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<document id="4292A008DA45FFDB14A5BD377EFBAD7D" ID-CLB-Dataset="56235" ID-DOI="10.11646/zootaxa.5262.1.1" ID-GBIF-Dataset="e24a8032-296e-442d-80ae-e0bf8c7237f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7795161" ID-ZooBank="F05805FD-BC5B-4236-BF04-C6AA5C37F2D7" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1680528957709" checkinUser="plazi" docAuthor="Hou, Yanmeng, Feng, Zegang &amp; Zhang, Feng" docDate="2023" docId="FD7487B1FFF9FFA8FF205F883985F86E" docLanguage="en" docName="zootaxa.5262.1.1.pdf" docOrigin="Zootaxa 5262 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5262.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Tyrannochthonius pictus Hou &amp; Feng &amp; Zhang 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="115" masterDocId="014DFFC9FF96FFDAFFB75F103A10FFB3" masterDocTitle="Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="158" masterPageNumber="1" pageNumber="112" updateTime="1701997099389" updateUser="plazi">
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<mods:title id="67ECBA93EBEBC0BCADB917B3001065E9">Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae)</mods:title>
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<mods:namePart id="ED89A190CD851415A027B42D965D7B93">Hou, Yanmeng</mods:namePart>
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<taxonomicName id="B2DD4D24FFF9FFB5FF205F883BD6FF01" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[151,454,152,179]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="pictus" status="sp. nov.">
<emphasis id="47A9EAB5FFF9FFB5FF205F883BD6FF01" bold="true" box="[151,454,152,179]" italics="true" pageId="111" pageNumber="112">Tyrannochthonius pictus</emphasis>
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AEṞƟṁae
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<figureCitation id="EDE62A22FFF9FFB5FF205FAC3B05FF65" box="[151,277,188,214]" captionStart-0="FIGURE 91" captionStart-1="FIGURE 92" captionStart-2="FIGURE93" captionStart-3="FIGURE 94" captionStart-4="FIGURE 95" captionStart-5="FIGURE 96" captionStartId-0="111.[151,250,1971,1995]" captionStartId-1="112.[151,250,1643,1667]" captionStartId-2="113.[151,250,1113,1137]" captionStartId-3="115.[151,250,1944,1968]" captionStartId-4="116.[151,250,1943,1967]" captionStartId-5="117.[151,250,1943,1967]" captionTargetBox-0="[174,1414,557,1947]" captionTargetBox-1="[151,1437,181,1619]" captionTargetBox-2="[178,1410,243,1071]" captionTargetBox-3="[160,1428,181,1920]" captionTargetBox-4="[185,1385,181,1915]" captionTargetBox-5="[175,1412,198,1919]" captionTargetId-0="figure-118@111.[174,1414,557,1947]" captionTargetId-1="figure-20@112.[151,1437,181,1619]" captionTargetId-2="figure-41@113.[166,1421,236,1090]" captionTargetId-3="figure-18@115.[160,1428,181,1920]" captionTargetId-4="figure-20@116.[179,1408,181,1919]" captionTargetId-5="figure-18@117.[175,1412,181,1919]" captionTargetPageId-0="111" captionTargetPageId-1="112" captionTargetPageId-2="113" captionTargetPageId-3="115" captionTargetPageId-4="116" captionTargetPageId-5="117" captionText-0="FIGURE 91. Gaoluo Cave, one of the type localities of Tyrannochthonius pictus sp. nov., A. Entrance; BC. Inside the cave entrance; DE.Areaswhere T.pictussp.nov. specimenswere collected;F.Live female of T.pictussp.nov. inits natural environment." captionText-1="FIGURE 92. Cangjun Cave, one of the type localities of Tyrannochthonius pictus sp. nov., A. Entrance; B. Inside the cave entrance; C. Exit (red arrow); DE.Areas where T. pictus sp. nov. specimens were collected; F. Live female of T. pictus sp. nov. in its natural environment." captionText-2="FIGURE93.Tyrannochthoniuspictussp.nov.,A.Holotype male,habitus (dorsalview);B.Paratypefemale,habitus (dorsal view)." captionText-3="FIGURE 94. Tyrannochthonius pictus sp. nov., holotype male (AF), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." captionText-4="FIGURE 95. Tyrannochthonius pictus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." captionText-5="FIGURE 96. Tyrannochthonius pictus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7795363" figureDoi-1="http://doi.org/10.5281/zenodo.7795365" figureDoi-2="http://doi.org/10.5281/zenodo.7795367" figureDoi-3="http://doi.org/10.5281/zenodo.7795369" figureDoi-4="http://doi.org/10.5281/zenodo.7795371" figureDoi-5="http://doi.org/10.5281/zenodo.7795373" httpUri-0="https://zenodo.org/record/7795363/files/figure.png" httpUri-1="https://zenodo.org/record/7795365/files/figure.png" httpUri-2="https://zenodo.org/record/7795367/files/figure.png" httpUri-3="https://zenodo.org/record/7795369/files/figure.png" httpUri-4="https://zenodo.org/record/7795371/files/figure.png" httpUri-5="https://zenodo.org/record/7795373/files/figure.png" pageId="111" pageNumber="112">Figs 9196</figureCitation>
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<emphasis id="47A9EAB5FFF9FFB5FF205FEF3B54FEAA" bold="true" box="[151,324,255,281]" pageId="111" pageNumber="112">Type material.</emphasis>
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♂ (Ps.-
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&amp;
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leg.
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(Ps.-
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HBUARA#2022-515-13), all with the same data as the holotype;
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16 ♂ (Ps.-
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HBUARA#2022-516-16),
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(Ps.-
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HBUARA#2022-516-29),
<location id="7002607CFFF9FFB5FE5B5EC7389DFE42" LSID="urn:lsid:plazi:treatment:FD7487B1FFF9FFA8FF205F883985F86E:7002607CFFF9FFB5FE5B5EC7389DFE42" box="[492,653,471,497]" latitude="25.74222" longLatPrecision="1" longitude="107.56771" municipality="Cangjun Cave" name="Jiabao Village" pageId="111" pageNumber="112">Jiabao Village</location>
,
<collectingMunicipality id="9506ACDDFFF9FFB5FD205EC73926FE42" box="[663,822,471,497]" pageId="111" pageNumber="112">Cangjun Cave</collectingMunicipality>
[
<geoCoordinate id="10E95060FFF9FFB5FCF55EC839E0FE40" box="[834,1008,472,499]" degrees="25" direction="north" minutes="44" orientation="latitude" pageId="111" pageNumber="112" precision="1" seconds="31.99" value="25.74222">25°4431.99″N</geoCoordinate>
,
<geoCoordinate id="10E95060FFF9FFB5FC4D5EC83EB4FE40" box="[1018,1188,472,499]" degrees="107" direction="east" minutes="34" orientation="longitude" pageId="111" pageNumber="112" precision="1" seconds="3.76" value="107.56771">107°343.76″E</geoCoordinate>
],
<elevation id="FEF0D194FFF9FFB5FB025EC73F27FE42" box="[1205,1335,471,497]" metricMagnitude="2" metricUnit="m" metricValue="9.48" pageId="111" pageNumber="112" unit="m" value="948.0">
<quantity id="B2259B42FFF9FFB5FB025EC73EEBFE42" box="[1205,1275,471,497]" metricMagnitude="2" metricUnit="m" metricValue="9.48" pageId="111" pageNumber="112" unit="m" value="948.0">948 m</quantity>
a.s.l.
</elevation>
, with the same collection date and collectors as the
<typeStatus id="AA668805FFF9FFB5FDD95EEB38C4FDA6" box="[622,724,507,533]" pageId="111" pageNumber="112" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="21A2662FFFF9FFB5FF2058A33F8DF85C" ID-DOI="http://doi.org/10.5281/zenodo.7795363" ID-Zenodo-Dep="7795363" httpUri="https://zenodo.org/record/7795363/files/figure.png" pageId="111" pageNumber="112" startId="111.[151,250,1971,1995]" targetBox="[174,1414,557,1947]" targetPageId="111">
<paragraph id="756236A7FFF9FFB5FF2058A33F8DF85C" blockId="111.[151,1437,1971,2031]" pageId="111" pageNumber="112">
<emphasis id="47A9EAB5FFF9FFB5FF2058A33B0FF878" bold="true" box="[151,287,1971,1995]" pageId="111" pageNumber="112">FIGURE 91.</emphasis>
Gaoluo Cave, one of the type localities of
<taxonomicName id="B2DD4D24FFF9FFB5FD7E58A339D2F878" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[713,962,1971,1995]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="pictus" status="sp. nov.">
<emphasis id="47A9EAB5FFF9FFB5FD7E58A339D2F878" box="[713,962,1971,1995]" italics="true" pageId="111" pageNumber="112">Tyrannochthonius pictus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFF9FFB5FC7058A33E07F878" bold="true" box="[967,1047,1971,1995]" pageId="111" pageNumber="112">
<taxonomicNameLabel id="5C9A57CEFFF9FFB5FC7058A33E07F878" box="[967,1047,1971,1995]" pageId="111" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Entrance; BC. Inside the cave entrance; DE.Areas where
<taxonomicName id="B2DD4D24FFF9FFB5FE2058C83BFAF85C" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[407,490,2008,2031]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="pictus" status="sp. nov.">
<emphasis id="47A9EAB5FFF9FFB5FE2058C83BFAF85C" box="[407,490,2008,2031]" italics="true" pageId="111" pageNumber="112">T.pictus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFF9FFB5FE5A58C7382BF85C" bold="true" box="[493,571,2007,2031]" pageId="111" pageNumber="112">
<taxonomicNameLabel id="5C9A57CEFFF9FFB5FE5A58C7382BF85C" box="[493,571,2007,2031]" pageId="111" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens were collected; F.Live female of
<taxonomicName id="B2DD4D24FFF9FFB5FC4658C83E54F85C" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[1009,1092,2008,2031]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="pictus" status="sp. nov.">
<emphasis id="47A9EAB5FFF9FFB5FC4658C83E54F85C" box="[1009,1092,2008,2031]" italics="true" pageId="111" pageNumber="112">T. pictus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFF9FFB5FBF058C73E85F85C" bold="true" box="[1095,1173,2007,2031]" pageId="111" pageNumber="112">
<taxonomicNameLabel id="5C9A57CEFFF9FFB5FBF058C73E85F85C" box="[1095,1173,2007,2031]" pageId="111" pageNumber="112" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in its natural environment.
</paragraph>
</caption>
<caption id="21A2662FFFE6FFAAFF20597B3BB4F978" ID-DOI="http://doi.org/10.5281/zenodo.7795365" ID-Zenodo-Dep="7795365" httpUri="https://zenodo.org/record/7795365/files/figure.png" pageId="112" pageNumber="113" startId="112.[151,250,1643,1667]" targetBox="[151,1437,181,1619]" targetPageId="112">
<paragraph id="756236A7FFE6FFAAFF20597B3BB4F978" blockId="112.[151,1436,1643,1739]" pageId="112" pageNumber="113">
<emphasis id="47A9EAB5FFE6FFAAFF20597B3B32F930" bold="true" box="[151,290,1643,1667]" pageId="112" pageNumber="113">FIGURE 92.</emphasis>
Cangjun Cave, one of the type localities of
<taxonomicName id="B2DD4D24FFE6FFAAFD45597B39FEF930" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[754,1006,1643,1667]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="pictus" status="sp. nov.">
<emphasis id="47A9EAB5FFE6FFAAFD45597B39FEF930" box="[754,1006,1643,1667]" italics="true" pageId="112" pageNumber="113">Tyrannochthonius pictus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFE6FFAAFC41597B3E58F930" bold="true" box="[1014,1096,1643,1667]" pageId="112" pageNumber="113">
<taxonomicNameLabel id="5C9A57CEFFE6FFAAFC41597B3E58F930" box="[1014,1096,1643,1667]" pageId="112" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Entrance; B. Inside the cave entrance; C. Exit (red arrow); DE. Areas where
<taxonomicName id="B2DD4D24FFE6FFAAFD35598038C7F914" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[642,727,1680,1703]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="pictus" status="sp. nov.">
<emphasis id="47A9EAB5FFE6FFAAFD35598038C7F914" box="[642,727,1680,1703]" italics="true" pageId="112" pageNumber="113">T. pictus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFE6FFAAFD6B599F393CF914" bold="true" box="[732,812,1679,1703]" pageId="112" pageNumber="113">
<taxonomicNameLabel id="5C9A57CEFFE6FFAAFD6B599F393CF914" box="[732,812,1679,1703]" pageId="112" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens were collected; F. Live female of
<taxonomicName id="B2DD4D24FFE6FFAAFB4559803F57F914" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[1266,1351,1680,1703]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="pictus" status="sp. nov.">
<emphasis id="47A9EAB5FFE6FFAAFB4559803F57F914" box="[1266,1351,1680,1703]" italics="true" pageId="112" pageNumber="113">T. pictus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFE6FFAAFAFB599F3F8CF914" bold="true" box="[1356,1436,1679,1703]" pageId="112" pageNumber="113">
<taxonomicNameLabel id="5C9A57CEFFE6FFAAFAFB599F3F8CF914" box="[1356,1436,1679,1703]" pageId="112" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in its natural environment.
</paragraph>
</caption>
<subSubSection id="3DC7652CFFE6FFAAFF7059E93990F884" pageId="112" pageNumber="113" type="etymology">
<paragraph id="756236A7FFE6FFAAFF7059E93990F884" blockId="112.[151,1436,1785,2028]" pageId="112" pageNumber="113">
<emphasis id="47A9EAB5FFE6FFAAFF7059E93B5EF8A0" bold="true" box="[199,334,1785,1811]" pageId="112" pageNumber="113">Etymology.</emphasis>
The specific name is derived from the Latin word “
<taxonomicName id="B2DD4D24FFE6FFAAFC2C59EA39CEF8A0" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[923,990,1786,1811]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="pictus">
<emphasis id="47A9EAB5FFE6FFAAFC2C59EA39CEF8A0" box="[923,990,1786,1811]" italics="true" pageId="112" pageNumber="113">pictus</emphasis>
</taxonomicName>
” (= figured), which refers to the dorsal surface of the cheliceral palm possessing rounded whitish patches.
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFE6FFAAFF7058513EF0F858" pageId="112" pageNumber="113" type="diagnosis">
<paragraph id="756236A7FFE6FFAAFF7058513EF0F858" blockId="112.[151,1436,1785,2028]" pageId="112" pageNumber="113">
<emphasis id="47A9EAB5FFE6FFAAFF7058513B94F8EC" bold="true" box="[199,388,1857,1887]" pageId="112" pageNumber="113">
Diagnosis (J
<specimenCount id="63DBFD2EFFE6FFAAFEEA58533B63F8EC" box="[349,371,1859,1887]" pageId="112" pageNumber="113" type="female"></specimenCount>
).
</emphasis>
Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, without epistome; posterior margin of carapace with 2 setae; tergite I with 2 setae, tergites IIIVIII each with 4 setae. Pedipalps slender, femur 7.297.71 (♂), 7.077.29 (
<specimenCount id="63DBFD2EFFE6FFAAFF1558BD3AA4F874" box="[162,180,1965,1991]" pageId="112" pageNumber="113" type="female"></specimenCount>
) times longer than broad; chela 8.068.50 (♂), 7.248.00 (
<specimenCount id="63DBFD2EFFE6FFAAFCF958BD3970F874" box="[846,864,1965,1991]" pageId="112" pageNumber="113" type="female"></specimenCount>
) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth smaller and denser than fixed chelal finger teeth.
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFE7FFA8FF705F863914F9E2" lastPageId="114" lastPageNumber="115" pageId="113" pageNumber="114" type="description">
<paragraph id="756236A7FFE7FFABFF705F8639E7FF03" blockId="113.[199,1263,150,213]" box="[199,1015,150,177]" pageId="113" pageNumber="114">
<emphasis id="47A9EAB5FFE7FFABFF705F863B49FF03" bold="true" box="[199,345,150,176]" pageId="113" pageNumber="114">Description.</emphasis>
Males (
<typeStatus id="AA668805FFE7FFABFE045F863807FF03" box="[435,535,150,176]" pageId="113" pageNumber="114" type="holotype">holotype</typeStatus>
and
<typeStatus id="AA668805FFE7FFABFDFA5F8638AAFF03" box="[589,698,150,176]" pageId="113" pageNumber="114" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="EDE62A22FFE7FFABFD675F86392CFF03" box="[720,828,150,176]" captionStart="FIGURE93" captionStartId="113.[151,250,1113,1137]" captionTargetBox="[178,1410,243,1071]" captionTargetId="figure-41@113.[166,1421,236,1090]" captionTargetPageId="113" captionText="FIGURE93.Tyrannochthoniuspictussp.nov.,A.Holotype male,habitus (dorsalview);B.Paratypefemale,habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795367" httpUri="https://zenodo.org/record/7795367/files/figure.png" pageId="113" pageNumber="114">Figs 93A</figureCitation>
,
<figureCitation id="EDE62A22FFE7FFABFCF05F863986FF03" box="[839,918,150,176]" captionStart="FIGURE 94" captionStartId="115.[151,250,1944,1968]" captionTargetBox="[160,1428,181,1920]" captionTargetId="figure-18@115.[160,1428,181,1920]" captionTargetPageId="115" captionText="FIGURE 94. Tyrannochthonius pictus sp. nov., holotype male (AF), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795369" httpUri="https://zenodo.org/record/7795369/files/figure.png" pageId="113" pageNumber="114">94AF</figureCitation>
,
<figureCitation id="EDE62A22FFE7FFABFC165F8639AFFF02" box="[929,959,150,177]" captionStart="FIGURE 95" captionStartId="116.[151,250,1943,1967]" captionTargetBox="[185,1385,181,1915]" captionTargetId="figure-20@116.[179,1408,181,1919]" captionTargetPageId="116" captionText="FIGURE 95. Tyrannochthonius pictus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795371" httpUri="https://zenodo.org/record/7795371/files/figure.png" pageId="113" pageNumber="114">95</figureCitation>
,
<figureCitation id="EDE62A22FFE7FFABFC7C5F8639FBFF03" box="[971,1003,150,176]" captionStart="FIGURE 96" captionStartId="117.[151,250,1943,1967]" captionTargetBox="[175,1412,198,1919]" captionTargetId="figure-18@117.[175,1412,181,1919]" captionTargetPageId="117" captionText="FIGURE 96. Tyrannochthonius pictus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795373" httpUri="https://zenodo.org/record/7795373/files/figure.png" pageId="113" pageNumber="114">96</figureCitation>
).
</paragraph>
<paragraph id="756236A7FFE7FFABFF705FAA3EFFFF67" blockId="113.[199,1263,150,213]" box="[199,1263,186,213]" pageId="113" pageNumber="114">Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</paragraph>
<caption id="21A2662FFFE7FFABFF205B493F8DFBC1" ID-DOI="http://doi.org/10.5281/zenodo.7795367" ID-Zenodo-Dep="7795367" box="[151,1437,1113,1138]" httpUri="https://zenodo.org/record/7795367/files/figure.png" pageId="113" pageNumber="114" startId="113.[151,250,1113,1137]" targetBox="[178,1410,243,1071]" targetPageId="113">
<paragraph id="756236A7FFE7FFABFF205B493F8DFBC1" blockId="113.[151,1437,1113,1138]" box="[151,1437,1113,1138]" pageId="113" pageNumber="114">
<emphasis id="47A9EAB5FFE7FFABFF205B493B0DFBC2" bold="true" box="[151,285,1113,1137]" pageId="113" pageNumber="114">FIGURE 93.</emphasis>
<taxonomicName id="B2DD4D24FFE7FFABFE975B4A3807FBC2" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[288,535,1114,1138]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="pictus" status="sp. nov.">
<emphasis id="47A9EAB5FFE7FFABFE975B4A3807FBC2" box="[288,535,1114,1138]" italics="true" pageId="113" pageNumber="114">Tyrannochthonius pictus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFE7FFABFDAE5B4A3877FBC1" bold="true" box="[537,615,1114,1138]" pageId="113" pageNumber="114">
<taxonomicNameLabel id="5C9A57CEFFE7FFABFDAE5B4A3877FBC1" box="[537,615,1114,1138]" pageId="113" pageNumber="114" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,A.Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view).
</paragraph>
</caption>
<paragraph id="756236A7FFE7FFABFF705BB038A8FA69" blockId="113.[151,1437,1184,2038]" pageId="113" pageNumber="114">
Cephalothorax (
<figureCitation id="EDE62A22FFE7FFABFEC85BB03BFCFB09" box="[383,492,1184,1210]" captionStart="FIGURE 94" captionStartId="115.[151,250,1944,1968]" captionTargetBox="[160,1428,181,1920]" captionTargetId="figure-18@115.[160,1428,181,1920]" captionTargetPageId="115" captionText="FIGURE 94. Tyrannochthonius pictus sp. nov., holotype male (AF), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795369" httpUri="https://zenodo.org/record/7795369/files/figure.png" pageId="113" pageNumber="114">Figs 94C</figureCitation>
,
<figureCitation id="EDE62A22FFE7FFABFE4B5BB03821FB08" box="[508,561,1184,1211]" captionStart="FIGURE 95" captionStartId="116.[151,250,1943,1967]" captionTargetBox="[185,1385,181,1915]" captionTargetId="figure-20@116.[179,1408,181,1919]" captionTargetPageId="116" captionText="FIGURE 95. Tyrannochthonius pictus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795371" httpUri="https://zenodo.org/record/7795371/files/figure.png" pageId="113" pageNumber="114">95A</figureCitation>
): carapace 1.021.08 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; without epistome; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta more than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 89 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
<figureCitation id="EDE62A22FFE7FFABFA9E5A8C3F9FFA04" box="[1321,1423,1436,1463]" captionStart="FIGURE 95" captionStartId="116.[151,250,1943,1967]" captionTargetBox="[185,1385,181,1915]" captionTargetId="figure-20@116.[179,1408,181,1919]" captionTargetPageId="116" captionText="FIGURE 95. Tyrannochthonius pictus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795371" httpUri="https://zenodo.org/record/7795371/files/figure.png" pageId="113" pageNumber="114">Fig. 95C</figureCitation>
); intercoxal tubercle absent; without sub-oral seta.
</paragraph>
<paragraph id="756236A7FFE7FFABFF705AF439D5F948" blockId="113.[151,1437,1184,2038]" pageId="113" pageNumber="114">
Chelicera (
<figureCitation id="EDE62A22FFE7FFABFEF55AF43BBEFA4D" box="[322,430,1508,1534]" captionStart="FIGURE 94" captionStartId="115.[151,250,1944,1968]" captionTargetBox="[160,1428,181,1920]" captionTargetId="figure-18@115.[160,1428,181,1920]" captionTargetPageId="115" captionText="FIGURE 94. Tyrannochthonius pictus sp. nov., holotype male (AF), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795369" httpUri="https://zenodo.org/record/7795369/files/figure.png" pageId="113" pageNumber="114">Figs 94D</figureCitation>
,
<figureCitation id="EDE62A22FFE7FFABFE0D5AF43BFEFA4C" box="[442,494,1508,1535]" captionStart="FIGURE 95" captionStartId="116.[151,250,1943,1967]" captionTargetBox="[185,1385,181,1915]" captionTargetId="figure-20@116.[179,1408,181,1919]" captionTargetPageId="116" captionText="FIGURE 95. Tyrannochthonius pictus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795371" httpUri="https://zenodo.org/record/7795371/files/figure.png" pageId="113" pageNumber="114">95B</figureCitation>
): large, about as long as carapace, 2.362.52 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 1316 teeth, distal one largest; movable finger with 1517 retrorse contiguous small teeth; galea represented by a very slight bump on movable finger (
<figureCitation id="EDE62A22FFE7FFABFF5B59883B45F900" box="[236,341,1688,1715]" captionStart="FIGURE 95" captionStartId="116.[151,250,1943,1967]" captionTargetBox="[185,1385,181,1915]" captionTargetId="figure-20@116.[179,1408,181,1919]" captionTargetPageId="116" captionText="FIGURE 95. Tyrannochthonius pictus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795371" httpUri="https://zenodo.org/record/7795371/files/figure.png" pageId="113" pageNumber="114">Fig. 95B</figureCitation>
). Serrula exterior with 1720 and serrula interior with 1315 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
<figureCitation id="EDE62A22FFE7FFABFCE459F039A8F948" box="[851,952,1760,1787]" captionStart="FIGURE 95" captionStartId="116.[151,250,1943,1967]" captionTargetBox="[185,1385,181,1915]" captionTargetId="figure-20@116.[179,1408,181,1919]" captionTargetPageId="116" captionText="FIGURE 95. Tyrannochthonius pictus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795371" httpUri="https://zenodo.org/record/7795371/files/figure.png" pageId="113" pageNumber="114">Fig. 95E</figureCitation>
).
</paragraph>
<paragraph id="756236A7FFE7FFA8FF7058143F41FE1E" blockId="113.[151,1437,1184,2038]" lastBlockId="114.[151,1437,151,2013]" lastPageId="114" lastPageNumber="115" pageId="113" pageNumber="114">
Pedipalp (
<figureCitation id="EDE62A22FFE7FFABFE8B58143BE1F8AD" box="[316,497,1796,1822]" captionStart="FIGURE 94" captionStartId="115.[151,250,1944,1968]" captionTargetBox="[160,1428,181,1920]" captionTargetId="figure-18@115.[160,1428,181,1920]" captionTargetPageId="115" captionText="FIGURE 94. Tyrannochthonius pictus sp. nov., holotype male (AF), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795369" httpUri="https://zenodo.org/record/7795369/files/figure.png" pageId="113" pageNumber="114">Figs 94A, B, E</figureCitation>
,
<figureCitation id="EDE62A22FFE7FFABFDB658143824F8AC" box="[513,564,1796,1823]" captionStart="FIGURE 95" captionStartId="116.[151,250,1943,1967]" captionTargetBox="[185,1385,181,1915]" captionTargetId="figure-20@116.[179,1408,181,1919]" captionTargetPageId="116" captionText="FIGURE 95. Tyrannochthonius pictus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795371" httpUri="https://zenodo.org/record/7795371/files/figure.png" pageId="113" pageNumber="114">95D</figureCitation>
,
<figureCitation id="EDE62A22FFE7FFABFDF45814388BF8AD" box="[579,667,1796,1822]" captionStart="FIGURE 96" captionStartId="117.[151,250,1943,1967]" captionTargetBox="[175,1412,198,1919]" captionTargetId="figure-18@117.[175,1412,181,1919]" captionTargetPageId="117" captionText="FIGURE 96. Tyrannochthonius pictus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795373" httpUri="https://zenodo.org/record/7795373/files/figure.png" pageId="113" pageNumber="114">96A, B</figureCitation>
): long and slender, trochanter 1.241.31, femur 7.297.71, patella 2.272.40, chela 8.068.50, hand 2.833.00 times longer than broad; femur 2.832.97 times longer than patella; movable chelal finger 1.761.79 times longer than hand and 0.620.63 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (
<figureCitation id="EDE62A22FFE7FFABFD5858603948F839" box="[751,856,1904,1930]" captionStart="FIGURE 94" captionStartId="115.[151,250,1944,1968]" captionTargetBox="[160,1428,181,1920]" captionTargetId="figure-18@115.[160,1428,181,1920]" captionTargetPageId="115" captionText="FIGURE 94. Tyrannochthonius pictus sp. nov., holotype male (AF), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795369" httpUri="https://zenodo.org/record/7795369/files/figure.png" pageId="113" pageNumber="114">Figs 94E</figureCitation>
,
<figureCitation id="EDE62A22FFE7FFABFCD358603988F838" box="[868,920,1904,1931]" captionStart="FIGURE 95" captionStartId="116.[151,250,1943,1967]" captionTargetBox="[185,1385,181,1915]" captionTargetId="figure-20@116.[179,1408,181,1919]" captionTargetPageId="116" captionText="FIGURE 95. Tyrannochthonius pictus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795371" httpUri="https://zenodo.org/record/7795371/files/figure.png" pageId="113" pageNumber="114">95D</figureCitation>
). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria,
<emphasis id="47A9EAB5FFE7FFABFAE158A83F7CF861" box="[1366,1388,1976,2002]" italics="true" pageId="113" pageNumber="114">ib</emphasis>
and
<emphasis id="47A9EAB5FFE7FFABFF2058CC3AA8F845" box="[151,184,2012,2038]" italics="true" pageId="113" pageNumber="114">isb</emphasis>
situated close together, submedially on dorsum of chelal hand;
<emphasis id="47A9EAB5FFE7FFABFC3358CC398EF845" box="[900,926,2012,2038]" italics="true" pageId="113" pageNumber="114">eb</emphasis>
,
<emphasis id="47A9EAB5FFE7FFABFC1A58CC39C2F845" box="[941,978,2012,2038]" italics="true" pageId="113" pageNumber="114">esb</emphasis>
and
<emphasis id="47A9EAB5FFE7FFABFBBE58CD3E33F845" box="[1033,1059,2013,2038]" italics="true" pageId="113" pageNumber="114">ist</emphasis>
forming a nearly straight oblique row at base of fixed chelal finger;
<emphasis id="47A9EAB5FFE4FFA8FDAA5F87383DFF03" box="[541,557,151,176]" italics="true" pageId="114" pageNumber="115">it</emphasis>
slightly distal to
<emphasis id="47A9EAB5FFE4FFA8FD455F873901FF03" box="[754,785,151,176]" italics="true" pageId="114" pageNumber="115">est</emphasis>
, situated subdistally;
<emphasis id="47A9EAB5FFE4FFA8FBB05F873E0BFF03" box="[1031,1051,151,176]" italics="true" pageId="114" pageNumber="115">et</emphasis>
slightly near to tip of fixed chelal finger, very close to chelal teeth;
<emphasis id="47A9EAB5FFE4FFA8FDB35FAB380EFF66" box="[516,542,187,213]" italics="true" pageId="114" pageNumber="115">dx</emphasis>
situated distal to
<emphasis id="47A9EAB5FFE4FFA8FD695FAB38E2FF67" box="[734,754,187,212]" italics="true" pageId="114" pageNumber="115">et</emphasis>
;
<emphasis id="47A9EAB5FFE4FFA8FCB75FAB3909FF66" box="[768,793,187,213]" italics="true" pageId="114" pageNumber="115">sb</emphasis>
closer to
<emphasis id="47A9EAB5FFE4FFA8FC345FAB3986FF67" box="[899,918,187,212]" italics="true" pageId="114" pageNumber="115">st</emphasis>
than to
<emphasis id="47A9EAB5FFE4FFA8FC5B5FAB39EAFF66" box="[1004,1018,187,213]" italics="true" pageId="114" pageNumber="115">b</emphasis>
;
<emphasis id="47A9EAB5FFE4FFA8FBBF5FAB3E06FF66" box="[1032,1046,187,213]" italics="true" pageId="114" pageNumber="115">b</emphasis>
and
<emphasis id="47A9EAB5FFE4FFA8FBFE5FAB3E41FF67" box="[1097,1105,187,212]" italics="true" pageId="114" pageNumber="115">t</emphasis>
situated subdistally,
<emphasis id="47A9EAB5FFE4FFA8FA8F5FAB3F50FF67" box="[1336,1344,187,212]" italics="true" pageId="114" pageNumber="115">t</emphasis>
situated distal to
<emphasis id="47A9EAB5FFE4FFA8FF425FCF3B13FF4A" box="[245,259,223,249]" italics="true" pageId="114" pageNumber="115">b</emphasis>
;
<emphasis id="47A9EAB5FFE4FFA8FEA65FCF3B20FF4B" box="[273,304,223,248]" italics="true" pageId="114" pageNumber="115">est</emphasis>
situated distal to
<emphasis id="47A9EAB5FFE4FFA8FE465FCF3BE9FF4B" box="[497,505,223,248]" italics="true" pageId="114" pageNumber="115">t</emphasis>
(
<figureCitation id="EDE62A22FFE4FFA8FDB05FCF387EFF4A" box="[519,622,223,249]" captionStart="FIGURE 96" captionStartId="117.[151,250,1943,1967]" captionTargetBox="[175,1412,198,1919]" captionTargetId="figure-18@117.[175,1412,181,1919]" captionTargetPageId="117" captionText="FIGURE 96. Tyrannochthonius pictus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795373" httpUri="https://zenodo.org/record/7795373/files/figure.png" pageId="114" pageNumber="115">Fig. 96A</figureCitation>
).A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to
<emphasis id="47A9EAB5FFE4FFA8FF4C5E133B05FEAF" box="[251,277,259,284]" italics="true" pageId="114" pageNumber="115">ist</emphasis>
). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 3032 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 4043 macrodenticles (smaller and denser than teeth on fixed chelal finger), slightly retrorse and pointed (
<figureCitation id="EDE62A22FFE4FFA8FB355E7F3EFBFE3A" box="[1154,1259,367,393]" captionStart="FIGURE 96" captionStartId="117.[151,250,1943,1967]" captionTargetBox="[175,1412,198,1919]" captionTargetId="figure-18@117.[175,1412,181,1919]" captionTargetPageId="117" captionText="FIGURE 96. Tyrannochthonius pictus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795373" httpUri="https://zenodo.org/record/7795373/files/figure.png" pageId="114" pageNumber="115">Fig. 96A</figureCitation>
). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
<figureCitation id="EDE62A22FFE4FFA8FB2B5E833F16FE1E" box="[1180,1286,403,429]" captionStart="FIGURE 94" captionStartId="115.[151,250,1944,1968]" captionTargetBox="[160,1428,181,1920]" captionTargetId="figure-18@115.[160,1428,181,1920]" captionTargetPageId="115" captionText="FIGURE 94. Tyrannochthonius pictus sp. nov., holotype male (AF), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795369" httpUri="https://zenodo.org/record/7795369/files/figure.png" pageId="114" pageNumber="115">Figs 94B</figureCitation>
,
<figureCitation id="EDE62A22FFE4FFA8FAA65E833F54FE1E" box="[1297,1348,403,429]" captionStart="FIGURE 96" captionStartId="117.[151,250,1943,1967]" captionTargetBox="[175,1412,198,1919]" captionTargetId="figure-18@117.[175,1412,181,1919]" captionTargetPageId="117" captionText="FIGURE 96. Tyrannochthonius pictus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795373" httpUri="https://zenodo.org/record/7795373/files/figure.png" pageId="114" pageNumber="115">96B</figureCitation>
).
</paragraph>
<paragraph id="756236A7FFE4FFA8FF705EA7396AFD8E" blockId="114.[151,1437,151,2013]" pageId="114" pageNumber="115">
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy IXII: 2: 24: 4: 4: 4: 4: 4: 4: 45: 4: T2T: 0. Sternal chaetotaxy IIIXII: 1215: 1213: 89: 89: 9: 9: 910: 9: 0: 2. Anterior genital operculum with 911 setae, genital opening slit-like, with 1517 marginal setae on each side,
<quantity id="B2259B42FFE4FFA8FDE95D3338D1FD8E" box="[606,705,547,573]" metricMagnitude="0" metricUnit="m" metricValue="1.1049" metricValueMax="1.1684" metricValueMin="1.0413999999999999" pageId="114" pageNumber="115" unit="in" value="43.5" valueMax="46.0" valueMin="41.0">4146 in</quantity>
total (
<figureCitation id="EDE62A22FFE4FFA8FCBE5D33397DFD8E" box="[777,877,547,573]" captionStart="FIGURE 94" captionStartId="115.[151,250,1944,1968]" captionTargetBox="[160,1428,181,1920]" captionTargetId="figure-18@115.[160,1428,181,1920]" captionTargetPageId="115" captionText="FIGURE 94. Tyrannochthonius pictus sp. nov., holotype male (AF), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795369" httpUri="https://zenodo.org/record/7795369/files/figure.png" pageId="114" pageNumber="115">Fig. 94F</figureCitation>
).
</paragraph>
<paragraph id="756236A7FFE4FFA8FF705D573E17FCA6" blockId="114.[151,1437,151,2013]" pageId="114" pageNumber="115">
Legs (
<figureCitation id="EDE62A22FFE4FFA8FEBB5D573B84FDD2" box="[268,404,583,609]" captionStart="FIGURE 96" captionStartId="117.[151,250,1943,1967]" captionTargetBox="[175,1412,198,1919]" captionTargetId="figure-18@117.[175,1412,181,1919]" captionTargetPageId="117" captionText="FIGURE 96. Tyrannochthonius pictus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795373" httpUri="https://zenodo.org/record/7795373/files/figure.png" pageId="114" pageNumber="115">Fig. 96C, D</figureCitation>
): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.812.00 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.312.37 times longer than tibia. Femoropatella of leg IV 3.874.00 times longer than deep; tibia 6.507.33 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.503.63 times longer than deep (TS= 0.250.34), tarsus 13.1715.00 times longer than deep and 2.592.72 times longer than metatarsus (TS= 0.240.31). Arolium slightly shorter than the claws, not divided; claws simple.
</paragraph>
<paragraph id="756236A7FFE4FFA8FF705C0F3BC8FC5E" blockId="114.[151,1437,151,2013]" pageId="114" pageNumber="115">
Adult females (
<typeStatus id="AA668805FFE4FFA8FEC15C0F3BF5FC8A" box="[374,485,799,825]" pageId="114" pageNumber="115" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="EDE62A22FFE4FFA8FE4B5C0F3872FC8A" box="[508,610,799,825]" captionStart="FIGURE 91" captionStartId="111.[151,250,1971,1995]" captionTargetBox="[174,1414,557,1947]" captionTargetId="figure-118@111.[174,1414,557,1947]" captionTargetPageId="111" captionText="FIGURE 91. Gaoluo Cave, one of the type localities of Tyrannochthonius pictus sp. nov., A. Entrance; BC. Inside the cave entrance; DE.Areaswhere T.pictussp.nov. specimenswere collected;F.Live female of T.pictussp.nov. inits natural environment." figureDoi="http://doi.org/10.5281/zenodo.7795363" httpUri="https://zenodo.org/record/7795363/files/figure.png" pageId="114" pageNumber="115">Figs 91F</figureCitation>
,
<figureCitation id="EDE62A22FFE4FFA8FDD85C0F388BFC8A" box="[623,667,799,825]" captionStart="FIGURE 92" captionStartId="112.[151,250,1643,1667]" captionTargetBox="[151,1437,181,1619]" captionTargetId="figure-20@112.[151,1437,181,1619]" captionTargetPageId="112" captionText="FIGURE 92. Cangjun Cave, one of the type localities of Tyrannochthonius pictus sp. nov., A. Entrance; B. Inside the cave entrance; C. Exit (red arrow); DE.Areas where T. pictus sp. nov. specimens were collected; F. Live female of T. pictus sp. nov. in its natural environment." figureDoi="http://doi.org/10.5281/zenodo.7795365" httpUri="https://zenodo.org/record/7795365/files/figure.png" pageId="114" pageNumber="115">92F</figureCitation>
,
<figureCitation id="EDE62A22FFE4FFA8FD105C0F38C8FC8A" box="[679,728,799,825]" captionStart="FIGURE93" captionStartId="113.[151,250,1113,1137]" captionTargetBox="[178,1410,243,1071]" captionTargetId="figure-41@113.[166,1421,236,1090]" captionTargetPageId="113" captionText="FIGURE93.Tyrannochthoniuspictussp.nov.,A.Holotype male,habitus (dorsalview);B.Paratypefemale,habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795367" httpUri="https://zenodo.org/record/7795367/files/figure.png" pageId="114" pageNumber="115">93B</figureCitation>
,
<figureCitation id="EDE62A22FFE4FFA8FD525C0F390AFC8A" box="[741,794,799,825]" captionStart="FIGURE 94" captionStartId="115.[151,250,1944,1968]" captionTargetBox="[160,1428,181,1920]" captionTargetId="figure-18@115.[160,1428,181,1920]" captionTargetPageId="115" captionText="FIGURE 94. Tyrannochthonius pictus sp. nov., holotype male (AF), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795369" httpUri="https://zenodo.org/record/7795369/files/figure.png" pageId="114" pageNumber="115">94G</figureCitation>
). Mostly same as males, but a little larger; chaetotaxy of coxae: P 34, I 3, II 4, III 5, IV 5; tergal chaetotaxy IXII: 2: 34: 4: 4: 4: 4: 4: 4: 45: 4: T2T: 0; sternal chaetotaxy IVXII: 1112: 810: 9: 89: 89: 810: 910: 0: 2; anterior genital operculum with 10 setae, posterior margin with 1215 marginal setae,
<quantity id="B2259B42FFE4FFA8FE225C9A3BE9FC16" box="[405,505,906,933]" metricMagnitude="-1" metricUnit="m" metricValue="5.968999999999999" metricValueMax="6.35" metricValueMin="5.588" pageId="114" pageNumber="115" unit="in" value="23.5" valueMax="25.0" valueMin="22.0">2225 in</quantity>
total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.133.86 times longer than deep (TS= 0.300.36), tarsus 11.8315.40 times longer than deep and 2.722.88 times longer than metatarsus (TS= 0.250.30).
</paragraph>
<paragraph id="756236A7FFE4FFA8FF705CE7383CFA82" blockId="114.[151,1437,151,2013]" pageId="114" pageNumber="115">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.802.09. Pedipalps: trochanter 0.200.21/0.160.17 (1.241.31), femur 0.981.08/0.130.14 (7.297.71), patella 0.340.38/0.150.16 (2.272.40), chela 1.451.57/0.180.19 (8.068.50), hand 0.510.55/0.180.19 (2.833.00), movable chelal finger length 0.910.98. Chelicera 0.520.55/0.210.23 (2.362.52), movable finger length 0.280.30. Carapace 0.470.54/0.460.51 (1.021.08). Leg I: trochanter 0.160.17/0.140.15 (1.071.21), femur 0.580.65/0.08 (7.258.13), patella 0.320.33/0.07 (4.574.71), tibia 0.270.29/0.050.06 (3.505.40), tarsus 0.640.68/0.050.06 (11.1713.60). Leg IV: trochanter 0.240.26/0.140.15 (1.671.86), femoropatella 0.87 0.95/0.220.24 (3.874.00), tibia 0.610.66/0.090.10 (6.507.33), metatarsus 0.280.29/0.08 (3.503.63), tarsus 0.730.79/0.050.06 (13.1715.00).</paragraph>
<paragraph id="756236A7FFE4FFA8FF705A2B3914F9E2" blockId="114.[151,1437,151,2013]" pageId="114" pageNumber="115">Females: body length 2.182.25. Pedipalps: trochanter 0.200.23/0.160.17 (1.251.44), femur 1.02 1.08/0.140.15 (7.077.29), patella 0.350.39/0.160.17 (2.192.44), chela 1.461.56/0.190.21 (7.248.00), hand 0.540.56/0.190.21 (2.672.84), movable chelal finger length 0.880.99. Chelicera 0.550.57/0.24 (2.292.38), movable finger length 0.290.32. Carapace 0.530.55/0.510.54 (1.001.04). Leg I: trochanter 0.16/0.140.15 (1.071.14), femur 0.580.66/0.08 (7.258.25), patella 0.310.33/0.070.08 (4.004.71), tibia 0.270.29/0.050.06 (4.675.80), tarsus 0.640.69/0.050.06 (10.6713.60). Leg IV: trochanter 0.240.27/0.140.16 (1.561.93), femoropatella 0.860.92/0.220.23 (3.744.00), tibia 0.580.63/0.090.10 (5.907.00), metatarsus 0.250.29/0.070.08 (3.133.86), tarsus 0.710.79/0.050.06 (11.8315.40).</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFE4FFA8FF70594B3BB5F80A" pageId="114" pageNumber="115" type="discussion">
<paragraph id="756236A7FFE4FFA8FF70594B383AF952" blockId="114.[151,1437,151,2013]" pageId="114" pageNumber="115">
<emphasis id="47A9EAB5FFE4FFA8FF70594B3B2BF9C6" bold="true" box="[199,315,1627,1653]" pageId="114" pageNumber="115">Remarks.</emphasis>
<taxonomicName id="B2DD4D24FFE4FFA8FEF5594B3846F9C7" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[322,598,1627,1653]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="pictus" status="sp. nov.">
<emphasis id="47A9EAB5FFE4FFA8FEF5594B3846F9C7" box="[322,598,1627,1653]" italics="true" pageId="114" pageNumber="115">Tyrannochthonius pictus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFE4FFA8FDEB594B38A6F9C6" bold="true" box="[604,694,1627,1653]" pageId="114" pageNumber="115">
<taxonomicNameLabel id="5C9A57CEFFE4FFA8FDEB594B38A6F9C6" box="[604,694,1627,1653]" pageId="114" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="B2DD4D24FFE4FFA8FCFF594B39BEF9C6" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[840,942,1627,1653]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="planus" status="sp. nov.">
<emphasis id="47A9EAB5FFE4FFA8FCFF594B39BEF9C6" box="[840,942,1627,1653]" italics="true" pageId="114" pageNumber="115">T. planus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFE4FFA8FC02594B3E1EF9C6" bold="true" box="[949,1038,1627,1653]" pageId="114" pageNumber="115">
<taxonomicNameLabel id="5C9A57CEFFE4FFA8FC02594B3E1EF9C6" box="[949,1038,1627,1653]" pageId="114" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in lacking an epistome on the carapace as well as intercalary teeth on both chelal fingers, but differs by having a greater number of teeth on the chelal fingers (fixed chelal finger with 3032
<emphasis id="47A9EAB5FFE4FFA8FDE459B33861F90F" box="[595,625,1699,1724]" italics="true" pageId="114" pageNumber="115">vs.</emphasis>
2021 teeth; movable chelal finger with 4043
<emphasis id="47A9EAB5FFE4FFA8FB3859B33EBEF90F" box="[1167,1198,1699,1724]" italics="true" pageId="114" pageNumber="115">vs.</emphasis>
2526 teeth) and the number of blades of rallum (8
<emphasis id="47A9EAB5FFE4FFA8FE5A59D7381CF953" box="[493,524,1735,1760]" italics="true" pageId="114" pageNumber="115">vs.</emphasis>
7).
</paragraph>
<paragraph id="756236A7FFE4FFA8FF7059FB3BB5F80A" blockId="114.[151,1437,151,2013]" pageId="114" pageNumber="115">
<taxonomicName id="B2DD4D24FFE4FFA8FF7059FB3BCAF8B7" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[199,474,1771,1797]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="pictus" status="sp. nov.">
<emphasis id="47A9EAB5FFE4FFA8FF7059FB3BCAF8B7" box="[199,474,1771,1797]" italics="true" pageId="114" pageNumber="115">Tyrannochthonius pictus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFE4FFA8FE5759FB3828F8B6" bold="true" box="[480,568,1771,1797]" pageId="114" pageNumber="115">
<taxonomicNameLabel id="5C9A57CEFFE4FFA8FE5759FB3828F8B6" box="[480,568,1771,1797]" pageId="114" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="B2DD4D24FFE4FFA8FCD459FB39C3F8B6" authorityName="Mahnert" authorityYear="2009" box="[867,979,1771,1797]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="akaelus">
<emphasis id="47A9EAB5FFE4FFA8FCD459FB39C3F8B6" box="[867,979,1771,1797]" italics="true" pageId="114" pageNumber="115">T. akaelus</emphasis>
</taxonomicName>
,
<taxonomicName id="B2DD4D24FFE4FFA8FC5759FB3E82F8B6" authorityName="Mahnert" authorityYear="2009" box="[992,1170,1771,1797]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="antridraconis">
<emphasis id="47A9EAB5FFE4FFA8FC5759FB3E82F8B6" box="[992,1170,1771,1797]" italics="true" pageId="114" pageNumber="115">T. antridraconis</emphasis>
</taxonomicName>
,
<taxonomicName id="B2DD4D24FFE4FFA8FB1759FB3F1CF8B6" authorityName="Gao, Wynne &amp; Fe.Zhang" authorityYear="2018" box="[1184,1292,1771,1797]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="chixing">
<emphasis id="47A9EAB5FFE4FFA8FB1759FB3F1CF8B6" box="[1184,1292,1771,1797]" italics="true" pageId="114" pageNumber="115">T. chixing</emphasis>
</taxonomicName>
,
<taxonomicName id="B2DD4D24FFE4FFA8FAAD59FB3ACDF89B" authorityName="Mahnert" authorityYear="2009" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="ganshuanensis">
<emphasis id="47A9EAB5FFE4FFA8FAAD59FB3ACDF89B" italics="true" pageId="114" pageNumber="115">T. ganshuanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="B2DD4D24FFE4FFA8FEA2581F3B70F89A" authorityName="Gao, Zhang &amp; Chen" authorityYear="2020" box="[277,352,1807,1833]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="zhai">
<emphasis id="47A9EAB5FFE4FFA8FEA2581F3B70F89A" box="[277,352,1807,1833]" italics="true" pageId="114" pageNumber="115">T. zhai</emphasis>
</taxonomicName>
by lacking intercalary teeth on the fixed chelal finger, from
<taxonomicName id="B2DD4D24FFE4FFA8FBBF581F3E68F89A" authorityName="Gao, Zhang &amp; Chen" authorityYear="2020" box="[1032,1144,1807,1833]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="harveyi">
<emphasis id="47A9EAB5FFE4FFA8FBBF581F3E68F89A" box="[1032,1144,1807,1833]" italics="true" pageId="114" pageNumber="115">T. harveyi</emphasis>
</taxonomicName>
by the number of setae at the anterior margin of the carapace (6
<emphasis id="47A9EAB5FFE4FFA8FDF458233871F8FF" box="[579,609,1843,1868]" italics="true" pageId="114" pageNumber="115">vs.</emphasis>
4), a slender palp (chela 8.068.50 (♂), 7.248.00 (
<specimenCount id="63DBFD2EFFE4FFA8FB1E58233EABF8FE" box="[1193,1211,1843,1869]" pageId="114" pageNumber="115" type="female"></specimenCount>
) times
<emphasis id="47A9EAB5FFE4FFA8FAA758233F3FF8FF" box="[1296,1327,1843,1868]" italics="true" pageId="114" pageNumber="115">vs.</emphasis>
7.50 (♂), 7.22 (
<specimenCount id="63DBFD2EFFE4FFA8FF6C58473AFDF8C2" box="[219,237,1879,1905]" pageId="114" pageNumber="115" type="female"></specimenCount>
) times longer than board, length min.
<quantity id="B2259B42FFE4FFA8FD1558463917F8C2" box="[674,775,1878,1905]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="114" pageNumber="115" unit="mm" value="1.45">1.45 mm</quantity>
<emphasis id="47A9EAB5FFE4FFA8FCB95847393CF8C3" box="[782,812,1879,1904]" italics="true" pageId="114" pageNumber="115">vs.</emphasis>
max.
<quantity id="B2259B42FFE4FFA8FCC4584739CBF8C2" box="[883,987,1879,1905]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="114" pageNumber="115" unit="mm" value="1.3">1.30 mm</quantity>
; palpal femur 7.297.71 (♂), 7.077.29 (
<specimenCount id="63DBFD2EFFE4FFA8FF15586B3AA4F826" box="[162,180,1915,1941]" pageId="114" pageNumber="115" type="female"></specimenCount>
) times
<emphasis id="47A9EAB5FFE4FFA8FEBD586B3B38F827" box="[266,296,1915,1940]" italics="true" pageId="114" pageNumber="115">vs.</emphasis>
6.29 (♂), 5.80 (
<specimenCount id="63DBFD2EFFE4FFA8FE57586B3BE2F826" box="[480,498,1915,1941]" pageId="114" pageNumber="115" type="female"></specimenCount>
) times longer than board, length min.
<quantity id="B2259B42FFE4FFA8FC15586B3E16F826" box="[930,1030,1915,1941]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="114" pageNumber="115" unit="mm" value="0.98">0.98 mm</quantity>
<emphasis id="47A9EAB5FFE4FFA8FBBA586B3E3BF827" box="[1037,1067,1915,1940]" italics="true" pageId="114" pageNumber="115">vs.</emphasis>
max.
<quantity id="B2259B42FFE4FFA8FBC6586B3EC6F826" box="[1137,1238,1915,1941]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="114" pageNumber="115" unit="mm" value="0.88">0.88 mm</quantity>
) (
<bibRefCitation id="114C4B56FFE4FFA8FB5A586B3F87F826" author="Mahnert, V." box="[1261,1431,1915,1941]" pageId="114" pageNumber="115" pagination="185 - 201" refId="ref77141" refString="Mahnert, V. (2009) New species of pseudoscorpions (Arachnida, Pseudoscorpiones, Chthoniidae, Chernetidae) from caves in China. Revue Suisse de Zoologie, 116 (2), 185 - 201. https: // doi. org / 10.5962 / bhl. part. 79492" type="journal article" year="2009">Mahnert 2009</bibRefCitation>
;
<bibRefCitation id="114C4B56FFE4FFA8FF20588F3B5EF80A" author="Gao, Z. Z. &amp; Wynne, J. J. &amp; Zhang, F." box="[151,334,1951,1977]" pageId="114" pageNumber="115" pagination="345 - 354" refId="ref76312" refString="Gao, Z. Z., Wynne, J. J. &amp; Zhang, F. (2018) Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones: Neobisiidae, Chthoniidae) from Guangxi, China. Journal of Arachnology, 46, 345 - 354. https: // doi. org / 10.1636 / JoA-S- 17 - 063.1" type="journal article" year="2018">
Gao
<emphasis id="47A9EAB5FFE4FFA8FF78588F3B19F80A" box="[207,265,1951,1977]" italics="true" pageId="114" pageNumber="115">et al.</emphasis>
2018
</bibRefCitation>
,
<bibRefCitation id="114C4B56FFE4FFA8FEEE588F3B86F80A" author="Gao, Z. Z. &amp; Zhang, F. &amp; Chen, H. M." box="[345,406,1951,1977]" pageId="114" pageNumber="115" pagination="572 - 580" refId="ref76374" refString="Gao, Z. Z., Zhang, F. &amp; Chen, H. M. (2020) Two new cave-dwelling species of Tyrannochthonius Chamberlin 1929 (Pseudoscorpiones: Chthoniidae) from the Guizhou karst, China. Zootaxa, 4853 (4), 572 - 580. https: // doi. org / 10.11646 / zootaxa. 4853.4.6" type="journal article" year="2020">2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFE4FFA8FF7058D23985F86E" box="[199,917,1986,2013]" pageId="114" pageNumber="115" type="distribution">
<paragraph id="756236A7FFE4FFA8FF7058D23985F86E" blockId="114.[151,1437,151,2013]" box="[199,917,1986,2013]" pageId="114" pageNumber="115">
<emphasis id="47A9EAB5FFE4FFA8FF7058D23B71F86F" bold="true" box="[199,353,1986,2012]" pageId="114" pageNumber="115">Distribution.</emphasis>
Known only from the Gaoluo and Cangjun caves.
</paragraph>
</subSubSection>
</treatment>
</document>