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<document id="7AC0C953BD3B55835E5B8B7E18F42393" ID-CLB-Dataset="56235" ID-DOI="10.11646/zootaxa.5262.1.1" ID-GBIF-Dataset="e24a8032-296e-442d-80ae-e0bf8c7237f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7795161" ID-ZooBank="F05805FD-BC5B-4236-BF04-C6AA5C37F2D7" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1680528957709" checkinUser="plazi" docAuthor="Hou, Yanmeng, Feng, Zegang &amp; Zhang, Feng" docDate="2023" docId="FD7487B1FFAAFF98FF205F8838E1FF4B" docLanguage="en" docName="zootaxa.5262.1.1.pdf" docOrigin="Zootaxa 5262 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5262.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Tyrannochthonius infirmus Hou &amp; Feng &amp; Zhang 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="67" masterDocId="014DFFC9FF96FFDAFFB75F103A10FFB3" masterDocTitle="Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="158" masterPageNumber="1" pageNumber="61" updateTime="1701997099389" updateUser="plazi">
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<mods:title id="6ADF5498D1EA7B9A1E84435722C16D9B">Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae)</mods:title>
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<mods:namePart id="DE34D0D39EA61A966ED8B36A3EE2D252">Hou, Yanmeng</mods:namePart>
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<mods:namePart id="B77CD1DA2C23C6306DCD47001C12E615">Feng, Zegang</mods:namePart>
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<taxonomicName id="B2DD4D24FFAAFFE6FF205F883BF6FF01" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[151,486,152,179]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="infirmus" status="sp. nov.">
<emphasis id="47A9EAB5FFAAFFE6FF205F883BF6FF01" bold="true" box="[151,486,152,179]" italics="true" pageId="60" pageNumber="61">Tyrannochthonius infirmus</emphasis>
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<taxonomicNameLabel id="5C9A57CEFFAAFFE6FE595F88385CFF01" box="[494,588,152,178]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
NjṄƟṁae
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<figureCitation id="EDE62A22FFAAFFE6FF205FAC3B04FF64" box="[151,276,188,215]" captionStart-0="FIGURE 47" captionStart-1="FIGURE48" captionStart-2="FIGURE 49" captionStart-3="FIGURE 50" captionStart-4="FIGURE 51" captionStartId-0="60.[151,250,1941,1965]" captionStartId-1="61.[151,250,1429,1453]" captionStartId-2="63.[151,250,1866,1890]" captionStartId-3="64.[151,250,1943,1967]" captionStartId-4="65.[151,250,1943,1967]" captionTargetBox-0="[164,1424,454,1918]" captionTargetBox-1="[163,1434,502,1406]" captionTargetBox-2="[151,1436,181,1842]" captionTargetBox-3="[179,1408,181,1919]" captionTargetBox-4="[179,1408,181,1919]" captionTargetId-0="figure-88@60.[164,1424,454,1918]" captionTargetId-1="figure-136@61.[150,1437,489,1406]" captionTargetId-2="figure-18@63.[151,1436,181,1842]" captionTargetId-3="figure-20@64.[179,1408,181,1919]" captionTargetId-4="figure-18@65.[179,1408,181,1919]" captionTargetPageId-0="60" captionTargetPageId-1="61" captionTargetPageId-2="63" captionTargetPageId-3="64" captionTargetPageId-4="65" captionText-0="FIGURE 47. Liujiadadong Cave, type locality of Tyrannochthonius infirmus sp. nov., A. Entrance; B. Inside the cave entrance; C. Area where T. infirmus sp. nov. specimens were collected; D. Live male of T. infirmus sp. nov. in its natural environment; E. Live female of T. infirmus sp. nov. in its natural environment." captionText-1="FIGURE48.Tyrannochthoniusinfirmussp.nov.,A.Holotypemale,habitus(dorsalview);B.Paratypefemale,habitus(dorsalview)." captionText-2="FIGURE 49. Tyrannochthonius infirmus sp. nov., holotype male (AF, H), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (lateral view), indicate eyespot; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." captionText-3="FIGURE 50. Tyrannochthonius infirmus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." captionText-4="FIGURE 51. Tyrannochthonius infirmus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7795267" figureDoi-1="http://doi.org/10.5281/zenodo.7795269" figureDoi-2="http://doi.org/10.5281/zenodo.7795271" figureDoi-3="http://doi.org/10.5281/zenodo.7795273" figureDoi-4="http://doi.org/10.5281/zenodo.7795275" httpUri-0="https://zenodo.org/record/7795267/files/figure.png" httpUri-1="https://zenodo.org/record/7795269/files/figure.png" httpUri-2="https://zenodo.org/record/7795271/files/figure.png" httpUri-3="https://zenodo.org/record/7795273/files/figure.png" httpUri-4="https://zenodo.org/record/7795275/files/figure.png" pageId="60" pageNumber="61">Figs 4751</figureCitation>
</paragraph>
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<emphasis id="47A9EAB5FFAAFFE6FF205E143B5AFEAD" bold="true" box="[151,330,260,286]" pageId="60" pageNumber="61">Type material.</emphasis>
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<typeStatus id="AA668805FFAAFFE6FEE25E143BADFEAD" box="[341,445,260,286]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
♂ (Ps.-
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):
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,
<collectingRegion id="B719F845FFAAFFE6FC285E143E61FEAD" box="[927,1137,260,286]" country="China" name="Guizhou" pageId="60" pageNumber="61">Guizhou Province</collectingRegion>
,
<collectingCounty id="9C034E2BFFAAFFE6FB365E143F32FEAD" box="[1153,1314,260,286]" pageId="60" pageNumber="61">Puan County</collectingCounty>
,
<collectingMunicipality id="9506ACDDFFAAFFE6FA865E143AC9FEF0" pageId="60" pageNumber="61">Qingshan Town</collectingMunicipality>
,
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,
<location id="7002607CFFAAFFE6FE1D5E383893FEF1" LSID="urn:lsid:plazi:treatment:FD7487B1FFAAFF98FF205F8838E1FF4B:7002607CFFAAFFE6FE1D5E383893FEF1" box="[426,643,296,322]" country="China" county="Pu'an County" latitude="25.468578" longLatPrecision="1" longitude="105.00544" municipality="Qingshan Town" name="Liujiadadong Cave" pageId="60" pageNumber="61" stateProvince="Guizhou">Liujiadadong Cave</location>
, under stones and clods in the deep zone (Temperature: 13°C, Humidity: 90%) [
<geoCoordinate id="10E95060FFAAFFE6FEBC5E5E3BBBFEDA" box="[267,427,334,361]" degrees="25" direction="north" minutes="28" orientation="latitude" pageId="60" pageNumber="61" precision="1" seconds="6.88" value="25.468578">25°286.88″N</geoCoordinate>
,
<geoCoordinate id="10E95060FFAAFFE6FE025E5E384FFEDA" box="[437,607,334,361]" degrees="105" direction="east" minutes="0" orientation="longitude" pageId="60" pageNumber="61" precision="1" seconds="19.57" value="105.00544">105°019.57″E</geoCoordinate>
],
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a.s.l.
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,
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<collectingDate id="1127E98FFFAAFFE6FCBD5E5C39BEFED5" box="[778,942,332,358]" pageId="60" pageNumber="61" value="2019-08-04">4 August 2019</collectingDate>
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,
<collectorName id="D8285371FFAAFFE6FC005E5D3E5CFED4" box="[951,1100,333,359]" pageId="60" pageNumber="61">Zegang Feng</collectorName>
,
<collectorName id="D8285371FFAAFFE6FBE15E5C3ED4FED5" box="[1110,1220,332,358]" pageId="60" pageNumber="61">Zhaoyi Li</collectorName>
&amp;
<collectorName id="D8285371FFAAFFE6FB525E5C3F7EFED5" box="[1253,1390,332,358]" pageId="60" pageNumber="61">Chen Zhang</collectorName>
leg.
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<typeStatus id="AA668805FFAAFFE6FF205E613B18FE38" box="[151,264,369,395]" pageId="60" pageNumber="61" type="paratype">Paratypes</typeStatus>
: 3 ♂ (Ps.-
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<specimenCode id="257B9EDCFFAAFFE6FD245E603956FE39" box="[659,838,368,394]" collectionCode="GZC" pageId="60" pageNumber="61">GZC190804-04</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="C5B53CFAFFAAFFE6FCEF5E603850FE1D" ID-GBIF-Occurrence="4071438316" collectingDate="2019-08-04" collectionCode="GZC" collectorName="Zegang Feng &amp; Zhaoyi Li &amp; Chen Zhang" country="China" county="Pu'an County" elevation="1559" latitude="25.468578" location="Xiajiehe Village" longLatPrecision="1" longitude="105.00544" municipality="Qingshan Town" pageId="60" pageNumber="61" specimenCode="MSWU-GZC190804-05, GZC190804-09" specimenCount="5" specimenCount-female="5" stateProvince="Guizhou" typeStatus="paratype">
<specimenCount id="63DBFD2EFFAAFFE6FCEF5E603991FE38" box="[856,897,368,395]" pageId="60" pageNumber="61" type="female">5 ♀</specimenCount>
(Ps.-
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<specimenCode id="257B9EDCFFAAFFE6FB6A5E603F80FE38" box="[1245,1424,368,395]" collectionCode="GZC" pageId="60" pageNumber="61">GZC190804-09</specimenCode>
), all with the same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
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<paragraph id="756236A7FFAAFFE6FF205885391AF845" blockId="60.[151,1437,1941,2038]" pageId="60" pageNumber="61">
<emphasis id="47A9EAB5FFAAFFE6FF2058853B0FF81D" bold="true" box="[151,287,1941,1966]" pageId="60" pageNumber="61">FIGURE 47.</emphasis>
Liujiadadong Cave, type locality of
<taxonomicName id="B2DD4D24FFAAFFE6FD27588639B0F81D" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[656,928,1942,1966]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="infirmus" status="sp. nov.">
<emphasis id="47A9EAB5FFAAFFE6FD27588639B0F81D" box="[656,928,1942,1966]" italics="true" pageId="60" pageNumber="61">Tyrannochthonius infirmus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFAAFFE6FC11588639E6F81D" bold="true" box="[934,1014,1942,1966]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="5C9A57CEFFAAFFE6FC11588639E6F81D" box="[934,1014,1942,1966]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Entrance; B. Inside the cave entrance; C. Area where
<taxonomicName id="B2DD4D24FFAAFFE6FE9958AA3B8BF861" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[302,411,1978,2002]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="infirmus" status="sp. nov.">
<emphasis id="47A9EAB5FFAAFFE6FE9958AA3B8BF861" box="[302,411,1978,2002]" italics="true" pageId="60" pageNumber="61">T. infirmus</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="5C9A57CEFFAAFFE6FE1658AA3BE1F861" box="[417,497,1978,2002]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens were collected; D. Live male of
<taxonomicName id="B2DD4D24FFAAFFE6FC1B58AA3E08F861" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[940,1048,1978,2002]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="infirmus" status="sp. nov.">
<emphasis id="47A9EAB5FFAAFFE6FC1B58AA3E08F861" box="[940,1048,1978,2002]" italics="true" pageId="60" pageNumber="61">T. infirmus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFAAFFE6FBA858AA3E7FF861" bold="true" box="[1055,1135,1978,2002]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="5C9A57CEFFAAFFE6FBA858AA3E7FF861" box="[1055,1135,1978,2002]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in its natural environment; E. Live female of
<taxonomicName id="B2DD4D24FFAAFFE6FE8458CE3B8FF845" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[307,415,2014,2038]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="infirmus" status="sp. nov.">
<emphasis id="47A9EAB5FFAAFFE6FE8458CE3B8FF845" box="[307,415,2014,2038]" italics="true" pageId="60" pageNumber="61">T. infirmus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFAAFFE6FE1158CE3BE7F845" bold="true" box="[422,503,2014,2038]" pageId="60" pageNumber="61">
<taxonomicNameLabel id="5C9A57CEFFAAFFE6FE1158CE3BE7F845" box="[422,503,2014,2038]" pageId="60" pageNumber="61" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in its natural environment.
</paragraph>
</caption>
<subSubSection id="3DC7652CFFABFFE7FF705F87383DFF66" pageId="61" pageNumber="62" type="etymology">
<paragraph id="756236A7FFABFFE7FF705F87383DFF66" blockId="61.[151,1436,151,465]" pageId="61" pageNumber="62">
<emphasis id="47A9EAB5FFABFFE7FF705F873B5EFF02" bold="true" box="[199,334,151,177]" pageId="61" pageNumber="62">Etymology.</emphasis>
The specific name is derived from the Latin word “
<taxonomicName id="B2DD4D24FFABFFE7FC2A5F8739EAFF02" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[925,1018,151,177]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="infirmus">
<emphasis id="47A9EAB5FFABFFE7FC2A5F8739EAFF02" box="[925,1018,151,177]" italics="true" pageId="61" pageNumber="62">infirmus</emphasis>
</taxonomicName>
”, meaning weak, which refers to the presence of a pair of weak eyespots.
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFABFFE7FF705FCE3BF2FE3A" pageId="61" pageNumber="62" type="diagnosis">
<paragraph id="756236A7FFABFFE7FF705FCE3BF2FE3A" blockId="61.[151,1436,151,465]" pageId="61" pageNumber="62">
<emphasis id="47A9EAB5FFABFFE7FF705FCE3B96FF4F" bold="true" box="[199,390,222,252]" pageId="61" pageNumber="62">
Diagnosis (J
<specimenCount id="63DBFD2EFFABFFE7FEE85FF03B65FF4F" box="[351,373,224,252]" pageId="61" pageNumber="62" type="female"></specimenCount>
).
</emphasis>
Moderately sized troglomorphic species with elongated appendages; carapace with a pair of anterior eyespots only; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I each with 4 setae. Pedipalps slender, femur 6.726.88 (♂), 6.476.94 (
<specimenCount id="63DBFD2EFFABFFE7FEAC5E5B3B3DFED6" box="[283,301,331,357]" pageId="61" pageNumber="62" type="female"></specimenCount>
) times longer than broad; chela 7.277.42 (♂), 6.286.83 (
<specimenCount id="63DBFD2EFFABFFE7FC6E5E5B39FBFED6" box="[985,1003,331,357]" pageId="61" pageNumber="62" type="female"></specimenCount>
) times longer than broad; both chelal fingers with intercalary teeth.
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFABFFE4FF705E823F63F86E" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62" type="description">
<paragraph id="756236A7FFABFFE7FF705E823E09FE1E" blockId="61.[151,1436,151,465]" box="[199,1049,402,429]" pageId="61" pageNumber="62">
<emphasis id="47A9EAB5FFABFFE7FF705E823B49FE1F" bold="true" box="[199,345,402,428]" pageId="61" pageNumber="62">Description.</emphasis>
Males (
<typeStatus id="AA668805FFABFFE7FE045E833807FE1E" box="[435,535,403,429]" pageId="61" pageNumber="62" type="holotype">holotype</typeStatus>
and
<typeStatus id="AA668805FFABFFE7FDFA5E8338AAFE1E" box="[589,698,403,429]" pageId="61" pageNumber="62" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="EDE62A22FFABFFE7FD675E83392CFE1E" box="[720,828,403,429]" captionStart="FIGURE48" captionStartId="61.[151,250,1429,1453]" captionTargetBox="[163,1434,502,1406]" captionTargetId="figure-136@61.[150,1437,489,1406]" captionTargetPageId="61" captionText="FIGURE48.Tyrannochthoniusinfirmussp.nov.,A.Holotypemale,habitus(dorsalview);B.Paratypefemale,habitus(dorsalview)." figureDoi="http://doi.org/10.5281/zenodo.7795269" httpUri="https://zenodo.org/record/7795269/files/figure.png" pageId="61" pageNumber="62">Figs 48A</figureCitation>
,
<figureCitation id="EDE62A22FFABFFE7FCF05E8339A8FE1E" box="[839,952,403,429]" captionStart="FIGURE 49" captionStartId="63.[151,250,1866,1890]" captionTargetBox="[151,1436,181,1842]" captionTargetId="figure-18@63.[151,1436,181,1842]" captionTargetPageId="63" captionText="FIGURE 49. Tyrannochthonius infirmus sp. nov., holotype male (AF, H), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (lateral view), indicate eyespot; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795271" httpUri="https://zenodo.org/record/7795271/files/figure.png" pageId="61" pageNumber="62">49AF, H</figureCitation>
,
<figureCitation id="EDE62A22FFABFFE7FC745E8239F1FE1E" box="[963,993,402,429]" captionStart="FIGURE 50" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@64.[179,1408,181,1919]" captionTargetPageId="64" captionText="FIGURE 50. Tyrannochthonius infirmus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795273" httpUri="https://zenodo.org/record/7795273/files/figure.png" pageId="61" pageNumber="62">50</figureCitation>
,
<figureCitation id="EDE62A22FFABFFE7FC5A5E823E1CFE1E" box="[1005,1036,402,429]" captionStart="FIGURE 51" captionStartId="65.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@65.[179,1408,181,1919]" captionTargetPageId="65" captionText="FIGURE 51. Tyrannochthonius infirmus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795275" httpUri="https://zenodo.org/record/7795275/files/figure.png" pageId="61" pageNumber="62">51</figureCitation>
).
</paragraph>
<paragraph id="756236A7FFABFFE7FF705EA73EFFFE62" blockId="61.[151,1436,151,465]" box="[199,1263,439,465]" pageId="61" pageNumber="62">Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</paragraph>
<caption id="21A2662FFFABFFE7FF205A853F8DFA1E" ID-DOI="http://doi.org/10.5281/zenodo.7795269" ID-Zenodo-Dep="7795269" box="[151,1437,1429,1453]" httpUri="https://zenodo.org/record/7795269/files/figure.png" pageId="61" pageNumber="62" startId="61.[151,250,1429,1453]" targetBox="[163,1434,502,1406]" targetPageId="61">
<paragraph id="756236A7FFABFFE7FF205A853F8DFA1E" blockId="61.[151,1437,1429,1453]" box="[151,1437,1429,1453]" pageId="61" pageNumber="62">
<emphasis id="47A9EAB5FFABFFE7FF205A853B0BFA1E" bold="true" box="[151,283,1429,1453]" pageId="61" pageNumber="62">FIGURE48.</emphasis>
<emphasis id="47A9EAB5FFABFFE7FEAA5A85383AFA1E" box="[285,554,1429,1453]" italics="true" pageId="61" pageNumber="62">Tyrannochthoniusinfirmus</emphasis>
<emphasis id="47A9EAB5FFABFFE7FD9B5A853868FA1E" bold="true" box="[556,632,1429,1453]" pageId="61" pageNumber="62">
<taxonomicNameLabel id="5C9A57CEFFABFFE7FD9B5A853868FA1E" box="[556,632,1429,1453]" pageId="61" pageNumber="62" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
,A.Holotypemale,habitus(dorsalview);B.Paratypefemale,habitus(dorsalview).
</paragraph>
</caption>
<paragraph id="756236A7FFABFFE7FF705ACB38A8F889" blockId="61.[151,1437,1499,2030]" pageId="61" pageNumber="62">
Cephalothorax (
<figureCitation id="EDE62A22FFABFFE7FECB5ACB381AFA45" box="[380,522,1499,1526]" captionStart="FIGURE 49" captionStartId="63.[151,250,1866,1890]" captionTargetBox="[151,1436,181,1842]" captionTargetId="figure-18@63.[151,1436,181,1842]" captionTargetPageId="63" captionText="FIGURE 49. Tyrannochthonius infirmus sp. nov., holotype male (AF, H), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (lateral view), indicate eyespot; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795271" httpUri="https://zenodo.org/record/7795271/files/figure.png" pageId="61" pageNumber="62">Figs 49C, D</figureCitation>
,
<figureCitation id="EDE62A22FFABFFE7FDA15ACB385CFA45" box="[534,588,1499,1526]" captionStart="FIGURE 50" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@64.[179,1408,181,1919]" captionTargetPageId="64" captionText="FIGURE 50. Tyrannochthonius infirmus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795273" httpUri="https://zenodo.org/record/7795273/files/figure.png" pageId="61" pageNumber="62">50A</figureCitation>
): carapace 0.970.99 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; anterior eyes reduced into eyespots, posterior eyes reduced completely; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 setae arranged s4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 1113 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
<figureCitation id="EDE62A22FFABFFE7FA9F59EB3F9FF8A5" box="[1320,1423,1787,1814]" captionStart="FIGURE 50" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@64.[179,1408,181,1919]" captionTargetPageId="64" captionText="FIGURE 50. Tyrannochthonius infirmus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795273" httpUri="https://zenodo.org/record/7795273/files/figure.png" pageId="61" pageNumber="62">Fig. 50C</figureCitation>
); intercoxal tubercle absent; without sub-oral seta.
</paragraph>
<paragraph id="756236A7FFABFFE4FF7058533879FF4A" blockId="61.[151,1437,1499,2030]" lastBlockId="62.[151,1437,150,2013]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
Chelicera (
<figureCitation id="EDE62A22FFABFFE7FEF558533BBBF8EE" box="[322,427,1859,1885]" captionStart="FIGURE 49" captionStartId="63.[151,250,1866,1890]" captionTargetBox="[151,1436,181,1842]" captionTargetId="figure-18@63.[151,1436,181,1842]" captionTargetPageId="63" captionText="FIGURE 49. Tyrannochthonius infirmus sp. nov., holotype male (AF, H), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (lateral view), indicate eyespot; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795271" httpUri="https://zenodo.org/record/7795271/files/figure.png" pageId="61" pageNumber="62">Figs 49E</figureCitation>
,
<figureCitation id="EDE62A22FFABFFE7FE0F58533BFBF8ED" box="[440,491,1859,1886]" captionStart="FIGURE 50" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@64.[179,1408,181,1919]" captionTargetPageId="64" captionText="FIGURE 50. Tyrannochthonius infirmus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795273" httpUri="https://zenodo.org/record/7795273/files/figure.png" pageId="61" pageNumber="62">50B</figureCitation>
): large, about as long as carapace, 2.292.30 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 1214 teeth, distal one largest; movable finger with 1213 retrorse contiguous small teeth; galea completely vestigial (
<figureCitation id="EDE62A22FFABFFE7FBC658C33EC8F85D" box="[1137,1240,2003,2030]" captionStart="FIGURE 50" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@64.[179,1408,181,1919]" captionTargetPageId="64" captionText="FIGURE 50. Tyrannochthonius infirmus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795273" httpUri="https://zenodo.org/record/7795273/files/figure.png" pageId="61" pageNumber="62">Fig. 50B</figureCitation>
). Serrula exterior with 1821 and serrula interior with 1315 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
<figureCitation id="EDE62A22FFA8FFE4FE415FCF384CFF4A" box="[502,604,222,249]" captionStart="FIGURE 50" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@64.[179,1408,181,1919]" captionTargetPageId="64" captionText="FIGURE 50. Tyrannochthonius infirmus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795273" httpUri="https://zenodo.org/record/7795273/files/figure.png" pageId="62" pageNumber="63">Fig. 50E</figureCitation>
).
</paragraph>
<paragraph id="756236A7FFA8FFE4FF705E133907FCEE" blockId="62.[151,1437,150,2013]" pageId="62" pageNumber="63">
Pedipalp (
<figureCitation id="EDE62A22FFA8FFE4FE8C5E133BE3FEAE" box="[315,499,259,285]" captionStart="FIGURE 49" captionStartId="63.[151,250,1866,1890]" captionTargetBox="[151,1436,181,1842]" captionTargetId="figure-18@63.[151,1436,181,1842]" captionTargetPageId="63" captionText="FIGURE 49. Tyrannochthonius infirmus sp. nov., holotype male (AF, H), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (lateral view), indicate eyespot; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795271" httpUri="https://zenodo.org/record/7795271/files/figure.png" pageId="62" pageNumber="63">Figs 49A, B, H</figureCitation>
,
<figureCitation id="EDE62A22FFA8FFE4FDB65E123824FEAE" box="[513,564,258,285]" captionStart="FIGURE 50" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@64.[179,1408,181,1919]" captionTargetPageId="64" captionText="FIGURE 50. Tyrannochthonius infirmus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795273" httpUri="https://zenodo.org/record/7795273/files/figure.png" pageId="62" pageNumber="63">50D</figureCitation>
,
<figureCitation id="EDE62A22FFA8FFE4FDF55E123889FEAE" box="[578,665,258,285]" captionStart="FIGURE 51" captionStartId="65.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@65.[179,1408,181,1919]" captionTargetPageId="65" captionText="FIGURE 51. Tyrannochthonius infirmus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795275" httpUri="https://zenodo.org/record/7795275/files/figure.png" pageId="62" pageNumber="63">51A, B</figureCitation>
): long and slender, trochanter 1.421751, femur 6.726.88, patella 2.092.32, chela 7.277.42, hand 2.642.67 times longer than broad; femur 2.502.63 times longer than patella; movable chelal finger 1.741.78 times longer than hand and 0.630.64 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (
<figureCitation id="EDE62A22FFA8FFE4FD595E7F3949FE3A" box="[750,857,367,393]" captionStart="FIGURE 49" captionStartId="63.[151,250,1866,1890]" captionTargetBox="[151,1436,181,1842]" captionTargetId="figure-18@63.[151,1436,181,1842]" captionTargetPageId="63" captionText="FIGURE 49. Tyrannochthonius infirmus sp. nov., holotype male (AF, H), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (lateral view), indicate eyespot; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795271" httpUri="https://zenodo.org/record/7795271/files/figure.png" pageId="62" pageNumber="63">Figs 49H</figureCitation>
,
<figureCitation id="EDE62A22FFA8FFE4FCD25E7E3989FE3A" box="[869,921,366,393]" captionStart="FIGURE 50" captionStartId="64.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@64.[179,1408,181,1919]" captionTargetPageId="64" captionText="FIGURE 50. Tyrannochthonius infirmus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795273" httpUri="https://zenodo.org/record/7795273/files/figure.png" pageId="62" pageNumber="63">50D</figureCitation>
). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria,
<emphasis id="47A9EAB5FFA8FFE4FAE15EA73F7CFE62" box="[1366,1388,439,465]" italics="true" pageId="62" pageNumber="63">ib</emphasis>
and
<emphasis id="47A9EAB5FFA8FFE4FF205ECB3AA8FE46" box="[151,184,475,501]" italics="true" pageId="62" pageNumber="63">isb</emphasis>
situated close together, submedially on dorsum of chelal hand;
<emphasis id="47A9EAB5FFA8FFE4FC335ECB398EFE46" box="[900,926,475,501]" italics="true" pageId="62" pageNumber="63">eb</emphasis>
,
<emphasis id="47A9EAB5FFA8FFE4FC1A5ECB39C2FE46" box="[941,978,475,501]" italics="true" pageId="62" pageNumber="63">esb</emphasis>
and
<emphasis id="47A9EAB5FFA8FFE4FBBE5ECB3E33FE47" box="[1033,1059,475,500]" italics="true" pageId="62" pageNumber="63">ist</emphasis>
forming a nearly straight oblique row at base of fixed chelal finger;
<emphasis id="47A9EAB5FFA8FFE4FDAA5EEF383DFDAB" box="[541,557,511,536]" italics="true" pageId="62" pageNumber="63">it</emphasis>
slightly distal to
<emphasis id="47A9EAB5FFA8FFE4FD455EEF3901FDAB" box="[754,785,511,536]" italics="true" pageId="62" pageNumber="63">est</emphasis>
, situated subdistally;
<emphasis id="47A9EAB5FFA8FFE4FBB05EEF3E0BFDAB" box="[1031,1051,511,536]" italics="true" pageId="62" pageNumber="63">et</emphasis>
slightly near to tip of fixed chelal finger, very close to chelal teeth;
<emphasis id="47A9EAB5FFA8FFE4FDBA5D333837FD8E" box="[525,551,547,573]" italics="true" pageId="62" pageNumber="63">dx</emphasis>
situated distal to
<emphasis id="47A9EAB5FFA8FFE4FD595D333912FD8F" box="[750,770,547,572]" italics="true" pageId="62" pageNumber="63">et</emphasis>
;
<emphasis id="47A9EAB5FFA8FFE4FCA65D33393AFD8E" box="[785,810,547,573]" italics="true" pageId="62" pageNumber="63">sb</emphasis>
closer to
<emphasis id="47A9EAB5FFA8FFE4FC2F5D3339B6FD8E" box="[920,934,547,573]" italics="true" pageId="62" pageNumber="63">b</emphasis>
than to
<emphasis id="47A9EAB5FFA8FFE4FBB55D333E05FD8F" box="[1026,1045,547,572]" italics="true" pageId="62" pageNumber="63">st</emphasis>
;
<emphasis id="47A9EAB5FFA8FFE4FB935D333E22FD8E" box="[1060,1074,547,573]" italics="true" pageId="62" pageNumber="63">b</emphasis>
and
<emphasis id="47A9EAB5FFA8FFE4FBDF5D333E60FD8F" box="[1128,1136,547,572]" italics="true" pageId="62" pageNumber="63">t</emphasis>
situated subdistally,
<emphasis id="47A9EAB5FFA8FFE4FAEC5D333F73FD8F" box="[1371,1379,547,572]" italics="true" pageId="62" pageNumber="63">t</emphasis>
situated midway between
<emphasis id="47A9EAB5FFA8FFE4FE385D573B8FFDD3" box="[399,415,583,608]" italics="true" pageId="62" pageNumber="63">it</emphasis>
and
<emphasis id="47A9EAB5FFA8FFE4FE645D573BE2FDD3" box="[467,498,583,608]" italics="true" pageId="62" pageNumber="63">est</emphasis>
;
<emphasis id="47A9EAB5FFA8FFE4FDB75D57380FFDD3" box="[512,543,583,608]" italics="true" pageId="62" pageNumber="63">est</emphasis>
situated distal to
<emphasis id="47A9EAB5FFA8FFE4FD575D5738FEFDD2" box="[736,750,583,609]" italics="true" pageId="62" pageNumber="63">b</emphasis>
(
<figureCitation id="EDE62A22FFA8FFE4FD4A5D573974FDD2" box="[765,868,582,609]" captionStart="FIGURE 51" captionStartId="65.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@65.[179,1408,181,1919]" captionTargetPageId="65" captionText="FIGURE 51. Tyrannochthonius infirmus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795275" httpUri="https://zenodo.org/record/7795275/files/figure.png" pageId="62" pageNumber="63">Fig. 51A</figureCitation>
). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to
<emphasis id="47A9EAB5FFA8FFE4FE4D5D7B3804FD37" box="[506,532,619,644]" italics="true" pageId="62" pageNumber="63">ist</emphasis>
). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 2831 macrodenticles, slightly retrorse and pointed, plus 20
<specimenCount id="63DBFD2EFFA8FFE4FB615DA33F7FFD7E" box="[1238,1391,691,717]" pageId="62" pageNumber="63" type="intercalary">21 intercalary</specimenCount>
microdenticles,
<quantity id="B2259B42FFA8FFE4FE9A5DC63B80FD42" box="[301,400,726,753]" metricMagnitude="0" metricUnit="m" metricValue="1.27" metricValueMax="1.3208" metricValueMin="1.2192" pageId="62" pageNumber="63" unit="in" value="50.0" valueMax="52.0" valueMin="48.0">4852 in</quantity>
total; movable chelal finger with 2526 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 18
<specimenCount id="63DBFD2EFFA8FFE4FD505DEB3991FCA6" box="[743,897,763,789]" pageId="62" pageNumber="63" type="intercalary">19 intercalary</specimenCount>
microdenticles and 810 vestigial, rounded and contiguous basal teeth,
<quantity id="B2259B42FFA8FFE4FE295C0E3811FC8A" box="[414,513,798,825]" metricMagnitude="0" metricUnit="m" metricValue="1.3462" metricValueMax="1.3716" metricValueMin="1.3208" pageId="62" pageNumber="63" unit="in" value="53.0" valueMax="54.0" valueMin="52.0">5254 in</quantity>
total (
<figureCitation id="EDE62A22FFA8FFE4FDFE5C0F38A1FC8A" box="[585,689,798,825]" captionStart="FIGURE 51" captionStartId="65.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@65.[179,1408,181,1919]" captionTargetPageId="65" captionText="FIGURE 51. Tyrannochthonius infirmus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795275" httpUri="https://zenodo.org/record/7795275/files/figure.png" pageId="62" pageNumber="63">Fig. 51A</figureCitation>
). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
<figureCitation id="EDE62A22FFA8FFE4FDD45C5338DDFCEE" box="[611,717,835,861]" captionStart="FIGURE 49" captionStartId="63.[151,250,1866,1890]" captionTargetBox="[151,1436,181,1842]" captionTargetId="figure-18@63.[151,1436,181,1842]" captionTargetPageId="63" captionText="FIGURE 49. Tyrannochthonius infirmus sp. nov., holotype male (AF, H), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (lateral view), indicate eyespot; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795271" httpUri="https://zenodo.org/record/7795271/files/figure.png" pageId="62" pageNumber="63">Figs 49B</figureCitation>
,
<figureCitation id="EDE62A22FFA8FFE4FD6F5C52391AFCEE" box="[728,778,834,861]" captionStart="FIGURE 51" captionStartId="65.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@65.[179,1408,181,1919]" captionTargetPageId="65" captionText="FIGURE 51. Tyrannochthonius infirmus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795275" httpUri="https://zenodo.org/record/7795275/files/figure.png" pageId="62" pageNumber="63">51B</figureCitation>
).
</paragraph>
<paragraph id="756236A7FFA8FFE4FF705C773F8DFC5E" blockId="62.[151,1437,150,2013]" pageId="62" pageNumber="63">
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy IXII: 4: 4: 4: 4: 4: 5: 56: 5: 5: 4: T2T: 0; tergites VIIIX each with an unpaired median seta. Sternal chaetotaxy IIIXII: 1213: 1213: 89: 10: 910: 810: 910: 910: 0: 2. Anterior genital operculum with 1011 setae, genital opening slit-like, with 2023 marginal setae on each side,
<quantity id="B2259B42FFA8FFE4FB075CC23EF8FC5E" box="[1200,1256,978,1005]" metricMagnitude="0" metricUnit="m" metricValue="1.3462" pageId="62" pageNumber="63" unit="in" value="53.0">53 in</quantity>
total (
<figureCitation id="EDE62A22FFA8FFE4FA9A5CC33F80FC5E" box="[1325,1424,979,1005]" captionStart="FIGURE 49" captionStartId="63.[151,250,1866,1890]" captionTargetBox="[151,1436,181,1842]" captionTargetId="figure-18@63.[151,1436,181,1842]" captionTargetPageId="63" captionText="FIGURE 49. Tyrannochthonius infirmus sp. nov., holotype male (AF, H), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (lateral view), indicate eyespot; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795271" httpUri="https://zenodo.org/record/7795271/files/figure.png" pageId="62" pageNumber="63">Fig. 49F</figureCitation>
).
</paragraph>
<paragraph id="756236A7FFA8FFE4FF705CE73992FB76" blockId="62.[151,1437,150,2013]" pageId="62" pageNumber="63">
Legs (
<figureCitation id="EDE62A22FFA8FFE4FEBB5CE73B84FBA2" box="[268,404,1014,1041]" captionStart="FIGURE 51" captionStartId="65.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@65.[179,1408,181,1919]" captionTargetPageId="65" captionText="FIGURE 51. Tyrannochthonius infirmus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795275" httpUri="https://zenodo.org/record/7795275/files/figure.png" pageId="62" pageNumber="63">Fig. 51C, D</figureCitation>
): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.922.09 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.442.59 times longer than tibia. Femoropatella of leg IV 3.593.85 times longer than deep; tibia 6.256.58 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.563.60 times longer than deep (TS= 0.28), tarsus 12.4313.83 times longer than deep and 2.422.59 times longer than metatarsus (TS= 0.360.37). Arolium slightly shorter than the claws, not divided; claws simple.
</paragraph>
<paragraph id="756236A7FFA8FFE4FF705BDF3B08FA2E" blockId="62.[151,1437,150,2013]" pageId="62" pageNumber="63">
Adult females (
<typeStatus id="AA668805FFA8FFE4FEC25BDF3BF4FB5A" box="[373,484,1231,1257]" pageId="62" pageNumber="63" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="EDE62A22FFA8FFE4FE4D5BDF3875FB5A" box="[506,613,1231,1257]" captionStart="FIGURE48" captionStartId="61.[151,250,1429,1453]" captionTargetBox="[163,1434,502,1406]" captionTargetId="figure-136@61.[150,1437,489,1406]" captionTargetPageId="61" captionText="FIGURE48.Tyrannochthoniusinfirmussp.nov.,A.Holotypemale,habitus(dorsalview);B.Paratypefemale,habitus(dorsalview)." figureDoi="http://doi.org/10.5281/zenodo.7795269" httpUri="https://zenodo.org/record/7795269/files/figure.png" pageId="62" pageNumber="63">Figs 48B</figureCitation>
,
<figureCitation id="EDE62A22FFA8FFE4FDC65BDF38B6FB5A" box="[625,678,1231,1257]" captionStart="FIGURE 49" captionStartId="63.[151,250,1866,1890]" captionTargetBox="[151,1436,181,1842]" captionTargetId="figure-18@63.[151,1436,181,1842]" captionTargetPageId="63" captionText="FIGURE 49. Tyrannochthonius infirmus sp. nov., holotype male (AF, H), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (lateral view), indicate eyespot; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795271" httpUri="https://zenodo.org/record/7795271/files/figure.png" pageId="62" pageNumber="63">49G</figureCitation>
). Mostly same as males, but a little larger; with same chaetotaxy of coxae as males; tergal chaetotaxy IXII: 4: 4: 4: 4: 4: 45: 56: 57: 6: 4: T2T: 0; sternal chaetotaxy IVXII: 1113: 9: 10: 910: 910: 1011: 10: 0: 2; anterior genital operculum with 10 setae, posterior margin with 1113 marginal setae,
<quantity id="B2259B42FFA8FFE4FF6A5A2B3B2FFAE6" box="[221,319,1339,1365]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" metricValueMax="5.842" metricValueMin="5.334" pageId="62" pageNumber="63" unit="in" value="22.0" valueMax="23.0" valueMin="21.0">2123 in</quantity>
total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.443.56 times longer than deep (TS= 0.280.29), tarsus 12.2913.50 times longer than deep and 2.532.77 times longer than metatarsus (TS= 0.350.38).
</paragraph>
<paragraph id="756236A7FFA8FFE4FF705AB73BFDF952" blockId="62.[151,1437,150,2013]" pageId="62" pageNumber="63">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.412.54. Pedipalps: trochanter 0.270.35/0.190.20 (1.421.75), femur 1.101.21/0.160.18 (6.726.88), patella 0.440.46/0.190.22 (2.092.32), chela 1.601.78/0.220.24 (7.277.42), hand 0.580.64/0.220.24 (2.64 2.67), movable chelal finger length 1.011.14. Chelicera 0.690.71/0.300.31 (2.292.30), movable finger length 0.370.40. Carapace 0.620.68/0.640.69 (0.970.99). Leg I: trochanter 0.190.21/0.150.17 (1.241.27), femur 0.670.75/0.090.11 (6.827.44), patella 0.320.39/0.09 (3.564.33), tibia 0.290.34/0.060.08 (4.254.83), tarsus 0.750.83/0.060.07 (11.8612.50). Leg IV: trochanter 0.310.33/0.170.19 (1.741.82), femoropatella 0.97 1.04/0.27 (3.593.85), tibia 0.750.79/0.12 (6.256.58), metatarsus 0.320.36/0.090.10 (3.563.60), tarsus 0.83 0.87/0.060.07 (12.4313.83).</paragraph>
<paragraph id="756236A7FFA8FFE4FF7059FB3F63F86E" blockId="62.[151,1437,150,2013]" pageId="62" pageNumber="63">Females: body length 2.502.67. Pedipalps: trochanter 0.210.24/0.18 (1.171.33), femur 1.101.18/0.17 (6.476.94), patella 0.45/0.20 (2.25), chela 1.571.64/0.240.25 (6.286.83), hand 0.590.61/0.240.25 (2.362.54), movable chelal finger length 1.001.04. Chelicera 0.690.71/0.320.33 (2.152.16), movable finger length 0.37 0.39. Carapace 0.660.67/0.690.70 (0.940.97). Leg I: trochanter 0.170.19/0.150.16 (1.061.27), femur 0.64 0.68/0.10 (6.406.80), patella 0.35/0.080.09 (3.894.38), tibia 0.280.32/0.070.08 (4.00), tarsus 0.700.77/0.07 (10.0011.00). Leg IV: trochanter 0.29/0.17 (1.71), femoropatella 0.950.97/0.250.26 (3.733.80), tibia 0.69 0.72/0.110.12 (6.006.27), metatarsus 0.310.32/0.09 (3.443.56), tarsus 0.810.86/0.060.07 (12.2913.50).</paragraph>
</subSubSection>
<caption id="21A2662FFFA9FFE5FF20585A3F5BF819" ID-DOI="http://doi.org/10.5281/zenodo.7795271" ID-Zenodo-Dep="7795271" httpUri="https://zenodo.org/record/7795271/files/figure.png" pageId="63" pageNumber="64" startId="63.[151,250,1866,1890]" targetBox="[151,1436,181,1842]" targetPageId="63">
<paragraph id="756236A7FFA9FFE5FF20585A3F5BF819" blockId="63.[151,1436,1866,1962]" pageId="63" pageNumber="64">
<emphasis id="47A9EAB5FFA9FFE5FF20585A3B0FF8D1" bold="true" box="[151,287,1866,1890]" pageId="63" pageNumber="64">FIGURE 49.</emphasis>
<taxonomicName id="B2DD4D24FFA9FFE5FE92585A3825F8D1" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[293,565,1866,1890]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="infirmus" status="sp. nov.">
<emphasis id="47A9EAB5FFA9FFE5FE92585A3825F8D1" box="[293,565,1866,1890]" italics="true" pageId="63" pageNumber="64">Tyrannochthonius infirmus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFA9FFE5FD8C585A389BF8D1" bold="true" box="[571,651,1866,1890]" pageId="63" pageNumber="64">
<taxonomicNameLabel id="5C9A57CEFFA9FFE5FD8C585A389BF8D1" box="[571,651,1866,1890]" pageId="63" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (AF, H), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (lateral view), indicate eyespot; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view).
</paragraph>
</caption>
<caption id="21A2662FFFD6FF9AFF2058873851F84B" ID-DOI="http://doi.org/10.5281/zenodo.7795273" ID-Zenodo-Dep="7795273" httpUri="https://zenodo.org/record/7795273/files/figure.png" pageId="64" pageNumber="65" startId="64.[151,250,1943,1967]" targetBox="[179,1408,181,1919]" targetPageId="64">
<paragraph id="756236A7FFD6FF9AFF2058873851F84B" blockId="64.[151,1436,1943,2040]" pageId="64" pageNumber="65">
<emphasis id="47A9EAB5FFD6FF9AFF2058873B31F81C" bold="true" box="[151,289,1943,1967]" pageId="64" pageNumber="65">FIGURE 50.</emphasis>
<taxonomicName id="B2DD4D24FFD6FF9AFE9F5888382AF803" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[296,570,1944,1968]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="infirmus" status="sp. nov.">
<emphasis id="47A9EAB5FFD6FF9AFE9F5888382AF803" box="[296,570,1944,1968]" italics="true" pageId="64" pageNumber="65">Tyrannochthonius infirmus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFD6FF9AFDF558883883F803" bold="true" box="[578,659,1944,1968]" pageId="64" pageNumber="65">
<taxonomicNameLabel id="5C9A57CEFFD6FF9AFDF558883883F803" box="[578,659,1944,1968]" pageId="64" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (AB, D); 0.10 mm (C, E).
</paragraph>
</caption>
<caption id="21A2662FFFD7FF9BFF2058873F70F860" ID-DOI="http://doi.org/10.5281/zenodo.7795275" ID-Zenodo-Dep="7795275" httpUri="https://zenodo.org/record/7795275/files/figure.png" pageId="65" pageNumber="66" startId="65.[151,250,1943,1967]" targetBox="[179,1408,181,1919]" targetPageId="65">
<paragraph id="756236A7FFD7FF9BFF2058873F70F860" blockId="65.[151,1436,1943,2004]" pageId="65" pageNumber="66">
<emphasis id="47A9EAB5FFD7FF9BFF2058873B31F81C" bold="true" box="[151,289,1943,1967]" pageId="65" pageNumber="66">FIGURE 51.</emphasis>
<taxonomicName id="B2DD4D24FFD7FF9BFE9E5888382CF803" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[297,572,1944,1968]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="infirmus" status="sp. nov.">
<emphasis id="47A9EAB5FFD7FF9BFE9E5888382CF803" box="[297,572,1944,1968]" italics="true" pageId="65" pageNumber="66">Tyrannochthonius infirmus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFD7FF9BFDF358883887F803" bold="true" box="[580,663,1944,1968]" pageId="65" pageNumber="66">
<taxonomicNameLabel id="5C9A57CEFFD7FF9BFDF358883887F803" box="[580,663,1944,1968]" pageId="65" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm.
</paragraph>
</caption>
<subSubSection id="3DC7652CFFD4FF98FF705F8639E5FF66" pageId="66" pageNumber="67" type="discussion">
<paragraph id="756236A7FFD4FF98FF705F8639E5FF66" blockId="66.[151,1437,150,249]" pageId="66" pageNumber="67">
<emphasis id="47A9EAB5FFD4FF98FF705F863B2BFF03" bold="true" box="[199,315,150,176]" pageId="66" pageNumber="67">Remarks.</emphasis>
<taxonomicName id="B2DD4D24FFD4FF98FEF45F863862FF03" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[323,626,150,176]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="66" pageNumber="67" phylum="Arthropoda" rank="species" species="infirmus" status="sp. nov.">
<emphasis id="47A9EAB5FFD4FF98FEF45F863862FF03" box="[323,626,150,176]" italics="true" pageId="66" pageNumber="67">Tyrannochthonius infirmus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FFD4FF98FDCC5F8638C6FF03" bold="true" box="[635,726,150,176]" pageId="66" pageNumber="67">
<taxonomicNameLabel id="5C9A57CEFFD4FF98FDCC5F8638C6FF03" box="[635,726,150,176]" pageId="66" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from other described Chinese troglomorphic
<taxonomicName id="B2DD4D24FFD4FF98FF795FAA3BE0FF67" box="[206,496,186,212]" pageId="66" pageNumber="67">
<emphasis id="47A9EAB5FFD4FF98FF795FAA3B88FF67" box="[206,408,186,212]" italics="true" pageId="66" pageNumber="67">Tyrannochthonius</emphasis>
species
</taxonomicName>
by the presence of a pair of anterior eyespots.
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFFD4FF98FF705FCE38E1FF4B" box="[199,753,222,249]" pageId="66" pageNumber="67" type="distribution">
<paragraph id="756236A7FFD4FF98FF705FCE38E1FF4B" blockId="66.[151,1437,150,249]" box="[199,753,222,249]" pageId="66" pageNumber="67">
<emphasis id="47A9EAB5FFD4FF98FF705FCE3B71FF4B" bold="true" box="[199,353,222,248]" pageId="66" pageNumber="67">Distribution.</emphasis>
Known only from the
<typeStatus id="AA668805FFD4FF98FDD75FCF3880FF4A" box="[608,656,223,249]" pageId="66" pageNumber="67">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>