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<mods:title id="A157F11CD8F79633FA1EA8B54FF35EDF">Two new species of Cyanopenthe Nikitsky from Taiwan (Coleoptera, Tetratomidae, PentHinae)</mods:title>
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<mods:namePart id="9DFD23109486A974A331DF3619B22A4E">Hsiao, Yun</mods:namePart>
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<mods:namePart id="95278B31149BB8F7A325C669E36183BB">Barclay, Maxwell V. L.</mods:namePart>
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<paragraph id="AD697F555D1CFFAFEAFCFF31FC27F1E2" blockId="10.[151,956,151,178]" box="[151,956,151,178]" pageId="10" pageNumber="588">
<heading id="F621C8395D1CFFAFEAFCFF31FC27F1E2" bold="true" box="[151,956,151,178]" fontSize="11" level="1" pageId="10" pageNumber="588" reason="1">
<emphasis id="9FA2A3475D1CFFAFEAFCFF31FC27F1E2" bold="true" box="[151,956,151,178]" pageId="10" pageNumber="588">
Key to the world fauna of the Genus
<taxonomicName id="6AD604D65D1CFFAFE808FF3EFC27F1E2" authority="Nikitsky, 1998" authorityName="Nikitsky" authorityYear="1998" box="[611,956,151,178]" class="Insecta" family="Tetratomidae" genus="Cyanopenthe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="588" phylum="Arthropoda" rank="genus">
<emphasis id="9FA2A3475D1CFFAFE808FF3EFC99F1E1" bold="true" box="[611,770,151,177]" italics="true" pageId="10" pageNumber="588">Cyanopenthe</emphasis>
<bibRefCitation id="C94702A45D1CFFAFE961FF31FC27F1E2" author="Nikitsky" box="[778,956,152,178]" pageId="10" pageNumber="588" refString="Nikitsky, N. B. (1998) Generic Classification of the Beetle Family Tetratomidae (Coleoptera, Tenebrionoidea) of the World, with Description of New Taxa. Series Faunistika (9). Pensoft, Sofia-Moscow, 80 pp." type="book" year="1998">Nikitsky, 1998</bibRefCitation>
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<paragraph id="AD697F555D1CFFAFEAFCFF77FA0DF040" blockId="10.[151,1436,222,625]" pageId="10" pageNumber="588">
<keyLead id="1627DC605D1CFFAFEAFCFF77FA0DF040" pageId="10" pageNumber="588">
1. Pronotum densely and coarsely punctate; scutellum bronzed or reddish bronzed, triangular, without distinct impression (
<figureCitation id="35ED63D05D1CFFAFEF1EFF77FF77F040" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1952,1975]" captionTargetBox="[394,1191,195,1931]" captionTargetId="figure@2.[378,1209,193,1932]" captionTargetPageId="2" captionText="FIGURES 1 6. Habitus, dorsal view: 1. Cyanopenthe metallica (Champion, 1916), female holotype; 2. C. thailandica Nikitsky, 1998, female holotype; 3 4. C. taiwana sp. nov., 3: male holotype, 4: female paratype; 5 6. C. leei sp. nov., 5: male paratype, 6: female holotype. Scale bars = 5 mm. (Figure 2 from Nikitsky, 1998)" httpUri="https://zenodo.org/record/237510/files/figure.png" pageId="10" pageNumber="588">Fig. 1</figureCitation>
).
<collectingCountry id="D5C13FC55D1CFFAFEB6AFF52FEDDF040" box="[257,326,251,272]" name="Bhutan" pageId="10" pageNumber="588">Bhutan</collectingCountry>
and North
<collectingCountry id="D5C13FC55D1CFFAFEBD9FF52FE78F040" box="[434,483,251,272]" name="India" pageId="10" pageNumber="588">India</collectingCountry>
.....................................................
<taxonomicName id="6AD604D65D1CFFAFEE01FF52FA0DF040" authority="Champion, 1916" authorityName="Champion" authorityYear="1916" box="[1130,1430,251,272]" class="Insecta" family="Tetratomidae" genus="Cyanopenthe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="588" phylum="Arthropoda" rank="species" species="metallica">
<emphasis id="9FA2A3475D1CFFAFEE01FF52FB45F040" box="[1130,1246,251,272]" italics="true" pageId="10" pageNumber="588">C. metallica</emphasis>
(
<bibRefCitation id="C94702A45D1CFFAFEE87FF52FA14F040" author="Champion" box="[1260,1423,251,272]" pageId="10" pageNumber="588" refString="Champion, G. C. (1916) Notes on Melandryidae. The Entomologist's monthly magazine, 52, 1 - 157." type="journal article" year="1916">Champion, 1916</bibRefCitation>
)
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<paragraph id="AD697F555D1CFFAFEAFCFEB0FA0EF01B" blockId="10.[151,1436,222,625]" pageId="10" pageNumber="588">
<keyLead id="1627DC605D1CFFAFEAFCFEB0FA0EF01B" pageId="10" pageNumber="588">- Pronotum sparsely and finely punctate; scutellum black, transverse, with rounded apex, with distinct, dark rounded impression in middle........................................................................................... 2</keyLead>
</paragraph>
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<keyStep id="162267F05D1CFFAFEAFCFEFAFA00F090" pageId="10" pageNumber="588">
<paragraph id="AD697F555D1CFFAFEAFCFEFAFA00F0D6" blockId="10.[151,1436,222,625]" pageId="10" pageNumber="588">
<keyLead id="1627DC605D1CFFAFEAFCFEFAFA00F0D6" pageId="10" pageNumber="588">
2. Antennomeres IIIV slightly more slender, projections of antennomeres VIIIIX longer than that of antennomere X (
<figureCitation id="35ED63D05D1CFFAFEF38FEFAFA15F038" box="[1363,1422,339,360]" captionStart="FIGURES 7 12" captionStartId="3.[151,264,897,920]" captionTargetBox="[161,1425,197,872]" captionTargetId="figure@3.[151,1436,193,877]" captionTargetPageId="3" captionText="FIGURES 7 12. Antenna: 7. Cyanopenthe metallica (Champion, 1916); 8. C. thailandica Nikitsky, 1998 (from Nikitsky, 1998); 9 10. C. taiwana sp. nov., 9: male, 10: female; 11 12. C. leei sp. nov., 11: male, 12: female. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/237511/files/figure.png" pageId="10" pageNumber="588">Fig. 8</figureCitation>
); paraproct shorter, about 1.2X as long as wide (
<figureCitation id="35ED63D05D1CFFAFE8F4FED8FD7CF0D6" box="[671,743,369,390]" captionStart="FIGURES 26 29" captionStartId="9.[151,264,865,888]" captionTargetBox="[316,1260,193,837]" captionTargetId="figure@9.[296,1291,193,844]" captionTargetPageId="9" captionText="FIGURES 26 29. Ovipositor, ventral view: 26. Cyanopenthe. metallica (Champion, 1916); 27. C. thailandica Nikitsky, 1998; 28. C. taiwana sp. nov.; 29. C. leei sp. nov. Scale bars = 0.5 mm. (Figures 27 from Nikitsky, 2005)" httpUri="https://zenodo.org/record/237516/files/figure.png" pageId="10" pageNumber="588">Fig. 27</figureCitation>
).
<collectingCountry id="D5C13FC55D1CFFAFE893FED8FCD6F0D6" box="[760,845,369,390]" name="Thailand" pageId="10" pageNumber="588">Thailand</collectingCountry>
...................
<taxonomicName id="6AD604D65D1CFFAFEE5AFED8FAD5F0D6" authority="Nikitsky, 1998" authorityName="Nikitsky" authorityYear="1998" box="[1073,1358,369,390]" class="Insecta" family="Tetratomidae" genus="Cyanopenthe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="588" phylum="Arthropoda" rank="species" species="thailandica">
<emphasis id="9FA2A3475D1CFFAFEE5AFED8FB21F0D6" box="[1073,1210,369,390]" italics="true" pageId="10" pageNumber="588">C. thailandica</emphasis>
<bibRefCitation id="C94702A45D1CFFAFEEAAFED8FAD5F0D6" author="Nikitsky" box="[1217,1358,369,390]" pageId="10" pageNumber="588" refString="Nikitsky, N. B. (1998) Generic Classification of the Beetle Family Tetratomidae (Coleoptera, Tenebrionoidea) of the World, with Description of New Taxa. Series Faunistika (9). Pensoft, Sofia-Moscow, 80 pp." type="book" year="1998">Nikitsky, 1998</bibRefCitation>
</taxonomicName>
(
<figureCitation id="35ED63D05D1CFFAFEF30FED8FA0EF0D6" box="[1371,1429,369,390]" captionStart="FIGURES 1 6" captionStartId="2.[151,264,1952,1975]" captionTargetBox="[394,1191,195,1931]" captionTargetId="figure@2.[378,1209,193,1932]" captionTargetPageId="2" captionText="FIGURES 1 6. Habitus, dorsal view: 1. Cyanopenthe metallica (Champion, 1916), female holotype; 2. C. thailandica Nikitsky, 1998, female holotype; 3 4. C. taiwana sp. nov., 3: male holotype, 4: female paratype; 5 6. C. leei sp. nov., 5: male paratype, 6: female holotype. Scale bars = 5 mm. (Figure 2 from Nikitsky, 1998)" httpUri="https://zenodo.org/record/237510/files/figure.png" pageId="10" pageNumber="588">Fig. 2</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="AD697F555D1CFFAFEAFCFE27FA00F090" blockId="10.[151,1436,222,625]" pageId="10" pageNumber="588">
<keyLead id="1627DC605D1CFFAFEAFCFE27FA00F090" pageId="10" pageNumber="588">
- Antennomeres IIIV slightly thicker, projections of antennomeres VIIIIX as long as or shorter than that of antennomere X; paraproct longer, about 1.41.6X width.
<collectingCountry id="D5C13FC55D1CFFAFE80CFE02FD36F090" box="[615,685,427,448]" name="Taiwan" pageId="10" pageNumber="588">Taiwan</collectingCountry>
............................................................. 3
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<keyStep id="162267F05D1CFFAFEAFCFE60FA07F321" pageId="10" pageNumber="588">
<paragraph id="AD697F555D1CFFAFEAFCFE60FA07F349" blockId="10.[151,1436,222,625]" pageId="10" pageNumber="588">
<keyLead id="1627DC605D1CFFAFEAFCFE60FA07F349" pageId="10" pageNumber="588">
3. Elytra and abdomen more rounded (
<figureCitation id="35ED63D05D1CFFAFE850FE60FD19F08E" box="[571,642,457,478]" captionStart="FIGURES 15 16" captionStartId="5.[151,264,1248,1271]" captionTargetBox="[157,1426,561,1220]" captionTargetId="figure@5.[151,1436,553,1227]" captionTargetPageId="5" captionText="FIGURES 15 16. Abdomen: 15. Cyanopenthe taiwana sp. nov., paratype female; 16. C. leei sp. nov., holotype female. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/237513/files/figure.png" pageId="10" pageNumber="588">Fig. 15</figureCitation>
), ratio of body width to length 1.0:1.41.5; anterolateral margin of pronotum more rounded (
<figureCitation id="35ED63D05D1CFFAFEB57FE4FFE18F0AB" box="[316,387,486,507]" captionStart="FIGURES 13 14" captionStartId="4.[151,264,1648,1671]" captionTargetBox="[382,1205,204,1620]" captionTargetId="figure@4.[370,1216,193,1627]" captionTargetPageId="4" captionText="FIGURES 13 14. Right half of pronotum: 13. Cyanopenthe taiwana sp. nov.; 14. C. leei sp. nov. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/237512/files/figure.png" pageId="10" pageNumber="588">Fig. 13</figureCitation>
); aedeagus: lateral margins of apical piece of tegmen slightly convergent distally, median lobe about 1.05X as long as tegmen; ovipositor: proctiger slightly wider in ventral view...............................
<taxonomicName id="6AD604D65D1CFFAFEEB4FDAAFAD3F348" box="[1247,1352,515,536]" class="Insecta" family="Tetratomidae" genus="Cyanopenthe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="588" phylum="Arthropoda" rank="species" species="taiwana" status="sp. nov.">
<emphasis id="9FA2A3475D1CFFAFEEB4FDAAFAD3F348" box="[1247,1352,515,536]" italics="true" pageId="10" pageNumber="588">C. taiwana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84911E3C5D1CFFAFEF24FDAAFA07F349" box="[1359,1436,515,537]" pageId="10" pageNumber="588" rank="species">
<emphasis id="9FA2A3475D1CFFAFEF24FDAAFA07F349" bold="true" box="[1359,1436,515,537]" pageId="10" pageNumber="588">sp. nov.</emphasis>
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</paragraph>
<paragraph id="AD697F555D1CFFAFEAFCFD88FA07F321" blockId="10.[151,1436,222,625]" pageId="10" pageNumber="588">
<keyLead id="1627DC605D1CFFAFEAFCFD88FA07F321" pageId="10" pageNumber="588">
- Elytra and abdomen more elongated (
<figureCitation id="35ED63D05D1CFFAFE822FD88FD0BF366" box="[585,656,545,566]" captionStart="FIGURES 15 16" captionStartId="5.[151,264,1248,1271]" captionTargetBox="[157,1426,561,1220]" captionTargetId="figure@5.[151,1436,553,1227]" captionTargetPageId="5" captionText="FIGURES 15 16. Abdomen: 15. Cyanopenthe taiwana sp. nov., paratype female; 16. C. leei sp. nov., holotype female. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/237513/files/figure.png" pageId="10" pageNumber="588">Fig. 16</figureCitation>
), ratio of body width to length 1.0:1.51.6; anterolateral margin of pronotum less rounded (
<figureCitation id="35ED63D05D1CFFAFEB55FD97FE1DF303" box="[318,390,574,595]" captionStart="FIGURES 13 14" captionStartId="4.[151,264,1648,1671]" captionTargetBox="[382,1205,204,1620]" captionTargetId="figure@4.[370,1216,193,1627]" captionTargetPageId="4" captionText="FIGURES 13 14. Right half of pronotum: 13. Cyanopenthe taiwana sp. nov.; 14. C. leei sp. nov. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/237512/files/figure.png" pageId="10" pageNumber="588">Fig. 14</figureCitation>
); aedeagus: lateral margins of apical piece of tegmen subparallel to slightly divergent distally, median lobe about 1.10X as long as tegmen; ovipositor: proctiger slightly more slender in ventral view................
<taxonomicName id="6AD604D65D1CFFAFEF60FDF5FAD2F321" box="[1291,1353,604,625]" class="Insecta" family="Tetratomidae" genus="Cyanopenthe" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="588" phylum="Arthropoda" rank="species" species="leei" status="sp. nov.">
<emphasis id="9FA2A3475D1CFFAFEF60FDF5FAD2F321" box="[1291,1353,604,625]" italics="true" pageId="10" pageNumber="588">C. leei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84911E3C5D1CFFAFEF24FDF2FA07F321" box="[1359,1436,603,625]" pageId="10" pageNumber="588" rank="species">
<emphasis id="9FA2A3475D1CFFAFEF24FDF2FA07F321" bold="true" box="[1359,1436,603,625]" pageId="10" pageNumber="588">sp. nov.</emphasis>
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