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<document ID-DOI="10.5733/afin.049.0208" ID-ISSN="2305-2562" ID-Zenodo-Dep="7911425" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1683578222239" checkinUser="felipe" docAuthor="Davies, Gregory B. P. &amp; Miller, Raymond M." docDate="2008" docId="03AEFB40FFF91C72FE4AFE5EBED3F978" docLanguage="en" docName="AfrInvertebr.49.2.131-158.pdf" docOrigin="African Invertebrates 49 (2)" docSource="http://www.bioone.org/doi/abs/10.5733/afin.049.0208" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.3:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="3" docTitle="Parapachycerina talea Davies &amp; Miller 2008, sp. n." docType="treatment" docVersion="5" lastPageNumber="149" masterDocId="FF978338FFE91C60FF82FF80BC47FFFF" masterDocTitle="Revision of the Afrotropical species of Parapachycerina (Diptera: Lauxaniidae)" masterLastPageNumber="131" masterPageNumber="131" pageNumber="147" updateTime="1683606976522" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Afrotropical species of Parapachycerina (Diptera: Lauxaniidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Davies, Gregory B. P.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Miller, Raymond M.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>49</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>131</mods:start>
<mods:end>131</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://www.bioone.org/doi/abs/10.5733/afin.049.0208</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5733/afin.049.0208</mods:identifier>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
<mods:identifier type="Zenodo-Dep">7911425</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03AEFB40FFF91C72FE4AFE5EBED3F978" httpUri="http://treatment.plazi.org/id/03AEFB40FFF91C72FE4AFE5EBED3F978" lastPageId="18" lastPageNumber="149" pageId="16" pageNumber="147">
<subSubSection box="[456,791,478,503]" pageId="16" pageNumber="147" type="nomenclature">
<paragraph blockId="16.[456,791,478,503]" box="[456,791,478,503]" pageId="16" pageNumber="147">
<heading box="[456,791,478,503]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="147" reason="2">
<taxonomicName authority="Davies &amp; Miller, 2008" authorityName="Davies &amp; Miller" authorityYear="2008" box="[456,725,478,503]" class="Insecta" family="Lauxaniidae" genus="Parapachycerina" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="147" phylum="Arthropoda" rank="species" species="talea" status="sp. n.">
<emphasis bold="true" box="[456,725,478,503]" pageId="16" pageNumber="147">Parapachycerina talea</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[731,791,478,503]" pageId="16" pageNumber="147" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="18" lastPageNumber="149" pageId="16" pageNumber="147" type="etymology">
<paragraph blockId="16.[144,1104,526,791]" pageId="16" pageNumber="147">
Etymology: From Latin
<taxonomicName authorityName="Davies &amp; Miller" authorityYear="2008" box="[423,479,526,550]" class="Insecta" family="Lauxaniidae" genus="Parapachycerina" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="147" phylum="Arthropoda" rank="species" species="talea">
<emphasis box="[423,479,526,550]" italics="true" pageId="16" pageNumber="147">talea</emphasis>
</taxonomicName>
(small rod), referring to posterior extension of ocellar spot that reaches onto the occiput.
</paragraph>
<paragraph blockId="16.[144,1104,526,791]" box="[144,264,598,623]" pageId="16" pageNumber="147">Diagnosis:</paragraph>
<paragraph blockId="16.[144,1104,526,791]" lastBlockId="17.[181,1142,206,871]" lastPageId="17" lastPageNumber="148" pageId="16" pageNumber="147">
Robust Malagasy species only known from female specimens. Ocellar triangle small and compact. Ocellar spot brown (not black) with narrow extension (prong) posteriad. Postpedicel with darkened apical end. Thick brownish edge to scutum/notopleuron. One sternopleural bristle. Profemoral ctenidium present. Anterior edge of wing fumose (specifically costal cell, marginal cell and anterior part of submarginal cell; no brownish smudges on anterior and posterior cross-veins). Undescribed taxon B closely resembles
<taxonomicName authorityName="Davies &amp; Miller" authorityYear="2008" box="[181,265,238,262]" class="Insecta" family="Lauxaniidae" genus="Parapachycerina" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="148" phylum="Arthropoda" rank="species" species="talea">
<emphasis box="[181,265,238,262]" italics="true" pageId="17" pageNumber="148">P. talea</emphasis>
</taxonomicName>
, see below for further details.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7911646" ID-Zenodo-Dep="7911646" box="[153,1095,1649,1669]" httpUri="https://zenodo.org/record/7911646/files/figure.png" pageId="16" pageNumber="147" startId="16.[153,192,1649,1669]" targetBox="[167,1067,841,1620]" targetPageId="16" targetType="figure">
<paragraph blockId="16.[153,1095,1649,1669]" box="[153,1095,1649,1669]" pageId="16" pageNumber="147">
Figs 2527. Inner ơ genitalia of
<taxonomicName authority="Davies &amp; Miller, 2008" authorityName="Davies &amp; Miller" authorityYear="2008" box="[447,527,1649,1669]" class="Insecta" family="Lauxaniidae" genus="Parapachycerina" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="147" phylum="Arthropoda" rank="species" species="lalitra" status="sp. n.">
<emphasis box="[447,527,1649,1669]" italics="true" pageId="16" pageNumber="147">P. lalitra</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[532,583,1649,1669]" pageId="16" pageNumber="147" rank="species">sp. n.</taxonomicNameLabel>
: (25, 26) dorsal and lateral views, (27) aedeagus wing.
</paragraph>
</caption>
<paragraph blockId="17.[181,1142,206,871]" box="[181,321,278,303]" pageId="17" pageNumber="148">Description:</paragraph>
<paragraph blockId="17.[181,1142,206,871]" pageId="17" pageNumber="148">
<emphasis box="[181,260,318,342]" italics="true" pageId="17" pageNumber="148">Colour</emphasis>
: Mainly yellow-orange. Scape and pedicel yellow. Distal half of postpedicel and area at aristal tumidity brown, rest of postpedicel yellow. Ocellar spot brown with characteristic brown medial prong extending posteriorly. Maxillary palpi yellow.Thorax yellow with rather faint orange-brown medial stripe on mesonotum; edges of notopleuron infuscated orange-brown. Abdomen brownish orange.
</paragraph>
<paragraph blockId="17.[181,1142,206,871]" pageId="17" pageNumber="148">
<emphasis box="[181,242,486,510]" italics="true" pageId="17" pageNumber="148">Head</emphasis>
: Orbital plates weakly differentiated. Basal half of scape recessed under anterior lip of postfrons. Vertex rounded. Ocellar triangle small, sides slightly longer than base.
</paragraph>
<paragraph blockId="17.[181,1142,206,871]" box="[181,927,558,583]" pageId="17" pageNumber="148">
<emphasis box="[181,262,558,582]" italics="true" pageId="17" pageNumber="148">Thorax</emphasis>
: Chaetation largely as per genus. One sternopleural bristle.
</paragraph>
<paragraph blockId="17.[181,1142,206,871]" pageId="17" pageNumber="148">
<emphasis box="[181,234,598,622]" italics="true" pageId="17" pageNumber="148">Legs</emphasis>
: Ctenidium present on profemur,
<emphasis box="[611,637,598,622]" italics="true" pageId="17" pageNumber="148">ca</emphasis>
9 setulae. Profemora posteriorly with ventral row of
<emphasis box="[260,286,630,654]" italics="true" pageId="17" pageNumber="148">ca</emphasis>
2 setae and dorsal row of
<emphasis box="[574,600,630,654]" italics="true" pageId="17" pageNumber="148">ca</emphasis>
4 setae. Mesofemur with medioanterior row of 5 setae.
</paragraph>
<paragraph blockId="17.[181,1142,206,871]" box="[181,1142,702,727]" pageId="17" pageNumber="148">
<emphasis box="[181,249,702,726]" italics="true" pageId="17" pageNumber="148">Wings</emphasis>
: Costal cell, marginal cell and anterior section of submarginal lightly infuscated.
</paragraph>
<paragraph blockId="17.[181,1142,206,871]" box="[181,599,742,767]" pageId="17" pageNumber="148">
<emphasis box="[181,287,742,766]" italics="true" pageId="17" pageNumber="148">Abdomen</emphasis>
: Male terminalia unknown.
</paragraph>
<paragraph blockId="17.[181,1142,206,871]" pageId="17" pageNumber="148">
<emphasis box="[181,343,782,806]" italics="true" pageId="17" pageNumber="148">Measurements</emphasis>
: ^(n=3): total length 3.8 (3.64.1); head length 0.5; head height 0.8; head width 1.1; thorax length 1.8 (1.71.9); abdomen length 1.5 (1.31.7); wing length 3.5 (3.33.8).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7911685" ID-Zenodo-Dep="7911685" httpUri="https://zenodo.org/record/7911685/files/figure.png" pageId="17" pageNumber="148" startId="17.[182,221,1600,1620]" targetBox="[202,1117,906,1569]" targetPageId="17" targetType="figure">
<paragraph blockId="17.[182,1142,1600,1668]" pageId="17" pageNumber="148">Figs 2830. Inner ơ genitalia of undescribed taxon A, Ivondro, Madagascar: (28, 29) dorsal and lateral views, (30) aedeagus wing. Note that the ventral area of the hypandrial apron is not illustrated due to the extreme transparency of the sclerite following over-treatment in KOH.</paragraph>
</caption>
<paragraph blockId="18.[144,1103,204,351]" box="[144,915,204,224]" pageId="18" pageNumber="149">
<materialsCitation box="[144,915,204,224]" collectingDate="1957-12-05" collectionCode="MHNB" collectorName="F. Keiser" country="Madagascar" location="Andasibe" pageId="18" pageNumber="149" specimenCount="1" typeStatus="holotype">
<typeStatus box="[144,232,204,224]" pageId="18" pageNumber="149">Holotype</typeStatus>
:
<collectingCountry box="[240,428,204,224]" name="Madagascar" pageId="18" pageNumber="149">^MADAGASCAR</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:03AEFB40FFF91C72FE4AFE5EBED3F978:8ED81C8DFFFB1C72FE31FF4CBE4EFF1F" box="[435,521,204,224]" country="Madagascar" name="Andasibe" pageId="18" pageNumber="149">Andasibe</location>
(= Périnet),
<date box="[635,729,204,224]" pageId="18" pageNumber="149" value="1957-12-05">
<collectingDate box="[635,729,204,224]" pageId="18" pageNumber="149" value="1957-12-05">5.xii.1957</collectingDate>
</date>
,
<collectorName box="[738,819,204,224]" pageId="18" pageNumber="149">F. Keiser</collectorName>
(
<collectionCode box="[832,905,204,224]" country="Switzerland" httpUri="http://grbio.org/cool/kt43-hjs1" name="Museum d'Histoire Naturelle de Bale" pageId="18" pageNumber="149">MHNB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="18.[144,1103,204,351]" pageId="18" pageNumber="149">
Other material examined:
<materialsCitation collectingDate="1958-01-18" collectingDateMax="1958-09-13" collectingDateMin="1958-01-18" collectionCode="MHNB" collectorName="F. Keiser" country="Madagascar" location="Vohiparara" pageId="18" pageNumber="149" specimenCount="1">
<collectingCountry box="[381,546,236,256]" name="Madagascar" pageId="18" pageNumber="149">MADAGASCAR</collectingCountry>
:
<location LSID="urn:lsid:plazi:treatment:03AEFB40FFF91C72FE4AFE5EBED3F978:8ED81C8DFFFB1C72FDA9FF6CBEF5FEFF" box="[555,690,236,256]" country="Madagascar" name="Vohiparara" pageId="18" pageNumber="149">1^Vohiparara</location>
,
<date box="[699,798,236,256]" pageId="18" pageNumber="149" value="1958-09-13">
<collectingDate box="[699,798,236,256]" pageId="18" pageNumber="149" value="1958-09-13">13.ix.1958</collectingDate>
</date>
,
<collectorName box="[808,889,236,256]" pageId="18" pageNumber="149">F. Keiser</collectorName>
(
<collectionCode box="[902,975,236,256]" country="Switzerland" httpUri="http://grbio.org/cool/kt43-hjs1" name="Museum d'Histoire Naturelle de Bale" pageId="18" pageNumber="149">MHNB</collectionCode>
); 1^Ambalamanakana,
<date box="[247,334,260,280]" pageId="18" pageNumber="149" value="1958-01-18">
<collectingDate box="[247,334,260,280]" pageId="18" pageNumber="149" value="1958-01-18">18.i.1958</collectingDate>
</date>
,
<collectorName box="[344,425,260,280]" pageId="18" pageNumber="149">F. Keiser</collectorName>
(
<collectionCode box="[437,510,260,280]" country="Switzerland" httpUri="http://grbio.org/cool/kt43-hjs1" name="Museum d'Histoire Naturelle de Bale" pageId="18" pageNumber="149">MHNB</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="18.[144,1103,204,351]" pageId="18" pageNumber="149">
Distribution: Restricted to
<collectingCountry box="[442,579,294,319]" name="Madagascar" pageId="18" pageNumber="149">Madagascar</collectingCountry>
. Known only from forested areas along eastern versant.
</paragraph>
<paragraph blockId="18.[144,1104,390,647]" box="[505,742,390,415]" pageId="18" pageNumber="149">Undescribed taxon A</paragraph>
<paragraph blockId="18.[144,1104,390,647]" pageId="18" pageNumber="149">
A male from east coast of
<collectingCountry box="[452,587,430,455]" name="Madagascar" pageId="18" pageNumber="149">Madagascar</collectingCountry>
(Ivondro,
<date box="[702,774,430,455]" pageId="18" pageNumber="149" value="1900-01">i.1900</date>
, C.Alluaud, MNHN) is extremely similar to
<taxonomicName authorityName="Davies &amp; Miller" authorityYear="2008" box="[324,422,462,486]" class="Insecta" family="Lauxaniidae" genus="Parapachycerina" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="149" phylum="Arthropoda" rank="species" species="lalitra">
<emphasis box="[324,422,462,486]" italics="true" pageId="18" pageNumber="149">P. lalitra</emphasis>
</taxonomicName>
externally, but differs in genitalic features in having shorter gonopods, a weaker hypandrial apron and lacking the mucro on the ventrolateral margin of the aedeagus (
<figureCitation box="[328,458,526,551]" captionStart="Figs 2830" captionStartId="17.[182,221,1600,1620]" captionTargetBox="[202,1117,906,1569]" captionTargetId="figure-249@17.[202,1117,906,1569]" captionTargetPageId="17" captionText="Figs 2830. Inner ơ genitalia of undescribed taxon A, Ivondro, Madagascar: (28, 29) dorsal and lateral views, (30) aedeagus wing. Note that the ventral area of the hypandrial apron is not illustrated due to the extreme transparency of the sclerite following over-treatment in KOH." figureDoi="http://doi.org/10.5281/zenodo.7911685" httpUri="https://zenodo.org/record/7911685/files/figure.png" pageId="18" pageNumber="149">Figs 2830</figureCitation>
). The single specimen available is in rather poor condition, and the genitalia were left too long in KOH during extractive treatment, resulting in the genitalic sclerites becoming largely translucent and difficult to view. Consequently, this taxon is left undescribed until further material becomes available.
</paragraph>
<paragraph blockId="18.[144,1104,694,1079]" box="[505,742,694,719]" pageId="18" pageNumber="149">Undescribed taxon B</paragraph>
<paragraph blockId="18.[144,1104,694,1079]" pageId="18" pageNumber="149">
A single female from the lush forest on Montagne dAmbre,
<collectingRegion box="[872,1007,734,759]" country="Madagascar" name="Antsiranana" pageId="18" pageNumber="149">Antsiranana</collectingRegion>
(Diego- Suarez;
<date box="[234,422,766,791]" pageId="18" pageNumber="149" value="1958-11-23" valueMax="1958-12-04" valueMin="1958-11-23">23.xi4.xii.1958</date>
, B. Stuckenberg, NMSA) appears to represent a new species. For want of more material, it is left undescribed. It resembles
<taxonomicName authorityName="Davies &amp; Miller" authorityYear="2008" box="[824,907,798,822]" class="Insecta" family="Lauxaniidae" genus="Parapachycerina" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="149" phylum="Arthropoda" rank="species" species="talea">
<emphasis box="[824,907,798,822]" italics="true" pageId="18" pageNumber="149">P. talea</emphasis>
</taxonomicName>
in being a robust, deep orange
<taxonomicName authorityName="Stuckenberg" authorityYear="1971" box="[288,482,830,854]" class="Insecta" family="Lauxaniidae" genus="Parapachycerina" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis box="[288,482,830,854]" italics="true" pageId="18" pageNumber="149">Parapachycerina</emphasis>
</taxonomicName>
with very lightly infuscated wings. It differs, however, in lacking the posterior extension from the ocellar spot, and in lacking a profemoral ctenidium (a feature which also separates it from the smaller
<taxonomicName authorityName="Davies &amp; Miller" authorityYear="2008" box="[836,935,894,918]" class="Insecta" family="Lauxaniidae" genus="Parapachycerina" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="149" phylum="Arthropoda" rank="species" species="lalitra">
<emphasis box="[836,935,894,918]" italics="true" pageId="18" pageNumber="149">P. lalitra</emphasis>
</taxonomicName>
). Other salient features of this specimen are: small ocellar triangle (with sides approximately equal); blackish ocellar spot; postpedicel mostly yellow but darkened apically; sides of notopleuron strongly infuscated; dark orange medial stripe on mesonotum; and 2 sternopleural bristles. The isolated Amber Mt (a forest-clad late Tertiary volcano) is well-known as a site of local endemism in
<collectingCountry box="[565,702,1054,1079]" name="Madagascar" pageId="18" pageNumber="149">Madagascar</collectingCountry>
.
</paragraph>
<paragraph blockId="18.[144,1104,1126,1671]" box="[505,742,1126,1151]" pageId="18" pageNumber="149">Undescribed taxon C</paragraph>
<paragraph blockId="18.[144,1104,1126,1671]" pageId="18" pageNumber="149">
<materialsCitation collectingDate="1973-11-15" collectionCode="MNHN" collectorName="La Grille &amp; L. Matile" country="Comoros" county="Guiri" elevation="875" location="The" municipality="La Grille" pageId="18" pageNumber="149" specimenCount="1" stateProvince="Grande Comore">
<location LSID="urn:lsid:plazi:treatment:03AEFB40FFF91C72FE4AFE5EBED3F978:8ED81C8DFFFB1C72FF28FB0EBC91FB58" box="[170,214,1166,1191]" country="Comoros" county="Guiri" municipality="La Grille" name="The" pageId="18" pageNumber="149" stateProvince="Grande Comore">The</location>
late
<location LSID="urn:lsid:plazi:treatment:03AEFB40FFF91C72FE4AFE5EBED3F978:8ED81C8DFFFB1C72FE8FFB0EBDD7FB58" box="[269,400,1166,1191]" country="Comoros" county="Guiri" municipality="La Grille" name="Loic Matile" pageId="18" pageNumber="149" stateProvince="Grande Comore">Loïc Matile</location>
collected a distinctive female specimen from
<collectingRegion box="[917,1100,1166,1191]" country="Comoros" name="Grande Comore" pageId="18" pageNumber="149">Grande Comore</collectingRegion>
, the largest island in the
<collectingCountry box="[404,507,1198,1223]" name="Comoros" pageId="18" pageNumber="149">Comoros</collectingCountry>
archipelago (
<collectingRegion box="[656,839,1198,1223]" country="Comoros" name="Grande Comore" pageId="18" pageNumber="149">Grande Comore</collectingRegion>
,
<collectorName box="[849,950,1198,1223]" pageId="18" pageNumber="149">
<collectingMunicipality box="[849,950,1198,1223]" pageId="18" pageNumber="149">La Grille</collectingMunicipality>
</collectorName>
(
<collectingCounty box="[964,1028,1198,1223]" pageId="18" pageNumber="149">Guiri</collectingCounty>
),
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.75" metricValueMax="9.0" metricValueMin="8.5" pageId="18" pageNumber="149" unit="m" value="875.0" valueMax="900.0" valueMin="850.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.75" metricValueMax="9.0" metricValueMin="8.5" pageId="18" pageNumber="149" unit="m" value="875.0" valueMax="900.0" valueMin="850.0">850 900 m</elevation>
</quantity>
,
<date box="[231,353,1230,1255]" pageId="18" pageNumber="149" value="1973-11-15">
<collectingDate box="[231,353,1230,1255]" pageId="18" pageNumber="149" value="1973-11-15">15.xi.1973</collectingDate>
</date>
,
<collectorName box="[366,474,1230,1255]" pageId="18" pageNumber="149">L. Matile</collectorName>
,
<collectionCode box="[486,577,1230,1255]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="18" pageNumber="149">MNHN</collectionCode>
). Immediately notable about this specimen are the large black spot on the mesopleuron, the weakly fumose anterior part of the wings, three broad, black mesoscutal vittae, and the black marks on the lateral sides of the occiput. The profemoral ctenidium is present. The arrangement, size and orientation of cephalic setation are as per the genus
</materialsCitation>
.
</paragraph>
<paragraph blockId="18.[144,1104,1126,1671]" pageId="18" pageNumber="149">
This specimen definitely seems to represent a new species, but until further specimens are procured, in particular males, it is left undescribed. The presence of
<taxonomicName authorityName="Stuckenberg" authorityYear="1971" box="[912,1104,1422,1446]" class="Insecta" family="Lauxaniidae" genus="Parapachycerina" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis box="[912,1104,1422,1446]" italics="true" pageId="18" pageNumber="149">Parapachycerina</emphasis>
</taxonomicName>
on Grande Comore hints at the possibility that the genus may yet be found on the
<collectingCountry box="[144,267,1486,1511]" name="Seychelles" pageId="18" pageNumber="149">Seychelles</collectingCountry>
. Based on superficial colouration similarity and the presence of a profemoral ctenidium, this taxon may be the sister-species of
<taxonomicName authorityName="Davies &amp; Miller" authorityYear="2008" box="[706,837,1518,1542]" class="Insecta" family="Lauxaniidae" genus="Parapachycerina" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="149" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis box="[706,837,1518,1542]" italics="true" pageId="18" pageNumber="149">P. infuscata</emphasis>
</taxonomicName>
. Interestingly, the three species with profemoral ctenidia are the two Malagasy species and
<taxonomicName authorityName="Davies &amp; Miller" authorityYear="2008" box="[901,1032,1550,1574]" class="Insecta" family="Lauxaniidae" genus="Parapachycerina" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="149" phylum="Arthropoda" rank="species" species="infuscata">
<emphasis box="[901,1032,1550,1574]" italics="true" pageId="18" pageNumber="149">P. infuscata</emphasis>
</taxonomicName>
of the African mainland. Taxon C is a geographical annectant step joining these ctenidiabearing
<taxonomicName authorityName="Stuckenberg" authorityYear="1971" box="[237,431,1614,1638]" class="Insecta" family="Lauxaniidae" genus="Parapachycerina" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="149" phylum="Arthropoda" rank="genus">
<emphasis box="[237,431,1614,1638]" italics="true" pageId="18" pageNumber="149">Parapachycerina</emphasis>
</taxonomicName>
taxa. Whether this is a coincidence, or represents cognate phylogenetic affinity, is still to be determined.
</paragraph>
</subSubSection>
</treatment>
</document>