249 lines
35 KiB
XML
249 lines
35 KiB
XML
<document id="F4A069A6355ACF02E95F0D99CB49187A" ID-DOI="10.5281/zenodo.5301059" ID-ISSN="0374-1036" ID-Zenodo-Dep="5301059" ID-ZooBank="EA465CB7-1796-4656-9D14-EF0964899EDAD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630099098575" checkinUser="marcus" docAuthor="Makranczy, György" docDate="2014" docId="03AECB0EDC49FFFBFEBB338C5389974E" docLanguage="en" docName="ActaEntMusNatPra.2014.54.2.539-554.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Thinodromus velutinus Makranczy 2014, comb. nov." docType="treatment" docVersion="3" lastPageNumber="547" masterDocId="FF97B376DC4CFFF3FFF33756520A915D" masterDocTitle="Review of the Thinodromus circulus species group (Coleoptera: Staphylinidae: Oxytelinae)" masterLastPageNumber="554" masterPageNumber="539" pageNumber="544" updateTime="1699149139555" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5E07043AF5372E16E22E260D99496193">Review of the Thinodromus circulus species group (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<mods:name id="6534EC91AE37CB7D789AFC5ACD7C390C" type="personal">
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<mods:namePart id="FD14A62F15EEC58A2127BE9909F9BBBD">Makranczy, György</mods:namePart>
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<mods:title id="2D2C7B915AB732AD2655DEED55306295">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="D3057BBA18702BAB25B308E42AEAD008">
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<mods:date id="740C743C75831692E77B27589F551A95">2014</mods:date>
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<mods:number id="187B03DB477F028953D0DE2CF63C1134">2014-12-15</mods:number>
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<mods:number id="8B2E17FD7F71485EBF92CF0171B54F75">54</mods:number>
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<treatment id="03AECB0EDC49FFFBFEBB338C5389974E" LSID="urn:lsid:plazi:treatment:03AECB0EDC49FFFBFEBB338C5389974E" httpUri="http://treatment.plazi.org/id/03AECB0EDC49FFFBFEBB338C5389974E" lastPageId="8" lastPageNumber="547" pageId="5" pageNumber="544">
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<subSubSection id="C31D2993DC49FFF6FEBB338C516195AC" box="[328,875,1242,1265]" pageId="5" pageNumber="544" type="nomenclature">
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<paragraph id="8BB87A18DC49FFF6FEBB338C516195AC" blockId="5.[328,875,1242,1265]" box="[328,875,1242,1265]" pageId="5" pageNumber="544">
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<heading id="D0F0CD74DC49FFF6FEBB338C516195AC" bold="true" box="[328,875,1242,1265]" centered="true" fontSize="8" level="2" pageId="5" pageNumber="544" reason="3">
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<emphasis id="B973A60ADC49FFF6FEBB338C516195AC" bold="true" box="[328,875,1242,1265]" pageId="5" pageNumber="544">
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<taxonomicName id="4C07019BDC49FFF6FEBB338C50E395AC" ID-CoL="56H6H" authority="(Sharp, 1887)" authorityName="Makranczy" authorityYear="2014" baseAuthorityName="Sharp" baseAuthorityYear="1887" box="[328,745,1242,1265]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="544" phylum="Arthropoda" rank="species" species="velutinus" status="comb. nov.">
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<emphasis id="B973A60ADC49FFF6FEBB338C504395AC" bold="true" box="[328,585,1242,1265]" italics="true" pageId="5" pageNumber="544">Thinodromus velutinus</emphasis>
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(
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<bibRefCitation id="EF9607E9DC49FFF6FDA4338C50E895AC" author="SHARP D. S." box="[599,738,1242,1265]" pageId="5" pageNumber="544" pagination="673 - 824" refId="ref7488" refString="SHARP D. S. 1887: Staphylinidae. Pp. 673 - 824, pls. 17 - 19. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. I. Part 2. Taylor & Francis, London, xvi + 824 pp., 19 pls." type="book chapter" year="1887">Sharp, 1887</bibRefCitation>
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||
)
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</taxonomicName>
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||
<taxonomicNameLabel id="A2401B71DC49FFF6FD03338C516195AC" box="[752,875,1242,1265]" pageId="5" pageNumber="544" rank="species">comb. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31D2993DC49FFF6FE0033AF50CA9451" box="[499,704,1273,1292]" pageId="5" pageNumber="544" type="description">
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<paragraph id="8BB87A18DC49FFF6FE0033AF50CA9451" blockId="5.[113,1077,1273,1352]" box="[499,704,1273,1292]" pageId="5" pageNumber="544">
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(
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<figureCitation id="133C669DDC49FFF6FE0B33AF50509451" box="[504,602,1273,1292]" captionStart="Figs 5–14. 5–10" captionStartId="4.[151,188,1497,1516]" captionTargetBox="[151,1100,223,1466]" captionTargetId="figure-9@4.[151,1103,221,1468]" captionTargetPageId="4" captionText="Figs 5–14. 5–10 – Thinodromus circulus (Bernhauer, 1922) (5 – mesotarsus; 6 – male sternite VIII; 7 – male tergite IX; 8 – male sternite IX; 9 – male tergite X; 10 – spermatheca); 11–12 – T. velutinus (Sharp, 1887) (11 – tergite X; 12 – spermatheca); 13–14 – T. chagosanus (Bernhauer, 1922) (13 – mesotarsus; 14 – platelike armature in hypopharynx). Scale bar: 0.06 mm for 14; 0.07 mm for 5, 13; 0.1 mm for 10–12; 0.12 mm for 8–9; 0.14 mm for 6–7." figureDoi="http://doi.org/10.5281/zenodo.5301063" httpUri="https://zenodo.org/record/5301063/files/figure.png" pageId="5" pageNumber="544">Figs 11–12</figureCitation>
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,
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<figureCitation id="133C669DDC49FFF6FD9033AF50909451" box="[611,666,1273,1292]" captionStart-0="Figs 15–19" captionStart-1="Figs 20–24. 20" captionStartId-0="6.[151,187,1483,1502]" captionStartId-1="7.[113,150,1480,1499]" captionTargetBox-0="[163,1115,220,1467]" captionTargetBox-1="[127,1077,222,1463]" captionTargetId-0="figure-10@6.[163,1116,219,1467]" captionTargetId-1="figure-9@7.[125,1078,219,1466]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Figs 15–19. Thinodromus velutinus (Sharp, 1887),female, SEM.15 – forebody; 16 – antennomeres 3–10; 17 – head and pronotum; 18 – mesotarsus; 19 – elytron. Scale bar:0.2 mm for 18; 0.25 mm for 16; 0.4 mm for 17, 19; 1.0 mm for 15." captionText-1="Figs 20–24. 20 – Thinodromus velutinus (Sharp, 1887), tergite VIII; 21–22 – T. circulus (Bernhauer, 1922): 21 – tergite VIII, 22 – aedeagus; 23–24 – T. chagosanus (Bernhauer, 1922): 23 – male sternite IX, 24 – aedeagus. Scale bar: 0.1 mm for 20–21; 0.15 mm for 22–24." figureDoi-0="http://doi.org/10.5281/zenodo.5301065" figureDoi-1="http://doi.org/10.5281/zenodo.5301069" httpUri-0="https://zenodo.org/record/5301065/files/figure.png" httpUri-1="https://zenodo.org/record/5301069/files/figure.png" pageId="5" pageNumber="544">15–20</figureCitation>
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,
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<figureCitation id="133C669DDC49FFF6FD5033AF50B39451" box="[675,697,1273,1292]" captionStart="Figs 31–35. 31" captionStartId="11.[113,150,1527,1546]" captionTargetBox="[114,1064,219,1466]" captionTargetId="figure-9@11.[113,1065,219,1466]" captionTargetPageId="11" captionText="Figs 31–35. 31 – Thinodromus velutinus (Sharp, 1887), female ringstructure. 32 – T. circulus (Bernhauer, 1922), female ringstructure. 33–34 – T. palustris (Bernhauer, 1922): 33 – male sternite IX, 34 – aedeagus. 35 – T.arcitenens (Fauvel, 1905), aedeagus. Scale bar: 0.1 mm for 31; 0.12 mm for 32, 35; 0.15 mm for 33; 0.2 mm for 34." figureDoi="http://doi.org/10.5281/zenodo.5301075" httpUri="https://zenodo.org/record/5301075/files/figure.png" pageId="5" pageNumber="544">31</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31D2993DC49FFF6FF82324D517D9415" pageId="5" pageNumber="544" type="reference_group">
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<paragraph id="8BB87A18DC49FFF6FF82324D517D9415" blockId="5.[113,1077,1273,1352]" pageId="5" pageNumber="544">
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<treatmentCitationGroup id="AB175D36DC49FFF6FF82324D517D9415" pageId="5" pageNumber="544">
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<treatmentCitation id="0AA65C09DC49FFF6FF82324D53DA9473" author="SHARP D. S." box="[113,464,1307,1326]" page="700" pageId="5" pageNumber="544" year="1887">
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<taxonomicName id="4C07019BDC49FFF6FF82324D53DA9473" ID-CoL="599Y3" authority="Sharp, 1887: 700" authorityName="Sharp" authorityPageNumber="700" authorityYear="1887" box="[113,464,1307,1326]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="544" phylum="Arthropoda" rank="species" species="velutinus">
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<emphasis id="B973A60ADC49FFF6FF82324D53329473" box="[113,312,1307,1326]" italics="true" pageId="5" pageNumber="544">Trogophloeus velutinus</emphasis>
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<bibRefCitation id="EF9607E9DC49FFF6FECD324D53DA9473" author="SHARP D. S." box="[318,464,1307,1326]" pageId="5" pageNumber="544" pagination="673 - 824" refId="ref7488" refString="SHARP D. S. 1887: Staphylinidae. Pp. 673 - 824, pls. 17 - 19. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. I. Part 2. Taylor & Francis, London, xvi + 824 pp., 19 pls." type="book chapter" year="1887">Sharp, 1887: 700</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
(original description),
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||
<treatmentCitation id="0AA65C09DC49FFF6FD66324D51C39473" author="BERNHAUER M. & SCHUBERT K." box="[661,969,1307,1326]" page="105" pageId="5" pageNumber="544" year="1911">
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||
<bibRefCitation id="EF9607E9DC49FFF6FD66324D51AB9473" author="BERNHAUER M. & SCHUBERT K." box="[661,929,1307,1326]" pageId="5" pageNumber="544" pagination="87 - 190" refId="ref7077" refString="BERNHAUER M. & SCHUBERT K. 1911: Staphylinidae II. In: SCHENKLING S. (ed.): Coleopterorum Catalogus, Pars 29. W. Junk, Berlin, Germany, pp. 87 - 190." type="book chapter" year="1911">BERNHAUER & SCHUBERT (1911)</bibRefCitation>
|
||
: 105
|
||
</treatmentCitation>
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||
(catalogue).
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||
<treatmentCitation id="0AA65C09DC49FFF6FF82326353EB9415" author="HERMAN L. H." box="[113,481,1333,1352]" page="394" pageId="5" pageNumber="544" year="1970">
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||
<taxonomicName id="4C07019BDC49FFF6FF82326353B39415" authority=": HERMAN (1970)" authorityName="HERMAN" authorityYear="1970" box="[113,441,1333,1352]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="544" phylum="Arthropoda" rank="species" species="velutinus">
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<emphasis id="B973A60ADC49FFF6FF823263532D9415" box="[113,295,1333,1352]" italics="true" pageId="5" pageNumber="544">Carpelimus velutinus</emphasis>
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||
:
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<bibRefCitation id="EF9607E9DC49FFF6FEC0326353B39415" author="HERMAN L. H." box="[307,441,1333,1352]" pageId="5" pageNumber="544" pagination="343 - 454" refId="ref7167" refString="HERMAN L. H. 1970: Phylogeny and reclassi? cation of the genera of the rove-beetle subfamily Oxytelinae of the World (Coleoptera, Staphylinidae). Bulletin of the American Museum of Natural History 142 (5): 343 - 454." type="journal article" year="1970">HERMAN (1970)</bibRefCitation>
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</taxonomicName>
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||
: 394
|
||
</treatmentCitation>
|
||
(catalogue),
|
||
<treatmentCitation id="0AA65C09DC49FFF6FDA2326351019415" author="HERMAN L. H." box="[593,779,1333,1352]" page="1709" pageId="5" pageNumber="544" year="2001">
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<bibRefCitation id="EF9607E9DC49FFF6FDA2326350DD9415" author="HERMAN L. H." box="[593,727,1333,1352]" pageId="5" pageNumber="544" pagination="1 - 4218" refId="ref7213" refString="HERMAN L. H. 2001: Catalog of the Staphylinidae (Insecta: Coleoptera). 1785 to the end of the second Millennium. Volume I - VII. Bulletin of the American Museum of Natural History 265: 1 - 4218." type="journal article" year="2001">HERMAN (2001)</bibRefCitation>
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||
: 1709
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</treatmentCitation>
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||
(catalogue).
|
||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31D2993DC49FFF6FF823209514194A5" pageId="5" pageNumber="544" type="materials_examined">
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<paragraph id="8BB87A18DC49FFF6FF823209516394D0" blockId="5.[113,1090,1375,1528]" pageId="5" pageNumber="544">
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<emphasis id="B973A60ADC49FFF6FF82320952E3942F" bold="true" box="[113,233,1375,1394]" pageId="5" pageNumber="544">
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<typeStatus id="54BCC4BADC49FFF6FF8232095294942F" box="[113,158,1375,1394]" pageId="5" pageNumber="544">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F3103A88DC49FFF6FF1E32095339942F" box="[237,307,1375,1394]" name="Panama" pageId="5" pageNumber="544">Panama</collectingCountry>
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,
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<collectingRegion id="49C3B4FADC49FFF6FEC83209538E942F" box="[315,388,1375,1394]" country="Panama" name="Chiriqui" pageId="5" pageNumber="544">Chiriquí</collectingRegion>
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, Volcán de
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<collectingRegion id="49C3B4FADC49FFF6FE1032095020942F" box="[483,554,1375,1394]" country="Panama" name="Chiriqui" pageId="5" pageNumber="544">Chiriquí</collectingRegion>
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[nowadays Barú]. The
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||
<typeStatus id="54BCC4BADC49FFF6FD1C3209511E942F" box="[751,788,1375,1394]" pageId="5" pageNumber="544">type</typeStatus>
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locality probably refers to SE slope of the vocano above Boquete, approx. 8°47ƍ40ƎN 82°26ƍ50ƎW – 8°49ƍ50ƎN 82°29ƍ23ƎW.
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||
</paragraph>
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||
<paragraph id="8BB87A18DC49FFF6FF8232C3514194A5" blockId="5.[113,1090,1375,1528]" pageId="5" pageNumber="544">
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<emphasis id="B973A60ADC49FFF6FF8232C3534694F5" bold="true" box="[113,332,1429,1448]" pageId="5" pageNumber="544">Type material examined.</emphasis>
|
||
<typeStatus id="54BCC4BADC49FFF6FEBC32C353AB94FB" box="[335,417,1429,1448]" pageId="5" pageNumber="544" type="holotype">HOLOTYPE</typeStatus>
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||
: J, glued on mounting card ‘
|
||
<taxonomicName id="4C07019BDC49FFF6FD7E32C3510D94F5" box="[653,775,1429,1448]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="544" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
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||
; velutinus.Type; D.S.;
|
||
<collectionCode id="ED16E2DDDC49FFF6FC3032C351D894F5" box="[963,978,1429,1448]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="544" type="Museum">V</collectionCode>
|
||
.de
|
||
<collectingRegion id="49C3B4FADC49FFF6FC0332C3563194F5" box="[1008,1083,1429,1448]" country="Panama" name="Chiriqui" pageId="5" pageNumber="544">Chiriqui</collectingRegion>
|
||
.;
|
||
<quantity id="4CFFD7FDDC49FFF6FF8232F952C8949F" box="[113,194,1455,1474]" metricMagnitude="2" metricUnit="m" metricValue="9.150096" metricValueMax="18.288" metricValueMin="0.012192" pageId="5" pageNumber="544" unit="ft" value="3002.0" valueMax="6000.0" valueMin="4.0">4-6000ft.</quantity>
|
||
Champion \ Type; Holo- [orange framed disc] \
|
||
<collectionCode id="ED16E2DDDC49FFF6FDAF32F95061949F" box="[604,619,1455,1474]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="544" type="Museum">V</collectionCode>
|
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. de
|
||
<collectingRegion id="49C3B4FADC49FFF6FD7F32F950D2949F" box="[652,728,1455,1474]" country="Panama" name="Chiriqui" pageId="5" pageNumber="544">Chiriqui</collectingRegion>
|
||
,;
|
||
<quantity id="4CFFD7FDDC49FFF6FD1032F9516F949F" box="[739,869,1455,1474]" metricMagnitude="3" metricUnit="m" metricValue="1.524" metricValueMax="1.8288" metricValueMin="1.2192" pageId="5" pageNumber="544" unit="ft" value="5000.0" valueMax="6000.0" valueMin="4000.0">
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<geoCoordinate id="EE331CDFDC49FFF6FCE632F95141949F" box="[789,843,1455,1474]" degrees="6,000" direction="west" orientation="longitude" pageId="5" pageNumber="544" precision="55" value="-6.0">-6,000</geoCoordinate>
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||
ft.
|
||
</quantity>
|
||
; Champion. \ B.C.A. Col. I. 2.;
|
||
<taxonomicName id="4C07019BDC49FFF6FF53329C53139480" box="[160,281,1482,1501]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="544" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
|
||
; velutinus,; Sharp. \ Sharp Coll.; 1905.-313. \
|
||
<typeStatus id="54BCC4BADC49FFF6FD5B329C510E9480" box="[680,772,1482,1501]" pageId="5" pageNumber="544" type="holotype">Holotypus</typeStatus>
|
||
;
|
||
<taxonomicName id="4C07019BDC49FFF6FCFE329C518C9480" box="[781,902,1482,1501]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="544" phylum="Arthropoda" rank="genus">Trogophloeus</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C07019BDC49FFF6FC7C329C561D9480" authority="Sharp" authorityName="Sharp" box="[911,1047,1482,1501]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="544" phylum="Arthropoda" rank="species" species="velutinus">velutinus Sharp</taxonomicName>
|
||
; ver. Makranczy, 2012 \
|
||
<taxonomicName id="4C07019BDC49FFF6FEE632B3538194A5" authorityName="Kraatz" authorityYear="1857" box="[277,395,1509,1528]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="544" phylum="Arthropoda" rank="genus">Thinodromus</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C07019BDC49FFF6FE6732B3502094A5" authority="(Sharp)" baseAuthorityName="Sharp" box="[404,554,1509,1528]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="544" phylum="Arthropoda" rank="species" species="velutinus">velutinus (Sharp)</taxonomicName>
|
||
; det. Makranczy, 2012’ (
|
||
<collectionCode id="ED16E2DDDC49FFF6FD0F32B3514B94A5" box="[764,833,1509,1528]" pageId="5" pageNumber="544">BMNH</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF782A90DC4AFFF5FF64329D566D94A4" ID-DOI="http://doi.org/10.5281/zenodo.5301065" ID-Zenodo-Dep="5301065" httpUri="https://zenodo.org/record/5301065/files/figure.png" pageId="6" pageNumber="545" startId="6.[151,187,1483,1502]" targetBox="[163,1115,220,1467]" targetPageId="6">
|
||
<paragraph id="8BB87A18DC4AFFF5FF64329D566D94A4" blockId="6.[151,1127,1483,1529]" pageId="6" pageNumber="545">
|
||
Figs 15–19.
|
||
<taxonomicName id="4C07019BDC4AFFF5FF0F329D50249483" authority="(Sharp, 1887)" authorityName="Makranczy" authorityYear="2014" baseAuthorityName="Sharp" baseAuthorityYear="1887" box="[252,558,1483,1502]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="545" phylum="Arthropoda" rank="species" species="velutinus">
|
||
<emphasis id="B973A60ADC4AFFF5FF0F329D53B29483" box="[252,440,1483,1502]" italics="true" pageId="6" pageNumber="545">Thinodromus velutinus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9607E9DC4AFFF5FE31329D50229483" author="SHARP D. S." box="[450,552,1483,1502]" pageId="6" pageNumber="545" pagination="673 - 824" refId="ref7488" refString="SHARP D. S. 1887: Staphylinidae. Pp. 673 - 824, pls. 17 - 19. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. I. Part 2. Taylor & Francis, London, xvi + 824 pp., 19 pls." type="book chapter" year="1887">Sharp, 1887</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,female, SEM.15 – forebody; 16 – antennomeres 3–10; 17 – head and pronotum; 18 – mesotarsus; 19 – elytron. Scale bar:0.2 mm for 18; 0.25 mm for 16; 0.4 mm for 17, 19; 1.0 mm for 15.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF782A90DC4BFFF4FF82329E53E3974C" ID-DOI="http://doi.org/10.5281/zenodo.5301069" ID-Zenodo-Dep="5301069" httpUri="https://zenodo.org/record/5301069/files/figure.png" pageId="7" pageNumber="546" startId="7.[113,150,1480,1499]" targetBox="[127,1077,222,1463]" targetPageId="7">
|
||
<paragraph id="8BB87A18DC4BFFF4FF82329E53E3974C" blockId="7.[113,1089,1480,1553]" pageId="7" pageNumber="546">
|
||
Figs 20–24. 20 –
|
||
<taxonomicName id="4C07019BDC4BFFF4FEF9329E50479486" authority="(Sharp, 1887)" authorityName="Makranczy" authorityYear="2014" baseAuthorityName="Sharp" baseAuthorityYear="1887" box="[266,589,1480,1499]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="546" phylum="Arthropoda" rank="species" species="velutinus">
|
||
<emphasis id="B973A60ADC4BFFF4FEF9329E53C59486" box="[266,463,1480,1499]" italics="true" pageId="7" pageNumber="546">Thinodromus velutinus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9607E9DC4BFFF4FE28329E504D9486" author="SHARP D. S." box="[475,583,1480,1499]" pageId="7" pageNumber="546" pagination="673 - 824" refId="ref7488" refString="SHARP D. S. 1887: Staphylinidae. Pp. 673 - 824, pls. 17 - 19. In: Biologia Centrali-Americana. Insecta. Coleoptera. Vol. I. Part 2. Taylor & Francis, London, xvi + 824 pp., 19 pls." type="book chapter" year="1887">Sharp, 1887</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, tergite VIII; 21–22 –
|
||
<taxonomicName id="4C07019BDC4BFFF4FCE2329E561B9486" authority="(Bernhauer, 1922)" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[785,1041,1480,1499]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="546" phylum="Arthropoda" rank="species" species="circulus">
|
||
<emphasis id="B973A60ADC4BFFF4FCE2329E51619486" box="[785,875,1480,1499]" italics="true" pageId="7" pageNumber="546">T. circulus</emphasis>
|
||
(Bernhauer, 1922)
|
||
</taxonomicName>
|
||
: 21 – tergite VIII, 22 – aedeagus; 23–24 –
|
||
<taxonomicName id="4C07019BDC4BFFF4FE5932B550C294AB" authority="(Bernhauer, 1922)" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[426,712,1507,1526]" class="Insecta" family="Staphylinidae" genus="Trogophloeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="546" phylum="Arthropoda" rank="species" species="chagosanus">
|
||
<emphasis id="B973A60ADC4BFFF4FE5932B5502E94AB" box="[426,548,1507,1526]" italics="true" pageId="7" pageNumber="546">T. chagosanus</emphasis>
|
||
(Bernhauer, 1922)
|
||
</taxonomicName>
|
||
: 23 – male sternite IX, 24 – aedeagus. Scale bar: 0.1 mm for 20–21; 0.15 mm for 22–24.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31D2993DC44FFFBFF64378A5019940E" pageId="8" pageNumber="547" type="description">
|
||
<paragraph id="8BB87A18DC44FFFBFF64378A5019940E" blockId="8.[151,1128,220,1555]" pageId="8" pageNumber="547">
|
||
<emphasis id="B973A60ADC44FFFBFF64378A534A91AE" bold="true" box="[151,320,220,243]" pageId="8" pageNumber="547">Redescription.</emphasis>
|
||
<emphasis id="B973A60ADC44FFFBFEBB378A53EF91AE" box="[328,485,220,243]" italics="true" pageId="8" pageNumber="547">Measurements</emphasis>
|
||
(in mm, n = 1): HW = 0.37; TW = 0.38; PW = 0.43; SW = 0.46; AW = 0.53; HL = 0.25; EL = 0.115; TL = 0.06; PL = 0.32; SL = 0.47; SC = 0.45; FB = 1.07; BL = 2.12.
|
||
<emphasis id="B973A60ADC44FFFBFE90364A5026906E" box="[355,556,284,307]" italics="true" pageId="8" pageNumber="547">Lustre and colour:</emphasis>
|
||
Body rather dull, especially head and pronotum, due to strong surface sculpture; rest of body with greasy lustre. Head and pronotum blackish dark brown, almost black. Elytra very dark brown, with very slight reddish tint in places. Abdomen blackish dark brown with slight reddish tint at apices of segments. Antennae reddish medium to dark brown, basal part noticeably lighter than apex. Legs and mouthparts reddish medium to dark brown, tibiae and tarsi somewhat lighter then femora.
|
||
<emphasis id="B973A60ADC44FFFBFC0736EA530F90AE" italics="true" pageId="8" pageNumber="547">Shape and sculpture:</emphasis>
|
||
Forebody as in
|
||
<figureCitation id="133C669DDC44FFFBFE43368A53F590AE" box="[432,511,476,499]" captionStart="Figs 15–19" captionStartId="6.[151,187,1483,1502]" captionTargetBox="[163,1115,220,1467]" captionTargetId="figure-10@6.[163,1116,219,1467]" captionTargetPageId="6" captionText="Figs 15–19. Thinodromus velutinus (Sharp, 1887),female, SEM.15 – forebody; 16 – antennomeres 3–10; 17 – head and pronotum; 18 – mesotarsus; 19 – elytron. Scale bar:0.2 mm for 18; 0.25 mm for 16; 0.4 mm for 17, 19; 1.0 mm for 15." figureDoi="http://doi.org/10.5281/zenodo.5301065" httpUri="https://zenodo.org/record/5301065/files/figure.png" pageId="8" pageNumber="547">Fig. 15</figureCitation>
|
||
. Head (
|
||
<figureCitation id="133C669DDC44FFFBFDA3368A509490AE" box="[592,670,476,499]" captionStart="Figs 15–19" captionStartId="6.[151,187,1483,1502]" captionTargetBox="[163,1115,220,1467]" captionTargetId="figure-10@6.[163,1116,219,1467]" captionTargetPageId="6" captionText="Figs 15–19. Thinodromus velutinus (Sharp, 1887),female, SEM.15 – forebody; 16 – antennomeres 3–10; 17 – head and pronotum; 18 – mesotarsus; 19 – elytron. Scale bar:0.2 mm for 18; 0.25 mm for 16; 0.4 mm for 17, 19; 1.0 mm for 15." figureDoi="http://doi.org/10.5281/zenodo.5301065" httpUri="https://zenodo.org/record/5301065/files/figure.png" pageId="8" pageNumber="547">Fig. 17</figureCitation>
|
||
) quite transverse, with large eyes and well developed, bulging temples approximately half the size of eyes. Neck well delineated (by different, alveolate microsculpture) but without distinct transversal groove. Antennae (
|
||
<figureCitation id="133C669DDC44FFFBFBC8354A52B9930E" captionStart="Figs 15–19" captionStartId="6.[151,187,1483,1502]" captionTargetBox="[163,1115,220,1467]" captionTargetId="figure-10@6.[163,1116,219,1467]" captionTargetPageId="6" captionText="Figs 15–19. Thinodromus velutinus (Sharp, 1887),female, SEM.15 – forebody; 16 – antennomeres 3–10; 17 – head and pronotum; 18 – mesotarsus; 19 – elytron. Scale bar:0.2 mm for 18; 0.25 mm for 16; 0.4 mm for 17, 19; 1.0 mm for 15." figureDoi="http://doi.org/10.5281/zenodo.5301065" httpUri="https://zenodo.org/record/5301065/files/figure.png" pageId="8" pageNumber="547">Fig. 16</figureCitation>
|
||
) rather short, antennomere 4 as long as broad, antennomere 5 1.10× longer than broad, antennomeres 9–10 1.18–1.21× broader than long. Pronotum strongly transverse, anterior corners broadly but not evenly rounded; corner still feebly marked. Posterior half of sides quite straight, posterior corners rounded and strongly obtuse-angled. Horseshoe-shaped impression slightly marked except posterior/median part where rather impressed; slight lateral depressions connected by it. Middle of disc bearing pair of shallow depressions. Marginal bead on pronotum from dorsal view seems absent, only traces on hind margin. Elytra (
|
||
<figureCitation id="133C669DDC44FFFBFF18344A5337926E" box="[235,317,796,819]" captionStart="Figs 15–19" captionStartId="6.[151,187,1483,1502]" captionTargetBox="[163,1115,220,1467]" captionTargetId="figure-10@6.[163,1116,219,1467]" captionTargetPageId="6" captionText="Figs 15–19. Thinodromus velutinus (Sharp, 1887),female, SEM.15 – forebody; 16 – antennomeres 3–10; 17 – head and pronotum; 18 – mesotarsus; 19 – elytron. Scale bar:0.2 mm for 18; 0.25 mm for 16; 0.4 mm for 17, 19; 1.0 mm for 15." figureDoi="http://doi.org/10.5281/zenodo.5301065" httpUri="https://zenodo.org/record/5301065/files/figure.png" pageId="8" pageNumber="547">Fig. 19</figureCitation>
|
||
) combined slightly broader than long, only gently dilated towards apex, with a pair of small, round impressions posteriad of scutellum; depressed area on elytron somewhat extending towards centre of disc. Posterior elytral margin (slightly oblique) with very thin bead and in outer 1/3 with small membranous lobe protruding. Apex of abdominal tergite VII with palisade fringe.
|
||
<emphasis id="B973A60ADC44FFFBFE0534CA515A92EE" box="[502,848,924,947]" italics="true" pageId="8" pageNumber="547">Punctation and microsculpture:</emphasis>
|
||
Head and pronotum with dense, deep, more or less umbilicate punctation with acinose microsculpture in between; only scattered tiny spots remaining with lustre. Interspaces only a fraction of puncture diameters, punctation only becoming sparser on middle of discs. Epistomal suture as shiny transversal line with slightly strigulate microsculpture. Elytral punctures smaller and more shallow, quite sparse in comparison, interspaces on average larger than puncture diameters. Surface rather more lustrous, microsculpture between punctures not so strong but noticeable. Abdomen with very small punctures and interspaces with imbricate microsculpture (more or less isodiametric cells) but not strong, so abdomen almost as lustrous as elytra.
|
||
<emphasis id="B973A60ADC44FFFBFC2D33CA566C95EE" box="[990,1126,1180,1203]" italics="true" pageId="8" pageNumber="547">Pubescence:</emphasis>
|
||
Body setation rather short, dense and equal sized on head and pronotum, a little longer on elytra, slightly less dense, with a few larger hairs at certain spots. Hairs very ¿ne but longer on abdomen, about as dense as on elytra; hairs on apices of segments rather long.
|
||
<emphasis id="B973A60ADC44FFFBFF64324A5046946E" box="[151,588,1308,1331]" italics="true" pageId="8" pageNumber="547">Primary and secondary sexual features:</emphasis>
|
||
Female tergite X as in
|
||
<figureCitation id="133C669DDC44FFFBFCB9324A5193946E" box="[842,921,1308,1331]" captionStart="Figs 5–14. 5–10" captionStartId="4.[151,188,1497,1516]" captionTargetBox="[151,1100,223,1466]" captionTargetId="figure-9@4.[151,1103,221,1468]" captionTargetPageId="4" captionText="Figs 5–14. 5–10 – Thinodromus circulus (Bernhauer, 1922) (5 – mesotarsus; 6 – male sternite VIII; 7 – male tergite IX; 8 – male sternite IX; 9 – male tergite X; 10 – spermatheca); 11–12 – T. velutinus (Sharp, 1887) (11 – tergite X; 12 – spermatheca); 13–14 – T. chagosanus (Bernhauer, 1922) (13 – mesotarsus; 14 – platelike armature in hypopharynx). Scale bar: 0.06 mm for 14; 0.07 mm for 5, 13; 0.1 mm for 10–12; 0.12 mm for 8–9; 0.14 mm for 6–7." figureDoi="http://doi.org/10.5281/zenodo.5301063" httpUri="https://zenodo.org/record/5301063/files/figure.png" pageId="8" pageNumber="547">Fig. 11</figureCitation>
|
||
, spermatheca as in
|
||
<figureCitation id="133C669DDC44FFFBFF64326A52EC940E" box="[151,230,1340,1363]" captionStart="Figs 5–14. 5–10" captionStartId="4.[151,188,1497,1516]" captionTargetBox="[151,1100,223,1466]" captionTargetId="figure-9@4.[151,1103,221,1468]" captionTargetPageId="4" captionText="Figs 5–14. 5–10 – Thinodromus circulus (Bernhauer, 1922) (5 – mesotarsus; 6 – male sternite VIII; 7 – male tergite IX; 8 – male sternite IX; 9 – male tergite X; 10 – spermatheca); 11–12 – T. velutinus (Sharp, 1887) (11 – tergite X; 12 – spermatheca); 13–14 – T. chagosanus (Bernhauer, 1922) (13 – mesotarsus; 14 – platelike armature in hypopharynx). Scale bar: 0.06 mm for 14; 0.07 mm for 5, 13; 0.1 mm for 10–12; 0.12 mm for 8–9; 0.14 mm for 6–7." figureDoi="http://doi.org/10.5281/zenodo.5301063" httpUri="https://zenodo.org/record/5301063/files/figure.png" pageId="8" pageNumber="547">Fig. 12</figureCitation>
|
||
, ringstructure as in
|
||
<figureCitation id="133C669DDC44FFFBFE4C326A5005940E" box="[447,527,1340,1363]" captionStart="Figs 31–35. 31" captionStartId="11.[113,150,1527,1546]" captionTargetBox="[114,1064,219,1466]" captionTargetId="figure-9@11.[113,1065,219,1466]" captionTargetPageId="11" captionText="Figs 31–35. 31 – Thinodromus velutinus (Sharp, 1887), female ringstructure. 32 – T. circulus (Bernhauer, 1922), female ringstructure. 33–34 – T. palustris (Bernhauer, 1922): 33 – male sternite IX, 34 – aedeagus. 35 – T.arcitenens (Fauvel, 1905), aedeagus. Scale bar: 0.1 mm for 31; 0.12 mm for 32, 35; 0.15 mm for 33; 0.2 mm for 34." figureDoi="http://doi.org/10.5281/zenodo.5301075" httpUri="https://zenodo.org/record/5301075/files/figure.png" pageId="8" pageNumber="547">Fig. 31</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D2993DC44FFFBFF64320A514194EE" pageId="8" pageNumber="547" type="diagnosis">
|
||
<paragraph id="8BB87A18DC44FFFBFF64320A514194EE" blockId="8.[151,1128,220,1555]" pageId="8" pageNumber="547">
|
||
<emphasis id="B973A60ADC44FFFBFF64320A539A942E" bold="true" box="[151,400,1372,1395]" pageId="8" pageNumber="547">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C07019BDC44FFFBFE66320A5082942E" authorityName="Makranczy" authorityYear="2014" baseAuthorityName="Sharp" baseAuthorityYear="1887" box="[405,648,1372,1395]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="547" phylum="Arthropoda" rank="species" species="velutinus">
|
||
<emphasis id="B973A60ADC44FFFBFE66320A5082942E" box="[405,648,1372,1395]" italics="true" pageId="8" pageNumber="547">Thinodromus velutinus</emphasis>
|
||
</taxonomicName>
|
||
is clearly allied to
|
||
<taxonomicName id="4C07019BDC44FFFBFCBD320A51B7942E" baseAuthorityName="Bernhauer" baseAuthorityYear="1922" box="[846,957,1372,1395]" class="Insecta" family="Staphylinidae" genus="Thinodromus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="547" phylum="Arthropoda" rank="species" species="circulus">
|
||
<emphasis id="B973A60ADC44FFFBFCBD320A51B7942E" box="[846,957,1372,1395]" italics="true" pageId="8" pageNumber="547">T. circulus</emphasis>
|
||
</taxonomicName>
|
||
, but is distinguishable by its strong punctation of the head and pronotum, and consequent great difference in lustre of head and pronotum compared to elytra and abdomen.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D2993DC44FFFBFF6432EA512A948E" box="[151,800,1468,1491]" pageId="8" pageNumber="547" type="distribution">
|
||
<paragraph id="8BB87A18DC44FFFBFF6432EA512A948E" blockId="8.[151,1128,220,1555]" box="[151,800,1468,1491]" pageId="8" pageNumber="547">
|
||
<emphasis id="B973A60ADC44FFFBFF6432EA5320948E" bold="true" box="[151,298,1468,1491]" pageId="8" pageNumber="547">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54BCC4BADC44FFFBFDEE32EA5041948E" box="[541,587,1468,1491]" pageId="8" pageNumber="547">type</typeStatus>
|
||
locality in
|
||
<collectingCountry id="F3103A88DC44FFFBFD3732EA5116948E" box="[708,796,1468,1491]" name="Panama" pageId="8" pageNumber="547">Panamá</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31D2993DC44FFFBFF64328A5389974E" pageId="8" pageNumber="547" type="discussion">
|
||
<paragraph id="8BB87A18DC44FFFBFF64328A5389974E" blockId="8.[151,1128,220,1555]" pageId="8" pageNumber="547">
|
||
<emphasis id="B973A60ADC44FFFBFF64328A530C94AE" bold="true" box="[151,262,1500,1523]" pageId="8" pageNumber="547">Remarks.</emphasis>
|
||
The
|
||
<typeStatus id="54BCC4BADC44FFFBFECF328A539394AE" box="[316,409,1500,1523]" pageId="8" pageNumber="547" type="holotype">holotype</typeStatus>
|
||
is the only known specimen of this species, there are no additional data about its biology.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |