206 lines
21 KiB
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206 lines
21 KiB
XML
<document id="54EA662F71188CB8A9326EF459B4058E" ID-DOI="10.5252/z2016n4a2" ID-ISSN="1638-9387" ID-Zenodo-Dep="4578343" ID-ZooBank="urn:lsid:zoobank.org:pub:BFFD82EF-50C9-42BF-8493-DF57591EA4FF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614790962589" checkinUser="felipe" docAuthor="Pyron, Robert Alexander, Ganesh, Sumaithangi Rajagopalan, Sayyed, Amit, Sharma, Vivek, Wallach, Van & Somaweera, Ruchira" docDate="2016" docId="03A82A478302FF9AFC0BFF62FBBC7EFC" docLanguage="en" docName="Zoosystema.38.4.453-506.pdf" docOrigin="Zoosystema 38 (4)" docSource="http://dx.doi.org/10.5252/z2016n4a2" docStyle="DocumentStyle:0AF8C315773078909029C6FC3CC05C6C.3:Zoosystema.2015-2017.journal_article" docStyleId="0AF8C315773078909029C6FC3CC05C6C" docStyleName="Zoosystema.2015-2017.journal_article" docStyleVersion="3" docTitle="Rhinophis porrectus Wall 1921" docType="treatment" docVersion="4" lastPageNumber="484" masterDocId="FF91523F831DFF85FFCFFFB5FFF97B72" masterDocTitle="A catalogue and systematic overview of the shield-tailed snakes (Serpentes: Uropeltidae)" masterLastPageNumber="506" masterPageNumber="453" pageNumber="484" updateTime="1698879475557" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="37EF7AE454144F63EFBDB597AE27B821">
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<mods:title id="8FD8BFDC6C82AB9CD02CE8011B72D95E">A catalogue and systematic overview of the shield-tailed snakes (Serpentes: Uropeltidae)</mods:title>
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<mods:name id="03356E4B60F708A8CF411869D0D9B51C" type="personal">
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<mods:namePart id="EDDC61E2911D353C89BB97475973AAF7">Pyron, Robert Alexander</mods:namePart>
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<mods:affiliation id="C8178BDECBFA9BFF3FBD0FE930CC4F88">Department of Biological Sciences, The George Washington University, 2029 G St. NW, Washington, D. C. 20052 (United States) rpyron @ colubroid. org</mods:affiliation>
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<mods:nameIdentifier id="56A59BDE1C9C319967233BC84ED955E5" type="email">rpyron@colubroid.org</mods:nameIdentifier>
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<mods:roleTerm id="82A1F29A30D6C6BF2060714B513D05D7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2186971CD59A79A3F41BEE394FD7F415">Ganesh, Sumaithangi Rajagopalan</mods:namePart>
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<mods:affiliation id="7D4859A494A124FC9B521BD933570573">Chennai Snake Park, Rajbhavan Post, Chennai, Tamil Nadu 600 022 (India)</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="5513D011C259AD637D93441574EF519B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E4584728CB68797316D9E1221EAC95CD">Sayyed, Amit</mods:namePart>
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<mods:affiliation id="46FBEEC0685EE57D6A3AF214ABE5E356">Wildlife Protection and Research Society, Satara, Maharashtra 415 001 (India)</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="06997BDB0ED1138A192601F8459BE7B6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="78F3E01F1B55150EFF8C16008440D74F">Sharma, Vivek</mods:namePart>
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<mods:affiliation id="FE64A5B94A9D427385FF9645D55586D0">Sanjeevni Nagar, Jabalpur, Madhya Pradesh 482 003 (India)</mods:affiliation>
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<mods:namePart id="A18C6D0ACBB7EAFC97F970EA874FAF7C">Wallach, Van</mods:namePart>
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<mods:affiliation id="BB2B98D580EEE21DB006C996CAFE3F85">Potter Park, Cambridge, Massachusetts 02138 (United States)</mods:affiliation>
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<mods:namePart id="D459AE395B44E1308B11C2CE32A0F985">Somaweera, Ruchira</mods:namePart>
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<mods:affiliation id="751C0D5D57E4B20D707D14FD4E03AAF1">CSIRO Land and Water, Floreat WA 6014 (Australia) Published on 30 December 2016</mods:affiliation>
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<mods:typeOfResource id="FFE93976181432D2B1232409B45A581A">text</mods:typeOfResource>
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<mods:title id="C2BC3DCB119896185A6D2757E87391B7">Zoosystema</mods:title>
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<mods:part id="EE5F01B256E4F8DD2E06119F19A83BB5">
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<mods:date id="FC63F3E35DFF47895B25E20C22512077">2016</mods:date>
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<mods:number id="3E96C2B979679F423A91C6126A64D608">2016-12-30</mods:number>
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<mods:detail id="AD394CF61EB70650E9EB8550EE065FB1" type="volume">
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<mods:number id="3D7FDF6F463555D76760D94CD5BC6300">38</mods:number>
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<mods:detail id="8C617347E9FAD54C7068DE4CEC7897BA" type="issue">
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<mods:number id="99FA1256730D1968DAB183D697EC4E03">4</mods:number>
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<mods:extent id="7CEB2FF561969D6900A1630E1CA80105" unit="page">
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<mods:start id="740499E495CAE3680FE326DD05060B21">453</mods:start>
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<mods:end id="E484A04B26B6FC13A1827A1749D0CF96">506</mods:end>
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<mods:url id="9BFC8C5D8B098FB7AA246C71DD0E2E51">http://dx.doi.org/10.5252/z2016n4a2</mods:url>
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<mods:classification id="E602DEF8FFDC82F84C368A94864003A4">journal article</mods:classification>
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<mods:identifier id="43A4427FFB59B60891E2CCB827132680" type="DOI">10.5252/z2016n4a2</mods:identifier>
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<mods:identifier id="32C3196F324A0B72377F883B5E82CD67" type="ISSN">1638-9387</mods:identifier>
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<mods:identifier id="74D192175006E146599301153511462C" type="Zenodo-Dep">4578343</mods:identifier>
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<mods:identifier id="1E23B34123ACA188607D99EE1F5DC7F6" type="ZooBank">urn:lsid:zoobank.org:pub:BFFD82EF-50C9-42BF-8493-DF57591EA4FF</mods:identifier>
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</mods:mods>
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<treatment id="03A82A478302FF9AFC0BFF62FBBC7EFC" LSID="urn:lsid:plazi:treatment:03A82A478302FF9AFC0BFF62FBBC7EFC" httpUri="http://treatment.plazi.org/id/03A82A478302FF9AFC0BFF62FBBC7EFC" lastPageNumber="484" pageId="31" pageNumber="484">
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<subSubSection id="C31BC8DA8302FF9AFC0BFF62FAE17B83" box="[964,1304,215,242]" pageId="31" pageNumber="484" type="nomenclature">
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<paragraph id="8BBE9B518302FF9AFC0BFF62FAE17B83" blockId="31.[964,1304,215,242]" box="[964,1304,215,242]" pageId="31" pageNumber="484">
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<heading id="D0F62C3D8302FF9AFC0BFF62FAE17B83" box="[964,1304,215,242]" centered="true" fontSize="9" level="2" pageId="31" pageNumber="484" reason="2">
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<taxonomicName id="4C01E0D28302FF9AFC0BFF62FAE17B83" authority="Wall, 1921" authorityName="Wall" authorityYear="1921" box="[964,1304,215,242]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="porrectus">
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<emphasis id="B97547438302FF9AFC0BFF62FB637B80" bold="true" box="[964,1178,215,242]" italics="true" pageId="31" pageNumber="484">Rhinophis porrectus</emphasis>
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Wall, 1921
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31BC8DA8302FF9AFCE2FEACFB767A43" box="[813,1167,281,305]" pageId="31" pageNumber="484" type="reference_group">
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<paragraph id="8BBE9B518302FF9AFCE2FEACFB767A43" blockId="31.[813,1167,281,305]" box="[813,1167,281,305]" pageId="31" pageNumber="484">
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<treatmentCitationGroup id="AB11BC7F8302FF9AFCE2FEACFB767A43" box="[813,1167,281,305]" pageId="31" pageNumber="484">
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<treatmentCitation id="0AA0BD408302FF9AFCE2FEACFB727A43" author="WALL F." box="[813,1163,281,305]" page="35" pageId="31" pageNumber="484" year="1921">
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<taxonomicName id="4C01E0D28302FF9AFCE2FEACFB727A43" authority="Wall, 1921 b: 35" authorityName="Wall" authorityPageNumber="35" authorityYear="1921" box="[813,1163,281,305]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="porrectus">
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<emphasis id="B97547438302FF9AFCE2FEACFC1A7A43" box="[813,995,281,305]" italics="true" pageId="31" pageNumber="484">Rhinophis porrectus</emphasis>
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<bibRefCitation id="EF90E6A08302FF9AFC26FEACFB727A43" author="WALL F." box="[1001,1163,281,305]" pageId="31" pageNumber="484" refId="ref51989" refString="WALL F. 1921 b. - Ophidia taprobanica or the snakes of Ceylon. H. R. Cottle, Colombo, 582 p." type="book" year="1921">Wall, 1921b: 35</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31BC8DA8302FF9AFCE2FEE7FBB17A80" pageId="31" pageNumber="484" type="materials_examined">
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<paragraph id="8BBE9B518302FF9AFCE2FEE7FC387AF6" blockId="31.[813,1455,338,388]" pageId="31" pageNumber="484">
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<materialsCitation id="3B69910C8302FF9AFCE2FEE7FC387AF6" collectionCode="TYPE, MATERIAL, BMNH" pageId="31" pageNumber="484" specimenCode="BMNH 1920.8" specimenCount="1" typeStatus="holotype">
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<collectionCode id="ED1003948302FF9AFCE2FEE7FC3F7A1B" box="[813,966,338,361]" pageId="31" pageNumber="484">TYPE MATERIAL</collectionCode>
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. —
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<typeStatus id="54BA25F38302FF9AFC38FEE7FBA37A18" box="[1015,1114,338,362]" pageId="31" pageNumber="484" type="holotype">Holotype</typeStatus>
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:
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<specimenCode id="DBA7332A8302FF9AFBAAFEE7FAF17A18" box="[1125,1288,338,362]" collectionCode="BMNH" pageId="31" pageNumber="484">BMNH 1920.8</specimenCode>
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.25.1 (=
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<collectionCode id="ED1003948302FF9AFA90FEE7FA567A1B" box="[1375,1455,338,361]" pageId="31" pageNumber="484">BMNH</collectionCode>
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1946.1.16.70).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BBE9B518302FF9AFCE2FE13FBB17A80" blockId="31.[811,1455,421,498]" pageId="31" pageNumber="484">
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<materialsCitation id="3B69910C8302FF9AFCE2FE13FBBD7A80" collectionCode="LOCALITY" country="Sri Lanka" location="On" municipality="Madurankuliya" pageId="31" pageNumber="484" specimenCount="1" stateProvince="Western" typeStatus="holotype">
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<typeStatus id="54BA25F38302FF9AFCE2FE13FCA47AC9" box="[813,861,422,445]" pageId="31" pageNumber="484">TYPE</typeStatus>
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<collectionCode id="ED1003948302FF9AFCADFE1EFC457ACE" box="[866,956,427,444]" pageId="31" pageNumber="484">LOCALITY</collectionCode>
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. —
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<location id="8EDECD8A8302FF9AFC2AFE10FBFF7ACF" LSID="urn:lsid:plazi:treatment:03A82A478302FF9AFC0BFF62FBBC7EFC:8EDECD8A8302FF9AFC2AFE10FBFF7ACF" box="[997,1030,421,445]" country="Sri Lanka" municipality="Madurankuliya" name="On" pageId="31" pageNumber="484" stateProvince="Western">On</location>
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the road
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<locationDeviation id="9A0C82738302FF9AFB92FE10FA877ACF" box="[1117,1406,421,445]" firstLocation="Puttalam" pageId="31" pageNumber="484" secondLocation="Chilaw">between Puttalam and Chilaw</locationDeviation>
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, near
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<collectingMunicipality id="6BDA012B8302FF9AFCE4FE0AFC3B7AA5" box="[811,962,447,471]" pageId="31" pageNumber="484">Madurankuliya</collectingMunicipality>
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<emphasis id="B97547438302FF9AFC04FE0AFC147AA5" box="[971,1005,447,471]" italics="true" pageId="31" pageNumber="484">fide</emphasis>
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<bibRefCitation id="EF90E6A08302FF9AFC3AFE0AFB837AAA" author="WILLEY A." box="[1013,1146,447,472]" pageId="31" pageNumber="484" pagination="81 - 89" refId="ref52407" refString="WILLEY A. 1903. - Some rare snakes of Ceylon. Spolia Zeylanica 1: 81 - 89." type="journal article" year="1903">Willey (1903</bibRefCitation>
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; the original collector), North-
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<collectingRegion id="49C555B38302FF9AFCE2FE6FFC237A80" box="[813,986,474,498]" country="Sri Lanka" name="Western" pageId="31" pageNumber="484">Western province</collectingRegion>
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,
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<collectingCountry id="F316DBC18302FF9AFC2BFE6FFBBD7A80" box="[996,1092,474,498]" name="Sri Lanka" pageId="31" pageNumber="484">Sri Lanka</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31BC8DA8302FF9AFCE2FDA6FB7D798C" pageId="31" pageNumber="484" type="distribution">
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<paragraph id="8BBE9B518302FF9AFCE2FDA6FB7D798C" blockId="31.[811,1457,531,766]" pageId="31" pageNumber="484">
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DISTRIBUTION. — This species is known only from the
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<typeStatus id="54BA25F38302FF9AFA87FDA6FA8B7959" box="[1352,1394,531,555]" pageId="31" pageNumber="484">type</typeStatus>
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specimen collected at Madurankuliya in the lowland dry zone of
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<collectingCountry id="F316DBC18302FF9AFA5CFD98FC937912" name="Sri Lanka" pageId="31" pageNumber="484">Sri Lanka</collectingCountry>
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(near sea level). Some sources (
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<bibRefCitation id="EF90E6A08302FF9AFB54FDFDFAE87912" author="SMITH M. A." box="[1179,1297,584,608]" pageId="31" pageNumber="484" refId="ref51495" refString="SMITH M. A. 1943. - The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. Vol. 3 (Serpentes). Taylor & Francis, London, 583 p." type="book" year="1943">Smith 1943</bibRefCitation>
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) record the
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<typeStatus id="54BA25F38302FF9AFA48FDFDFA487912" box="[1415,1457,584,608]" pageId="31" pageNumber="484">type</typeStatus>
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locality as Maradankadawala, an area further interior in the lowland dry zone, but this is an error. The original collector (
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<bibRefCitation id="EF90E6A08302FF9AFAE5FDC9FA5E79E7" author="WILLEY A." box="[1322,1447,636,661]" pageId="31" pageNumber="484" pagination="81 - 89" refId="ref52407" refString="WILLEY A. 1903. - Some rare snakes of Ceylon. Spolia Zeylanica 1: 81 - 89." type="journal article" year="1903">Willey 1903</bibRefCitation>
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) reports the specimen as originating from Madurankuliya. More recently,
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<bibRefCitation id="EF90E6A08302FF9AFC45FD04FB5879B8" author="KUMARASINGHE A. & PRADEEP W. A. A. D. G. & PEABOTUWAGE P. I. K. & WICKRAMAARACHCHI R. G. A. T. S. & SOMARATHNE S. & PERERA B. N. H. & ABEYAWARDANE U. T. I. & WIJESINGHE M. R. & KARUNARATHNA D. M. S. S." box="[906,1185,689,714]" pageId="31" pageNumber="484" pagination="3 - 20" refId="ref50050" refString="KUMARASINGHE A., PRADEEP W. A. A. D. G., PEABOTUWAGE P. I. K., WICKRAMAARACHCHI R. G. A. T. S., SOMARATHNE S., PERERA B. N. H., ABEYAWARDANE U. T. I., WIJESINGHE M. R. & KARUNARATHNA D. M. S. S. 2013. - An attempt to reduce impacts of limestone quarries through biodiversity assessment and translocation: a case study at the Holcim Limestone Quarry Site in Puttalam, Sri Lanka. Asian Journal of Conservation Biology 2: 3 - 20." type="journal article" year="2013">
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Kumarasinghe
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<emphasis id="B97547438302FF9AFBECFD07FBAC79BB" box="[1059,1109,689,713]" italics="true" pageId="31" pageNumber="484">et al.</emphasis>
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(2013)
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</bibRefCitation>
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reported a specimen from Eluwankulama Aruwakkalu Forest,
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<emphasis id="B97547438302FF9AFB40FD78FB667996" box="[1167,1183,717,740]" italics="true" pageId="31" pageNumber="484">c.</emphasis>
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<quantity id="4CF936B48302FF9AFB69FD79FB1E7996" box="[1190,1255,716,740]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="31" pageNumber="484" unit="km" value="35.0">35 km</quantity>
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North of Puttalam, identified as
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<taxonomicName id="4C01E0D28302FF9AFC65FD53FB87798C" authorityName="Wall" authorityYear="1921" box="[938,1150,742,766]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="porrectus">
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<emphasis id="B97547438302FF9AFC65FD53FBF1798C" box="[938,1032,742,766]" italics="true" pageId="31" pageNumber="484">Rhinophis</emphasis>
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cf.
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<emphasis id="B97547438302FF9AFBE3FD52FB87798C" box="[1068,1150,743,766]" italics="true" pageId="31" pageNumber="484">porrectus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31BC8DA8302FF9AFCE2FC83FB377F5C" pageId="31" pageNumber="484" type="description">
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<paragraph id="8BBE9B518302FF9AFCE2FC83FC44783C" blockId="31.[811,1457,822,1071]" box="[813,957,822,848]" pageId="31" pageNumber="484">DESCRIPTION</paragraph>
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<paragraph id="8BBE9B518302FF9AFCE2FCE0FB377F5C" blockId="31.[811,1457,822,1071]" pageId="31" pageNumber="484">
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The size, scalation, and coloration are indistinguishable from
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<taxonomicName id="4C01E0D28302FF9AFCE4FCC0FC0278FD" authorityName="Muller" authorityYear="1832" box="[811,1019,885,911]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="punctatus">
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<emphasis id="B97547438302FF9AFCE4FCC0FC0278FD" box="[811,1019,885,911]" italics="true" pageId="31" pageNumber="484">Rhinophis punctatus</emphasis>
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</taxonomicName>
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(see below), from which it is distinguishable only by the higher number of ventral scales (281 vs 236-246) and a much thicker body (see
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<bibRefCitation id="EF90E6A08302FF9AFB37FC00FA7978BD" author="SMITH M. A." box="[1272,1408,949,975]" pageId="31" pageNumber="484" refId="ref51495" refString="SMITH M. A. 1943. - The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. Vol. 3 (Serpentes). Taylor & Francis, London, 583 p." type="book" year="1943">Smith 1943</bibRefCitation>
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). As with
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<taxonomicName id="4C01E0D28302FF9AFCADFC60FC1B789C" authorityName="Muller" authorityYear="1832" box="[866,994,981,1006]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="punctatus">
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<emphasis id="B97547438302FF9AFCADFC60FC1B789C" box="[866,994,981,1006]" italics="true" pageId="31" pageNumber="484">R. punctatus</emphasis>
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</taxonomicName>
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(see below), color pattern a blackish-brown dorsum with white margins, a white stripe on either side of the body, and a brown vertebral stripe.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31BC8DA8302FF9AFCE2FBE0FBBC7EFC" pageId="31" pageNumber="484" type="discussion">
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<paragraph id="8BBE9B518302FF9AFCE2FBE0FC687F1F" blockId="31.[811,1457,1109,1422]" box="[813,913,1109,1135]" pageId="31" pageNumber="484">REMARKS</paragraph>
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<paragraph id="8BBE9B518302FF9AFCE2FBC0FBBC7EFC" blockId="31.[811,1457,1109,1422]" pageId="31" pageNumber="484">
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It is unclear whether or not this is a distinct species from
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<taxonomicName id="4C01E0D28302FF9AFCE0FB21FC067FDC" authorityName="Muller" authorityYear="1832" box="[815,1023,1172,1198]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="punctatus">
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<emphasis id="B97547438302FF9AFCE0FB21FC067FDC" box="[815,1023,1172,1198]" italics="true" pageId="31" pageNumber="484">Rhinophis punctatus</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="EF90E6A08302FF9AFBC2FB21FB687FDC" author="SMITH M. A." box="[1037,1169,1172,1198]" pageId="31" pageNumber="484" refId="ref51495" refString="SMITH M. A. 1943. - The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. Vol. 3 (Serpentes). Taylor & Francis, London, 583 p." type="book" year="1943">Smith 1943</bibRefCitation>
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;
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<bibRefCitation id="EF90E6A08302FF9AFB53FB21FAEF7FDD" author="GANS C." box="[1180,1302,1172,1199]" pageId="31" pageNumber="484" pagination="1 - 29" refId="ref48814" refString="GANS C. 1966. - Liste der rezenten Reptilien und Amphibien. Uropeltidae. Das Tierreich 84: 1 - 29." type="journal article" year="1966">Gans 1966</bibRefCitation>
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). We continue to treat it as such, given the large difference in ventral-scale count, the largest such count of any known uropeltid species. It is also geographically distinct from
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<taxonomicName id="4C01E0D28302FF9AFB27FB41FA9E7E7F" authorityName="Muller" authorityYear="1832" box="[1256,1383,1268,1293]" class="Reptilia" family="Uropeltidae" genus="Rhinophis" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="484" phylum="Chordata" rank="species" species="punctatus">
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<emphasis id="B97547438302FF9AFB27FB41FA9E7E7F" box="[1256,1383,1268,1293]" italics="true" pageId="31" pageNumber="484">R. punctatus</emphasis>
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</taxonomicName>
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, being found in the Northwest dry zone vs the central hills in the wet zone (but see
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<bibRefCitation id="EF90E6A08302FF9AFC2AFA86FA9C7E3F" author="KARUNARATHNA D. M. S. S. & AMARASINGHE A. A. T." box="[997,1381,1331,1357]" pageId="31" pageNumber="484" pagination="69 - 76" refId="ref49898" refString="KARUNARATHNA D. M. S. S. & AMARASINGHE A. A. T. 2011. - A preliminary survey of the reptile fauna in Nilgala forest and its vicinity, Monaragala District, Sri Lanka. Taprobanica 3: 69 - 76." type="journal article" year="2011">Karunarathna & Amarasinghe 2011</bibRefCitation>
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). More specimens are needed to examine the distinctiveness of these species. See
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<bibRefCitation id="EF90E6A08302FF9AFC61FAC6FBB87EFC" author="WALL F." box="[942,1089,1395,1422]" pageId="31" pageNumber="484" refId="ref51989" refString="WALL F. 1921 b. - Ophidia taprobanica or the snakes of Ceylon. H. R. Cottle, Colombo, 582 p." type="book" year="1921">Wall (1921b)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |