544 lines
86 KiB
XML
544 lines
86 KiB
XML
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<mods:title id="D2CDF8331F6ADEFB2A788B12AA509189">A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species</mods:title>
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<mods:namePart id="8B06BA2927476AFCB19A56C071993D2C">Le, Son X.</mods:namePart>
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<mods:affiliation id="AA941D281B2835433CE72ACC5F749D6B">Institute of Tropical Ecology, Vietnam-Russia Tropical Centre, 63 Nguyen Van Huyen Rd, Cau Giay District, Hanoi, Vietnam. lesonenv 86 @ gmail. com; https: // orcid. org / 0000 - 0003 - 1648 - 6773 & VNU University of Sciences, Vietnam National University, 334 Nguyen Trai, Thanh Xuan District, Hanoi, Vietnam.</mods:affiliation>
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<mods:namePart id="BC349347F3988C8AEE2645134E97EB4C">Schileyko, Arkady A.</mods:namePart>
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<mods:affiliation id="7F138BFB0FA47803FAECDCD41F252904">Zoological Museum of the Moscow Lomonosov State University, Bolshaya Nikitskaja Str. 2, Moscow, 125009, Russia. schileyko 1965 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6139 - 5240</mods:affiliation>
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<mods:affiliation id="0BBC0C0A797EAD5B192C0CBBB81A8CE9">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Cau Giay District, Hanoi, Vietnam. ducanh. iebr @ gmail. com; https: // orcid. org / 0000 - 0001 - 9273 - 0040 & Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18, Hoangquocviet Rd., Cau Giay District, Hanoi, Vietnam.</mods:affiliation>
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<taxonomicName id="4C3884D5E940FF8EFF3DBA3DFCF6FA91" authority="Attems, 1938" authorityName="Attems" authorityYear="1938" box="[151,786,1316,1343]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="432" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
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<emphasis id="B94C2344E940FF8EFF3DBA3DFD83FA90" bold="true" box="[151,615,1316,1343]" italics="true" pageId="21" pageNumber="432">Scolopocryptops broelemanni esulcata</emphasis>
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<bibRefCitation id="EFA982A7E940FF8EFDC5BA3DFCF6FA91" author="Attems, C." box="[623,786,1316,1342]" pageId="21" pageNumber="432" pagination="187 - 353" refId="ref14199" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in franzosisch Indochina gesammelten Myriopoden. Memoires du Museum national d'histoire naturelle, New Series, 6 (2), 187 - 353." type="journal article" year="1938">Attems, 1938</bibRefCitation>
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<figureCitation id="1303E3D3E940FF8EFF3DBA51FEF0FACD" box="[151,276,1352,1378]" captionStart-0="FIGURES 14" captionStart-1="FIGURES 15" captionStart-2="FIGURE 16" captionStartId-0="23.[152,270,1863,1887]" captionStartId-1="24.[152,270,1593,1617]" captionStartId-2="25.[151,250,1943,1967]" captionTargetBox-0="[151,1434,181,1839]" captionTargetBox-1="[151,1435,184,1566]" captionTargetBox-2="[193,1394,181,1919]" captionTargetId-0="figure-18@23.[151,1436,181,1840]" captionTargetId-1="figure-33@24.[151,1436,181,1569]" captionTargetId-2="figure-18@25.[193,1394,181,1919]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-2="25" captionText-0="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." captionText-1="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." captionText-2="FIGURE 16. Distribution of Scolopocryptops broelemanni esulcata Attems, 1938." figureDoi-0="http://doi.org/10.5281/zenodo.7539972" figureDoi-1="http://doi.org/10.5281/zenodo.7539974" figureDoi-2="http://doi.org/10.5281/zenodo.7539976" httpUri-0="https://zenodo.org/record/7539972/files/figure.png" httpUri-1="https://zenodo.org/record/7539974/files/figure.png" httpUri-2="https://zenodo.org/record/7539976/files/figure.png" pageId="21" pageNumber="432">Figs 14–16</figureCitation>
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<emphasis id="B94C2344E940FF8EFF3DBA94FDC4FA0A" box="[151,544,1421,1445]" italics="true" pageId="21" pageNumber="432">Scolopocryptops broelemanni esulcata</emphasis>
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<bibRefCitation id="EFA982A7E940FF8EFD8DBA94FD01FA0A" author="Attems, C." box="[551,741,1421,1445]" pageId="21" pageNumber="432" pagination="187 - 353" refId="ref14199" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in franzosisch Indochina gesammelten Myriopoden. Memoires du Museum national d'histoire naturelle, New Series, 6 (2), 187 - 353." type="journal article" year="1938">Attems, 1938: 338</bibRefCitation>
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<taxonomicName id="4C3884D5E940FF8EFF3DBAB5FD09FA6B" authority=": Attems, 1953: 138" authorityName="Attems" authorityPageNumber="138" authorityYear="1953" box="[151,749,1451,1476]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="432" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
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<emphasis id="B94C2344E940FF8EFF3DBAB5FDC4FA6B" box="[151,544,1452,1476]" italics="true" pageId="21" pageNumber="432">Scolopocryptops broelemanni esulcata</emphasis>
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<bibRefCitation id="EFA982A7E940FF8EFD84BAB5FD09FA6B" author="Attems, C." box="[558,749,1451,1476]" pageId="21" pageNumber="432" pagination="133 - 230" refId="ref14238" refString="Attems, C. (1953) Myriopoden von Indochina. Expedition von Dr. Dawydoff. C. (1938 - 1939). Memoires du Museum National d'Histoire Naturelle, Nouvelle Serie, Serie A, Zoologie, 5 (3), 133 - 230." type="journal article" year="1953">Attems, 1953: 138</bibRefCitation>
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<emphasis id="B94C2344E940FF8EFF3DBAD2FDE5FA4D" box="[151,513,1482,1506]" italics="true" pageId="21" pageNumber="432">Dinocryptops broelemanni esulcata</emphasis>
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:
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<bibRefCitation id="EFA982A7E940FF8EFDA5BAD3FD39FA4D" author="Schileyko, A. A." box="[527,733,1482,1506]" pageId="21" pageNumber="432" pagination="73 - 87" refId="ref15892" refString="Schileyko, A. A. (1995) The Scolopendromorph centipedes of Vietnam (Chilopoda: Scolopendromorpha). Part 2. Arthropoda Selecta, 4, 73 - 87." type="journal article" year="1995">Schileyko, 1995: 73</bibRefCitation>
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<taxonomicName id="4C3884D5E940FF8EFF3DBAF3FD39F9AE" ID-CoL="7H3CH" authority=": Schileyko, 2007: 92" authorityName=": Schileyko" authorityPageNumber="92" authorityYear="2007" box="[151,733,1513,1537]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="432" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
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<emphasis id="B94C2344E940FF8EFF3DBAF3FDE5F9AE" box="[151,513,1513,1537]" italics="true" pageId="21" pageNumber="432">Dinocryptops broelemanni esulcata</emphasis>
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<bibRefCitation id="EFA982A7E940FF8EFDA5BAF0FD39F9AE" author="Schileyko, A. A." box="[527,733,1513,1537]" pageId="21" pageNumber="432" pagination="71 - 95" refId="ref16026" refString="Schileyko, A. A. (2007) The Scolopendromorph centipedes (Chilopoda) of Vietnam, with contributions to the faunas of Cambodia and Laos. Part 3. Arthropoda Selecta, 16, 71 - 95." type="journal article" year="2007">Schileyko, 2007: 92</bibRefCitation>
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<emphasis id="B94C2344E940FF8EFF3DB936FEA7F9E6" bold="true" box="[151,323,1583,1609]" pageId="21" pageNumber="432">Locus typicus.</emphasis>
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<collectingRegion id="49FC31B4E940FF8EFD8FB929FCE8F9E6" box="[549,780,1583,1610]" country="Vietnam" name="Lam Dong" pageId="21" pageNumber="432">Lam Dong Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BE3652CE940FF8EFCB3B929FC79F9E6" box="[793,925,1583,1610]" pageId="21" pageNumber="432">Lang Biang</collectingMunicipality>
|
||
(= Langbiang)
|
||
<location id="8EE7A98DE940FF8EFBE5B936FB5BF9E6" LSID="urn:lsid:plazi:treatment:03914E40E940FF81FF3DBA3DFDC1FD62:8EE7A98DE940FF8EFBE5B936FB5BF9E6" box="[1103,1215,1583,1609]" country="Vietnam" county="Southern" municipality="Lang Biang" name="Mountain" pageId="21" pageNumber="432" stateProvince="Lam Dong">Mountain</location>
|
||
,
|
||
<elevation id="00151865E940FF8EFB61B936FABAF9E6" box="[1227,1374,1583,1610]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="21" pageNumber="432" unit="m" value="1500.0">
|
||
<quantity id="4CC052B3E940FF8EFB61B936FAC5F9E5" box="[1227,1313,1583,1610]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="21" pageNumber="432" unit="m" value="1500.0">1500 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
, coll.
|
||
<collectorName id="26CD9A80E940FF8EFF3DB94AFED5F9C2" box="[151,305,1619,1645]" pageId="21" pageNumber="432">C. Dawydoff.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C322ACDDE940FF8EFF6DB96EFD24F91A" pageId="21" pageNumber="432" type="materials_examined">
|
||
<paragraph id="8B87FF56E940FF8EFF6DB96EFD24F91A" blockId="21.[151,1437,1583,2042]" pageId="21" pageNumber="432">
|
||
<emphasis id="B94C2344E940FF8EFF6DB96EFED2F93E" bold="true" box="[199,310,1655,1681]" pageId="21" pageNumber="432">Material.</emphasis>
|
||
<materialsCitation id="3B50F50BE940FF8EFE96B96EFD58F91A" ID-GBIF-Occurrence="4015518363" collectionCode="NHMW" location="LAM DONG Province" pageId="21" pageNumber="432" specimenCount="1" specimenCount-adult="1" stateProvince="Lam Dong" typeStatus="syntype">
|
||
<collectingRegion id="49FC31B4E940FF8EFE96B96EFDDBF93E" box="[316,575,1655,1681]" country="Vietnam" name="Lam Dong" pageId="21" pageNumber="432">LAM DONG Province</collectingRegion>
|
||
: collected in locus typicus, (sub)adult
|
||
<typeStatus id="548341F4E940FF8EFC4FB961FBA3F93D" box="[997,1095,1656,1682]" pageId="21" pageNumber="432" type="syntype">syntypes</typeStatus>
|
||
1 and 2 (MY
|
||
<quantity id="4CC052B3E940FF8EFB74B96EFAD5F93E" box="[1246,1329,1655,1681]" metricMagnitude="1" metricUnit="m" metricValue="5.24002" pageId="21" pageNumber="432" unit="in" value="2063.0">2063 in</quantity>
|
||
<collectionCode id="ED296793E940FF8EFA9DB96EFA71F93E" box="[1335,1429,1655,1681]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="432" type="Museum">NHMW</collectionCode>
|
||
) and
|
||
<specimenCount id="9D3E34DFE940FF8EFF6DB982FEF0F91A" box="[199,276,1691,1717]" pageId="21" pageNumber="432" type="adult">1 adult</specimenCount>
|
||
non-type spm (MY
|
||
<quantity id="4CC052B3E940FF8EFE5DB982FDA8F91A" box="[503,588,1691,1717]" metricMagnitude="2" metricUnit="m" metricValue="2.213356" pageId="21" pageNumber="432" unit="in" value="8714.0">8714 in</quantity>
|
||
<collectionCode id="ED296793E940FF8EFDF9B982FD50F91A" box="[595,692,1691,1717]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="432" type="Museum">NHMW</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C322ACDDE940FF8DFF6DB9A6FD4FFBF6" lastPageId="22" lastPageNumber="433" pageId="21" pageNumber="432" type="diagnosis">
|
||
<paragraph id="8B87FF56E940FF8EFF6DB9A6FE6CF952" blockId="21.[151,1437,1583,2042]" pageId="21" pageNumber="432">
|
||
<emphasis id="B94C2344E940FF8EFF6DB9A6FEA6F976" bold="true" box="[199,322,1727,1753]" pageId="21" pageNumber="432">Diagnosis.</emphasis>
|
||
The only Vietnamese
|
||
<taxonomicName id="4C3884D5E940FF8EFD94B9A6FD1DF976" authorityName="Newport" authorityYear="1844" box="[574,761,1727,1753]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="432" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94C2344E940FF8EFD94B9A6FD1DF976" box="[574,761,1727,1753]" italics="true" pageId="21" pageNumber="432">Scolopocryptops</emphasis>
|
||
</taxonomicName>
|
||
with spiracles on LBS 7 (diagnostic character of the former genus
|
||
<taxonomicName id="4C3884D5E940FF8EFF4AB9FDFE9CF952" authorityName="Crabill" authorityYear="1953" box="[224,376,1764,1789]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="432" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94C2344E940FF8EFF4AB9FDFE9CF952" box="[224,376,1764,1789]" italics="true" pageId="21" pageNumber="432">Dinocryptops</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E940FF8EFF6DB81EFBDAF87A" blockId="21.[151,1437,1583,2042]" pageId="21" pageNumber="432">
|
||
<emphasis id="B94C2344E940FF8EFF6DB81EFE15F88E" bold="true" box="[199,497,1799,1825]" pageId="21" pageNumber="432">The original description</emphasis>
|
||
(direct translation by the second author). “Body length
|
||
<quantity id="4CC052B3E940FF8EFB3AB81EFB03F88D" box="[1168,1255,1799,1826]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="21" pageNumber="432" unit="mm" value="24.0">24 mm</quantity>
|
||
. Cephalic plate marginate laterally. Tergite 1 with well-developed anterior transverse suture, anterior margin of tergite 1 covered by cephalic plate. Ultimate tergite lacks median suture, with a single rounded depression in posterior third; this tergite has definite lateral margination and characteristic small pointed process at each posterior corner. Legs 1–21 with one tarsal spur, 1–20 with two tibial spurs, leg 21 with a single tibial one, leg 22 lacking spurs. In all other characters/ moments corresponds to the detailed description of
|
||
<taxonomicName id="4C3884D5E940FF8EFD70B8A2FC67F87A" authorityName="Kraepelin" authorityYear="1903" box="[730,899,1979,2005]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="21" pageNumber="432" phylum="Arthropoda" rank="species" species="broelemanni">
|
||
<emphasis id="B94C2344E940FF8EFD70B8A2FC67F87A" box="[730,899,1979,2005]" italics="true" pageId="21" pageNumber="432">S. broelemanni</emphasis>
|
||
</taxonomicName>
|
||
by Brolemann”.
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E940FF8EFF6DB8C6FC79F856" blockId="21.[151,1437,1583,2042]" box="[199,925,2015,2042]" pageId="21" pageNumber="432">
|
||
<emphasis id="B94C2344E940FF8EFF6DB8C6FE11F856" bold="true" box="[199,501,2015,2041]" pageId="21" pageNumber="432">Composite re-description</emphasis>
|
||
<materialsCitation id="3B50F50BE940FF8EFE56B8C6FC7DF856" ID-GBIF-Occurrence="4015518346" box="[508,921,2015,2042]" collectionCode="NHMW" pageId="21" pageNumber="432" specimenCount="1" typeStatus="syntype">
|
||
of
|
||
<specimenCount id="9D3E34DFE940FF8EFDB1B8F9FD4AF855" box="[539,686,2016,2042]" pageId="21" pageNumber="432" type="generic" typeStatus="syntypes">two syntypes</specimenCount>
|
||
MY 2063 (
|
||
<collectionCode id="ED296793E940FF8EFC84B8C6FC75F856" box="[814,913,2015,2041]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="432" type="Museum">NHMW</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E943FF8DFF6DBF8EFA99FF7A" blockId="22.[151,1437,150,1977]" pageId="22" pageNumber="433">
|
||
Antennae) composed of 17 articles (in not damaged antenna) of them 2 basal ones dorsally (
|
||
<figureCitation id="1303E3D3E943FF8DFB15BF8EFACEFF1E" box="[1215,1322,151,177]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="22" pageNumber="433">Figs 14A</figureCitation>
|
||
,
|
||
<figureCitation id="1303E3D3E943FF8DFA9EBF8FFA81FF1E" box="[1332,1381,150,177]" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="22" pageNumber="433">15A</figureCitation>
|
||
) and 1 basal ventrally (
|
||
<figureCitation id="1303E3D3E943FF8DFEF5BFA2FE2DFF7A" box="[351,457,187,213]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="22" pageNumber="433">Figs 14B</figureCitation>
|
||
,
|
||
<figureCitation id="1303E3D3E943FF8DFE7FBFA3FDE7FF7A" box="[469,515,186,213]" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="22" pageNumber="433">15B</figureCitation>
|
||
) with the single long setae, following articles densely covered by minute setae.
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E943FF8DFF6DBFC6FDB8FEB2" blockId="22.[151,1437,150,1977]" pageId="22" pageNumber="433">
|
||
Cephalic plate virtually as long as wide, very sparsely setose, with very definite lateral and less developed posterior margination (
|
||
<figureCitation id="1303E3D3E943FF8DFE3DBE1AFE0EFEB2" box="[407,490,259,285]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="22" pageNumber="433">Figs 14</figureCitation>
|
||
AF, 15A).
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E943FF8DFF6DBE3EFE22FE02" blockId="22.[151,1437,150,1977]" pageId="22" pageNumber="433">
|
||
Forcipular segment (
|
||
<figureCitation id="1303E3D3E943FF8DFE1BBE3EFDF8FEEE" box="[433,540,295,321]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="22" pageNumber="433">Figs 14B</figureCitation>
|
||
,
|
||
<figureCitation id="1303E3D3E943FF8DFD82BE3FFDBFFEEE" box="[552,603,294,321]" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="22" pageNumber="433">15B</figureCitation>
|
||
): anterior half of coxosternite with a few hardly visible transverse sutures (median one is not recognizable). Anterior margin of coxosternite visibly sclerotised and definitely divided by a median diastema into two standard low lobes, so it does not look straight; process of trochanteroprefemur short; tarsungulum long, pointed.
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E943FF8DFF6DBEAEFEF2FDCE" blockId="22.[151,1437,150,1977]" pageId="22" pageNumber="433">
|
||
Tergite 1 with anterior transverse suture, its anterior margin covered by cephalic plate (
|
||
<figureCitation id="1303E3D3E943FF8DFB3BBEAEFB18FE7E" box="[1169,1276,439,465]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="22" pageNumber="433">Figs 14A</figureCitation>
|
||
,
|
||
<figureCitation id="1303E3D3E943FF8DFAADBEAFFADFFE7E" box="[1287,1339,438,465]" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="22" pageNumber="433">15A</figureCitation>
|
||
). Tergite 2 short, half (or somewhat less) as long as tergite 3. Tergal paramedian sutures not recognizable (
|
||
<figureCitation id="1303E3D3E943FF8DFB4CBEC2FAB4FE5A" box="[1254,1360,475,501]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="22" pageNumber="433">Figs 14C</figureCitation>
|
||
,
|
||
<figureCitation id="1303E3D3E943FF8DFAF7BEC3FA6BFE5A" box="[1373,1423,474,501]" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="22" pageNumber="433">15C</figureCitation>
|
||
). Lateral margination incomplete on tergites 4–10(11) (
|
||
<figureCitation id="1303E3D3E943FF8DFD55BEE6FC8CFDB6" box="[767,872,511,537]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="22" pageNumber="433">Fig. 14D</figureCitation>
|
||
) and nearly complete on following tergites, only tergite 23 marginate completely (
|
||
<figureCitation id="1303E3D3E943FF8DFDA1BD3AFD97FD92" box="[523,627,546,573]" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="22" pageNumber="433">Fig. 15D</figureCitation>
|
||
). Tergite 23 nearly as long as wide, its posterior margin much convex in the middle.
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E943FF8DFF6DBD72FBACFD06" blockId="22.[151,1437,150,1977]" pageId="22" pageNumber="433">
|
||
Sternites lacking sutures, with very shallow rounded depression in the center (
|
||
<figureCitation id="1303E3D3E943FF8DFBECBD72FB48FD2A" box="[1094,1196,618,645]" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="22" pageNumber="433">Fig. 15E</figureCitation>
|
||
). Sternite 23 tongueshaped, its rounded posterior margin slightly concave in the middle (
|
||
<figureCitation id="1303E3D3E943FF8DFC32BD96FBE4FD06" box="[920,1024,655,681]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="22" pageNumber="433">Figs 14E</figureCitation>
|
||
,
|
||
<figureCitation id="1303E3D3E943FF8DFBA6BD97FBDEFD06" box="[1036,1082,654,681]" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="22" pageNumber="433">15F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E943FF8DFF6DBDAAFC2BFD62" blockId="22.[151,1437,150,1977]" box="[199,975,690,717]" pageId="22" pageNumber="433">
|
||
LBS 7 with a pair of standardly developed spiracles (
|
||
<figureCitation id="1303E3D3E943FF8DFCB0BDAAFC61FD62" box="[794,901,691,717]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="22" pageNumber="433">Figs 14D</figureCitation>
|
||
,
|
||
<figureCitation id="1303E3D3E943FF8DFC3BBDABFC26FD62" box="[913,962,690,717]" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="22" pageNumber="433">15E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E943FF8DFF6DBDCEFF31FCF2" blockId="22.[151,1437,150,1977]" pageId="22" pageNumber="433">
|
||
Coxopleuron (
|
||
<figureCitation id="1303E3D3E943FF8DFECDBDCEFE29FD5E" box="[359,461,727,753]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="22" pageNumber="433">Figs 14E</figureCitation>
|
||
,
|
||
<figureCitation id="1303E3D3E943FF8DFE72BDCFFDE0FD5E" box="[472,516,726,753]" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="22" pageNumber="433">15F</figureCitation>
|
||
) covered by irregularly located coxal pores of various sizes. Coxopleural processes conical, very short (much shorter than sternite 23) and diverging slightly, they visibly extended over the posterior margin of tergite 23 (
|
||
<figureCitation id="1303E3D3E943FF8DFE2CBC06FE0BFC96" box="[390,495,798,825]" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="22" pageNumber="433">Fig. 15D</figureCitation>
|
||
). Posterior margin of pleuron forming an acute angle, its tip definitely pointed (
|
||
<figureCitation id="1303E3D3E943FF8DFAC5BC06FF2CFCF2" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="22" pageNumber="433">Fig. 14E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E943FF8DFF6DBC7EFB42FC0A" blockId="22.[151,1437,150,1977]" pageId="22" pageNumber="433">
|
||
Legs 1–21 with monopartite tarsus (
|
||
<figureCitation id="1303E3D3E943FF8DFDFDBC7EFD5FFC2E" box="[599,699,870,897]" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="22" pageNumber="433">Fig. 15E</figureCitation>
|
||
); legs 1–21 with one tarsal and two tibial spurs and 22 lacking any spur (
|
||
<figureCitation id="1303E3D3E943FF8DFF7DBC92FEDEFC0A" box="[215,314,907,933]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="22" pageNumber="433">Fig. 14E</figureCitation>
|
||
) legs 1–22 with two pretarsal accessory spines. All legs practically not setose.
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E943FF8DFF6DBCB6FCB6FC42" blockId="22.[151,1437,150,1977]" pageId="22" pageNumber="433">
|
||
Ultimate legs not setose; prefemur (
|
||
<figureCitation id="1303E3D3E943FF8DFDC3BCB6FD24FC66" box="[617,704,943,969]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="22" pageNumber="433">Figs 14</figureCitation>
|
||
CE) with two spinous processes of standard both structure and disposition—the larger ventral and smaller dorso-medial ones.
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E943FF8DFF6DBCEFFC70FB9A" blockId="22.[151,1437,150,1977]" pageId="22" pageNumber="433">
|
||
<emphasis id="B94C2344E943FF8DFF6DBCEFFEA9FBBF" bold="true" box="[199,333,1014,1040]" pageId="22" pageNumber="433">Variability.</emphasis>
|
||
In
|
||
<typeStatus id="548341F4E943FF8DFEDABCEEFE23FBBE" box="[368,455,1015,1041]" pageId="22" pageNumber="433" type="syntype">syntype</typeStatus>
|
||
1 both sternite 23 and the coxopleural process are relatively longer than in
|
||
<typeStatus id="548341F4E943FF8DFAAFBCEEFAB8FBBE" box="[1285,1372,1015,1041]" pageId="22" pageNumber="433" type="syntype">syntype</typeStatus>
|
||
2 (cf.
|
||
<figureCitation id="1303E3D3E943FF8DFF3DBB02FF1FFB9A" box="[151,251,1051,1077]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="22" pageNumber="433">Figs 14E</figureCitation>
|
||
and
|
||
<figureCitation id="1303E3D3E943FF8DFE98BB03FE85FB9A" box="[306,353,1050,1077]" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="22" pageNumber="433">15F</figureCitation>
|
||
). The ultimate legs are attached only in
|
||
<typeStatus id="548341F4E943FF8DFC8BBB02FC9CFB9A" box="[801,888,1051,1077]" pageId="22" pageNumber="433" type="syntype">syntype</typeStatus>
|
||
1.
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E943FF8DFF6DBB26FD4FFBF6" blockId="22.[151,1437,150,1977]" box="[199,683,1087,1113]" pageId="22" pageNumber="433">
|
||
<emphasis id="B94C2344E943FF8DFF6DBB26FEFEFBF6" bold="true" box="[199,282,1087,1113]" pageId="22" pageNumber="433">Range.</emphasis>
|
||
Known from the
|
||
<typeStatus id="548341F4E943FF8DFE4BBB26FDF5FBF6" box="[481,529,1087,1113]" pageId="22" pageNumber="433">type</typeStatus>
|
||
locality only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C322ACDDE943FF81FF6DBB7AFDC1FD62" lastPageId="26" lastPageNumber="437" pageId="22" pageNumber="433" type="discussion">
|
||
<paragraph id="8B87FF56E943FF8DFF6DBB7AFBAFFA32" blockId="22.[151,1437,150,1977]" pageId="22" pageNumber="433">
|
||
<emphasis id="B94C2344E943FF8DFF6DBB7AFEDFFBD2" bold="true" box="[199,315,1123,1149]" pageId="22" pageNumber="433">Remarks.</emphasis>
|
||
<taxonomicName id="4C3884D5E943FF8DFEEDBB7AFDB8FBD2" authorityName=": Schileyko" authorityYear="2007" box="[327,604,1123,1149]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
|
||
<emphasis id="B94C2344E943FF8DFEEDBB7AFDB8FBD2" box="[327,604,1123,1149]" italics="true" pageId="22" pageNumber="433">S. broelemanni esulcata</emphasis>
|
||
</taxonomicName>
|
||
is known only from
|
||
<specimenCount id="9D3E34DFE943FF8DFCF4BB7AFBF2FBD2" box="[862,1046,1123,1149]" pageId="22" pageNumber="433" type="generic">three specimens</specimenCount>
|
||
kept in NHMW, collected by C. Dawydoff in the Lang Biang Mountains (
|
||
<figureCitation id="1303E3D3E943FF8DFDD7BB9EFD30FB0E" box="[637,724,1159,1185]" captionStart="FIGURE 16" captionStartId="25.[151,250,1943,1967]" captionTargetBox="[193,1394,181,1919]" captionTargetId="figure-18@25.[193,1394,181,1919]" captionTargetPageId="25" captionText="FIGURE 16. Distribution of Scolopocryptops broelemanni esulcata Attems, 1938." figureDoi="http://doi.org/10.5281/zenodo.7539976" httpUri="https://zenodo.org/record/7539976/files/figure.png" pageId="22" pageNumber="433">Fig. 16</figureCitation>
|
||
).
|
||
<taxonomicName id="4C3884D5E943FF8DFD46BB9EFBE4FB0E" authorityName=": Schileyko" authorityYear="2007" box="[748,1024,1159,1185]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
|
||
<emphasis id="B94C2344E943FF8DFD46BB9EFBE4FB0E" box="[748,1024,1159,1185]" italics="true" pageId="22" pageNumber="433">S. broelemanni esulcata</emphasis>
|
||
</taxonomicName>
|
||
is the only Vietnamese member of the former genus
|
||
<taxonomicName id="4C3884D5E943FF8DFECCBBB2FE1AFB6B" authorityName="Crabill" authorityYear="1953" box="[358,510,1195,1220]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94C2344E943FF8DFECCBBB2FE1AFB6B" box="[358,510,1195,1220]" italics="true" pageId="22" pageNumber="433">Dinocryptops</emphasis>
|
||
</taxonomicName>
|
||
which name has been synonymised with
|
||
<taxonomicName id="4C3884D5E943FF8DFC4DBBB2FB46FB6A" authorityName="Newport" authorityYear="1844" box="[999,1186,1195,1221]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94C2344E943FF8DFC4DBBB2FB46FB6A" box="[999,1186,1195,1221]" italics="true" pageId="22" pageNumber="433">Scolopocryptops</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFA982A7E943FF8DFB78BBB2FEB1FB46" author="Edgecombe, G. D. & Vahtera, V. & Stock, S. R. & Kallonen, A. & Xiao, X. & Rack, A. & Giribet, G." pageId="22" pageNumber="433" pagination="768 - 786" refId="ref15077" refString="Edgecombe, G. D., Vahtera, V., Stock, S. R., Kallonen, A., Xiao, X., Rack, A. & Giribet, G. (2012) A scolopocryptopid centipede (Chilopoda: Scolopendromorpha) from Mexican amber: synchrotron microtomography and phylogenetic placement using a combined morphological and molecular data set. Zoological Journal of the Linnean Society, 166, 768 - 786. https: // doi. org / 10.1111 / j. 1096 - 3642.2012.00860. x" type="journal article" year="2012">
|
||
Edgecombe
|
||
<emphasis id="B94C2344E943FF8DFACBBBB2FA71FB6A" box="[1377,1429,1195,1221]" italics="true" pageId="22" pageNumber="433">et al</emphasis>
|
||
. (2012: 777–778)
|
||
</bibRefCitation>
|
||
based on both morphology and molecular analysis (this synonymy has been omitted by
|
||
<bibRefCitation id="EFA982A7E943FF8DFAE1BBD6FEF0FAA2" author="Bonato, L. & Chagas-Jr., A. & Edgecombe, G. D. & Lewis, J. G. E. & Minelli, A. & Pereira, L. A. & Shelley, R. M. & Stoev, P. & Zapparoli, M." pageId="22" pageNumber="433" refId="ref14353" refString="Bonato, L., Chagas-Jr., A., Edgecombe, G. D., Lewis, J. G. E., Minelli, A., Pereira, L. A., Shelley, R. M., Stoev, P. & Zapparoli, M. (2016) ChiloBase 2.0 - A World Catalogue of Centipedes (Chilopoda). Available from: https: // chilobase. biologia. unipd. it (accessed 18 February 2021)" type="url" year="2016">
|
||
Bonato
|
||
<emphasis id="B94C2344E943FF8DFF3DBBEAFF34FAA2" box="[151,208,1267,1293]" italics="true" pageId="22" pageNumber="433">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
). The study of digital images of both
|
||
<typeStatus id="548341F4E943FF8DFD13BBEAFCFFFAA2" box="[697,795,1267,1293]" pageId="22" pageNumber="433" type="syntype">syntypes</typeStatus>
|
||
showed that they have well developed (not rudimentary) spiracles on LBS 7 (
|
||
<figureCitation id="1303E3D3E943FF8DFE20BA0EFE1EFA9E" box="[394,506,1303,1329]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="22" pageNumber="433">Figs 14D</figureCitation>
|
||
,
|
||
<figureCitation id="1303E3D3E943FF8DFDA3BA0FFDD0FA9E" box="[521,564,1302,1329]" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="22" pageNumber="433">15E</figureCitation>
|
||
)—the only diagnostic character of former
|
||
<taxonomicName id="4C3884D5E943FF8DFB85BA0EFB23FA9F" authorityName="Crabill" authorityYear="1953" box="[1071,1223,1303,1328]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B94C2344E943FF8DFB85BA0EFB23FA9F" box="[1071,1223,1303,1328]" italics="true" pageId="22" pageNumber="433">Dinocryptops</emphasis>
|
||
</taxonomicName>
|
||
. New data on the occasional presence of spiracles on LBS 7 (
|
||
<figureCitation id="1303E3D3E943FF8DFD30BA22FD0BFAFA" box="[666,751,1339,1365]" captionStart="FIGURES 2" captionStartId="5.[152,270,1727,1751]" captionTargetBox="[161,1427,185,1702]" captionTargetId="figure-18@5.[159,1429,181,1703]" captionTargetPageId="5" captionText="FIGURES 2A–F. Scolopocryptops rubiginosus C.L. Koch, 1878; SVR.PO.020 A head and LBS 1, dorsally; B head and forcipular segment, ventrally; C LBS 6–10, dorsally; D LBS 6–10, ventrally; Rc 6738 (ZMMU) E head and forcipular segment, ventrally; Rc 6686 (ZMMU) F LBS 7, 8 (right side), laterally." figureDoi="http://doi.org/10.5281/zenodo.7539942" httpUri="https://zenodo.org/record/7539942/files/figure.png" pageId="22" pageNumber="433">Fig. 2F</figureCitation>
|
||
,
|
||
<figureCitation id="1303E3D3E943FF8DFD55BA22FCC6FAFA" box="[767,802,1339,1365]" captionStart="FIGURES 6" captionStartId="11.[152,270,1943,1967]" captionTargetBox="[242,1346,184,1917]" captionTargetId="figure-18@11.[241,1346,181,1919]" captionTargetPageId="11" captionText="FIGURES 6A–F. Scolopocryptops spinicaudus Wood, 1862; SVR.PO.202 A LBS 23 and right ultimate leg, laterally; B LBS 21–23 and ultimate legs, ventrally; C right ultimate leg, medially; Rc 7497 (ZMMU) D LBS 7 and 8, laterally; Rc 7225 (ZMMU) E left ultimate leg, dorsally; Rc 7497 (ZMMU) F LBS 13–16, dorsally." figureDoi="http://doi.org/10.5281/zenodo.7539954" httpUri="https://zenodo.org/record/7539954/files/figure.png" pageId="22" pageNumber="433">6D</figureCitation>
|
||
) in both
|
||
<taxonomicName id="4C3884D5E943FF8DFC38BA22FBD2FAFA" authorityName="C. L. Koch" authorityYear="1878" box="[914,1078,1339,1365]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="species" species="rubiginosus">
|
||
<emphasis id="B94C2344E943FF8DFC38BA22FBD2FAFA" box="[914,1078,1339,1365]" italics="true" pageId="22" pageNumber="433">S. rubiginosus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3884D5E943FF8DFBDEBA22FAFEFAFA" authorityName="Wood" authorityYear="1862" box="[1140,1306,1339,1365]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="species" species="spinicaudus">
|
||
<emphasis id="B94C2344E943FF8DFBDEBA22FAFEFAFA" box="[1140,1306,1339,1365]" italics="true" pageId="22" pageNumber="433">S. spinicaudus</emphasis>
|
||
</taxonomicName>
|
||
reduce the difference between them and
|
||
<emphasis id="B94C2344E943FF8DFE42BA46FCE4FAD6" box="[488,768,1375,1401]" italics="true" pageId="22" pageNumber="433">
|
||
<taxonomicName id="4C3884D5E943FF8DFE42BA46FD18FAD6" authorityName=": Schileyko" authorityYear="2007" box="[488,764,1375,1401]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">S. broelemanni esulcata</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Thus, a character such as the number of spiracles must be used with much care (when used alone) to reliably identify
|
||
<taxonomicName id="4C3884D5E943FF8DFC9BBA9AFBAFFA32" box="[817,1099,1411,1437]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B94C2344E943FF8DFC9BBA9AFC08FA32" box="[817,1004,1411,1437]" italics="true" pageId="22" pageNumber="433">Scolopocryptops</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E943FF8DFF6DBABFFCE9FA6E" blockId="22.[151,1437,150,1977]" box="[199,781,1446,1473]" pageId="22" pageNumber="433">
|
||
<emphasis id="B94C2344E943FF8DFF6DBABFFCE9FA6E" bold="true" box="[199,781,1446,1473]" pageId="22" pageNumber="433">
|
||
Discussion on validity of
|
||
<taxonomicName id="4C3884D5E943FF8DFE5ABABEFCE2FA6E" authorityName=": Schileyko" authorityYear="2007" box="[496,774,1446,1473]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
|
||
<emphasis id="B94C2344E943FF8DFE5ABABEFCE2FA6E" bold="true" box="[496,774,1446,1473]" italics="true" pageId="22" pageNumber="433">S. broelemanni esulcata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E943FF8DFF6DBAD2FB8EF982" blockId="22.[151,1437,150,1977]" pageId="22" pageNumber="433">Available data on this subspecies are scarce; the original description is very short and incomplete, lacking both illustrations and information about some diagnostic characters (number of glabrous basal antennal articles, presence/conditions of tergal sutures, pilosity of distal articles of the ultimate legs etc.).</paragraph>
|
||
<paragraph id="8B87FF56E943FF8DFF6DB92EFD99F816" blockId="22.[151,1437,150,1977]" pageId="22" pageNumber="433">
|
||
Judging from the digital images, the NHMW specimens fit the descriptions of
|
||
<taxonomicName id="4C3884D5E943FF8DFB92B92EFA78F9FE" authority="Kraepelin, 1903" authorityName="Kraepelin" authorityYear="1903" box="[1080,1436,1591,1617]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="species" species="broelemanni">
|
||
<emphasis id="B94C2344E943FF8DFB92B92EFB05F9FE" box="[1080,1249,1591,1617]" italics="true" pageId="22" pageNumber="433">S. broelemanni</emphasis>
|
||
<bibRefCitation id="EFA982A7E943FF8DFB42B92EFA78F9FE" author="Kraepelin, K." box="[1256,1436,1591,1617]" pageId="22" pageNumber="433" pagination="1 - 276" refId="ref15456" refString="Kraepelin, K. (1903) Revision der Scolopendriden. Mitteilungen aus dem naturhistorischen Museum in Hamburg, 20, 1 - 276." type="journal article" year="1903">Kraepelin, 1903</bibRefCitation>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFA982A7E943FF8DFF10B942FE47F9DA" author="Kraepelin, K." box="[186,419,1627,1653]" pageId="22" pageNumber="433" pagination="1 - 276" refId="ref15456" refString="Kraepelin, K. (1903) Revision der Scolopendriden. Mitteilungen aus dem naturhistorischen Museum in Hamburg, 20, 1 - 276." type="journal article" year="1903">Kraepelin (1903: 77)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFA982A7E943FF8DFE7DB942FD56F9DA" author="Attems, C." box="[471,690,1626,1653]" pageId="22" pageNumber="433" refId="ref14161" refString="Attems, C. (1930) Myriopoda. 2. Scolopendromorpha. Das Tierreich. Vol. 54. Walter de Gruyter et Co. Publishing House, Berlin, 308 pp. [in German]" type="book" year="1930">Attems (1930: 256)</bibRefCitation>
|
||
well. In the original description of
|
||
<taxonomicName id="4C3884D5E943FF8DFB91B942FEFFF936" authority="Attems (1938: 338)" authorityName="Attems" authorityPageNumber="338" authorityYear="1938" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
|
||
<emphasis id="B94C2344E943FF8DFB91B942FAA2F9DA" box="[1083,1350,1627,1653]" italics="true" pageId="22" pageNumber="433">S. broelemanni esulcata</emphasis>
|
||
<bibRefCitation id="EFA982A7E943FF8DFAE1B942FEFFF936" author="Attems, C." pageId="22" pageNumber="433" pagination="187 - 353" refId="ref14199" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in franzosisch Indochina gesammelten Myriopoden. Memoires du Museum national d'histoire naturelle, New Series, 6 (2), 187 - 353." type="journal article" year="1938">Attems (1938: 338)</bibRefCitation>
|
||
</taxonomicName>
|
||
indicated, that this subspecies is, on the whole, very similar to
|
||
<taxonomicName id="4C3884D5E943FF8DFC74B966FB62F936" authorityName="Kraepelin" authorityYear="1903" box="[990,1158,1663,1689]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="species" species="broelemanni">
|
||
<emphasis id="B94C2344E943FF8DFC74B966FB62F936" box="[990,1158,1663,1689]" italics="true" pageId="22" pageNumber="433">S. broelemanni</emphasis>
|
||
</taxonomicName>
|
||
, differing from the latter mainly in the absence (vs. presence) of the median suture on the ultimate tergite. This character is present in various groups of scolopendromorphs (
|
||
<taxonomicName id="4C3884D5E943FF8DFE58B9DEFD41F94E" baseAuthorityName="Leach" baseAuthorityYear="1814" box="[498,677,1735,1761]" class="Chilopoda" family="Scolopendridae" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="family">Scolopendridae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3884D5E943FF8DFD05B9DEFCB1F94E" authorityName="Bollman" authorityYear="1893" box="[687,853,1735,1761]" class="Chilopoda" family="Plutoniumidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="family">Plutoniumidae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3884D5E943FF8DFCCAB9DEFBA1F94E" box="[864,1093,1735,1761]" class="Chilopoda" family="Scolopocryptopidae" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="subFamily" subFamily="Scolopocryptopinae">Scolopocryptopinae</taxonomicName>
|
||
), being species-specific for the overwhelming majority of their species. Digital images of three NHMW specimens do not recognize this suture (
|
||
<figureCitation id="1303E3D3E943FF8DFAC5B9F2FF2EF886" captionStart="FIGURES 15" captionStartId="24.[152,270,1593,1617]" captionTargetBox="[151,1435,184,1566]" captionTargetId="figure-33@24.[151,1436,181,1569]" captionTargetPageId="24" captionText="FIGURES 15A–F. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C LBS 6–11, dorsally; D LBS 20–23, dorsally; E LBS 5–11 (right side), laterally; F LBS 21–23, ventrally." figureDoi="http://doi.org/10.5281/zenodo.7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="22" pageNumber="433">Fig. 15D</figureCitation>
|
||
), so they are fully consistent with
|
||
<taxonomicName id="4C3884D5E943FF8DFDFDB816FC8CF886" authorityName="Attems" authorityYear="1938" box="[599,872,1807,1833]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
|
||
<emphasis id="B94C2344E943FF8DFDFDB816FC8CF886" box="[599,872,1807,1833]" italics="true" pageId="22" pageNumber="433">S. broelemanni esulcata</emphasis>
|
||
</taxonomicName>
|
||
. There is also a significant geographical distance between the range of
|
||
<taxonomicName id="4C3884D5E943FF8DFE3FB82AFD4CF8E2" authorityName="Attems" authorityYear="1938" box="[405,680,1843,1869]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
|
||
<emphasis id="B94C2344E943FF8DFE3FB82AFD4CF8E2" box="[405,680,1843,1869]" italics="true" pageId="22" pageNumber="433">S. broelemanni esulcata</emphasis>
|
||
</taxonomicName>
|
||
(Southern
|
||
<collectingCountry id="F32FBFC6E943FF8DFC81B82AFC6AF8E2" box="[811,910,1843,1869]" name="Vietnam" pageId="22" pageNumber="433">Vietnam</collectingCountry>
|
||
), and the nominotypical subspecies described from “Chou-San” (Zhoushan, formerly romanized as Chusan,
|
||
<collectingRegion id="49FC31B4E943FF8DFCFCB84EFBC1F8DE" box="[854,1061,1879,1905]" country="China" name="Zhejiang" pageId="22" pageNumber="433">Zhejiang Province</collectingRegion>
|
||
in East
|
||
<collectingCountry id="F32FBFC6E943FF8DFB2BB84EFB2DF8DE" box="[1153,1225,1879,1905]" name="China" pageId="22" pageNumber="433">China</collectingCountry>
|
||
). Summing up, we consider
|
||
<taxonomicName id="4C3884D5E943FF8DFF56B862FD46F83A" authority="Attems, 1938" authorityName="Attems" authorityYear="1938" box="[252,674,1915,1941]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="22" pageNumber="433" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
|
||
<emphasis id="B94C2344E943FF8DFF56B862FDE3F83A" box="[252,519,1915,1941]" italics="true" pageId="22" pageNumber="433">S. broelemanni esulcata</emphasis>
|
||
<bibRefCitation id="EFA982A7E943FF8DFDA6B862FD46F83A" author="Attems, C." box="[524,674,1915,1941]" pageId="22" pageNumber="433" pagination="187 - 353" refId="ref14199" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in franzosisch Indochina gesammelten Myriopoden. Memoires du Museum national d'histoire naturelle, New Series, 6 (2), 187 - 353." type="journal article" year="1938">Attems, 1938</bibRefCitation>
|
||
</taxonomicName>
|
||
to be a valid subspecies, but its taxonomic status may be changed by examining representative relevant material.
|
||
</paragraph>
|
||
<caption id="DF47AFDEE942FF8CFF32B85EFCA6F864" ID-DOI="http://doi.org/10.5281/zenodo.7539972" ID-Zenodo-Dep="7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="23" pageNumber="434" startId="23.[152,270,1863,1887]" targetBox="[151,1434,181,1839]" targetPageId="23">
|
||
<paragraph id="8B87FF56E942FF8CFF32B85EFCA6F864" blockId="23.[151,1436,1863,1996]" pageId="23" pageNumber="434">
|
||
<emphasis id="B94C2344E942FF8CFF32B85EFE8DF8F0" bold="true" box="[152,361,1863,1887]" pageId="23" pageNumber="434">FIGURES 14A–G.</emphasis>
|
||
<taxonomicName id="4C3884D5E942FF8CFEDBB85EFC71F8CF" authority="Attems, 1938" authorityName="Attems" authorityYear="1938" box="[369,917,1863,1888]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="23" pageNumber="434" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
|
||
<emphasis id="B94C2344E942FF8CFEDBB85EFD1AF8F0" box="[369,766,1863,1887]" italics="true" pageId="23" pageNumber="434">Scolopocryptops broelemanni esulcata</emphasis>
|
||
<bibRefCitation id="EFA982A7E942FF8CFCADB851FC71F8CF" author="Attems, C." box="[775,917,1864,1888]" pageId="23" pageNumber="434" pagination="187 - 353" refId="ref14199" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in franzosisch Indochina gesammelten Myriopoden. Memoires du Museum national d'histoire naturelle, New Series, 6 (2), 187 - 353." type="journal article" year="1938">Attems, 1938</bibRefCitation>
|
||
</taxonomicName>
|
||
; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek)
|
||
<emphasis id="B94C2344E942FF8CFF40B872FF18F82C" bold="true" box="[234,252,1899,1923]" pageId="23" pageNumber="434">A</emphasis>
|
||
head + LBS 1–3, dorsally;
|
||
<emphasis id="B94C2344E942FF8CFDBBB875FDC6F82B" bold="true" box="[529,546,1900,1924]" pageId="23" pageNumber="434">B</emphasis>
|
||
head and forcipular segment, ventrally;
|
||
<emphasis id="B94C2344E942FF8CFC13B872FC2FF82C" bold="true" box="[953,971,1899,1923]" pageId="23" pageNumber="434">C</emphasis>
|
||
general view, dorsally;
|
||
<emphasis id="B94C2344E942FF8CFB16B875FB2AF82B" bold="true" box="[1212,1230,1900,1924]" pageId="23" pageNumber="434">D</emphasis>
|
||
LBS 6–9 (left side), laterally;
|
||
<emphasis id="B94C2344E942FF8CFF5DB889FEECF807" bold="true" box="[247,264,1936,1960]" pageId="23" pageNumber="434">E</emphasis>
|
||
LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW)
|
||
<emphasis id="B94C2344E942FF8CFAF2B889FA83F807" bold="true" box="[1368,1383,1936,1960]" pageId="23" pageNumber="434">F</emphasis>
|
||
head + LBS 1–3, dorsally;
|
||
<emphasis id="B94C2344E942FF8CFED8B8AAFE62F864" bold="true" box="[370,390,1971,1995]" pageId="23" pageNumber="434">G</emphasis>
|
||
label of syntypes written by Chagas Junior.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF47AFDEE94DFF83FF32B920FD0AF936" ID-DOI="http://doi.org/10.5281/zenodo.7539974" ID-Zenodo-Dep="7539974" httpUri="https://zenodo.org/record/7539974/files/figure.png" pageId="24" pageNumber="435" startId="24.[152,270,1593,1617]" targetBox="[151,1435,184,1566]" targetPageId="24">
|
||
<paragraph id="8B87FF56E94DFF83FF32B920FD0AF936" blockId="24.[151,1437,1593,1690]" pageId="24" pageNumber="435">
|
||
<emphasis id="B94C2344E94DFF83FF32B920FE87F9FE" bold="true" box="[152,355,1593,1617]" pageId="24" pageNumber="435">FIGURES 15A–F.</emphasis>
|
||
<taxonomicName id="4C3884D5E94DFF83FEC6B920FC75F9FD" authority="Attems, 1938" authorityName="Attems" authorityYear="1938" box="[364,913,1593,1618]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="24" pageNumber="435" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
|
||
<emphasis id="B94C2344E94DFF83FEC6B920FD1DF9FE" box="[364,761,1593,1617]" italics="true" pageId="24" pageNumber="435">Scolopocryptops broelemanni esulcata</emphasis>
|
||
<bibRefCitation id="EFA982A7E94DFF83FCA9B923FC75F9FD" author="Attems, C." box="[771,913,1594,1618]" pageId="24" pageNumber="435" pagination="187 - 353" refId="ref14199" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in franzosisch Indochina gesammelten Myriopoden. Memoires du Museum national d'histoire naturelle, New Series, 6 (2), 187 - 353." type="journal article" year="1938">Attems, 1938</bibRefCitation>
|
||
</taxonomicName>
|
||
; Syntype 2, MY 2063 (NHMW) (photo by Oliver Macek)
|
||
<emphasis id="B94C2344E94DFF83FF41B944FF19F9DA" bold="true" box="[235,253,1629,1653]" pageId="24" pageNumber="435">A</emphasis>
|
||
head + LBS 1–3, dorsally;
|
||
<emphasis id="B94C2344E94DFF83FDBFB947FDC2F9D9" bold="true" box="[533,550,1630,1654]" pageId="24" pageNumber="435">B</emphasis>
|
||
head and forcipular segment, ventrally;
|
||
<emphasis id="B94C2344E94DFF83FC15B944FC35F9DA" bold="true" box="[959,977,1629,1653]" pageId="24" pageNumber="435">C</emphasis>
|
||
LBS 6–11, dorsally;
|
||
<emphasis id="B94C2344E94DFF83FB00B947FB58F9D9" bold="true" box="[1194,1212,1630,1654]" pageId="24" pageNumber="435">D</emphasis>
|
||
LBS 20–23, dorsally;
|
||
<emphasis id="B94C2344E94DFF83FF3DB99BFF4CF935" bold="true" box="[151,168,1666,1690]" pageId="24" pageNumber="435">E</emphasis>
|
||
LBS 5–11 (right side), laterally;
|
||
<emphasis id="B94C2344E94DFF83FE52B99BFDE3F935" bold="true" box="[504,519,1666,1690]" pageId="24" pageNumber="435">F</emphasis>
|
||
LBS 21–23, ventrally.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF47AFDEE94CFF82FF3DB88EFC05F81F" ID-DOI="http://doi.org/10.5281/zenodo.7539976" ID-Zenodo-Dep="7539976" box="[151,993,1943,1968]" httpUri="https://zenodo.org/record/7539976/files/figure.png" pageId="25" pageNumber="436" startId="25.[151,250,1943,1967]" targetBox="[193,1394,181,1919]" targetPageId="25">
|
||
<paragraph id="8B87FF56E94CFF82FF3DB88EFC05F81F" blockId="25.[151,993,1943,1968]" box="[151,993,1943,1968]" pageId="25" pageNumber="436">
|
||
<emphasis id="B94C2344E94CFF82FF3DB88EFEC4F800" bold="true" box="[151,288,1943,1967]" pageId="25" pageNumber="436">FIGURE 16.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="4C3884D5E94CFF82FE68B881FC3AF81F" authority="Attems, 1938" authorityName="Attems" authorityYear="1938" box="[450,990,1944,1968]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="25" pageNumber="436" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
|
||
<emphasis id="B94C2344E94CFF82FE68B881FCAFF81F" box="[450,843,1944,1968]" italics="true" pageId="25" pageNumber="436">Scolopocryptops broelemanni esulcata</emphasis>
|
||
<bibRefCitation id="EFA982A7E94CFF82FCF8B881FC3AF81F" author="Attems, C." box="[850,990,1944,1968]" pageId="25" pageNumber="436" pagination="187 - 353" refId="ref14199" refString="Attems, C. (1938) Die von Dr. C. Dawydoff in franzosisch Indochina gesammelten Myriopoden. Memoires du Museum national d'histoire naturelle, New Series, 6 (2), 187 - 353." type="journal article" year="1938">Attems, 1938</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B87FF56E94FFF81FF6DBF8EFB9FFE5A" blockId="26.[151,1437,150,717]" pageId="26" pageNumber="437">
|
||
In 2005, NHMW specimens were examined by Amazons Chagas Junior, who identified them as “
|
||
<taxonomicName id="4C3884D5E94FFF81FAAEBF8EFEF3FF7A" authorityName="Kraepelin" authorityYear="1903" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="brolemanni">
|
||
<emphasis id="B94C2344E94FFF81FAAEBF8EFEF3FF7A" italics="true" pageId="26" pageNumber="437">Dinocryptops brolemanni</emphasis>
|
||
</taxonomicName>
|
||
(sic!) (
|
||
<bibRefCitation id="EFA982A7E94FFF81FEC7BFA2FDCCFF7A" author="Kraepelin, K." box="[365,552,187,213]" pageId="26" pageNumber="437" pagination="1 - 276" refId="ref15456" refString="Kraepelin, K. (1903) Revision der Scolopendriden. Mitteilungen aus dem naturhistorischen Museum in Hamburg, 20, 1 - 276." type="journal article" year="1903">Kraepelin, 1903</bibRefCitation>
|
||
)” (
|
||
<figureCitation id="1303E3D3E94FFF81FDE7BFA2FD5CFF7A" box="[589,696,187,213]" captionStart="FIGURES 14" captionStartId="23.[152,270,1863,1887]" captionTargetBox="[151,1434,181,1839]" captionTargetId="figure-18@23.[151,1436,181,1840]" captionTargetPageId="23" captionText="FIGURES 14A–G. Scolopocryptops broelemanni esulcata Attems, 1938; Syntype 1, MY 2063 (NHMW) (photo by Oliver Macek) A head + LBS 1–3, dorsally; B head and forcipular segment, ventrally; C general view, dorsally; D LBS 6–9 (left side), laterally; E LBS 22, 23, legs 21, 22 and basal articles of ultimate legs, ventrally; non-type specimen, MY 8714 (NHMW) F head + LBS 1–3, dorsally; G label of syntypes written by Chagas Junior." figureDoi="http://doi.org/10.5281/zenodo.7539972" httpUri="https://zenodo.org/record/7539972/files/figure.png" pageId="26" pageNumber="437">Fig. 14G</figureCitation>
|
||
). In his PhD thesis Chagas (2008) proposed the synonymy of
|
||
<taxonomicName id="4C3884D5E94FFF81FA2DBFA2FE6DFF56" authorityName="Attems" authorityYear="1938" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
|
||
<emphasis id="B94C2344E94FFF81FA2DBFA2FE6DFF56" italics="true" pageId="26" pageNumber="437">S. broelemanni esulcata</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="4C3884D5E94FFF81FE05BFC6FD0AFF56" box="[431,750,223,249]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="broelemanni">
|
||
<emphasis id="B94C2344E94FFF81FE05BFC6FD0AFF56" box="[431,750,223,249]" italics="true" pageId="26" pageNumber="437">S. broelemanni broelemanni</emphasis>
|
||
</taxonomicName>
|
||
and considered (p. 100) both of them as representative(s) of some new (but unnamed) genus—“Gen. Nov. 1” (any nomenclatural acts proposed there have no formal status— disclaimer to dissertation). He referred to both of these subspecies as “Gen. Nov. 1
|
||
<taxonomicName id="4C3884D5E94FFF81FBEBBE3EFF07FECA" authority="(Kraepelin, 1903) Comb." authorityName="Comb." baseAuthorityName="Kraepelin" baseAuthorityYear="1903" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="broelemanni">
|
||
<emphasis id="B94C2344E94FFF81FBEBBE3EFB2AFEEE" box="[1089,1230,295,321]" italics="true" pageId="26" pageNumber="437">broelemanni</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA982A7E94FFF81FB76BE3EFA70FEEE" author="Kraepelin, K." box="[1244,1428,295,321]" pageId="26" pageNumber="437" pagination="1 - 276" refId="ref15456" refString="Kraepelin, K. (1903) Revision der Scolopendriden. Mitteilungen aus dem naturhistorischen Museum in Hamburg, 20, 1 - 276." type="journal article" year="1903">Kraepelin, 1903</bibRefCitation>
|
||
) Comb.
|
||
</taxonomicName>
|
||
nova” (p. 101), but also as “
|
||
<taxonomicName id="4C3884D5E94FFF81FD99BE52FC86FECA" authorityName="Kraepelin" authorityYear="1903" box="[563,866,331,357]" class="Chilopoda" family="Scolopocryptopidae" genus="Dinocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="species" species="broelemanni">
|
||
<emphasis id="B94C2344E94FFF81FD99BE52FC86FECA" box="[563,866,331,357]" italics="true" pageId="26" pageNumber="437">Dinocryptops broelemanni</emphasis>
|
||
</taxonomicName>
|
||
” (pp. 25, 27 etc). Considering both subspecies to be the same form, Chagas (2008: 101) gave a composite description of the NHMW specimens as well as
|
||
<specimenCount id="9D3E34DFE94FFF81FADEBE76FEE5FE02" pageId="26" pageNumber="437" type="generic">one specimen</specimenCount>
|
||
of
|
||
<taxonomicName id="4C3884D5E94FFF81FE82BE8AFCE8FE02" box="[296,780,403,429]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="broelemanni">
|
||
<emphasis id="B94C2344E94FFF81FE82BE8AFCE8FE02" box="[296,780,403,429]" italics="true" pageId="26" pageNumber="437">Scolopocryptops broelemanni broelemanni</emphasis>
|
||
</taxonomicName>
|
||
from “
|
||
<collectingRegion id="49FC31B4E94FFF81FCF6BE8AFBD6FE02" box="[860,1074,403,429]" country="China" name="Guangdong" pageId="26" pageNumber="437">Guangdong Sheng</collectingRegion>
|
||
, Nanking” (actually Nanking or Nanjing is disposed in
|
||
<collectingRegion id="49FC31B4E94FFF81FE32BEAEFDBDFE7E" box="[408,601,439,465]" country="China" name="Jiangsu" pageId="26" pageNumber="437">Jiangsu Province</collectingRegion>
|
||
, not in
|
||
<collectingRegion id="49FC31B4E94FFF81FD06BEAEFC7FFE7E" box="[684,923,439,465]" country="China" name="Guangdong" pageId="26" pageNumber="437">Guangdong Province</collectingRegion>
|
||
) kept in National Museum of Natural History (Smithsonian Institution,
|
||
<collectingRegion id="49FC31B4E94FFF81FE1EBEC2FD98FE5A" box="[436,636,475,501]" country="United States of America" name="District of Columbia" pageId="26" pageNumber="437">Washington D. C.</collectingRegion>
|
||
) (No 31325, “Cook & Loomis,
|
||
<date id="FF86D996E94FFF81FC49BEC2FB81FE5A" box="[995,1125,475,501]" pageId="26" pageNumber="437" value="1919-10-23">23-X-1919</date>
|
||
”).
|
||
</paragraph>
|
||
<paragraph id="8B87FF56E94FFF81FF6DBEE6FDC1FD62" blockId="26.[151,1437,150,717]" pageId="26" pageNumber="437">
|
||
This description is sufficiently detailed, but there are no data on the presence or absence of a median suture on tergite 23, a diagnostic character of
|
||
<taxonomicName id="4C3884D5E94FFF81FD98BD3AFCA5FD92" authorityName="Attems" authorityYear="1938" box="[562,833,547,573]" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
|
||
<emphasis id="B94C2344E94FFF81FD98BD3AFCA5FD92" box="[562,833,547,573]" italics="true" pageId="26" pageNumber="437">S. broelemanni esulcata</emphasis>
|
||
</taxonomicName>
|
||
(it is clearly seen in the corresponding figure 27f on p. 183 that tergite 23 is devoid of this suture). In discussing the validity of
|
||
<taxonomicName id="4C3884D5E94FFF81FC4EBD5EFF2EFD2A" authority="Chagas (2008: 138)" authorityName="Chagas" authorityPageNumber="138" authorityYear="2008" class="Chilopoda" family="Scolopocryptopidae" genus="Scolopocryptops" kingdom="Animalia" order="Scolopendromorpha" pageId="26" pageNumber="437" phylum="Arthropoda" rank="subSpecies" species="broelemanni" subSpecies="esulcata">
|
||
<emphasis id="B94C2344E94FFF81FC4EBD5EFB15FDCE" box="[996,1265,583,609]" italics="true" pageId="26" pageNumber="437">S. broelemanni esulcata</emphasis>
|
||
Chagas (2008: 138)
|
||
</taxonomicName>
|
||
wrote that both subspecies share common characters such as a spiracle on LBS 7 and lateral margination of the cephalic plate, but again omitted the diagnostic character mentioned above. Thus the synonymy proposed by Chagas (2008) remains in question.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |