pageNumber="7">Jerusalem</collectingRegion>
: damaged
<typeStatus id="54F91C8D5D6FFFB8CADF12F89BD8FD42" box="[900,998,721,747]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
<specimenCount id="9D4469A65D6FFFB8CAB612FB9C3CFD45" box="[1005,1026,722,748]" count="1" pageId="6" pageNumber="7" type="female">♀</specimenCount>
mounted
<emphasis id="B9367E3D5D6FFFB8CD2912FB9C85FD42" box="[1138,1211,722,747]" italics="true" pageId="6" pageNumber="7">in toto</emphasis>
on slide
</materialsCitation>
.
</paragraph>
<paragraph id="8BFDA22F5D6FFFB8C99C12DC9DA9FCFE" blockId="6.[151,1437,541,2008]" pageId="6" pageNumber="7">
<emphasis id="B9367E3D5D6FFFB8C99C12DC9A05FCA6" bold="true" box="[199,571,757,783]" pageId="6" pageNumber="7">Partial redescription of female.</emphasis>
Total body length from tip of rostrum to posterior margin of caudal rami 277 μm [290–320 μm according to
<bibRefCitation id="EFD3DFDE5D6FFFB8C8AD13339AA8FC9A" author="Masry, D." box="[502,662,793,820]" pageId="6" pageNumber="7" pagination="229 - 258" refId="ref28101" refString="Masry, D. (1970) Ecological study of some sandy beaches along the Israeli Mediterranean coast, with a description of the interstitial harpacticoids (Crustacea, Copepoda). Cahiers de Biologie marine, 11, 229 - 258. https: // doi. org / 10.21411 / CBM. A. 4943 D 377" type="journal article" year="1970">Masry (1970)</bibRefCitation>
]. Hyaline frills unconfirmed but probably like in other species of the genus (thoracic ones weakly developed and crenulated, abdominal ones consisting of rectangular digitate lappets).
</paragraph>
<paragraph id="8BFDA22F5D6FFFB8C99C13489D39FB86" blockId="6.[151,1437,541,2008]" pageId="6" pageNumber="7">
Caudal ramus (
<figureCitation id="1379BEAA5D6FFFB8C834134899FEFCD2" box="[367,448,865,891]" captionStart="FIGURE 1" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[196,1390,196,1899]" captionTargetId="figure-17@7.[184,1404,179,1908]" captionTargetPageId="7" captionText="FIGURE 1. Arenopontia problematica Masry, 1970 (♀): (A) P1, anterior [coxa omitted]; (B) P2, anterior [protopod omitted]; (C) P3, anterior [protopod omitted]; (D) P4, anterior [protopod largely omitted]; (E) antennule, dorsal; (F) right caudal ramus, lateral; (G) antenna [coxa omitted]." figureDoi="http://doi.org/10.5281/zenodo.10953648" httpUri="https://zenodo.org/record/10953648/files/figure.png" pageId="6" pageNumber="7">Fig. 1F</figureCitation>
) short, 1.7 times longer than wide (measured in lateral view from anterior margin to apex of spinous process), tapering posteriorly; with a pore laterally; outer distal corner produced into posteriorly directed recurved spinous process, accompanied at base by outer spinular row; dorsal surface without spur-like process but with row of strong spinules near inner margin. Armature consisting of seven setae; seta I small; setae II and III long and naked; seta IV short, located between seta
<collectionCode id="ED533AEA5D6FFFB8CBFC13DB9A85FBA5" box="[679,699,1010,1036]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="7" type="Museum">V</collectionCode>
and spinous process; seta
<collectionCode id="ED533AEA5D6FFFB8CABF13DB9BC6FBA5" box="[996,1016,1010,1036]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="7" type="Museum">V</collectionCode>
long and with fracture plane; seta
<collectionCode id="ED533AEA5D6FFFB8CC2413DB9DA3FBA5" box="[1407,1437,1010,1036]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="7">VI</collectionCode>
small, naked and located at inner distal corner; seta VII distinctly foliaceous and tri-articulate at base.
</paragraph>
<paragraph id="8BFDA22F5D6FFFB8C99C1410996BFB4A" blockId="6.[151,1437,541,2008]" pageId="6" pageNumber="7">
Antennule (
<figureCitation id="1379BEAA5D6FFFB8C8151410999AFBFA" box="[334,420,1081,1107]" captionStart="FIGURE 1" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[196,1390,196,1899]" captionTargetId="figure-17@7.[184,1404,179,1908]" captionTargetPageId="7" captionText="FIGURE 1. Arenopontia problematica Masry, 1970 (♀): (A) P1, anterior [coxa omitted]; (B) P2, anterior [protopod omitted]; (C) P3, anterior [protopod omitted]; (D) P4, anterior [protopod largely omitted]; (E) antennule, dorsal; (F) right caudal ramus, lateral; (G) antenna [coxa omitted]." figureDoi="http://doi.org/10.5281/zenodo.10953648" httpUri="https://zenodo.org/record/10953648/files/figure.png" pageId="6" pageNumber="7">Fig. 1E</figureCitation>
) short, six-segmented. Segment 1 with a short seta near anterodistal margin. Segment 2 longest, about twice longer than maximum width. Segment 4 with long aesthetasc (L: 25 μm) fused at base with seta. Distal segment with seven naked setae (none of which clearly spatulate) and apical acrothek consisting of short aesthetasc (L: 15 μm) and two slender setae. Armature formula: 1-[1], 2-[7 + 1 plumose], 3-[4], 4-[(1 + ae)], 5-[1], 6-[7 + acrothek].
</paragraph>
<paragraph id="8BFDA22F5D6FFFB8C99C14C79CCAFA15" blockId="6.[151,1437,541,2008]" pageId="6" pageNumber="7">
Antenna (
<figureCitation id="1379BEAA5D6FFFB8C86214C499A8FAAE" box="[313,406,1261,1287]" captionStart="FIGURE 1" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[196,1390,196,1899]" captionTargetId="figure-17@7.[184,1404,179,1908]" captionTargetPageId="7" captionText="FIGURE 1. Arenopontia problematica Masry, 1970 (♀): (A) P1, anterior [coxa omitted]; (B) P2, anterior [protopod omitted]; (C) P3, anterior [protopod omitted]; (D) P4, anterior [protopod largely omitted]; (E) antennule, dorsal; (F) right caudal ramus, lateral; (G) antenna [coxa omitted]." figureDoi="http://doi.org/10.5281/zenodo.10953648" httpUri="https://zenodo.org/record/10953648/files/figure.png" pageId="6" pageNumber="7">Fig. 1G</figureCitation>
). Coxa (not figured) small, without ornamentation. Allobasis about 2.6 times as long as maximum width; original basis-endopod boundary marked by partial transverse spinule row; with fine spinules along exopodal margin as figured. Exopod one-segmented, elongate, with short naked apical seta (about 1.5 times longer than exopod). Free endopod with two spinular rows on anterior surface and finer spinules at outer distal corner; medial armature consisting of two short spines; apical armature consisting of two spines and three geniculate setae, strongest of which with spinules around geniculation and fused basally to tiny accessory seta.
</paragraph>
<paragraph id="8BFDA22F5D6FFFB8C99C15EC9DADFA76" blockId="6.[151,1437,541,2008]" box="[199,1427,1477,1504]" pageId="6" pageNumber="7">The detailed morphology of the mouthparts and maxillipeds could not be discerned in the mounted specimen.</paragraph>
<paragraph id="8BFDA22F5D6FFFB8C99C15C09ABEF972" blockId="6.[151,1437,541,2008]" pageId="6" pageNumber="7">
P1 (
<figureCitation id="1379BEAA5D6FFFB8C9AE15C0996EF9AA" box="[245,336,1513,1539]" captionStart="FIGURE 1" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[196,1390,196,1899]" captionTargetId="figure-17@7.[184,1404,179,1908]" captionTargetPageId="7" captionText="FIGURE 1. Arenopontia problematica Masry, 1970 (♀): (A) P1, anterior [coxa omitted]; (B) P2, anterior [protopod omitted]; (C) P3, anterior [protopod omitted]; (D) P4, anterior [protopod largely omitted]; (E) antennule, dorsal; (F) right caudal ramus, lateral; (G) antenna [coxa omitted]." figureDoi="http://doi.org/10.5281/zenodo.10953648" httpUri="https://zenodo.org/record/10953648/files/figure.png" pageId="6" pageNumber="7">Fig. 1A</figureCitation>
). Basis with spinular row near bases of endopod and exopod; anterior surface with a small inner seta. Exopod three-segmented; exp-1 and -2 with spinules around outer margin; exp-1 longest, with long bare outer spine; exp-2 without outer element; exp-3 with short unipinnate outer spine, a long curved unipinnate spine and one geniculate seta distally, and one inner, apically penicillate seta subdistally. Endopod two-segmented, prehensile; enp-1 6.9 times longer than wide, and about 1.68 times longer than exopod; with a serrate inner seta in proximal third, and three pairs of spinules along outer margin; enp-2 slightly longer than wide, with a short unipinnate spine, a geniculate claw and a small inner spinule.
</paragraph>
<paragraph id="8BFDA22F5D6FFFB6C99C16CC9A50FF7C" blockId="6.[151,1437,541,2008]" lastBlockId="8.[151,1436,151,213]" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
P2–P4 (
<figureCitation id="1379BEAA5D6FFFB8C87916CC99A1F956" box="[290,415,1765,1791]" captionStart="FIGURE 1" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[196,1390,196,1899]" captionTargetId="figure-17@7.[184,1404,179,1908]" captionTargetPageId="7" captionText="FIGURE 1. Arenopontia problematica Masry, 1970 (♀): (A) P1, anterior [coxa omitted]; (B) P2, anterior [protopod omitted]; (C) P3, anterior [protopod omitted]; (D) P4, anterior [protopod largely omitted]; (E) antennule, dorsal; (F) right caudal ramus, lateral; (G) antenna [coxa omitted]." figureDoi="http://doi.org/10.5281/zenodo.10953648" httpUri="https://zenodo.org/record/10953648/files/figure.png" pageId="6" pageNumber="7">Fig. 1B–D</figureCitation>
). Bases with a spinular row near base of endopod (P3–P4; not figured for P3) and outer distal corner (P2, P4); outer basal seta absent (P2), plumose (P3) or naked (P4). Exopods three-segmented; segments with spinular ornamentation as figured; inner distal seta of exp-3 sparsely bipinnate, all other elements unipinnate; hyaline frills of exp-1 and -2 well developed; exp-2 with lateral pore halfway down inner margin length; P3–P4 exp-3 with anterior surface pore. Endopods two-segmented; P2–P4 enp-1 about 1.7, 1.9 and 3.3 times longer than their respective distal endopodal segments, with few spinules as figured. P2 enp-2 with a long, apically serrate, posteriorly oriented seta near inner margin and a long, sparsely pinnate distal seta. P3 enp-2 with a long bipinnate seta distally. P4 enp-2 minute, with a basally fused, apically serrate seta, and long unipinnate seta at outer distal corner. Spine and seta formula as follows:
</paragraph>
<caption id="DF3DF2A75D6EFFB9C9CC17A799C0F847" ID-DOI="http://doi.org/10.5281/zenodo.10953648" ID-Zenodo-Dep="10953648" httpUri="https://zenodo.org/record/10953648/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,1934,1958]" targetBox="[196,1390,196,1899]" targetPageId="7" targetType="figure">
<paragraph id="8BFDA22F5D6EFFB9C9CC17A799C0F847" blockId="7.[151,1436,1934,2031]" pageId="7" pageNumber="8">
<emphasis id="B9367E3D5D6EFFB9C9CC17A7992AF80F" bold="true" box="[151,276,1934,1958]" pageId="7" pageNumber="8">FIGURE 1.</emphasis>
<taxonomicName id="4C42D9AC5D6EFFB9C84117A69A95F80E" authority="Masry, 1970" authorityName="Masry" authorityYear="1970" box="[282,683,1934,1959]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="problematica">
<emphasis id="B9367E3D5D6EFFB9C84117A69A1BF80F" box="[282,549,1934,1958]" italics="true" pageId="7" pageNumber="8">Arenopontia problematica</emphasis>
<bibRefCitation id="EFD3DFDE5D6EFFB9CB7717A69A95F80E" author="Masry, D." box="[556,683,1935,1959]" pageId="7" pageNumber="8" pagination="229 - 258" refId="ref28101" refString="Masry, D. (1970) Ecological study of some sandy beaches along the Israeli Mediterranean coast, with a description of the interstitial harpacticoids (Crustacea, Copepoda). Cahiers de Biologie marine, 11, 229 - 258. https: // doi. org / 10.21411 / CBM. A. 4943 D 377" type="journal article" year="1970">Masry, 1970</bibRefCitation>
</taxonomicName>
(♀): (A) P1, anterior [coxa omitted]; (B) P2, anterior [protopod omitted]; (C) P3, anterior [protopod omitted]; (D) P4, anterior [protopod largely omitted]; (E) antennule, dorsal; (F) right caudal ramus, lateral; (G) antenna [coxa omitted].
</paragraph>
</caption>
<paragraph id="8BFDA22F5D61FFB6CBAA11219CDEFE3C" pageId="8" pageNumber="9">
<table id="F942508F5D610041C83B11219CCFFE3C" box="[352,1265,264,405]" gridcols="3" gridrows="4" pageId="8" pageNumber="9">
<tr id="3572A06D5D610041C83B11219CCFFE89" box="[352,1265,264,288]" gridrow="0" pageId="8" pageNumber="9" rowspan-0="1">
<th id="76A3C9115D610041CBAA11219B7EFE89" box="[753,832,264,288]" gridcol="1" gridrow="0" pageId="8" pageNumber="9">Exopod</th>
<th id="76A3C9115D610041CDCD11219CCFFE89" box="[1174,1265,264,288]" gridcol="2" gridrow="0" pageId="8" pageNumber="9">Endopod</th>
</tr>
<tr id="3572A06D5D610041C83B11199CCFFEE1" box="[352,1265,304,328]" gridrow="1" pageId="8" pageNumber="9">
<th id="76A3C9115D610041C83B11199944FEE1" box="[352,378,304,328]" gridcol="0" gridrow="1" pageId="8" pageNumber="9">P2</th>
<td id="76A3C9115D610041CBAA11199B7EFEE1" box="[753,832,304,328]" gridcol="1" gridrow="1" pageId="8" pageNumber="9">0.0.021</td>
<td id="76A3C9115D610041CDCD11199CCFFEE1" box="[1174,1265,304,328]" gridcol="2" gridrow="1" pageId="8" pageNumber="9">0.110</td>
</tr>
<tr id="3572A06D5D610041C83B117E9CCFFEC6" box="[352,1265,343,367]" gridrow="2" pageId="8" pageNumber="9">
<th id="76A3C9115D610041C83B117E9944FEC6" box="[352,378,343,367]" gridcol="0" gridrow="2" pageId="8" pageNumber="9">P3</th>
<td id="76A3C9115D610041CBAA117E9B7EFEC6" box="[753,832,343,367]" gridcol="1" gridrow="2" pageId="8" pageNumber="9">0.0.021</td>
<td id="76A3C9115D610041CDCD117E9CCFFEC6" box="[1174,1265,343,367]" gridcol="2" gridrow="2" pageId="8" pageNumber="9">0.010</td>
</tr>
<tr id="3572A06D5D610041C83B11549CCFFE3C" box="[352,1265,381,405]" gridrow="3" pageId="8" pageNumber="9">
<th id="76A3C9115D610041C83B11549944FE3C" box="[352,378,381,405]" gridcol="0" gridrow="3" pageId="8" pageNumber="9">P4</th>
<td id="76A3C9115D610041CBAA11549B7EFE3C" box="[753,832,381,405]" gridcol="1" gridrow="3" pageId="8" pageNumber="9">0.0.021</td>
<td id="76A3C9115D610041CDCD11549CCFFE3C" box="[1174,1265,381,405]" gridcol="2" gridrow="3" pageId="8" pageNumber="9">0.020</td>
</tr>
</table>
</paragraph>
<paragraph id="8BFDA22F5D61FFB6C99C11ED9CCCFE77" blockId="8.[151,1437,452,1307]" box="[199,1266,452,479]" pageId="8" pageNumber="9">
The structure of the P5 and genital field could not be observed in the badly preserved
<typeStatus id="54F91C8D5D61FFB6CDD311ED9CD0FE77" box="[1160,1262,452,478]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C358F1A45D61FFB6C99C11C19CFAFAB3" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8BFDA22F5D61FFB6C99C11C19A4CFBCF" blockId="8.[151,1437,452,1307]" pageId="8" pageNumber="9">
<emphasis id="B9367E3D5D61FFB6C99C11C19905FDAB" bold="true" box="[199,315,488,514]" pageId="8" pageNumber="9">Remarks.</emphasis>
Our re-examination of the
<typeStatus id="54F91C8D5D61FFB6CB3311C19AF4FDAB" box="[616,714,488,514]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
showed that
<bibRefCitation id="EFD3DFDE5D61FFB6CA0011C19C38FDAB" author="Masry, D." box="[859,1030,488,514]" pageId="8" pageNumber="9" pagination="229 - 258" refId="ref28101" refString="Masry, D. (1970) Ecological study of some sandy beaches along the Israeli Mediterranean coast, with a description of the interstitial harpacticoids (Crustacea, Copepoda). Cahiers de Biologie marine, 11, 229 - 258. https: // doi. org / 10.21411 / CBM. A. 4943 D 377" type="journal article" year="1970">Masry’s (1970)</bibRefCitation>
illustrations and text contain several significant errors, invalidating most of his criteria proposed to separate
<taxonomicName id="4C42D9AC5D61FFB6CAEE12249C56FD8F" authorityName="Masry" authorityYear="1970" box="[949,1128,524,551]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="problematica">
<emphasis id="B9367E3D5D61FFB6CAEE12249C56FD8F" box="[949,1128,524,551]" italics="true" pageId="8" pageNumber="9">A. problematica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C42D9AC5D61FFB6CDC512249D7AFD8F" authorityName="Kunz" authorityYear="1937" box="[1182,1348,524,551]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="B9367E3D5D61FFB6CDC512249D7AFD8F" box="[1182,1348,524,551]" italics="true" pageId="8" pageNumber="9">A. subterranea</emphasis>
</taxonomicName>
: (1) the author claimed that the female antennule is only five-segmented with a single, very long aesthetasc arising from the annulated apical segment; in reality, the antennule in
<taxonomicName id="4C42D9AC5D61FFB6CA4C127C9BF5FDC7" authorityName="Masry" authorityYear="1970" box="[791,971,596,623]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="problematica">
<emphasis id="B9367E3D5D61FFB6CA4C127C9BF5FDC7" box="[791,971,596,623]" italics="true" pageId="8" pageNumber="9">A. problematica</emphasis>
</taxonomicName>
displays the typical arenopontiid pattern, being six-segmented with aesthetacs on segments 4 and 6 and no subdivisions of the apical segment (
<figureCitation id="1379BEAA5D61FFB6CC5412519D5BFD3B" box="[1295,1381,632,658]" captionStart="FIGURE 1" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[196,1390,196,1899]" captionTargetId="figure-17@7.[184,1404,179,1908]" captionTargetPageId="7" captionText="FIGURE 1. Arenopontia problematica Masry, 1970 (♀): (A) P1, anterior [coxa omitted]; (B) P2, anterior [protopod omitted]; (C) P3, anterior [protopod omitted]; (D) P4, anterior [protopod largely omitted]; (E) antennule, dorsal; (F) right caudal ramus, lateral; (G) antenna [coxa omitted]." figureDoi="http://doi.org/10.5281/zenodo.10953648" httpUri="https://zenodo.org/record/10953648/files/figure.png" pageId="8" pageNumber="9">Fig. 1E</figureCitation>
); (2) the antennary exopod, claimed to be absent, was overlooked (
<figureCitation id="1379BEAA5D61FFB6CA1A12B59BA4FD1F" box="[833,922,668,694]" captionStart="FIGURE 1" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[196,1390,196,1899]" captionTargetId="figure-17@7.[184,1404,179,1908]" captionTargetPageId="7" captionText="FIGURE 1. Arenopontia problematica Masry, 1970 (♀): (A) P1, anterior [coxa omitted]; (B) P2, anterior [protopod omitted]; (C) P3, anterior [protopod omitted]; (D) P4, anterior [protopod largely omitted]; (E) antennule, dorsal; (F) right caudal ramus, lateral; (G) antenna [coxa omitted]." figureDoi="http://doi.org/10.5281/zenodo.10953648" httpUri="https://zenodo.org/record/10953648/files/figure.png" pageId="8" pageNumber="9">Fig. 1G</figureCitation>
); (3) the innermost seta on P1 exp-3 is distally penicillate and not pinnate (
<figureCitation id="1379BEAA5D61FFB6C88B12E99A14FD73" box="[464,554,704,730]" captionStart="FIGURE 1" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[196,1390,196,1899]" captionTargetId="figure-17@7.[184,1404,179,1908]" captionTargetPageId="7" captionText="FIGURE 1. Arenopontia problematica Masry, 1970 (♀): (A) P1, anterior [coxa omitted]; (B) P2, anterior [protopod omitted]; (C) P3, anterior [protopod omitted]; (D) P4, anterior [protopod largely omitted]; (E) antennule, dorsal; (F) right caudal ramus, lateral; (G) antenna [coxa omitted]." figureDoi="http://doi.org/10.5281/zenodo.10953648" httpUri="https://zenodo.org/record/10953648/files/figure.png" pageId="8" pageNumber="9">Fig. 1A</figureCitation>
); (4) the inner serrate seta on P1 enp-1 was overlooked (
<figureCitation id="1379BEAA5D61FFB6CDFC12E99D3FFD73" box="[1191,1281,704,730]" captionStart="FIGURE 1" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[196,1390,196,1899]" captionTargetId="figure-17@7.[184,1404,179,1908]" captionTargetPageId="7" captionText="FIGURE 1. Arenopontia problematica Masry, 1970 (♀): (A) P1, anterior [coxa omitted]; (B) P2, anterior [protopod omitted]; (C) P3, anterior [protopod omitted]; (D) P4, anterior [protopod largely omitted]; (E) antennule, dorsal; (F) right caudal ramus, lateral; (G) antenna [coxa omitted]." figureDoi="http://doi.org/10.5281/zenodo.10953648" httpUri="https://zenodo.org/record/10953648/files/figure.png" pageId="8" pageNumber="9">Fig. 1A</figureCitation>
); (5) contrary to Masry’s Figure 62, the P2 basis does not have an outer seta (
<figureCitation id="1379BEAA5D61FFB6CA0312CD9B90FD57" box="[856,942,740,766]" captionStart="FIGURE 1" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[196,1390,196,1899]" captionTargetId="figure-17@7.[184,1404,179,1908]" captionTargetPageId="7" captionText="FIGURE 1. Arenopontia problematica Masry, 1970 (♀): (A) P1, anterior [coxa omitted]; (B) P2, anterior [protopod omitted]; (C) P3, anterior [protopod omitted]; (D) P4, anterior [protopod largely omitted]; (E) antennule, dorsal; (F) right caudal ramus, lateral; (G) antenna [coxa omitted]." figureDoi="http://doi.org/10.5281/zenodo.10953648" httpUri="https://zenodo.org/record/10953648/files/figure.png" pageId="8" pageNumber="9">Fig. 1B</figureCitation>
); (6) P2 enp-2 has one apical seta instead of two (the short outer one illustrated in his Figure 62 is a spinule (
<figureCitation id="1379BEAA5D61FFB6CA3213219B83FC8B" box="[873,957,776,802]" captionStart="FIGURE 1" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[196,1390,196,1899]" captionTargetId="figure-17@7.[184,1404,179,1908]" captionTargetPageId="7" captionText="FIGURE 1. Arenopontia problematica Masry, 1970 (♀): (A) P1, anterior [coxa omitted]; (B) P2, anterior [protopod omitted]; (C) P3, anterior [protopod omitted]; (D) P4, anterior [protopod largely omitted]; (E) antennule, dorsal; (F) right caudal ramus, lateral; (G) antenna [coxa omitted]." figureDoi="http://doi.org/10.5281/zenodo.10953648" httpUri="https://zenodo.org/record/10953648/files/figure.png" pageId="8" pageNumber="9">Fig. 1B</figureCitation>
)); (7) P3 enp-2 has one apical seta instead of two, the short outer one figured by Masry being a spinule (and in reality much shorter;
<figureCitation id="1379BEAA5D61FFB6CDC513059CCBFCEF" box="[1182,1269,812,838]" captionStart="FIGURE 1" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[196,1390,196,1899]" captionTargetId="figure-17@7.[184,1404,179,1908]" captionTargetPageId="7" captionText="FIGURE 1. Arenopontia problematica Masry, 1970 (♀): (A) P1, anterior [coxa omitted]; (B) P2, anterior [protopod omitted]; (C) P3, anterior [protopod omitted]; (D) P4, anterior [protopod largely omitted]; (E) antennule, dorsal; (F) right caudal ramus, lateral; (G) antenna [coxa omitted]." figureDoi="http://doi.org/10.5281/zenodo.10953648" httpUri="https://zenodo.org/record/10953648/files/figure.png" pageId="8" pageNumber="9">Fig. 1C</figureCitation>
); (8) P4 exp-3 has only one outer spine instead of two (
<figureCitation id="1379BEAA5D61FFB6CB3513799AF6FCC3" box="[622,712,848,874]" captionStart="FIGURE 1" captionStartId="7.[151,250,1934,1958]" captionTargetBox="[196,1390,196,1899]" captionTargetId="figure-17@7.[184,1404,179,1908]" captionTargetPageId="7" captionText="FIGURE 1. Arenopontia problematica Masry, 1970 (♀): (A) P1, anterior [coxa omitted]; (B) P2, anterior [protopod omitted]; (C) P3, anterior [protopod omitted]; (D) P4, anterior [protopod largely omitted]; (E) antennule, dorsal; (F) right caudal ramus, lateral; (G) antenna [coxa omitted]." figureDoi="http://doi.org/10.5281/zenodo.10953648" httpUri="https://zenodo.org/record/10953648/files/figure.png" pageId="8" pageNumber="9">Fig. 1D</figureCitation>
);
<bibRefCitation id="EFD3DFDE5D61FFB6CBBA13789B42FCC3" author="Masry, D." box="[737,892,848,875]" pageId="8" pageNumber="9" pagination="229 - 258" refId="ref28101" refString="Masry, D. (1970) Ecological study of some sandy beaches along the Israeli Mediterranean coast, with a description of the interstitial harpacticoids (Crustacea, Copepoda). Cahiers de Biologie marine, 11, 229 - 258. https: // doi. org / 10.21411 / CBM. A. 4943 D 377" type="journal article" year="1970">Masry (1970)</bibRefCitation>
lists two elements in the armature formula and illustrates two in his Figure 64; for this reason
<taxonomicName id="4C42D9AC5D61FFB6CBF2135C9B62FC27" authorityName="Masry" authorityYear="1970" box="[681,860,884,911]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="problematica">
<emphasis id="B9367E3D5D61FFB6CBF2135C9B62FC27" box="[681,860,884,911]" italics="true" pageId="8" pageNumber="9">A. problematica</emphasis>
</taxonomicName>
keys out to the wrong codon in Wells’ (2007: 188) key; and (9) caudal seta VII is clearly foliaceous. In addition, his claim that the male P6 bears three spines is almost certainly wrong since the maximum number of armature elements observed in any arenopontiid is two. The female P5 displays a deep incision along its free distal margin; this unique morphology could not be confirmed due to the bad condition of the
<typeStatus id="54F91C8D5D61FFB6C820142D99DFFBB7" box="[379,481,1028,1054]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
.
<bibRefCitation id="EFD3DFDE5D61FFB6C8B2142C9ABFFBB7" author="Masry, D." box="[489,641,1028,1055]" pageId="8" pageNumber="9" pagination="229 - 258" refId="ref28101" refString="Masry, D. (1970) Ecological study of some sandy beaches along the Israeli Mediterranean coast, with a description of the interstitial harpacticoids (Crustacea, Copepoda). Cahiers de Biologie marine, 11, 229 - 258. https: // doi. org / 10.21411 / CBM. A. 4943 D 377" type="journal article" year="1970">Masry (1970)</bibRefCitation>
also states that the posterior margins of all body somites are smooth but this is highly unlikely given that the hyaline frills of the abdominal somites have rectangular digitate or nondigitate lappets in all other members of the family.
</paragraph>
<paragraph id="8BFDA22F5D61FFB6C99C14589CFAFAB3" blockId="8.[151,1437,452,1307]" pageId="8" pageNumber="9">
<taxonomicName id="4C42D9AC5D61FFB6C99C145899D0FB23" authorityName="Masry" authorityYear="1970" box="[199,494,1136,1163]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="problematica">
<emphasis id="B9367E3D5D61FFB6C99C145899D0FB23" box="[199,494,1136,1163]" italics="true" pageId="8" pageNumber="9">Arenopontia problematica</emphasis>
</taxonomicName>
is extremely close to
<taxonomicName id="4C42D9AC5D61FFB6CBB114589BAFFB23" authorityName="Kunz" authorityYear="1937" box="[746,913,1136,1163]" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="B9367E3D5D61FFB6CBB114589BAFFB23" box="[746,913,1136,1163]" italics="true" pageId="8" pageNumber="9">A. subterranea</emphasis>
</taxonomicName>
, differing only in body size (the former being smaller), the morphology of the female P5 (unknown in the male) and the relatively shorter P2–P4 enp-1 and caudal ramus. A proper redescription of the fifth legs and the male are required before potential conspecificity with
<taxonomicName id="4C42D9AC5D61FFB6CCDF14909920FB5F" authorityName="Kunz" authorityYear="1937" class="Copepoda" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="B9367E3D5D61FFB6CCDF14909920FB5F" italics="true" pageId="8" pageNumber="9">A. subterranea</emphasis>
</taxonomicName>
can be ruled out. The species is known only from the Israeli coast and has not been recorded again since its original description. According to
<bibRefCitation id="EFD3DFDE5D61FFB6CB6115289AEBFAB3" author="Masry, D." box="[570,725,1280,1307]" pageId="8" pageNumber="9" pagination="229 - 258" refId="ref28101" refString="Masry, D. (1970) Ecological study of some sandy beaches along the Israeli Mediterranean coast, with a description of the interstitial harpacticoids (Crustacea, Copepoda). Cahiers de Biologie marine, 11, 229 - 258. https: // doi. org / 10.21411 / CBM. A. 4943 D 377" type="journal article" year="1970">Masry (1970)</bibRefCitation>
, females (95%) greatly outnumbered males.
</paragraph>
</subSubSection>
</treatment>
</document>
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