treatments-xml/data/FD/74/87/FD7487B1FF84FFC2FF205CBE38E1FBEA.xml

<document id="0183687307C9A38C083B1CE8D4AB46AE" ID-CLB-Dataset="56235" ID-DOI="10.11646/zootaxa.5262.1.1" ID-GBIF-Dataset="e24a8032-296e-442d-80ae-e0bf8c7237f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7795161" ID-ZooBank="F05805FD-BC5B-4236-BF04-C6AA5C37F2D7" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1680528957709" checkinUser="plazi" docAuthor="Hou, Yanmeng, Feng, Zegang &amp; Zhang, Feng" docDate="2023" docId="FD7487B1FF84FFC2FF205CBE38E1FBEA" docLanguage="en" docName="zootaxa.5262.1.1.pdf" docOrigin="Zootaxa 5262 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5262.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Tyrannochthonius arificus Hou &amp; Feng &amp; Zhang 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="25" masterDocId="014DFFC9FF96FFDAFFB75F103A10FFB3" masterDocTitle="Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="158" masterPageNumber="1" pageNumber="19" updateTime="1701997099389" updateUser="plazi">
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<mods:title id="F58BD3250794DE605475AAD79052D2F3">Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae)</mods:title>
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<mods:namePart id="AF3E0523F8730AF02A3859B94E0B22D7">Hou, Yanmeng</mods:namePart>
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<mods:namePart id="CD9D6775A94A5A95FBCFF6C2110E30D9">Feng, Zegang</mods:namePart>
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<mods:namePart id="60E4DE402D2F706D6AA3AA8501AAB798">Zhang, Feng</mods:namePart>
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<heading id="2E2A81CBFF84FFC8FF205CBE3905FC74" bold="true" box="[151,789,942,969]" fontSize="11" level="2" pageId="18" pageNumber="19" reason="4">
<emphasis id="47A9EAB5FF84FFC8FF205CBE3905FC74" bold="true" box="[151,789,942,969]" pageId="18" pageNumber="19">
<taxonomicName id="B2DD4D24FF84FFC8FF205CBE3BCBFC7B" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[151,475,942,969]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="arificus" status="sp. nov.">
<emphasis id="47A9EAB5FF84FFC8FF205CBE3BCBFC7B" bold="true" box="[151,475,942,969]" italics="true" pageId="18" pageNumber="19">Tyrannochthonius arificus</emphasis>
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<taxonomicNameLabel id="5C9A57CEFF84FFC8FE545CBF3850FC7A" box="[483,576,943,969]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
‹ȐƮƟṁae›
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<paragraph id="756236A7FF84FFC8FF205CC33B04FC5E" blockId="18.[151,789,942,1005]" box="[151,276,979,1005]" pageId="18" pageNumber="19">
<figureCitation id="EDE62A22FF84FFC8FF205CC33B04FC5E" box="[151,276,979,1005]" captionStart-0="FIGURE 12" captionStart-1="FIGURE13" captionStart-2="FIGURE 14" captionStart-3="FIGURE 15" captionStart-4="FIGURE 16" captionStartId-0="19.[151,250,1693,1717]" captionStartId-1="20.[151,250,1431,1455]" captionStartId-2="21.[151,250,1889,1913]" captionStartId-3="22.[151,250,1943,1967]" captionStartId-4="23.[151,250,1943,1967]" captionTargetBox-0="[148,1439,273,1670]" captionTargetBox-1="[158,1436,541,1407]" captionTargetBox-2="[177,1411,181,1865]" captionTargetBox-3="[179,1408,181,1908]" captionTargetBox-4="[179,1408,181,1919]" captionTargetId-0="figure-59@19.[148,1439,273,1670]" captionTargetId-1="figure-199@20.[151,1436,525,1408]" captionTargetId-2="figure-18@21.[177,1411,181,1865]" captionTargetId-3="figure-20@22.[179,1408,181,1919]" captionTargetId-4="figure-18@23.[179,1408,181,1919]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionTargetPageId-3="22" captionTargetPageId-4="23" captionText-0="FIGURE 12. Hei Cave, type locality of Tyrannochthonius arificus sp. nov., A. Cave location (red arrow); B. Entrance; C. Inside the cave entrance; D–E. Some narrow tunnels in the cave; F. Area where T. arificus sp. nov. specimens were collected." captionText-1="FIGURE13.Tyrannochthoniusarificussp.nov.,A.Holotypemale,habitus(dorsalview);B.Paratypefemale,habitus(dorsalview)." captionText-2="FIGURE 14. Tyrannochthonius arificus sp. nov., holotype male (A–C, E–F, H), paratype male (D), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (dorsal view), indicate epistome with one seta flanking base; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." captionText-3="FIGURE 15. Tyrannochthonius arificus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.10 mm." captionText-4="FIGURE 16. Tyrannochthonius arificus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.10 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7795190" figureDoi-1="http://doi.org/10.5281/zenodo.7795192" figureDoi-2="http://doi.org/10.5281/zenodo.7795196" figureDoi-3="http://doi.org/10.5281/zenodo.7795198" figureDoi-4="http://doi.org/10.5281/zenodo.7795200" httpUri-0="https://zenodo.org/record/7795190/files/figure.png" httpUri-1="https://zenodo.org/record/7795192/files/figure.png" httpUri-2="https://zenodo.org/record/7795196/files/figure.png" httpUri-3="https://zenodo.org/record/7795198/files/figure.png" httpUri-4="https://zenodo.org/record/7795200/files/figure.png" pageId="18" pageNumber="19">Figs 12–16</figureCitation>
</paragraph>
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<paragraph id="756236A7FF84FFC8FF205B0B3E83FB76" blockId="18.[151,1437,1050,2013]" pageId="18" pageNumber="19">
<emphasis id="47A9EAB5FF84FFC8FF205B0B3B54FB87" bold="true" box="[151,324,1050,1077]" pageId="18" pageNumber="19">Type material.</emphasis>
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<typeStatus id="AA668805FF84FFC8FEFD5B0B3BA2FB86" box="[330,434,1051,1077]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
♂ (Ps.- 
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- 
<specimenCode id="257B9EDCFF84FFC8FCD15B0A3990FB86" box="[870,896,1050,1077]" collectionCode="MHBU-HBUARA" pageId="18" pageNumber="19">01</specimenCode>
): 
<collectingCountry id="0DCA7637FF84FFC8FC255B0B39C8FB86" box="[914,984,1051,1077]" name="China" pageId="18" pageNumber="19">China</collectingCountry>
, 
<collectingRegion id="B719F845FF84FFC8FC555B0B3EBFFB86" box="[994,1199,1051,1077]" country="China" name="Guizhou" pageId="18" pageNumber="19">Guizhou Province</collectingRegion>
, 
<collectingCounty id="9C034E2BFF84FFC8FB0F5B0B3F77FB86" box="[1208,1383,1051,1077]" pageId="18" pageNumber="19">Anlong County</collectingCounty>
, 
<collectingMunicipality id="9506ACDDFF84FFC8FAC65B0B3B03FBEA" pageId="18" pageNumber="19">Dushan Town</collectingMunicipality>
, 
<location id="7002607CFF84FFC8FEA95B2F3BA8FBEA" LSID="urn:lsid:plazi:treatment:FD7487B1FF84FFC2FF205CBE38E1FBEA:7002607CFF84FFC8FEA95B2F3BA8FBEA" box="[286,440,1087,1113]" country="China" county="Anlong County" latitude="25.312933" longLatPrecision="1" longitude="105.600746" municipality="Dushan Town" name="Polao Village" pageId="18" pageNumber="19" stateProvince="Guizhou">Polao Village</location>
, 
<location id="7002607CFF84FFC8FE755B2F383DFBEA" LSID="urn:lsid:plazi:treatment:FD7487B1FF84FFC2FF205CBE38E1FBEA:7002607CFF84FFC8FE755B2F383DFBEA" box="[450,557,1087,1113]" country="China" county="Anlong County" latitude="25.312933" longLatPrecision="1" longitude="105.600746" municipality="Dushan Town" name="Hei Cave" pageId="18" pageNumber="19" stateProvince="Guizhou">Hei Cave</location>
, under stones and clods in the deep zone (Temperature: 16°C, Humidity: 65%) [
<geoCoordinate id="10E95060FF84FFC8FF295B743B5CFBCC" box="[158,332,1124,1151]" degrees="25" direction="north" minutes="18" orientation="latitude" pageId="18" pageNumber="19" precision="1" seconds="46.56" value="25.312933">25°18′46.56″N</geoCoordinate>
, 
<geoCoordinate id="10E95060FF84FFC8FEEF5B743812FBCC" box="[344,514,1124,1151]" degrees="105" direction="east" minutes="36" orientation="longitude" pageId="18" pageNumber="19" precision="1" seconds="2.69" value="105.600746">105°36′2.69″E</geoCoordinate>
], 
<elevation id="FEF0D194FF84FFC8FDA05B7338BCFBCE" box="[535,684,1123,1149]" metricMagnitude="3" metricUnit="m" metricValue="1.067" pageId="18" pageNumber="19" unit="m" value="1067.0">
<quantity id="B2259B42FF84FFC8FDA05B73387DFBCE" box="[535,621,1123,1149]" metricMagnitude="3" metricUnit="m" metricValue="1.067" pageId="18" pageNumber="19" unit="m" value="1067.0">1067 m</quantity>
a.s.l.
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, 
<date id="01631067FF84FFC8FD0D5B733974FBCE" box="[698,868,1123,1149]" pageId="18" pageNumber="19" value="2022-08-06">
<collectingDate id="1127E98FFF84FFC8FD0D5B733974FBCE" box="[698,868,1123,1149]" pageId="18" pageNumber="19" value="2022-08-06">6 August 2022</collectingDate>
</date>
, 
<collectorName id="D8285371FF84FFC8FCD85B733E05FBCE" box="[879,1045,1123,1149]" pageId="18" pageNumber="19">Yanmeng Hou</collectorName>
, 
<collectorName id="D8285371FF84FFC8FB955B733E84FBCE" box="[1058,1172,1123,1149]" pageId="18" pageNumber="19">Lu Zhang</collectorName>
, 
<collectorName id="D8285371FF84FFC8FB155B733F2AFBCE" box="[1186,1338,1123,1149]" pageId="18" pageNumber="19">Jianzhou Sun</collectorName>
&amp; 
<collectorName id="D8285371FF84FFC8FAD75B733AEDFB12" pageId="18" pageNumber="19">Wenlong Fan</collectorName>
leg.
</materialsCitation>
<materialsCitation id="C5B53CFAFF84FFC8FE8E5B973E9FFB76" ID-GBIF-Occurrence="4071438340" collectingDate="2022-08-06" collectionCode="MHBU-HBUARA" collectorName="Yanmeng Hou &amp; Lu Zhang &amp; Jianzhou Sun &amp; Wenlong Fan" country="China" county="Anlong County" elevation="1067" latitude="25.312933" location="Polao Village" longLatPrecision="1" longitude="105.600746" municipality="Dushan Town" pageId="18" pageNumber="19" specimenCode="MHBU-HBUARA#2022-506, 02" specimenCount="3" specimenCount-female="3" stateProvince="Guizhou" typeStatus="paratype">
<typeStatus id="AA668805FF84FFC8FE8E5B973BBAFB12" box="[313,426,1159,1185]" pageId="18" pageNumber="19" type="paratype">Paratypes</typeStatus>
: 3 ♂ (Ps.- 
<specimenCode id="257B9EDCFF84FFC8FDAC5B963961FB12" box="[539,881,1158,1185]" collectionCode="MHBU-HBUARA" pageId="18" pageNumber="19">MHBU-HBUARA#2022-506</specimenCode>
- 
<specimenCode id="257B9EDCFF84FFC8FCCF5B963984FB12" box="[888,916,1158,1185]" collectionCode="MHBU-HBUARA" pageId="18" pageNumber="19">02</specimenCode>
–HBUARA#2022-506-04), 
<specimenCount id="63DBFD2EFF84FFC8FB665B973EEEFB12" box="[1233,1278,1159,1185]" pageId="18" pageNumber="19" type="female">3 ♀</specimenCount>
(Ps.-MSWU-HBUARA#2022-506-05–HBUARA#2022-506-07), all with the same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFF84FFC8FF705BDF3838FABE" pageId="18" pageNumber="19" type="etymology">
<paragraph id="756236A7FF84FFC8FF705BDF3838FABE" blockId="18.[151,1437,1050,2013]" pageId="18" pageNumber="19">
<emphasis id="47A9EAB5FF84FFC8FF705BDF3B5EFB5A" bold="true" box="[199,334,1231,1257]" pageId="18" pageNumber="19">Etymology.</emphasis>
The specific name is derived from the Latin word “ 
<taxonomicName id="B2DD4D24FF84FFC8FC3D5BDF39F0FB5A" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[906,992,1231,1257]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="arificus">
<emphasis id="47A9EAB5FF84FFC8FC3D5BDF39F0FB5A" box="[906,992,1231,1257]" italics="true" pageId="18" pageNumber="19">arificus</emphasis>
</taxonomicName>
”, meaning arid, which refers to the species that lives in a dry environment.
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFF84FFC8FF705A063F8DFA2E" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="756236A7FF84FFC8FF705A063F8DFA2E" blockId="18.[151,1437,1050,2013]" pageId="18" pageNumber="19">
<emphasis id="47A9EAB5FF84FFC8FF705A063B94FA87" bold="true" box="[199,388,1302,1332]" pageId="18" pageNumber="19">
Diagnosis (J 
<specimenCount id="63DBFD2EFF84FFC8FEEA5A083B63FA87" box="[349,371,1304,1332]" pageId="18" pageNumber="19" type="female">♀</specimenCount>
).
</emphasis>
Small-sized troglomorphic species with slightly elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, rounded and obtuse; posterior margin of carapace with 2 setae; tergites I–Ⅴ each with 4 setae. Palpal femur 5.27–5.55 (♂), 5.36–5.50 (
<specimenCount id="63DBFD2EFF84FFC8FB585A4F3F11FACA" box="[1263,1281,1375,1401]" pageId="18" pageNumber="19" type="female">♀</specimenCount>
) times longer than broad; chela 6.29–6.62 (♂), 6.07–6.20 (
<specimenCount id="63DBFD2EFF84FFC8FD275A9338B2FA2E" box="[656,674,1411,1437]" pageId="18" pageNumber="19" type="female">♀</specimenCount>
) times longer than broad; both chelal fingers with intercalary teeth.
</paragraph>
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<subSubSection id="3DC7652CFF84FFC2FF705AB63F8DFD1A" lastPageId="24" lastPageNumber="25" pageId="18" pageNumber="19" type="description">
<paragraph id="756236A7FF84FFC8FF705AB63E09FA72" blockId="18.[151,1437,1050,2013]" box="[199,1049,1446,1473]" pageId="18" pageNumber="19">
<emphasis id="47A9EAB5FF84FFC8FF705AB63B49FA73" bold="true" box="[199,345,1446,1472]" pageId="18" pageNumber="19">Description.</emphasis>
Males (
<typeStatus id="AA668805FF84FFC8FE045AB73807FA72" box="[435,535,1447,1473]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
and 
<typeStatus id="AA668805FF84FFC8FDFA5AB738AAFA72" box="[589,698,1447,1473]" pageId="18" pageNumber="19" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="EDE62A22FF84FFC8FD675AB7392BFA72" box="[720,827,1447,1473]" captionStart="FIGURE13" captionStartId="20.[151,250,1431,1455]" captionTargetBox="[158,1436,541,1407]" captionTargetId="figure-199@20.[151,1436,525,1408]" captionTargetPageId="20" captionText="FIGURE13.Tyrannochthoniusarificussp.nov.,A.Holotypemale,habitus(dorsalview);B.Paratypefemale,habitus(dorsalview)." figureDoi="http://doi.org/10.5281/zenodo.7795192" httpUri="https://zenodo.org/record/7795192/files/figure.png" pageId="18" pageNumber="19">Figs 13A</figureCitation>
, 
<figureCitation id="EDE62A22FF84FFC8FCF05AB739A8FA72" box="[839,952,1447,1473]" captionStart="FIGURE 14" captionStartId="21.[151,250,1889,1913]" captionTargetBox="[177,1411,181,1865]" captionTargetId="figure-18@21.[177,1411,181,1865]" captionTargetPageId="21" captionText="FIGURE 14. Tyrannochthonius arificus sp. nov., holotype male (A–C, E–F, H), paratype male (D), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (dorsal view), indicate epistome with one seta flanking base; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795196" httpUri="https://zenodo.org/record/7795196/files/figure.png" pageId="18" pageNumber="19">14A–F, H</figureCitation>
, 
<figureCitation id="EDE62A22FF84FFC8FC745AB639F1FA72" box="[963,993,1446,1473]" captionStart="FIGURE 15" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-20@22.[179,1408,181,1919]" captionTargetPageId="22" captionText="FIGURE 15. Tyrannochthonius arificus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7795198" httpUri="https://zenodo.org/record/7795198/files/figure.png" pageId="18" pageNumber="19">15</figureCitation>
, 
<figureCitation id="EDE62A22FF84FFC8FC5A5AB73E1CFA72" box="[1005,1036,1447,1473]" captionStart="FIGURE 16" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@23.[179,1408,181,1919]" captionTargetPageId="23" captionText="FIGURE 16. Tyrannochthonius arificus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7795200" httpUri="https://zenodo.org/record/7795200/files/figure.png" pageId="18" pageNumber="19">16</figureCitation>
).
</paragraph>
<paragraph id="756236A7FF84FFC8FF705ADB3EFFFA56" blockId="18.[151,1437,1050,2013]" box="[199,1263,1483,1509]" pageId="18" pageNumber="19">Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</paragraph>
<paragraph id="756236A7FF84FFC8FF705AFF39D3F89A" blockId="18.[151,1437,1050,2013]" pageId="18" pageNumber="19">
Cephalothorax (
<figureCitation id="EDE62A22FF84FFC8FECB5AFF381AF9BA" box="[380,522,1519,1545]" captionStart="FIGURE 14" captionStartId="21.[151,250,1889,1913]" captionTargetBox="[177,1411,181,1865]" captionTargetId="figure-18@21.[177,1411,181,1865]" captionTargetPageId="21" captionText="FIGURE 14. Tyrannochthonius arificus sp. nov., holotype male (A–C, E–F, H), paratype male (D), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (dorsal view), indicate epistome with one seta flanking base; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795196" httpUri="https://zenodo.org/record/7795196/files/figure.png" pageId="18" pageNumber="19">Figs 14C, D</figureCitation>
, 
<figureCitation id="EDE62A22FF84FFC8FDA15AFE385BF9BA" box="[534,587,1518,1545]" captionStart="FIGURE 15" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-20@22.[179,1408,181,1919]" captionTargetPageId="22" captionText="FIGURE 15. Tyrannochthonius arificus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7795198" httpUri="https://zenodo.org/record/7795198/files/figure.png" pageId="18" pageNumber="19">15A</figureCitation>
): carapace 0.93–0.95 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small, rounded and obtuse; with 17–18 setae arranged s3–4s: 4: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to setae of ocular row, second pair situated lateral to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 3–4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 10 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
<figureCitation id="EDE62A22FF84FFC8FE90581F3B9DF89A" box="[295,397,1806,1833]" captionStart="FIGURE 15" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-20@22.[179,1408,181,1919]" captionTargetPageId="22" captionText="FIGURE 15. Tyrannochthonius arificus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7795198" httpUri="https://zenodo.org/record/7795198/files/figure.png" pageId="18" pageNumber="19">Fig. 15C</figureCitation>
); intercoxal tubercle absent; without sub-oral seta.
</paragraph>
<paragraph id="756236A7FF84FFC9FF7058233B83FF4A" blockId="18.[151,1437,1050,2013]" lastBlockId="19.[151,1437,151,249]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
Chelicera (
<figureCitation id="EDE62A22FF84FFC8FEF558233BBBF8FE" box="[322,427,1843,1869]" captionStart="FIGURE 14" captionStartId="21.[151,250,1889,1913]" captionTargetBox="[177,1411,181,1865]" captionTargetId="figure-18@21.[177,1411,181,1865]" captionTargetPageId="21" captionText="FIGURE 14. Tyrannochthonius arificus sp. nov., holotype male (A–C, E–F, H), paratype male (D), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (dorsal view), indicate epistome with one seta flanking base; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795196" httpUri="https://zenodo.org/record/7795196/files/figure.png" pageId="18" pageNumber="19">Figs 14E</figureCitation>
, 
<figureCitation id="EDE62A22FF84FFC8FE0F58223BFBF8FE" box="[440,491,1842,1869]" captionStart="FIGURE 15" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-20@22.[179,1408,181,1919]" captionTargetPageId="22" captionText="FIGURE 15. Tyrannochthonius arificus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7795198" httpUri="https://zenodo.org/record/7795198/files/figure.png" pageId="18" pageNumber="19">15B</figureCitation>
): large, about as long as carapace, 2.05–2.20 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on dorsal side. Both fingers well provided with teeth, fixed finger with 11–12 teeth, distal one largest; movable finger with 11–13 retrorse contiguous small teeth; galea completely vestigial (
<figureCitation id="EDE62A22FF84FFC8FC0C58D33E31F86E" box="[955,1057,1986,2013]" captionStart="FIGURE 15" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-20@22.[179,1408,181,1919]" captionTargetPageId="22" captionText="FIGURE 15. Tyrannochthonius arificus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7795198" httpUri="https://zenodo.org/record/7795198/files/figure.png" pageId="18" pageNumber="19">Fig. 15B</figureCitation>
). Serrula exterior with 18–20 and serrula interior with 11–13 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
<figureCitation id="EDE62A22FF85FFC9FE965FCF3B96FF4A" box="[289,390,222,249]" captionStart="FIGURE 15" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-20@22.[179,1408,181,1919]" captionTargetPageId="22" captionText="FIGURE 15. Tyrannochthonius arificus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7795198" httpUri="https://zenodo.org/record/7795198/files/figure.png" pageId="19" pageNumber="20">Fig. 15E</figureCitation>
).
</paragraph>
<caption id="21A2662FFF85FFC9FF20598D3F59F96A" ID-DOI="http://doi.org/10.5281/zenodo.7795190" ID-Zenodo-Dep="7795190" httpUri="https://zenodo.org/record/7795190/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1693,1717]" targetBox="[148,1439,273,1670]" targetPageId="19">
<paragraph id="756236A7FF85FFC9FF20598D3F59F96A" blockId="19.[151,1436,1693,1754]" pageId="19" pageNumber="20">
<emphasis id="47A9EAB5FF85FFC9FF20598D3B0FF906" bold="true" box="[151,287,1693,1717]" pageId="19" pageNumber="20">FIGURE 12.</emphasis>
Hei Cave, type locality of 
<taxonomicName id="B2DD4D24FF85FFC9FD9D598D3923F906" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[554,819,1693,1717]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="arificus" status="sp. nov.">
<emphasis id="47A9EAB5FF85FFC9FD9D598D3923F906" box="[554,819,1693,1717]" italics="true" pageId="19" pageNumber="20">Tyrannochthonius arificus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FF85FFC9FC8E598D3998F906" bold="true" box="[825,904,1693,1717]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="5C9A57CEFF85FFC9FC8E598D3998F906" box="[825,904,1693,1717]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Cave location (red arrow); B. Entrance; C. Inside the cave entrance; D–E. Some narrow tunnels in the cave; F. Area where 
<taxonomicName id="B2DD4D24FF85FFC9FCCC59D239F1F96A" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[891,993,1729,1753]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="arificus" status="sp. nov.">
<emphasis id="47A9EAB5FF85FFC9FCCC59D239F1F96A" box="[891,993,1729,1753]" italics="true" pageId="19" pageNumber="20">T. arificus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FF85FFC9FC5F59D13E28F96A" bold="true" box="[1000,1080,1729,1753]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="5C9A57CEFF85FFC9FC5F59D13E28F96A" box="[1000,1080,1729,1753]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens were collected.
</paragraph>
</caption>
<paragraph id="756236A7FF85FFCEFF7058173844FE46" blockId="19.[151,1437,1799,2042]" lastBlockId="20.[151,1437,151,501]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
Pedipalp (
<figureCitation id="EDE62A22FF85FFC9FE8058173BF4F891" box="[311,484,1799,1826]" captionStart="FIGURE 14" captionStartId="21.[151,250,1889,1913]" captionTargetBox="[177,1411,181,1865]" captionTargetId="figure-18@21.[177,1411,181,1865]" captionTargetPageId="21" captionText="FIGURE 14. Tyrannochthonius arificus sp. nov., holotype male (A–C, E–F, H), paratype male (D), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (dorsal view), indicate epistome with one seta flanking base; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795196" httpUri="https://zenodo.org/record/7795196/files/figure.png" pageId="19" pageNumber="20">Figs 14A, B, H</figureCitation>
, 
<figureCitation id="EDE62A22FF85FFC9FE5958173851F891" box="[494,577,1799,1826]" captionStart="FIGURE 16" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@23.[179,1408,181,1919]" captionTargetPageId="23" captionText="FIGURE 16. Tyrannochthonius arificus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7795200" httpUri="https://zenodo.org/record/7795200/files/figure.png" pageId="19" pageNumber="20">16A, B</figureCitation>
): long and slender, trochanter 1.15–1.25, femur 5.27–5.55, patella 2.15–2.17, chela 6.29–6.62, hand 2.07–2.15 times longer than broad; femur 2.18–2.23 times longer than patella; movable chelal finger 2.10–2.14 times longer than hand and 0.69–0.70 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (
<figureCitation id="EDE62A22FF85FFC9FDCA586338F9F83D" box="[637,745,1907,1934]" captionStart="FIGURE 14" captionStartId="21.[151,250,1889,1913]" captionTargetBox="[177,1411,181,1865]" captionTargetId="figure-18@21.[177,1411,181,1865]" captionTargetPageId="21" captionText="FIGURE 14. Tyrannochthonius arificus sp. nov., holotype male (A–C, E–F, H), paratype male (D), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (dorsal view), indicate epistome with one seta flanking base; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795196" httpUri="https://zenodo.org/record/7795196/files/figure.png" pageId="19" pageNumber="20">Figs 14H</figureCitation>
, 
<figureCitation id="EDE62A22FF85FFC9FD4258633939F83D" box="[757,809,1907,1934]" captionStart="FIGURE 15" captionStartId="22.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1908]" captionTargetId="figure-20@22.[179,1408,181,1919]" captionTargetPageId="22" captionText="FIGURE 15. Tyrannochthonius arificus sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7795198" httpUri="https://zenodo.org/record/7795198/files/figure.png" pageId="19" pageNumber="20">15D</figureCitation>
). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger and hand. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria, 
<emphasis id="47A9EAB5FF85FFC9FA3158AB3F8CF866" box="[1414,1436,1979,2005]" italics="true" pageId="19" pageNumber="20">ib</emphasis>
and 
<emphasis id="47A9EAB5FF85FFC9FF7158CF3AF7F84A" box="[198,231,2015,2041]" italics="true" pageId="19" pageNumber="20">isb</emphasis>
situated close together, submedially on dorsum of chelal hand; 
<emphasis id="47A9EAB5FF85FFC9FC1058CF39D1F84A" box="[935,961,2015,2041]" italics="true" pageId="19" pageNumber="20">eb</emphasis>
, 
<emphasis id="47A9EAB5FF85FFC9FC7858CF39E4F84A" box="[975,1012,2015,2041]" italics="true" pageId="19" pageNumber="20">esb</emphasis>
and 
<emphasis id="47A9EAB5FF85FFC9FB9E58F03E53F84A" box="[1065,1091,2016,2041]" italics="true" pageId="19" pageNumber="20">ist</emphasis>
forming a straight oblique row at base of fixed chelal finger; 
<emphasis id="47A9EAB5FF82FFCEFE565F873BE1FF03" box="[481,497,151,176]" italics="true" pageId="20" pageNumber="21">it</emphasis>
slightly distal to 
<emphasis id="47A9EAB5FF82FFCEFD075F8738DFFF03" box="[688,719,151,176]" italics="true" pageId="20" pageNumber="21">est</emphasis>
, situated subdistally; 
<emphasis id="47A9EAB5FF82FFCEFC755F8739C6FF03" box="[962,982,151,176]" italics="true" pageId="20" pageNumber="21">et</emphasis>
slightly near to tip of fixed chelal finger, very close to chelal teeth; 
<emphasis id="47A9EAB5FF82FFCEFE0F5FAB3BC2FF66" box="[440,466,187,213]" italics="true" pageId="20" pageNumber="21">dx</emphasis>
situated distal to 
<emphasis id="47A9EAB5FF82FFCEFD225FAB38B9FF67" box="[661,681,187,212]" italics="true" pageId="20" pageNumber="21">et</emphasis>
; 
<emphasis id="47A9EAB5FF82FFCEFD005FAB38C0FF66" box="[695,720,187,213]" italics="true" pageId="20" pageNumber="21">sb</emphasis>
closer to 
<emphasis id="47A9EAB5FF82FFCEFC8C5FAB395EFF67" box="[827,846,187,212]" italics="true" pageId="20" pageNumber="21">st</emphasis>
than to 
<emphasis id="47A9EAB5FF82FFCEFC115FAB39A4FF66" box="[934,948,187,213]" italics="true" pageId="20" pageNumber="21">b</emphasis>
; 
<emphasis id="47A9EAB5FF82FFCEFC755FAB39C0FF66" box="[962,976,187,213]" italics="true" pageId="20" pageNumber="21">b</emphasis>
and 
<emphasis id="47A9EAB5FF82FFCEFBB25FAB3E1DFF67" box="[1029,1037,187,212]" italics="true" pageId="20" pageNumber="21">t</emphasis>
situated subdistally, 
<emphasis id="47A9EAB5FF82FFCEFB435FAB3EECFF67" box="[1268,1276,187,212]" italics="true" pageId="20" pageNumber="21">t</emphasis>
situated distal to 
<emphasis id="47A9EAB5FF82FFCEFF025FCF3AD3FF4A" box="[181,195,223,249]" italics="true" pageId="20" pageNumber="21">b</emphasis>
and at same level as 
<emphasis id="47A9EAB5FF82FFCEFE0F5FCF3BC7FF4B" box="[440,471,223,248]" italics="true" pageId="20" pageNumber="21">est</emphasis>
(
<figureCitation id="EDE62A22FF82FFCEFE5F5FCF3841FF4A" box="[488,593,223,249]" captionStart="FIGURE 16" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@23.[179,1408,181,1919]" captionTargetPageId="23" captionText="FIGURE 16. Tyrannochthonius arificus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7795200" httpUri="https://zenodo.org/record/7795200/files/figure.png" pageId="20" pageNumber="21">Fig. 16A</figureCitation>
). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to 
<emphasis id="47A9EAB5FF82FFCEFF4D5E133B04FEAF" box="[250,276,259,284]" italics="true" pageId="20" pageNumber="21">ist</emphasis>
). Both chelal fingers with a row of teeth, heterodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 30–32 macrodenticles, slightly retrorse and pointed, plus 25– 
<specimenCount id="63DBFD2EFF82FFCEFC6B5E5A3E69FED6" box="[988,1145,330,357]" pageId="20" pageNumber="21" type="intercalary">26 intercalary</specimenCount>
microdenticles, 
<quantity id="B2259B42FF82FFCEFA805E5A3F8CFED6" box="[1335,1436,330,357]" metricMagnitude="0" metricUnit="m" metricValue="1.4350999999999998" metricValueMax="1.4478" metricValueMin="1.4224" pageId="20" pageNumber="21" unit="in" value="56.5" valueMax="57.0" valueMin="56.0">56–57 in</quantity>
total; movable chelal finger with 12–13 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus 10– 
<specimenCount id="63DBFD2EFF82FFCEFE555E833869FE1E" box="[482,633,403,429]" pageId="20" pageNumber="21" type="intercalary">11 intercalary</specimenCount>
microdenticles and 17–19 vestigial, rounded and contiguous basal teeth, 
<quantity id="B2259B42FF82FFCEFF205EA73AC2FE62" box="[151,210,439,465]" metricMagnitude="0" metricUnit="m" metricValue="1.0413999999999999" pageId="20" pageNumber="21" unit="in" value="41.0">41 in</quantity>
total (
<figureCitation id="EDE62A22FF82FFCEFEAA5EA73B97FE62" box="[285,391,439,465]" captionStart="FIGURE 16" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@23.[179,1408,181,1919]" captionTargetPageId="23" captionText="FIGURE 16. Tyrannochthonius arificus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7795200" httpUri="https://zenodo.org/record/7795200/files/figure.png" pageId="20" pageNumber="21">Fig. 16A</figureCitation>
). Chelal fingers slightly curved in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
<figureCitation id="EDE62A22FF82FFCEFE175ECB381AFE46" box="[416,522,475,501]" captionStart="FIGURE 14" captionStartId="21.[151,250,1889,1913]" captionTargetBox="[177,1411,181,1865]" captionTargetId="figure-18@21.[177,1411,181,1865]" captionTargetPageId="21" captionText="FIGURE 14. Tyrannochthonius arificus sp. nov., holotype male (A–C, E–F, H), paratype male (D), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (dorsal view), indicate epistome with one seta flanking base; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795196" httpUri="https://zenodo.org/record/7795196/files/figure.png" pageId="20" pageNumber="21">Figs 14B</figureCitation>
, 
<figureCitation id="EDE62A22FF82FFCEFDA25ECB3857FE46" box="[533,583,475,501]" captionStart="FIGURE 16" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@23.[179,1408,181,1919]" captionTargetPageId="23" captionText="FIGURE 16. Tyrannochthonius arificus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7795200" httpUri="https://zenodo.org/record/7795200/files/figure.png" pageId="20" pageNumber="21">16B</figureCitation>
).
</paragraph>
<caption id="21A2662FFF82FFCEFF205A873F8DFA1C" ID-DOI="http://doi.org/10.5281/zenodo.7795192" ID-Zenodo-Dep="7795192" box="[151,1437,1431,1456]" httpUri="https://zenodo.org/record/7795192/files/figure.png" pageId="20" pageNumber="21" startId="20.[151,250,1431,1455]" targetBox="[158,1436,541,1407]" targetPageId="20">
<paragraph id="756236A7FF82FFCEFF205A873F8DFA1C" blockId="20.[151,1437,1431,1456]" box="[151,1437,1431,1456]" pageId="20" pageNumber="21">
<emphasis id="47A9EAB5FF82FFCEFF205A873B0CFA1C" bold="true" box="[151,284,1431,1455]" pageId="20" pageNumber="21">FIGURE13.</emphasis>
<taxonomicName id="B2DD4D24FF82FFCEFEA95A873834FA03" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[286,548,1431,1456]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="arificus" status="sp. nov.">
<emphasis id="47A9EAB5FF82FFCEFEA95A873834FA03" box="[286,548,1431,1456]" italics="true" pageId="20" pageNumber="21">Tyrannochthonius arificus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FF82FFCEFD915A883863FA03" bold="true" box="[550,627,1432,1456]" pageId="20" pageNumber="21">
<taxonomicNameLabel id="5C9A57CEFF82FFCEFD915A883863FA03" box="[550,627,1432,1456]" pageId="20" pageNumber="21" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
,A.Holotype male,habitus (dorsal view);B.Paratype female,habitus (dorsal view).
</paragraph>
</caption>
<paragraph id="756236A7FF82FFCEFF705ACE3F4BF9D7" blockId="20.[151,1437,1502,2032]" pageId="20" pageNumber="21">
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 5: 5: 5: 5: 3: T2T: 0, tergites VI–IX each with an unpaired median seta. Sternal chaetotaxy III–XII: 13–15: 13–14: 10: 9: 9: 9: 9–10: 9: 0: 2. Anterior genital operculum with 10–11 setae, genital opening slit-like, with 11–14 marginal setae on each side, 
<quantity id="B2259B42FF82FFCEFBF759593EB3F9D7" box="[1088,1187,1609,1636]" metricMagnitude="-1" metricUnit="m" metricValue="9.017" metricValueMax="9.144" metricValueMin="8.89" pageId="20" pageNumber="21" unit="in" value="35.5" valueMax="36.0" valueMin="35.0">35–36 in</quantity>
total (
<figureCitation id="EDE62A22FF82FFCEFB5C595A3F5FF9D7" box="[1259,1359,1610,1636]" captionStart="FIGURE 14" captionStartId="21.[151,250,1889,1913]" captionTargetBox="[177,1411,181,1865]" captionTargetId="figure-18@21.[177,1411,181,1865]" captionTargetPageId="21" captionText="FIGURE 14. Tyrannochthonius arificus sp. nov., holotype male (A–C, E–F, H), paratype male (D), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (dorsal view), indicate epistome with one seta flanking base; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795196" httpUri="https://zenodo.org/record/7795196/files/figure.png" pageId="20" pageNumber="21">Fig. 14F</figureCitation>
).
</paragraph>
<paragraph id="756236A7FF82FFCEFF70597E3992F88F" blockId="20.[151,1437,1502,2032]" pageId="20" pageNumber="21">
Legs (
<figureCitation id="EDE62A22FF82FFCEFEBB597E3B84F93B" box="[268,404,1646,1672]" captionStart="FIGURE 16" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-18@23.[179,1408,181,1919]" captionTargetPageId="23" captionText="FIGURE 16. Tyrannochthonius arificus sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.7795200" httpUri="https://zenodo.org/record/7795200/files/figure.png" pageId="20" pageNumber="21">Fig. 16C, D</figureCitation>
): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.76–1.89 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.22–2.37 times longer than tibia. Femoropatella of leg IV 2.84–3.06 times longer than deep; tibia 4.78–5.00 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.17 times longer than deep (TS= 0.26–0.37), tarsus 11.75–14.67 times longer than deep and 2.32–2.47 times longer than metatarsus (TS= 0.25–0.34). Arolium slightly shorter than the claws, not divided; claws simple.
</paragraph>
<paragraph id="756236A7FF82FFCEFF7058563EF0F843" blockId="20.[151,1437,1502,2032]" pageId="20" pageNumber="21">
Adult females (
<typeStatus id="AA668805FF82FFCEFEC558563BF1F8D3" box="[370,481,1862,1888]" pageId="20" pageNumber="21" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="EDE62A22FF82FFCEFE415856384FF8D3" box="[502,607,1862,1888]" captionStart="FIGURE13" captionStartId="20.[151,250,1431,1455]" captionTargetBox="[158,1436,541,1407]" captionTargetId="figure-199@20.[151,1436,525,1408]" captionTargetPageId="20" captionText="FIGURE13.Tyrannochthoniusarificussp.nov.,A.Holotypemale,habitus(dorsalview);B.Paratypefemale,habitus(dorsalview)." figureDoi="http://doi.org/10.5281/zenodo.7795192" httpUri="https://zenodo.org/record/7795192/files/figure.png" pageId="20" pageNumber="21">Figs 13B</figureCitation>
, 
<figureCitation id="EDE62A22FF82FFCEFDDD5856388EF8D3" box="[618,670,1862,1888]" captionStart="FIGURE 14" captionStartId="21.[151,250,1889,1913]" captionTargetBox="[177,1411,181,1865]" captionTargetId="figure-18@21.[177,1411,181,1865]" captionTargetPageId="21" captionText="FIGURE 14. Tyrannochthonius arificus sp. nov., holotype male (A–C, E–F, H), paratype male (D), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (dorsal view), indicate epistome with one seta flanking base; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795196" httpUri="https://zenodo.org/record/7795196/files/figure.png" pageId="20" pageNumber="21">14G</figureCitation>
). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; with same tergal chaetotaxy as males; sternal chaetotaxy IV–XII: 14–15: 10: 9–10: 9: 9–10: 9–11: 8–9: 0: 2; anterior genital operculum with 10–11 setae, posterior margin with 14 marginal setae, 
<quantity id="B2259B42FF82FFCEFB67589D3F23F81B" box="[1232,1331,1933,1960]" metricMagnitude="-1" metricUnit="m" metricValue="6.223" metricValueMax="6.35" metricValueMin="6.096" pageId="20" pageNumber="21" unit="in" value="24.5" valueMax="25.0" valueMin="24.0">24–25 in</quantity>
total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.00–3.50 times longer than deep (TS= 0.33), tarsus 12.00–14.67 times longer than deep and 2.29–2.44 times longer than metatarsus (TS= 0.27–0.29).
</paragraph>
<caption id="21A2662FFF83FFCFFF205871388FF856" ID-DOI="http://doi.org/10.5281/zenodo.7795196" ID-Zenodo-Dep="7795196" httpUri="https://zenodo.org/record/7795196/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,250,1889,1913]" targetBox="[177,1411,181,1865]" targetPageId="21">
<paragraph id="756236A7FF83FFCFFF205871388FF856" blockId="21.[151,1437,1889,2021]" pageId="21" pageNumber="22">
<emphasis id="47A9EAB5FF83FFCFFF2058713B31F8CA" bold="true" box="[151,289,1889,1913]" pageId="21" pageNumber="22">FIGURE 14.</emphasis>
<taxonomicName id="B2DD4D24FF83FFCFFE9E58713825F8CA" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[297,565,1889,1913]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arificus" status="sp. nov.">
<emphasis id="47A9EAB5FF83FFCFFE9E58713825F8CA" box="[297,565,1889,1913]" italics="true" pageId="21" pageNumber="22">Tyrannochthonius arificus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FF83FFCFFD8A5872389FF8C9" bold="true" box="[573,655,1890,1914]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="5C9A57CEFF83FFCFFD8A5872389FF8C9" box="[573,655,1890,1914]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (A–C, E–F, H), paratype male (D), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Carapace (dorsal view), indicate epistome with one seta flanking base; E. Left chelicera (dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view); H. Left pedipalp (minus chela, dorsal view).
</paragraph>
</caption>
<caption id="21A2662FFF80FFCCFF2058873B20F844" ID-DOI="http://doi.org/10.5281/zenodo.7795198" ID-Zenodo-Dep="7795198" httpUri="https://zenodo.org/record/7795198/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,250,1943,1967]" targetBox="[179,1408,181,1908]" targetPageId="22">
<paragraph id="756236A7FF80FFCCFF2058873B20F844" blockId="22.[151,1436,1943,2040]" pageId="22" pageNumber="23">
<emphasis id="47A9EAB5FF80FFCCFF2058873B31F81C" bold="true" box="[151,289,1943,1967]" pageId="22" pageNumber="23">FIGURE 15.</emphasis>
<taxonomicName id="B2DD4D24FF80FFCCFE9E58883824F803" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[297,564,1944,1968]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="arificus" status="sp. nov.">
<emphasis id="47A9EAB5FF80FFCCFE9E58883824F803" box="[297,564,1944,1968]" italics="true" pageId="22" pageNumber="23">Tyrannochthonius arificus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FF80FFCCFD8A5888389FF803" bold="true" box="[573,655,1944,1968]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="5C9A57CEFF80FFCCFD8A5888389FF803" box="[573,655,1944,1968]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.10 mm.
</paragraph>
</caption>
<caption id="21A2662FFF81FFCDFF2058873F70F860" ID-DOI="http://doi.org/10.5281/zenodo.7795200" ID-Zenodo-Dep="7795200" httpUri="https://zenodo.org/record/7795200/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1943,1967]" targetBox="[179,1408,181,1919]" targetPageId="23">
<paragraph id="756236A7FF81FFCDFF2058873F70F860" blockId="23.[151,1436,1943,2004]" pageId="23" pageNumber="24">
<emphasis id="47A9EAB5FF81FFCDFF2058873B32F81C" bold="true" box="[151,290,1943,1967]" pageId="23" pageNumber="24">FIGURE 16.</emphasis>
<taxonomicName id="B2DD4D24FF81FFCDFE9D58883826F803" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[298,566,1944,1968]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="arificus" status="sp. nov.">
<emphasis id="47A9EAB5FF81FFCDFE9D58883826F803" box="[298,566,1944,1968]" italics="true" pageId="23" pageNumber="24">Tyrannochthonius arificus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FF81FFCDFD8858883881F803" bold="true" box="[575,657,1944,1968]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="5C9A57CEFF81FFCDFD8858883881F803" box="[575,657,1944,1968]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.10 mm.
</paragraph>
</caption>
<paragraph id="756236A7FF8EFFC2FF705F873F24FE1E" blockId="24.[151,1437,151,1113]" pageId="24" pageNumber="25">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 1.31–1.40. Pedipalps: trochanter 0.15/0.12–0.13 (1.15–1.25), femur 0.58–0.61/0.11 (5.27–5.55), patella 0.26–0.28/0.12–0.13 (2.15–2.17), chela 0.86–0.88/0.13–0.14 (6.29–6.62), hand 0.28–0.29/0.13–0.14 (2.07–2.15), movable chelal finger length 0.60–0.61. Chelicera 0.41–0.44/0.20 (2.05–2.20), movable finger length 0.23–0.24. Carapace 0.41–0.43/0.43–0.46 (0.93–0.95). Leg I: trochanter 0.12/0.10 (1.20), femur 0.36–0.37/0.06–0.07 (5.14– 6.17), patella 0.19–0.21/0.05 (3.80–4.20), tibia 0.18–0.19/0.04 (4.50–4.75), tarsus 0.40–0.45/0.03–0.04 (11.25– 13.33). Leg IV: trochanter 0.16–0.17/0.11–0.12 (1.33–1.55), femoropatella 0.52–0.54/0.17–0.19 (2.84–3.06), tibia 0.40–0.43/0.08–0.09 (4.78–5.00), metatarsus 0.19/0.06 (3.17), tarsus 0.44–0.47/0.03–0.04 (11.75–14.67).</paragraph>
<paragraph id="756236A7FF8EFFC2FF705EA73F8DFD1A" blockId="24.[151,1437,151,1113]" pageId="24" pageNumber="25">Females: body length 1.62–1.69. Pedipalps: trochanter 0.16–0.17/0.12–0.14 (1.21–1.33), femur 0.59–0.66/0.11– 0.12 (5.36–5.50), patella 0.30–0.32/0.13–0.15 (2.13–2.31), chela 0.91–0.93/0.15 (6.07–6.20), hand 0.31/0.15 (2.07), movable chelal finger length 0.62–0.63. Chelicera 0.45–0.47/0.21–0.23 (2.04–2.14), movable finger length 0.24–0.25. Carapace 0.45–0.48/0.46–0.49 (0.98). Leg I: trochanter 0.12–0.13/0.10–0.11 (1.18–1.20), femur 0.37– 0.39/0.06–0.07 (5.57–6.17), patella 0.19–0.22/0.06 (3.17–3.67), tibia 0.19–0.20/0.04–0.05 (4.00–4.75), tarsus 0.44– 0.45/0.04 (11.00–11.25). Leg IV: trochanter 0.18/0.11 (1.64), femoropatella 0.50–0.56/0.17–0.19 (2.94–2.95), tibia 0.39–0.45/0.08–0.09 (4.88–5.00), metatarsus 0.18–0.21/0.06 (3.00–3.50), tarsus 0.44–0.48/0.03–0.04 (12.00–14.67).</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFF8EFFC2FF705DA339E9FB86" pageId="24" pageNumber="25" type="discussion">
<paragraph id="756236A7FF8EFFC2FF705DA33EA7FCEE" blockId="24.[151,1437,151,1113]" pageId="24" pageNumber="25">
<emphasis id="47A9EAB5FF8EFFC2FF705DA33B2BFD7E" bold="true" box="[199,315,691,717]" pageId="24" pageNumber="25">Remarks.</emphasis>
<taxonomicName id="B2DD4D24FF8EFFC2FEF55DA33879FD7E" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[322,617,691,717]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="arificus" status="sp. nov.">
<emphasis id="47A9EAB5FF8EFFC2FEF55DA33879FD7E" box="[322,617,691,717]" italics="true" pageId="24" pageNumber="25">Tyrannochthonius arificus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FF8EFFC2FDC75DA338DAFD7E" bold="true" box="[624,714,691,717]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="5C9A57CEFF8EFFC2FDC75DA338DAFD7E" box="[624,714,691,717]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to 
<taxonomicName id="B2DD4D24FF8EFFC2FCEA5DA33E1EFD7E" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[861,1038,691,717]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="yanshanensis" status="sp. nov.">
<emphasis id="47A9EAB5FF8EFFC2FCEA5DA33E1EFD7E" box="[861,1038,691,717]" italics="true" pageId="24" pageNumber="25">T. yanshanensis</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FF8EFFC2FBA25DA33E7FFD7E" bold="true" box="[1045,1135,691,717]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="5C9A57CEFF8EFFC2FBA25DA33E7FFD7E" box="[1045,1135,691,717]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having intercalary teeth on both chelal fingers and a small, rounded and obtuse epistome, but differs by the larger body size (body length 1.62–1.69 (
<specimenCount id="63DBFD2EFF8EFFC2FEAE5DEB3B3BFCA6" box="[281,299,763,789]" pageId="24" pageNumber="25" type="female">♀</specimenCount>
) 
<emphasis id="47A9EAB5FF8EFFC2FE895DEB3B4CFCA7" box="[318,348,763,788]" italics="true" pageId="24" pageNumber="25">vs.</emphasis>
1.29–1.33 (
<specimenCount id="63DBFD2EFF8EFFC2FE515DEB3BE8FCA6" box="[486,504,763,789]" pageId="24" pageNumber="25" type="female">♀</specimenCount>
) mm; chela 6.07–6.20 (
<specimenCount id="63DBFD2EFF8EFFC2FCBD5DEB390CFCA6" box="[778,796,763,789]" pageId="24" pageNumber="25" type="female">♀</specimenCount>
) 
<emphasis id="47A9EAB5FF8EFFC2FC995DEB395DFCA7" box="[814,845,763,788]" italics="true" pageId="24" pageNumber="25">vs.</emphasis>
5.69–5.75 (
<specimenCount id="63DBFD2EFF8EFFC2FC615DEB39F8FCA6" box="[982,1000,763,789]" pageId="24" pageNumber="25" type="female">♀</specimenCount>
) times longer than board), the number of setae on tergite Ⅴ (4 
<emphasis id="47A9EAB5FF8EFFC2FE2B5C0F3BAAFC8B" box="[412,442,799,824]" italics="true" pageId="24" pageNumber="25">vs.</emphasis>
5) and tergite Ⅹ (3 
<emphasis id="47A9EAB5FF8EFFC2FD245C0F38A2FC8B" box="[659,690,799,824]" italics="true" pageId="24" pageNumber="25">vs.</emphasis>
5), more intercalary teeth on the fixed chelal finger (25–26 
<emphasis id="47A9EAB5FF8EFFC2FAFC5C0F3F7AFC8B" box="[1355,1386,799,824]" italics="true" pageId="24" pageNumber="25">vs.</emphasis>
14), the number of blades of coxal spines (10 
<emphasis id="47A9EAB5FF8EFFC2FDD25C533893FCEF" box="[613,643,835,860]" italics="true" pageId="24" pageNumber="25">vs.</emphasis>
7–8) and the number of blades of rallum (8 
<emphasis id="47A9EAB5FF8EFFC2FBC45C533E82FCEF" box="[1139,1170,835,860]" italics="true" pageId="24" pageNumber="25">vs.</emphasis>
7).
</paragraph>
<paragraph id="756236A7FF8EFFC2FF705C7738D6FC16" blockId="24.[151,1437,151,1113]" pageId="24" pageNumber="25">
<taxonomicName id="B2DD4D24FF8EFFC2FF705C773BFDFC32" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[199,493,871,897]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="arificus" status="sp. nov.">
<emphasis id="47A9EAB5FF8EFFC2FF705C773BFDFC32" box="[199,493,871,897]" italics="true" pageId="24" pageNumber="25">Tyrannochthonius arificus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FF8EFFC2FE455C77385AFC32" bold="true" box="[498,586,871,897]" pageId="24" pageNumber="25">
<taxonomicNameLabel id="5C9A57CEFF8EFFC2FE455C77385AFC32" box="[498,586,871,897]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from 
<taxonomicName id="B2DD4D24FF8EFFC2FCC35C7739F5FC32" authorityName="Mahnert" authorityYear="2009" box="[884,997,871,897]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="akaelus">
<emphasis id="47A9EAB5FF8EFFC2FCC35C7739F5FC32" box="[884,997,871,897]" italics="true" pageId="24" pageNumber="25">T. akaelus</emphasis>
</taxonomicName>
, 
<taxonomicName id="B2DD4D24FF8EFFC2FC465C773EA0FC32" authorityName="Mahnert" authorityYear="2009" box="[1009,1200,871,897]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="ganshuanensis">
<emphasis id="47A9EAB5FF8EFFC2FC465C773EA0FC32" box="[1009,1200,871,897]" italics="true" pageId="24" pageNumber="25">T. ganshuanensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="B2DD4D24FF8EFFC2FB535C773F42FC32" authorityName="Gao, Zhang &amp; Chen" authorityYear="2020" box="[1252,1362,871,897]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="harveyi">
<emphasis id="47A9EAB5FF8EFFC2FB535C773F42FC32" box="[1252,1362,871,897]" italics="true" pageId="24" pageNumber="25">T. harveyi</emphasis>
</taxonomicName>
by the number of setae on the tergites I–II (4 
<emphasis id="47A9EAB5FF8EFFC2FDF25C9B3873FC17" box="[581,611,907,932]" italics="true" pageId="24" pageNumber="25">vs.</emphasis>
2), from
</paragraph>
<paragraph id="756236A7FF8EFFC2FF705CBF39E9FB86" blockId="24.[151,1437,151,1113]" pageId="24" pageNumber="25">
<taxonomicName id="B2DD4D24FF8EFFC2FF705CBF3B23FC7A" authorityName="Gao, Wynne &amp; Fe.Zhang" authorityYear="2018" box="[199,307,943,969]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chixing">
<emphasis id="47A9EAB5FF8EFFC2FF705CBF3B23FC7A" box="[199,307,943,969]" italics="true" pageId="24" pageNumber="25">T. chixing</emphasis>
</taxonomicName>
and 
<taxonomicName id="B2DD4D24FF8EFFC2FEDF5CBF3BA1FC7A" authorityName="Gao, Zhang &amp; Chen" authorityYear="2020" box="[360,433,943,969]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="zhai">
<emphasis id="47A9EAB5FF8EFFC2FEDF5CBF3BA1FC7A" box="[360,433,943,969]" italics="true" pageId="24" pageNumber="25">T. zhai</emphasis>
</taxonomicName>
by the number of setae at the anterior margin of the carapace (6 
<emphasis id="47A9EAB5FF8EFFC2FBC35CBF3E82FC7B" box="[1140,1170,943,968]" italics="true" pageId="24" pageNumber="25">vs.</emphasis>
4), a shorter palp (chela length max. 0.93 
<emphasis id="47A9EAB5FF8EFFC2FEEF5CC33B66FC5F" box="[344,374,979,1004]" italics="true" pageId="24" pageNumber="25">vs.</emphasis>
min. 
<quantity id="B2259B42FF8EFFC2FE025CC3380BFC5E" box="[437,539,979,1005]" metricMagnitude="-3" metricUnit="m" metricValue="1.21" pageId="24" pageNumber="25" unit="mm" value="1.21">1.21 mm</quantity>
; palpal femur length max. 0.66 
<emphasis id="47A9EAB5FF8EFFC2FCCE5CC33988FC5F" box="[889,920,979,1004]" italics="true" pageId="24" pageNumber="25">vs.</emphasis>
min. 
<quantity id="B2259B42FF8EFFC2FC615CC33E2AFC5E" box="[982,1082,979,1005]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="24" pageNumber="25" unit="mm" value="0.84">0.84 mm</quantity>
) and the presence of intercalary teeth on both chelal fingers, from 
<taxonomicName id="B2DD4D24FF8EFFC2FDA75CE738D3FBA2" authorityName="Mahnert" authorityYear="2009" box="[528,707,1015,1041]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="antridraconis">
<emphasis id="47A9EAB5FF8EFFC2FDA75CE738D3FBA2" box="[528,707,1015,1041]" italics="true" pageId="24" pageNumber="25">T. antridraconis</emphasis>
</taxonomicName>
by a smaller size (body length max. 1.40 
<emphasis id="47A9EAB5FF8EFFC2FB255CE73EA1FBA3" box="[1170,1201,1015,1040]" italics="true" pageId="24" pageNumber="25">vs.</emphasis>
min. 
<quantity id="B2259B42FF8EFFC2FB475CE73F46FBA2" box="[1264,1366,1015,1041]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="24" pageNumber="25" unit="mm" value="1.8">1.80 mm</quantity>
; chela length max. 0.88 
<emphasis id="47A9EAB5FF8EFFC2FEED5B0B3B69FB87" box="[346,377,1051,1076]" italics="true" pageId="24" pageNumber="25">vs.</emphasis>
min. 
<quantity id="B2259B42FF8EFFC2FE0E5B0B380EFB86" box="[441,542,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="24" pageNumber="25" unit="mm" value="1.7">1.70 mm</quantity>
) (
<bibRefCitation id="114C4B56FF8EFFC2FD825B0B38F0FB86" author="Mahnert, V." box="[565,736,1051,1077]" pageId="24" pageNumber="25" pagination="185 - 201" refId="ref77141" refString="Mahnert, V. (2009) New species of pseudoscorpions (Arachnida, Pseudoscorpiones, Chthoniidae, Chernetidae) from caves in China. Revue Suisse de Zoologie, 116 (2), 185 - 201. https: // doi. org / 10.5962 / bhl. part. 79492" type="journal article" year="2009">Mahnert 2009</bibRefCitation>
; 
<bibRefCitation id="114C4B56FF8EFFC2FD5C5B0B39B1FB86" author="Gao, Z. Z. &amp; Wynne, J. J. &amp; Zhang, F." box="[747,929,1051,1077]" pageId="24" pageNumber="25" pagination="345 - 354" refId="ref76312" refString="Gao, Z. Z., Wynne, J. J. &amp; Zhang, F. (2018) Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones: Neobisiidae, Chthoniidae) from Guangxi, China. Journal of Arachnology, 46, 345 - 354. https: // doi. org / 10.1636 / JoA-S- 17 - 063.1" type="journal article" year="2018">
Gao 
<emphasis id="47A9EAB5FF8EFFC2FC945B0B394DFB86" box="[803,861,1051,1077]" italics="true" pageId="24" pageNumber="25">et al.</emphasis>
2018
</bibRefCitation>
, 
<bibRefCitation id="114C4B56FF8EFFC2FC1A5B0B39FAFB86" author="Gao, Z. Z. &amp; Zhang, F. &amp; Chen, H. M." box="[941,1002,1051,1077]" pageId="24" pageNumber="25" pagination="572 - 580" refId="ref76374" refString="Gao, Z. Z., Zhang, F. &amp; Chen, H. M. (2020) Two new cave-dwelling species of Tyrannochthonius Chamberlin 1929 (Pseudoscorpiones: Chthoniidae) from the Guizhou karst, China. Zootaxa, 4853 (4), 572 - 580. https: // doi. org / 10.11646 / zootaxa. 4853.4.6" type="journal article" year="2020">2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFF8EFFC2FF705B2E38E1FBEA" box="[199,753,1086,1113]" pageId="24" pageNumber="25" type="distribution">
<paragraph id="756236A7FF8EFFC2FF705B2E38E1FBEA" blockId="24.[151,1437,151,1113]" box="[199,753,1086,1113]" pageId="24" pageNumber="25">
<emphasis id="47A9EAB5FF8EFFC2FF705B2E3B71FBEB" bold="true" box="[199,353,1086,1112]" pageId="24" pageNumber="25">Distribution.</emphasis>
Known only from the 
<typeStatus id="AA668805FF8EFFC2FDD75B2F3880FBEA" box="[608,656,1087,1113]" pageId="24" pageNumber="25">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>