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<document id="72E58E2620C4E22A694BC4D10C3C1D6A" ID-DOI="10.11646/zootaxa.5483.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13209307" ID-ZooBank="AF0272DB-5588-411D-9EAE-DED4785BF170" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722786113536" checkinUser="plazi" docAuthor="WOOD, T. J." docDate="2024" docId="612B87FCAC17432B0B83FD318F3B0588" docLanguage="en" docName="zootaxa.5483.1.1.pdf" docOrigin="Zootaxa 5483 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5483.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Andrena (Aciandrena) elbensis WOOD, 2024, spec. nov." docType="treatment" docVersion="2" lastPageNumber="81" masterDocId="9D12FF84AC59437B0B14FFCB8E650641" masterDocTitle="Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)" masterLastPageNumber="150" masterPageNumber="1" pageNumber="79" updateTime="1722787353532" updateUser="ExternalLinkService">
<mods:mods id="99406F0FE2CF1A986FF3712F68F545A1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="D045C732C6ED3EC23BB997225CC816DF">Further revisions to the Palaearctic Andrena fauna (Hymenoptera: Andrenidae)</mods:title>
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<mods:namePart id="02CC18F29CF9D8F62B5D83CA41BFA735">WOOD, T. J.</mods:namePart>
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<mods:title id="5BE91EF6359D6DC44A568070B919A1E8">Zootaxa</mods:title>
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<mods:date id="2FFE69A2E38A731C115F076FAA9E5CEF">2024</mods:date>
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<mods:number id="72D77B3C4E53EA0CD9F72230F927D0B0">2024-07-22</mods:number>
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</mods:location>
<mods:classification id="4236FB544F16CAF98D058BF68FC812CF">journal article</mods:classification>
<mods:identifier id="33E3A588BD05617738F5FF97EB200D93" type="DOI">10.11646/zootaxa.5483.1.1</mods:identifier>
<mods:identifier id="CAD42E16386546DD92CE9BACB7DBB6A6" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="CC7C6850B4495CF98BFBF59CEFA808C2" type="Zenodo-Dep">13209307</mods:identifier>
<mods:identifier id="62F2F8D1CEA890513DB5B3C77DDFED17" type="ZooBank">AF0272DB-5588-411D-9EAE-DED4785BF170</mods:identifier>
</mods:mods>
<treatment id="612B87FCAC17432B0B83FD318F3B0588" ID-DOI="http://doi.org/10.5281/zenodo.13209493" ID-Zenodo-Dep="13209493" LSID="urn:lsid:plazi:treatment:612B87FCAC17432B0B83FD318F3B0588" httpUri="http://treatment.plazi.org/id/612B87FCAC17432B0B83FD318F3B0588" lastPageId="80" lastPageNumber="81" pageId="78" pageNumber="79">
<subSubSection id="A1986561AC1743350B83FD318CF50554" box="[151,656,762,789]" pageId="78" pageNumber="79" type="nomenclature">
<paragraph id="E93D36EAAC1743350B83FD318CF50554" blockId="78.[151,656,762,825]" box="[151,656,762,789]" pageId="78" pageNumber="79">
<heading id="B2758186AC1743350B83FD318CF50554" bold="true" box="[151,656,762,789]" fontSize="11" level="2" pageId="78" pageNumber="79" reason="4">
<emphasis id="DBF6EAF8AC1743350B83FD318CF50554" bold="true" box="[151,656,762,789]" pageId="78" pageNumber="79">
<taxonomicName id="2E824D69AC1743350B83FD318C740554" authorityName="WOOD" authorityYear="2024" box="[151,529,762,789]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="elbensis" status="sp. nov." subGenus="Aciandrena">
<emphasis id="DBF6EAF8AC1743350B83FD318F640554" bold="true" box="[151,257,762,789]" italics="true" pageId="78" pageNumber="79">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC1743350A06FD318FC50554" bold="true" box="[274,416,762,789]" italics="true" pageId="78" pageNumber="79">Aciandrena</emphasis>
)
<emphasis id="DBF6EAF8AC1743350AA5FD318C740554" bold="true" box="[433,529,762,789]" italics="true" pageId="78" pageNumber="79">elbensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C0C55783AC174335090DFD308CF50554" box="[537,656,763,789]" pageId="78" pageNumber="79" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1743350B83FCD48F200578" box="[151,325,798,825]" pageId="78" pageNumber="79" type="description">
<paragraph id="E93D36EAAC1743350B83FCD48F200578" blockId="78.[151,656,762,825]" box="[151,325,798,825]" pageId="78" pageNumber="79">
<figureCitation id="71B92A6FAC1743350B83FCD48F270578" box="[151,322,798,825]" captionStart="FIGURE 52" captionStartId="79.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-16@79.[151,1436,181,1621]" captionTargetPageId="79" captionText="FIGURE 52. Andrena (Aciandrena) elbensis spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209417" httpUri="https://zenodo.org/record/13209417/files/figure.png" pageId="78" pageNumber="79">Figures 52A–F</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1743350B83FCAD8E9405E6" pageId="78" pageNumber="79" type="materials_examined">
<paragraph id="E93D36EAAC1743350B83FCAD8E9405E6" blockId="78.[151,1437,870,1905]" pageId="78" pageNumber="79">
<materialsCitation id="59EA3CB7AC1743350B83FCAD8E9405E6" collectingDate="1937-01" collectorName="H. Priesner" country="Egypt" location="Gebel Elba" pageId="78" pageNumber="79" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="DBF6EAF8AC1743350B83FCAD8FD705C1" bold="true" box="[151,434,870,896]" pageId="78" pageNumber="79">
<typeStatus id="36398848AC1743350B83FCAD8F5F05C1" box="[151,314,870,896]" pageId="78" pageNumber="79" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="9195767AAC1743350A5EFCAD8FC805C1" box="[330,429,870,896]" name="Egypt" pageId="78" pageNumber="79">EGYPT</collectingCountry>
:
</emphasis>
<location id="EC5D6031AC1743350AA9FCAC8C5805C0" LSID="urn:lsid:plazi:treatment:612B87FCAC17432B0B83FD318F3B0588:EC5D6031AC1743350AA9FCAC8C5805C0" box="[445,573,871,897]" country="Egypt" name="Gebel Elba" pageId="78" pageNumber="79">Gebel Elba</location>
[Gabal Elba, inferred 22.2001
<superScript id="1EF79BA2AC17433508B7FCAC8DA705C0" attach="right" box="[931,962,871,897]" fontSize="6" pageId="78" pageNumber="79">oN</superScript>
, 36.3321
<superScript id="1EF79BA2AC1743350F38FCAC8A2F05C0" attach="right" box="[1068,1098,871,897]" fontSize="6" pageId="78" pageNumber="79">oE</superScript>
],
<date id="9D3C102AAC1743350F75FCAC8ACF05C0" box="[1121,1194,871,897]" pageId="78" pageNumber="79" value="1937-01">
<collectingDate id="8D78E9C2AC1743350F75FCAC8ACF05C0" box="[1121,1194,871,897]" pageId="78" pageNumber="79" value="1937-01">i.1937</collectingDate>
</date>
,
<specimenCount id="FF84FD63AC1743350FAEFCAC8AAA05C0" box="[1210,1231,871,897]" count="1" pageId="78" pageNumber="79" type="female">♀</specimenCount>
, leg.
<collectorName id="4477533CAC1743350E01FCAC8BFC05C0" box="[1301,1433,871,897]" pageId="78" pageNumber="79">H. Priesner</collectorName>
, OÖLM.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC1743350BD3FC658FEF0130" pageId="78" pageNumber="79" type="description">
<paragraph id="E93D36EAAC1743350BD3FC658C9E02A8" blockId="78.[151,1437,870,1905]" pageId="78" pageNumber="79">
<emphasis id="DBF6EAF8AC1743350BD3FC658FA30588" bold="true" box="[199,454,942,969]" pageId="78" pageNumber="79">Description: Female:</emphasis>
Body length: 6.5 mm (
<figureCitation id="71B92A6FAC17433509CEFC648D010588" box="[730,868,942,969]" captionStart="FIGURE 52" captionStartId="79.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-16@79.[151,1436,181,1621]" captionTargetPageId="79" captionText="FIGURE 52. Andrena (Aciandrena) elbensis spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209417" httpUri="https://zenodo.org/record/13209417/files/figure.png" pageId="78" pageNumber="79">Figure 52A</figureCitation>
).
<emphasis id="DBF6EAF8AC174335086FFC648DA50588" box="[891,960,943,969]" italics="true" pageId="78" pageNumber="79">Head:</emphasis>
Dark, 1.1 times wider than long (
<figureCitation id="71B92A6FAC1743350E45FC648EAF05AC" captionStart="FIGURE 52" captionStartId="79.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-16@79.[151,1436,181,1621]" captionTargetPageId="79" captionText="FIGURE 52. Andrena (Aciandrena) elbensis spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209417" httpUri="https://zenodo.org/record/13209417/files/figure.png" pageId="78" pageNumber="79">Figure 52B</figureCitation>
). Clypeus domed, smooth and shining over its entire surface, irregularly punctate with large to small punctures, punctures separated by 1–3 puncture diameters (
<figureCitation id="71B92A6FAC17433509D5FC3C8D2D0250" box="[705,840,1014,1041]" captionStart="FIGURE 52" captionStartId="79.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-16@79.[151,1436,181,1621]" captionTargetPageId="79" captionText="FIGURE 52. Andrena (Aciandrena) elbensis spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209417" httpUri="https://zenodo.org/record/13209417/files/figure.png" pageId="78" pageNumber="79">Figure 52C</figureCitation>
). Process of labrum trapezoidal-rectangular, slightly wider than long, apical margin straight. Gena slightly narrower than width of compound eye; ocelloccipital distance subequal to diameter of lateral ocellus. Foveae dorsally broad, occupying ⅔ of space between compound eye and lateral ocellus, ventrally narrowing to narrower than width of flagellum at level of antennal insertions; foveae filled with white hairs. Face, gena, vertex, and scape with short white hairs, none equalling length of scape. Scape dark, A2–12 ventrally and dorsally extensively lightened bright orange; A3 exceeding A4, slightly shorter than A4+5, A4 and A5 sub-square, shorter than long, A6–12 quadrate.
</paragraph>
<paragraph id="E93D36EAAC1743350BD3FB388B050034" blockId="78.[151,1437,870,1905]" pageId="78" pageNumber="79">
<emphasis id="DBF6EAF8AC1743350BD3FB388F2D034D" box="[199,328,1267,1292]" italics="true" pageId="78" pageNumber="79">
<taxonomicName id="2E824D69AC1743350BD3FB388F27034D" box="[199,322,1267,1292]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
:
</emphasis>
Scutum and scutellum smooth and shining, with fine shagreen limited to anterior margin of scutum; scutum irregularly punctate, punctures variable, separated by 0.5–2 puncture, scutellum more irregularly punctate, punctures separated by 0.5–3 puncture diameters (
<figureCitation id="71B92A6FAC17433509D7FAF08D2D0314" box="[707,840,1338,1365]" captionStart="FIGURE 52" captionStartId="79.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-16@79.[151,1436,181,1621]" captionTargetPageId="79" captionText="FIGURE 52. Andrena (Aciandrena) elbensis spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209417" httpUri="https://zenodo.org/record/13209417/files/figure.png" pageId="78" pageNumber="79">Figure 52D</figureCitation>
). Pronotum rounded. Mesepisternum and dorsolateral parts of propodeum with fine granular microreticulation, weakly shining; propodeal triangle weakly defined, internal surface covered with fine granular shagreen, weakly shining, with weak and obscure rugae basally (
<figureCitation id="71B92A6FAC1743350E1FFA488BF503DC" box="[1291,1424,1410,1437]" captionStart="FIGURE 52" captionStartId="79.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-16@79.[151,1436,181,1621]" captionTargetPageId="79" captionText="FIGURE 52. Andrena (Aciandrena) elbensis spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209417" httpUri="https://zenodo.org/record/13209417/files/figure.png" pageId="78" pageNumber="79">Figure 52E</figureCitation>
). Mesepisternum with long white hairs, none equalling length of scape, scutum and scutellum with shorter scattered semi-squamous hairs, most concentrated laterally. Propodeal corbicula incomplete, weakly formed, dorsal fringe composed of whitish-yellowish plumose hairs, internal surface with scattered simple white hairs. Legs basally brownish, tarsi and hind tibiae lightened orange, pubescence light brown. Flocculus incomplete, composed of light brownish plumose hairs; femoral and tibial scopae composed of simple yellowish hairs. Hind tarsal claws without inner tooth. Wings hyaline, stigma and venation orange, nervulus interstitial to marginally slightly antefurcal.
</paragraph>
<paragraph id="E93D36EAAC1743350BD3F9B48CD5010C" blockId="78.[151,1437,870,1905]" pageId="78" pageNumber="79">
<emphasis id="DBF6EAF8AC1743350BD3F9B48F2000D9" box="[199,325,1663,1688]" italics="true" pageId="78" pageNumber="79">Metasoma:</emphasis>
Tergal discs dark, tergal margins broadly lightened reddish-brown (
<figureCitation id="71B92A6FAC1743350F2FF9B48AD900D8" box="[1083,1212,1662,1689]" captionStart="FIGURE 52" captionStartId="79.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-16@79.[151,1436,181,1621]" captionTargetPageId="79" captionText="FIGURE 52. Andrena (Aciandrena) elbensis spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209417" httpUri="https://zenodo.org/record/13209417/files/figure.png" pageId="78" pageNumber="79">Figure 52F</figureCitation>
). Tergal discs finely shagreened to microreticulate, weakly shining, more or less impunctate, with obscure fine and scattered punctures. T2–4 apically with dense white hairbands, broadly interrupted on T2, complete on T3–4, obscuring underlying surface.Apical fringe of T5 and hairs flanking pygidial plate whitish-golden. Pygidial plate large, rounded triangular, apex slightly domed, impunctate, basal half with surface densely but shallowly punctate, punctures separated by 0.5 puncture diameters, interspaces weakly shining.
</paragraph>
<paragraph id="E93D36EAAC1743350BD3F89C8FEF0130" blockId="78.[151,1437,870,1905]" box="[199,394,1879,1905]" pageId="78" pageNumber="79">
<emphasis id="DBF6EAF8AC1743350BD3F89C8F680130" bold="true" box="[199,269,1879,1905]" pageId="78" pageNumber="79">Male:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<caption id="BDFD6662AC1643340B83F9A68A7000E8" ID-DOI="http://doi.org/10.5281/zenodo.13209417" ID-Zenodo-Dep="13209417" httpUri="https://zenodo.org/record/13209417/files/figure.png" pageId="79" pageNumber="80" startId="79.[151,250,1645,1669]" targetBox="[151,1436,181,1621]" targetPageId="79" targetType="figure">
<paragraph id="E93D36EAAC1643340B83F9A68A7000E8" blockId="79.[151,1436,1645,1706]" pageId="79" pageNumber="80">
<emphasis id="DBF6EAF8AC1643340B83F9A68F4700C4" bold="true" box="[151,290,1645,1669]" pageId="79" pageNumber="80">FIGURE 52.</emphasis>
<taxonomicName id="2E824D69AC1643340A3EF9A68C0D00C4" authorityName="WOOD" authorityYear="2024" box="[298,616,1645,1670]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="elbensis" status="sp. nov." subGenus="Aciandrena">
<emphasis id="DBF6EAF8AC1643340A3EF9A68FE500C4" box="[298,384,1645,1669]" italics="true" pageId="79" pageNumber="80">Andrena</emphasis>
(
<emphasis id="DBF6EAF8AC1643340A85F9A68C6300C4" box="[401,518,1645,1669]" italics="true" pageId="79" pageNumber="80">Aciandrena</emphasis>
)
<emphasis id="DBF6EAF8AC1643340902F9A68C0D00C4" box="[534,616,1645,1669]" italics="true" pageId="79" pageNumber="80">elbensis</emphasis>
</taxonomicName>
<emphasis id="DBF6EAF8AC1643340964F9A58CBC00C7" bold="true" box="[624,729,1646,1670]" pageId="79" pageNumber="80">
<taxonomicNameLabel id="C0C55783AC1643340964F9A58CBC00C7" box="[624,729,1646,1670]" pageId="79" pageNumber="80" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view.
</paragraph>
</caption>
<subSubSection id="A1986561AC16432B0BD3F91C8BE30578" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80" type="diagnosis">
<paragraph id="E93D36EAAC16432B0BD3F91C8A3B075C" blockId="79.[151,1437,1751,2030]" lastBlockId="80.[151,1437,151,969]" lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
<emphasis id="DBF6EAF8AC1643340BD3F91C8F2700B0" bold="true" box="[199,322,1751,1777]" pageId="79" pageNumber="80">Diagnosis.</emphasis>
It is important to note that the current definition of the subgenus
<taxonomicName id="2E824D69AC1643340F5CF9138B1A00B3" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" box="[1096,1407,1752,1778]" pageId="79" pageNumber="80" rank="subGenus" subGenus="Aciandrena">
<emphasis id="DBF6EAF8AC1643340F5CF9138AAF00B3" box="[1096,1226,1752,1778]" italics="true" pageId="79" pageNumber="80">Aciandrena</emphasis>
Warncke, 1968
</taxonomicName>
is unclear (
<bibRefCitation id="8D134B1BAC1643340BECF9378FBE0157" author="Pisanty, G. &amp; Richter, R. &amp; Martin, T. &amp; Dettman, J. &amp; Cardinal, S." box="[248,475,1788,1814]" pageId="79" pageNumber="80" refId="ref90124" refString="Pisanty, G., Richter, R., Martin, T., Dettman, J. &amp; Cardinal, S. (2022 b) Molecular phylogeny, historical biogeography and revised classification of andrenine bees (Hymenoptera: Andrenidae). Molecular Phylogenetics and Evolution, 170, 107151. https: // doi. org / 10.1016 / j. ympev. 2021.107151" type="journal volume" year="2022">
Pisanty
<emphasis id="DBF6EAF8AC1643340A46F9368FE60157" box="[338,387,1788,1814]" italics="true" pageId="79" pageNumber="80">et al</emphasis>
. 2022b
</bibRefCitation>
). Revisions are ongoing (S. Bossert, T.J. Wood, G. Pisanty,
<emphasis id="DBF6EAF8AC1643340F67F9378B510157" box="[1139,1332,1788,1814]" italics="true" pageId="79" pageNumber="80">unpublished data</emphasis>
), and the diagnoses presented here should be seen in this context.
<taxonomicName id="2E824D69AC164334081EF8EB8DAC017B" authorityName="WOOD" authorityYear="2024" box="[778,969,1824,1850]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="elbensis">
<emphasis id="DBF6EAF8AC164334081EF8EB8DAC017B" box="[778,969,1824,1850]" italics="true" pageId="79" pageNumber="80">Andrena elbensis</emphasis>
</taxonomicName>
can be recognised as part of the subgenus
<taxonomicName id="2E824D69AC1643340B83F88F8F7C011F" authorityName="Warncke" authorityYear="1968" box="[151,281,1860,1886]" pageId="79" pageNumber="80" rank="subGenus" subGenus="Aciandrena">
<emphasis id="DBF6EAF8AC1643340B83F88F8F7C011F" box="[151,281,1860,1886]" italics="true" pageId="79" pageNumber="80">Aciandrena</emphasis>
</taxonomicName>
due to the small body size, propodeal triangle not defined by lateral carinae, surface covered by fine granular shagreen, with weak and obscure rugae basally, facial foveae narrower ventrally than dorsally but which are not so narrow as to be linear, domed and striation-free clypeus (also lacking striations on the supraclypeal area and lower paraocular areas), and hind tibial claws lacking an inner tooth. Due to the tarsi and hind tibiae that are lightened orange, the short, sparse, and semi-squamous hairs on the scutum and scutellum, and thick white hairbands on the terga it can be placed close to
<taxonomicName id="2E824D69AC09432B0955FF5C8D1E06F0" authority="Warncke, 1974" authorityName="Warncke" authorityYear="1974" box="[577,891,151,177]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pellucida">
<emphasis id="DBF6EAF8AC09432B0955FF5C8CAC06F0" box="[577,713,151,177]" italics="true" pageId="80" pageNumber="81">A. pellucida</emphasis>
<bibRefCitation id="8D134B1BAC09432B09C5FF5C8D1E06F0" author="Warncke, K." box="[721,891,151,177]" pageId="80" pageNumber="81" pagination="3 - 54" refId="ref91760" refString="Warncke, K. (1974) Beitrag zur Kenntnis und Verbreitung der Sandbienen in Nordafrika (Hymenoptera, Apoidea, Andrena). Mitteilungen aus dem Zoologischen Museum in Berlin, 50, 3 - 54." type="journal article" year="1974">Warncke, 1974</bibRefCitation>
</taxonomicName>
(eastern
<collectingCountry id="9195767AAC09432B08F1FF5C8A4306F0" box="[997,1062,151,177]" name="Libya" pageId="80" pageNumber="81">Libya</collectingCountry>
and northern
<collectingCountry id="9195767AAC09432B0FD2FF5C8B6806F0" box="[1222,1293,151,177]" name="Egypt" pageId="80" pageNumber="81">Egypt</collectingCountry>
),
<taxonomicName id="2E824D69AC09432B0E36FF5C8F5A0694" authority="Warncke, 1974" authorityName="Warncke" authorityYear="1974" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pavonia">
<emphasis id="DBF6EAF8AC09432B0E36FF5C8BF906F1" box="[1314,1436,151,177]" italics="true" pageId="80" pageNumber="81">A. pavonia</emphasis>
<bibRefCitation id="8D134B1BAC09432B0B83FF708F5A0694" author="Warncke, K." box="[151,319,187,213]" pageId="80" pageNumber="81" pagination="3 - 54" refId="ref91760" refString="Warncke, K. (1974) Beitrag zur Kenntnis und Verbreitung der Sandbienen in Nordafrika (Hymenoptera, Apoidea, Andrena). Mitteilungen aus dem Zoologischen Museum in Berlin, 50, 3 - 54." type="journal article" year="1974">Warncke, 1974</bibRefCitation>
</taxonomicName>
(eastern
<collectingCountry id="9195767AAC09432B0AB6FF708F930694" box="[418,502,187,213]" name="Algeria" pageId="80" pageNumber="81">Algeria</collectingCountry>
and
<collectingCountry id="9195767AAC09432B093FFF708CE60694" box="[555,643,187,213]" name="Tunisia" pageId="80" pageNumber="81">Tunisia</collectingCountry>
),
<taxonomicName id="2E824D69AC09432B0980FF708DD40694" authority="Wood, 2021" authorityName="Wood" authorityYear="2021" box="[660,945,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="palmyriae">
<emphasis id="DBF6EAF8AC09432B0980FF708D410694" box="[660,804,187,213]" italics="true" pageId="80" pageNumber="81">A. palmyriae</emphasis>
Wood, 2021
</taxonomicName>
(
<collectingCountry id="9195767AAC09432B08ABFF708D9A0694" box="[959,1023,187,213]" name="Syria" pageId="80" pageNumber="81">Syria</collectingCountry>
), and
<taxonomicName id="2E824D69AC09432B0F54FF708B490694" authority="Pisanty, 2022" authorityName="Pisanty" authorityYear="2022" box="[1088,1324,187,213]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="xera">
<emphasis id="DBF6EAF8AC09432B0F54FF708AF40695" box="[1088,1169,187,213]" italics="true" pageId="80" pageNumber="81">A. xera</emphasis>
Pisanty, 2022
</taxonomicName>
(southern
<collectingCountry id="9195767AAC09432B0B83FF148EBF06B8" box="[151,218,223,249]" name="Israel" pageId="80" pageNumber="81">Israel</collectingCountry>
). These desert-dwelling
<taxonomicName id="2E824D69AC09432B0AFCFF148C0F06B8" authorityName="Warncke" authorityYear="1968" box="[488,618,223,249]" pageId="80" pageNumber="106" rank="subGenus" subGenus="Aciandrena">
<emphasis id="DBF6EAF8AC09432B0AFCFF148C0F06B8" box="[488,618,223,249]" italics="true" pageId="80" pageNumber="81">Aciandrena</emphasis>
</taxonomicName>
have highly local distributions, and
<taxonomicName id="2E824D69AC09432B08E9FF148A1006B8" authorityName="WOOD" authorityYear="2024" box="[1021,1141,223,249]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="elbensis">
<emphasis id="DBF6EAF8AC09432B08E9FF148A1006B8" box="[1021,1141,223,249]" italics="true" pageId="80" pageNumber="81">A. elbensis</emphasis>
</taxonomicName>
is strongly geographically separated from the nearest populations in northern
<collectingCountry id="9195767AAC09432B09C5FEC88D71075C" box="[721,788,259,285]" name="Egypt" pageId="80" pageNumber="81">Egypt</collectingCountry>
(Borg el-Arab,
<taxonomicName id="2E824D69AC09432B08D2FEC88A28075C" authorityName="Warncke" authorityYear="1974" box="[966,1101,259,285]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pellucida">
<emphasis id="DBF6EAF8AC09432B08D2FEC88A28075C" box="[966,1101,259,285]" italics="true" pageId="80" pageNumber="81">A. pellucida</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="E93D36EAAC09432B0BD3FEEC8BE30578" blockId="80.[151,1437,151,969]" pageId="80" pageNumber="81">
Morphological separation can be made by the interstitial nervulus (antefurcal in
<taxonomicName id="2E824D69AC09432B0F4AFEEC8ABD0701" authorityName="Warncke" authorityYear="1974" box="[1118,1240,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pavonia">
<emphasis id="DBF6EAF8AC09432B0F4AFEEC8ABD0701" box="[1118,1240,295,321]" italics="true" pageId="80" pageNumber="81">A. pavonia</emphasis>
</taxonomicName>
,
<taxonomicName id="2E824D69AC09432B0FFDFEEC8B5E0701" authorityName="Pisanty" authorityYear="2022" box="[1257,1339,295,321]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="xera">
<emphasis id="DBF6EAF8AC09432B0FFDFEEC8B5E0701" box="[1257,1339,295,321]" italics="true" pageId="80" pageNumber="81">A. xera</emphasis>
</taxonomicName>
, weakly antefurcal in
<taxonomicName id="2E824D69AC09432B0A26FE808FA00724" authorityName="Wood" authorityYear="2021" box="[306,453,331,357]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="palmyriae">
<emphasis id="DBF6EAF8AC09432B0A26FE808FA00724" box="[306,453,331,357]" italics="true" pageId="80" pageNumber="81">A. palmyriae</emphasis>
</taxonomicName>
), smooth and shining scutum and scutellum (
<figureCitation id="71B92A6FAC09432B08CDFE808A040724" box="[985,1121,330,357]" captionStart="FIGURE 52" captionStartId="79.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-16@79.[151,1436,181,1621]" captionTargetPageId="79" captionText="FIGURE 52. Andrena (Aciandrena) elbensis spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209417" httpUri="https://zenodo.org/record/13209417/files/figure.png" pageId="80" pageNumber="81">Figure 52D</figureCitation>
; shagreened in
<taxonomicName id="2E824D69AC09432B0E0DFE808BF00725" authorityName="Warncke" authorityYear="1974" box="[1305,1429,331,357]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pavonia">
<emphasis id="DBF6EAF8AC09432B0E0DFE808BF00725" box="[1305,1429,331,357]" italics="true" pageId="80" pageNumber="81">A. pavonia</emphasis>
</taxonomicName>
, laterally shagreened and medially shining in
<taxonomicName id="2E824D69AC09432B0986FEA48C8607C9" authorityName="Pisanty" authorityYear="2022" box="[658,739,367,393]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="xera">
<emphasis id="DBF6EAF8AC09432B0986FEA48C8607C9" box="[658,739,367,393]" italics="true" pageId="80" pageNumber="81">A. xera</emphasis>
</taxonomicName>
), head only 1.1 times wider than long (
<figureCitation id="71B92A6FAC09432B0FB0FEA48B4D07C8" box="[1188,1320,366,393]" captionStart="FIGURE 52" captionStartId="79.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-16@79.[151,1436,181,1621]" captionTargetPageId="79" captionText="FIGURE 52. Andrena (Aciandrena) elbensis spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209417" httpUri="https://zenodo.org/record/13209417/files/figure.png" pageId="80" pageNumber="81">Figure 52B</figureCitation>
;
<figureCitation id="71B92A6FAC09432B0E21FEA48B3D07C8" box="[1333,1368,367,393]" captionStart="FIGURE 1" captionStartId="6.[151,250,1218,1242]" captionTargetBox="[151,1436,237,1194]" captionTargetId="figure-45@6.[151,1436,237,1194]" captionTargetPageId="6" captionText="FIGURE 1. Andrena (Avandrena) avara Warncke, 1967 sensu stricto. A. Label details; B. Habitus, lateral view; C. Face, frontal view; D. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209309" httpUri="https://zenodo.org/record/13209309/files/figure.png" pageId="80" pageNumber="81">1.2</figureCitation>
times wider than long in
<taxonomicName id="2E824D69AC09432B0A7EFE588F9507EC" authorityName="Warncke" authorityYear="1974" box="[362,496,403,429]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pellucida">
<emphasis id="DBF6EAF8AC09432B0A7EFE588F9507EC" box="[362,496,403,429]" italics="true" pageId="80" pageNumber="81">A. pellucida</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC09432B0932FE588C1307ED" authorityName="Pisanty" authorityYear="2022" box="[550,630,403,429]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="xera">
<emphasis id="DBF6EAF8AC09432B0932FE588C1307ED" box="[550,630,403,429]" italics="true" pageId="80" pageNumber="81">A. xera</emphasis>
</taxonomicName>
, 1.3 times wider than long in
<taxonomicName id="2E824D69AC09432B08D0FE588A3607EC" authorityName="Wood" authorityYear="2021" box="[964,1107,403,429]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="palmyriae">
<emphasis id="DBF6EAF8AC09432B08D0FE588A3607EC" box="[964,1107,403,429]" italics="true" pageId="80" pageNumber="81">A. palmyriae</emphasis>
</taxonomicName>
and
<taxonomicName id="2E824D69AC09432B0F9EFE588B6707ED" authorityName="Warncke" authorityYear="1974" box="[1162,1282,403,429]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pavonia">
<emphasis id="DBF6EAF8AC09432B0F9EFE588B6707ED" box="[1162,1282,403,429]" italics="true" pageId="80" pageNumber="81">A. pavonia</emphasis>
</taxonomicName>
), and process of the labrum with anterior margin straight (deeply notched in
<taxonomicName id="2E824D69AC09432B0849FE7C8D880790" authorityName="Wood" authorityYear="2021" box="[861,1005,439,465]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="palmyriae">
<emphasis id="DBF6EAF8AC09432B0849FE7C8D880790" box="[861,1005,439,465]" italics="true" pageId="80" pageNumber="81">A. palmyriae</emphasis>
</taxonomicName>
, weakly notched in
<taxonomicName id="2E824D69AC09432B0FC0FE7C8B400791" authorityName="Pisanty" authorityYear="2022" box="[1236,1317,439,465]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="xera">
<emphasis id="DBF6EAF8AC09432B0FC0FE7C8B400791" box="[1236,1317,439,465]" italics="true" pageId="80" pageNumber="81">A. xera</emphasis>
</taxonomicName>
).
<taxonomicName id="2E824D69AC09432B0E2AFE7C8E9507B4" authorityName="WOOD" authorityYear="2024" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="elbensis">
<emphasis id="DBF6EAF8AC09432B0E2AFE7C8E9507B4" italics="true" pageId="80" pageNumber="81">Andrena elbensis</emphasis>
</taxonomicName>
can be further separated from
<taxonomicName id="2E824D69AC09432B0928FE108CA507B4" authorityName="Warncke" authorityYear="1974" box="[572,704,475,501]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pellucida">
<emphasis id="DBF6EAF8AC09432B0928FE108CA507B4" box="[572,704,475,501]" italics="true" pageId="80" pageNumber="81">A. pellucida</emphasis>
</taxonomicName>
by the facial foveae which dorsally occupy ⅔ of the space between the compound eye and a lateral ocellus and are filled with white hairs (in
<taxonomicName id="2E824D69AC09432B08F5FE348A0C0458" authorityName="Warncke" authorityYear="1974" box="[993,1129,511,537]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pellucida">
<emphasis id="DBF6EAF8AC09432B08F5FE348A0C0458" box="[993,1129,511,537]" italics="true" pageId="80" pageNumber="81">A. pellucida</emphasis>
</taxonomicName>
facial foveae occupying ½ space between the compound eye and a lateral ocellus, filled with brown hairs), the scutum which is irregularly punctate by punctures of variable size, punctures separated by 0.5–2 puncture diameters (in
<taxonomicName id="2E824D69AC09432B0F9CFD8C8B680420" authorityName="Warncke" authorityYear="1974" box="[1160,1293,583,609]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pellucida">
<emphasis id="DBF6EAF8AC09432B0F9CFD8C8B680420" box="[1160,1293,583,609]" italics="true" pageId="80" pageNumber="81">A. pellucida</emphasis>
</taxonomicName>
punctures of scutum sparser, separated by 2–3 puncture diameters), scutellum more irregularly punctate, punctures separated by 0.5–3 puncture diameters (in
<taxonomicName id="2E824D69AC09432B0AF6FD448C0C04E8" authorityName="Warncke" authorityYear="1974" box="[482,617,655,681]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pellucida">
<emphasis id="DBF6EAF8AC09432B0AF6FD448C0C04E8" box="[482,617,655,681]" italics="true" pageId="80" pageNumber="81">A. pellucida</emphasis>
</taxonomicName>
scutellum more regularly punctate, punctures separated by 1–2 puncture diameters), T2–4 with thick apical hairbands of white hairs (
<figureCitation id="71B92A6FAC09432B0820FD788DD1048C" box="[820,948,690,717]" captionStart="FIGURE 52" captionStartId="79.[151,250,1645,1669]" captionTargetBox="[151,1436,181,1621]" captionTargetId="figure-16@79.[151,1436,181,1621]" captionTargetPageId="79" captionText="FIGURE 52. Andrena (Aciandrena) elbensis spec. nov. female. A. Habitus, lateral view; B. Face, frontal view; C. Clypeus, frontal view; D. Scutum, dorsal view; E. Propodeum, dorsal view; F. Terga, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13209417" httpUri="https://zenodo.org/record/13209417/files/figure.png" pageId="80" pageNumber="81">Figure 52F</figureCitation>
; broadly interrupted on T2) that obscure the underlying surface (in
<taxonomicName id="2E824D69AC09432B0A80FD1C8C7F04B0" authorityName="Warncke" authorityYear="1974" box="[404,538,727,753]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pellucida">
<emphasis id="DBF6EAF8AC09432B0A80FD1C8C7F04B0" box="[404,538,727,753]" italics="true" pageId="80" pageNumber="81">A. pellucida</emphasis>
</taxonomicName>
with only weak and obscure apical hairbands that do not obscure the underlying surface), and by the clypeus which is strongly domed and smooth and shining over its entire surface (in
<taxonomicName id="2E824D69AC09432B0E02FD308BF90554" authorityName="Warncke" authorityYear="1974" box="[1302,1436,763,789]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="pellucida">
<emphasis id="DBF6EAF8AC09432B0E02FD308BF90554" box="[1302,1436,763,789]" italics="true" pageId="80" pageNumber="81">A. pellucida</emphasis>
</taxonomicName>
with the clypeus comparatively flat, only weakly domed, shiny medially but with shagreenation present laterally).
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC09432B0BD3FC888D4705C0" pageId="80" pageNumber="81" type="etymology">
<paragraph id="E93D36EAAC09432B0BD3FC888D4705C0" blockId="80.[151,1437,151,969]" pageId="80" pageNumber="81">
<emphasis id="DBF6EAF8AC09432B0BD3FC888F2B051C" bold="true" box="[199,334,835,861]" pageId="80" pageNumber="81">Etymology.</emphasis>
The name is taken from the mountain Gabal Elba in south-eastern
<collectingCountry id="9195767AAC09432B0F3FFC888A0B051C" box="[1067,1134,835,861]" name="Egypt" pageId="80" pageNumber="81">Egypt</collectingCountry>
(literally “Box Mountain”) with the Latin suffix
<emphasis id="DBF6EAF8AC09432B0A97FCAC8FA005C1" box="[387,453,871,896]" italics="true" pageId="80" pageNumber="81">-ensis</emphasis>
, indicating “originating from”.
</paragraph>
</subSubSection>
<subSubSection id="A1986561AC09432B0BD3FC418F3B0588" pageId="80" pageNumber="81" type="distribution">
<paragraph id="E93D36EAAC09432B0BD3FC418F3B0588" blockId="80.[151,1437,151,969]" pageId="80" pageNumber="81">
<emphasis id="DBF6EAF8AC09432B0BD3FC418F0405E5" bold="true" box="[199,353,906,932]" pageId="80" pageNumber="81">Distribution.</emphasis>
South-eastern
<collectingCountry id="9195767AAC09432B091AFC408C3405E4" box="[526,593,907,933]" name="Egypt" pageId="80" pageNumber="81">Egypt</collectingCountry>
(
<emphasis id="DBF6EAF8AC09432B0971FC408CA405E4" box="[613,705,907,933]" italics="true" pageId="80" pageNumber="81">de facto</emphasis>
), though this area is currently disputed and is claimed by both
<collectingCountry id="9195767AAC09432B0B83FC648EBF0588" box="[151,218,943,969]" name="Egypt" pageId="80" pageNumber="81">Egypt</collectingCountry>
and
<collectingCountry id="9195767AAC09432B0A05FC648F3E0588" box="[273,347,943,969]" name="Sudan" pageId="80" pageNumber="81">Sudan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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