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<document id="5771DADEC8AB30090AC68D7BED1D909B" ID-CLB-Dataset="56235" ID-DOI="10.11646/zootaxa.5262.1.1" ID-GBIF-Dataset="e24a8032-296e-442d-80ae-e0bf8c7237f2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7795161" ID-ZooBank="F05805FD-BC5B-4236-BF04-C6AA5C37F2D7" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1680528957709" checkinUser="plazi" docAuthor="Hou, Yanmeng, Feng, Zegang &amp; Zhang, Feng" docDate="2023" docId="FD7487B1FF11FF53FF205F8838E1F8FE" docLanguage="en" docName="zootaxa.5262.1.1.pdf" docOrigin="Zootaxa 5262 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5262.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.16:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="16" docTitle="Tyrannochthonius quattuor Hou &amp; Feng &amp; Zhang 2023, sp. nov." docType="treatment" docVersion="8" lastPageNumber="138" masterDocId="014DFFC9FF96FFDAFFB75F103A10FFB3" masterDocTitle="Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae)" masterLastPageNumber="158" masterPageNumber="1" pageNumber="136" updateTime="1701997099389" updateUser="plazi">
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<mods:title id="CE38C7BF7C1D9DDAF1588A1EC386E03A">Diversity of cave-dwelling pseudoscorpions from Guizhou in China, with the description of twenty-four new species of the genus Tyrannochthonius (Pseudoscorpiones, Chthoniidae)</mods:title>
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<mods:name id="472BADC0FBFC9171625FAE7B49E2D6B2" type="personal">
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<mods:namePart id="7D8257EF4903A9B5D6E569399128F282">Hou, Yanmeng</mods:namePart>
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<mods:namePart id="A0C4BD91AC88FC4E4C0AAE1827C2947B">Feng, Zegang</mods:namePart>
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<mods:namePart id="DA63A8E5269EC944B3AEF3A702B2C247">Zhang, Feng</mods:namePart>
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<mods:title id="EB8E19007A25DA5EAB344EAF7041325F">Zootaxa</mods:title>
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<treatment id="FD7487B1FF11FF53FF205F8838E1F8FE" ID-DOI="http://doi.org/10.5281/zenodo.7795467" ID-GBIF-Taxon="209487191" ID-Zenodo-Dep="7795467" LSID="urn:lsid:plazi:treatment:FD7487B1FF11FF53FF205F8838E1F8FE" httpUri="http://treatment.plazi.org/id/FD7487B1FF11FF53FF205F8838E1F8FE" lastPageId="137" lastPageNumber="138" pageId="135" pageNumber="136">
<subSubSection id="3DC7652CFF11FF5DFF205F883937FF03" box="[151,807,152,179]" pageId="135" pageNumber="136" type="nomenclature">
<paragraph id="756236A7FF11FF5DFF205F883937FF03" blockId="135.[151,807,152,215]" box="[151,807,152,179]" pageId="135" pageNumber="136">
<heading id="2E2A81CBFF11FF5DFF205F883937FF03" bold="true" box="[151,807,152,179]" fontSize="11" level="2" pageId="135" pageNumber="136" reason="4">
<emphasis id="47A9EAB5FF11FF5DFF205F883937FF03" bold="true" box="[151,807,152,179]" pageId="135" pageNumber="136">
<taxonomicName id="B2DD4D24FF11FF5DFF205F883BF6FF00" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[151,486,152,179]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="quattuor" status="sp. nov.">
<emphasis id="47A9EAB5FF11FF5DFF205F883BF6FF00" bold="true" box="[151,486,152,179]" italics="true" pageId="135" pageNumber="136">Tyrannochthonius quattuor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5C9A57CEFF11FF5DFE595F88385CFF01" box="[494,588,152,178]" pageId="135" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
‹四ƭƟṁae›
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFF11FF5DFF205FAC3B3DFF64" box="[151,301,188,215]" pageId="135" pageNumber="136" type="description">
<paragraph id="756236A7FF11FF5DFF205FAC3B3DFF64" blockId="135.[151,807,152,215]" box="[151,301,188,215]" pageId="135" pageNumber="136">
<figureCitation id="EDE62A22FF11FF5DFF205FAC3B3DFF64" box="[151,301,188,215]" captionStart-0="FIGURE 111" captionStart-1="FIGURE 112" captionStart-2="FIGURE 113" captionStart-3="FIGURE 114" captionStart-4="FIGURE 115" captionStartId-0="135.[151,250,1488,1512]" captionStartId-1="136.[151,250,1222,1246]" captionStartId-2="138.[151,250,1943,1967]" captionStartId-3="139.[151,250,1943,1967]" captionStartId-4="140.[151,250,1943,1967]" captionTargetBox-0="[151,1436,526,1464]" captionTargetBox-1="[172,1414,353,1197]" captionTargetBox-2="[179,1408,181,1920]" captionTargetBox-3="[179,1383,181,1919]" captionTargetBox-4="[179,1408,181,1919]" captionTargetId-0="figure-114@135.[151,1436,526,1464]" captionTargetId-1="figure-120@136.[161,1427,345,1199]" captionTargetId-2="figure-20@138.[179,1408,181,1920]" captionTargetId-3="figure-18@139.[179,1408,181,1919]" captionTargetId-4="figure-20@140.[179,1408,181,1919]" captionTargetPageId-0="135" captionTargetPageId-1="136" captionTargetPageId-2="138" captionTargetPageId-3="139" captionTargetPageId-4="140" captionText-0="FIGURE 111. Guanjiadadong Cave, type locality of Tyrannochthonius quattuor sp. nov., A. Cave location (red arrow); B–C. Inside the cave entrance; D. Area where T. quattuor sp. nov. specimens were collected." captionText-1="FIGURE 112. Tyrannochthonius quattuor sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." captionText-2="FIGURE 113. Tyrannochthonius quattuor sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." captionText-3="FIGURE 114. Tyrannochthonius quattuor sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." captionText-4="FIGURE 115. Tyrannochthonius quattuor sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7795421" figureDoi-1="http://doi.org/10.5281/zenodo.7795423" figureDoi-2="http://doi.org/10.5281/zenodo.7795427" figureDoi-3="http://doi.org/10.5281/zenodo.7795431" figureDoi-4="http://doi.org/10.5281/zenodo.7795433" httpUri-0="https://zenodo.org/record/7795421/files/figure.png" httpUri-1="https://zenodo.org/record/7795423/files/figure.png" httpUri-2="https://zenodo.org/record/7795427/files/figure.png" httpUri-3="https://zenodo.org/record/7795431/files/figure.png" httpUri-4="https://zenodo.org/record/7795433/files/figure.png" pageId="135" pageNumber="136">Figs 111–115</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFF11FF5DFF205E143F37FE1D" pageId="135" pageNumber="136" type="materials_examined">
<paragraph id="756236A7FF11FF5DFF205E143F37FE1D" blockId="135.[151,1437,260,503]" pageId="135" pageNumber="136">
<emphasis id="47A9EAB5FF11FF5DFF205E143B5AFEAD" bold="true" box="[151,330,260,286]" pageId="135" pageNumber="136">Type material.</emphasis>
<materialsCitation id="C5B53CFAFF11FF5DFEE15E143BA6FE39" ID-GBIF-Occurrence="4071438343" collectingDate="2022-07-28" collectionCode="MHBU-HBUARA" collectorName="Yanmeng Hou &amp; Lu Zhang &amp; Jianzhou Sun &amp; Wenlong Fan" country="China" county="Dafang County" elevation="75" latitude="27.078781" location="Guanjiadadong Cave" longLatPrecision="1" longitude="105.6528" municipality="Yangchang Town" pageId="135" pageNumber="136" specimenCode="MHBU-HBUARA#2022-484, 01" specimenCount="1" stateProvince="Guizhou" typeStatus="holotype">
<typeStatus id="AA668805FF11FF5DFEE15E143BAEFEAD" box="[342,446,260,286]" pageId="135" pageNumber="136" type="holotype">Holotype</typeStatus>
♂ (Ps.-
<specimenCode id="257B9EDCFF11FF5DFDAF5E143965FEAD" box="[536,885,260,286]" collectionCode="MHBU-HBUARA" pageId="135" pageNumber="136">MHBU-HBUARA#2022-484</specimenCode>
-
<specimenCode id="257B9EDCFF11FF5DFCCA5E143987FEAD" box="[893,919,260,286]" collectionCode="MHBU-HBUARA" pageId="135" pageNumber="136">01</specimenCode>
):
<collectingCountry id="0DCA7637FF11FF5DFC195E1439E4FEAD" box="[942,1012,260,286]" name="China" pageId="135" pageNumber="136">China</collectingCountry>
,
<collectingRegion id="B719F845FF11FF5DFBB35E143EC6FEAD" box="[1028,1238,260,286]" country="China" name="Guizhou" pageId="135" pageNumber="136">Guizhou Province</collectingRegion>
,
<collectingCounty id="9C034E2BFF11FF5DFB515E143F89FEAD" box="[1254,1433,260,286]" pageId="135" pageNumber="136">Dafang County</collectingCounty>
,
<collectingMunicipality id="9506ACDDFF11FF5DFF205E383B4CFEF0" box="[151,348,296,323]" pageId="135" pageNumber="136">Yangchang Town</collectingMunicipality>
,
<location id="7002607CFF11FF5DFED15E383843FEF1" LSID="urn:lsid:plazi:treatment:FD7487B1FF11FF53FF205F8838E1F8FE:7002607CFF11FF5DFED15E383843FEF1" box="[358,595,296,322]" country="China" county="Dafang County" latitude="27.078781" longLatPrecision="1" longitude="105.6528" municipality="Yangchang Town" name="Guanjiadadong Cave" pageId="135" pageNumber="136" stateProvince="Guizhou">Guanjiadadong Cave</location>
, under stones within 50–100 m from the cave entrance (Temperature: 17°C, Humidity: 85%) [
<geoCoordinate id="10E95060FF11FF5DFEEB5E5E3BECFEDA" box="[348,508,334,361]" degrees="27" direction="north" minutes="4" orientation="latitude" pageId="135" pageNumber="136" precision="1" seconds="43.61" value="27.078781">27°4′43.61″N</geoCoordinate>
,
<geoCoordinate id="10E95060FF11FF5DFDB05E5E38D0FEDA" box="[519,704,334,361]" degrees="105" direction="east" minutes="39" orientation="longitude" pageId="135" pageNumber="136" precision="1" seconds="10.09" value="105.6528">105°39′10.09″E</geoCoordinate>
],
<elevation id="FEF0D194FF11FF5DFD665E5C3973FED5" box="[721,867,332,359]" metricMagnitude="3" metricUnit="m" metricValue="1.555" pageId="135" pageNumber="136" unit="m" value="1555.0">
<quantity id="B2259B42FF11FF5DFD665E5C3936FED4" box="[721,806,332,359]" metricMagnitude="3" metricUnit="m" metricValue="1.555" pageId="135" pageNumber="136" unit="m" value="1555.0">1555 m</quantity>
a.s.l.
</elevation>
,
<date id="01631067FF11FF5DFCD85E5C3E13FED5" box="[879,1027,332,358]" pageId="135" pageNumber="136" value="2022-07-28">
<collectingDate id="1127E98FFF11FF5DFCD85E5C3E13FED5" box="[879,1027,332,358]" pageId="135" pageNumber="136" value="2022-07-28">28 July 2022</collectingDate>
</date>
,
<collectorName id="D8285371FF11FF5DFBBA5E5D3EA1FED4" box="[1037,1201,333,359]" pageId="135" pageNumber="136">Yanmeng Hou</collectorName>
,
<collectorName id="D8285371FF11FF5DFB0A5E5D3F3DFED5" box="[1213,1325,332,359]" pageId="135" pageNumber="136">Lu Zhang</collectorName>
,
<collectorName id="D8285371FF11FF5DFA8E5E5C3AD3FE39" pageId="135" pageNumber="136">Jianzhou Sun</collectorName>
&amp;
<collectorName id="D8285371FF11FF5DFF5C5E603B93FE38" box="[235,387,368,395]" pageId="135" pageNumber="136">Wenlong Fan</collectorName>
leg.
</materialsCitation>
<materialsCitation id="C5B53CFAFF11FF5DFE085E613F34FE1D" ID-GBIF-Occurrence="4071438339" collectingDate="2022-07-28" collectionCode="MSWU-HBUARA, MHBU-HBUARA" collectorName="Yanmeng Hou &amp; Lu Zhang &amp; Jianzhou Sun &amp; Wenlong Fan" country="China" county="Dafang County" elevation="75" latitude="27.078781" location="Guanjiadadong Cave" longLatPrecision="1" longitude="105.6528" municipality="Yangchang Town" pageId="135" pageNumber="136" specimenCode="MHBU-HBUARA#2022-484, 02, MSWU-HBUARA#2022-484, 04" specimenCount="9" specimenCount-female="9" stateProvince="Guizhou" typeStatus="paratype">
<typeStatus id="AA668805FF11FF5DFE085E613820FE38" box="[447,560,369,395]" pageId="135" pageNumber="136" type="paratype">Paratypes</typeStatus>
: 2 ♂ (Ps.-
<specimenCode id="257B9EDCFF11FF5DFD285E6039E5FE39" box="[671,1013,368,394]" collectionCode="MHBU-HBUARA" pageId="135" pageNumber="136">MHBU-HBUARA#2022-484</specimenCode>
-
<specimenCode id="257B9EDCFF11FF5DFC4A5E603E09FE39" box="[1021,1049,368,394]" collectionCode="MHBU-HBUARA" pageId="135" pageNumber="136">02</specimenCode>
&amp; HBUARA#2022-484-03),
<specimenCount id="63DBFD2EFF11FF5DFAC75E603F8CFE38" box="[1392,1436,368,395]" pageId="135" pageNumber="136" type="female">9 ♀</specimenCount>
(Ps.-
<specimenCode id="257B9EDCFF11FF5DFF735E84380FFE1D" box="[196,543,404,430]" collectionCode="MSWU-HBUARA" pageId="135" pageNumber="136">MSWU-HBUARA#2022-484</specimenCode>
-
<specimenCode id="257B9EDCFF11FF5DFD915E843852FE1D" box="[550,578,404,430]" collectionCode="MSWU-HBUARA" pageId="135" pageNumber="136">04</specimenCode>
–HBUARA#2022-484-12), all with the same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFF11FF5DFF705EA838A6FE44" pageId="135" pageNumber="136" type="etymology">
<paragraph id="756236A7FF11FF5DFF705EA838A6FE44" blockId="135.[151,1437,260,503]" pageId="135" pageNumber="136">
<emphasis id="47A9EAB5FF11FF5DFF705EA83B5EFE61" bold="true" box="[199,334,440,466]" pageId="135" pageNumber="136">Etymology.</emphasis>
The specific name is derived from the Latin word “
<taxonomicName id="B2DD4D24FF11FF5DFC145EA93E13FE61" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[931,1027,441,466]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="quattuor">
<emphasis id="47A9EAB5FF11FF5DFC145EA93E13FE61" box="[931,1027,441,466]" italics="true" pageId="135" pageNumber="136">quattuor</emphasis>
</taxonomicName>
”, meaning four, which refers to the tergites (except tergite XII) each with four setae.
</paragraph>
</subSubSection>
<caption id="21A2662FFF11FF5DFF205AC03E1AF9BF" ID-DOI="http://doi.org/10.5281/zenodo.7795421" ID-Zenodo-Dep="7795421" httpUri="https://zenodo.org/record/7795421/files/figure.png" pageId="135" pageNumber="136" startId="135.[151,250,1488,1512]" targetBox="[151,1436,526,1464]" targetPageId="135">
<paragraph id="756236A7FF11FF5DFF205AC03E1AF9BF" blockId="135.[151,1437,1488,1548]" pageId="135" pageNumber="136">
<emphasis id="47A9EAB5FF11FF5DFF205AC03B3BFA5B" bold="true" box="[151,299,1488,1512]" pageId="135" pageNumber="136">FIGURE 111.</emphasis>
Guanjiadadong Cave, type locality of
<taxonomicName id="B2DD4D24FF11FF5DFD0F5AC039DCFA5B" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[696,972,1488,1512]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="quattuor" status="sp. nov.">
<emphasis id="47A9EAB5FF11FF5DFD0F5AC039DCFA5B" box="[696,972,1488,1512]" italics="true" pageId="135" pageNumber="136">Tyrannochthonius quattuor</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FF11FF5DFC635AC03E35FA5B" bold="true" box="[980,1061,1488,1512]" pageId="135" pageNumber="136">
<taxonomicNameLabel id="5C9A57CEFF11FF5DFC635AC03E35FA5B" box="[980,1061,1488,1512]" pageId="135" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Cave location (red arrow); B–C. Inside the cave entrance; D. Area where
<taxonomicName id="B2DD4D24FF11FF5DFD855AE538B2F9BF" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[562,674,1525,1548]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="135" pageNumber="136" phylum="Arthropoda" rank="species" species="quattuor" status="sp. nov.">
<emphasis id="47A9EAB5FF11FF5DFD855AE538B2F9BF" box="[562,674,1525,1548]" italics="true" pageId="135" pageNumber="136">T. quattuor</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FF11FF5DFD1F5AE438E9F9BF" bold="true" box="[680,761,1524,1548]" pageId="135" pageNumber="136">
<taxonomicNameLabel id="5C9A57CEFF11FF5DFD1F5AE438E9F9BF" box="[680,761,1524,1548]" pageId="135" pageNumber="136" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
specimens were collected.
</paragraph>
</caption>
<subSubSection id="3DC7652CFF11FF5DFF70592A3BC1F8BA" pageId="135" pageNumber="136" type="diagnosis">
<paragraph id="756236A7FF11FF5DFF70592A3BC1F8BA" blockId="135.[151,1437,1594,2017]" pageId="135" pageNumber="136">
<emphasis id="47A9EAB5FF11FF5DFF70592A3B97F9E4" bold="true" box="[199,391,1594,1623]" pageId="135" pageNumber="136">
Diagnosis (J
<specimenCount id="63DBFD2EFF11FF5DFED7592B3B66F9E4" box="[352,374,1595,1623]" pageId="135" pageNumber="136" type="female">♀</specimenCount>
).
</emphasis>
Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I–XI each with 4 setae (except the females). Pedipalps slender, femur 7.80–8.40 (♂), 7.33–7.75 (
<specimenCount id="63DBFD2EFF11FF5DFE7359B63BC6F973" box="[452,470,1702,1728]" pageId="135" pageNumber="136" type="female">♀</specimenCount>
) times longer than broad; chela 7.91–8.25 (♂), 6.89–7.25 (
<specimenCount id="63DBFD2EFF11FF5DFBDD59B63E6CF973" box="[1130,1148,1702,1728]" pageId="135" pageNumber="136" type="female">♀</specimenCount>
) times longer than broad; both chelal fingers without intercalary teeth, movable chelal finger teeth markedly smaller than fixed chelal finger teeth and distinctly retrorse.
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFF11FF53FF70580239A5FA2E" lastPageId="137" lastPageNumber="138" pageId="135" pageNumber="136" type="description">
<paragraph id="756236A7FF11FF5DFF7058023E3BF89E" blockId="135.[151,1437,1594,2017]" box="[199,1067,1810,1837]" pageId="135" pageNumber="136">
<emphasis id="47A9EAB5FF11FF5DFF7058023B49F89F" bold="true" box="[199,345,1810,1836]" pageId="135" pageNumber="136">Description.</emphasis>
Males (
<typeStatus id="AA668805FF11FF5DFE0458023807F89F" box="[435,535,1810,1836]" pageId="135" pageNumber="136" type="holotype">holotype</typeStatus>
and
<typeStatus id="AA668805FF11FF5DFDFA580238AAF89F" box="[589,698,1810,1836]" pageId="135" pageNumber="136" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="EDE62A22FF11FF5DFD6758023958F89F" box="[720,840,1810,1836]" captionStart="FIGURE 112" captionStartId="136.[151,250,1222,1246]" captionTargetBox="[172,1414,353,1197]" captionTargetId="figure-120@136.[161,1427,345,1199]" captionTargetPageId="136" captionText="FIGURE 112. Tyrannochthonius quattuor sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795423" httpUri="https://zenodo.org/record/7795423/files/figure.png" pageId="135" pageNumber="136">Figs 112A</figureCitation>
,
<figureCitation id="EDE62A22FF11FF5DFCE3580239BFF89F" box="[852,943,1810,1836]" captionStart="FIGURE 113" captionStartId="138.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-20@138.[179,1408,181,1920]" captionTargetPageId="138" captionText="FIGURE 113. Tyrannochthonius quattuor sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795427" httpUri="https://zenodo.org/record/7795427/files/figure.png" pageId="135" pageNumber="136">113A–F</figureCitation>
,
<figureCitation id="EDE62A22FF11FF5DFC0C580239F7F89F" box="[955,999,1810,1836]" captionStart="FIGURE 114" captionStartId="139.[151,250,1943,1967]" captionTargetBox="[179,1383,181,1919]" captionTargetId="figure-18@139.[179,1408,181,1919]" captionTargetPageId="139" captionText="FIGURE 114. Tyrannochthonius quattuor sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795431" httpUri="https://zenodo.org/record/7795431/files/figure.png" pageId="135" pageNumber="136">114</figureCitation>
,
<figureCitation id="EDE62A22FF11FF5DFC4558023E0FF89E" box="[1010,1055,1810,1837]" captionStart="FIGURE 115" captionStartId="140.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@140.[179,1408,181,1919]" captionTargetPageId="140" captionText="FIGURE 115. Tyrannochthonius quattuor sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795433" httpUri="https://zenodo.org/record/7795433/files/figure.png" pageId="135" pageNumber="136">115</figureCitation>
).
</paragraph>
<paragraph id="756236A7FF11FF5DFF7058263EFFF8E3" blockId="135.[151,1437,1594,2017]" box="[199,1263,1846,1873]" pageId="135" pageNumber="136">Colour: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.</paragraph>
<paragraph id="756236A7FF11FF52FF70584A3F67FEF2" blockId="135.[151,1437,1594,2017]" lastBlockId="136.[151,1436,151,321]" lastPageId="136" lastPageNumber="137" pageId="135" pageNumber="136">
Cephalothorax (
<figureCitation id="EDE62A22FF11FF5DFECB584A3BE6F8C7" box="[380,502,1882,1908]" captionStart="FIGURE 113" captionStartId="138.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-20@138.[179,1408,181,1920]" captionTargetPageId="138" captionText="FIGURE 113. Tyrannochthonius quattuor sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795427" httpUri="https://zenodo.org/record/7795427/files/figure.png" pageId="135" pageNumber="136">Figs 113C</figureCitation>
,
<figureCitation id="EDE62A22FF11FF5DFDB5584A3854F8C7" box="[514,580,1882,1908]" captionStart="FIGURE 114" captionStartId="139.[151,250,1943,1967]" captionTargetBox="[179,1383,181,1919]" captionTargetId="figure-18@139.[179,1408,181,1919]" captionTargetPageId="139" captionText="FIGURE 114. Tyrannochthonius quattuor sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795431" httpUri="https://zenodo.org/record/7795431/files/figure.png" pageId="135" pageNumber="136">114A</figureCitation>
): carapace 1.02–1.04 times longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome small and pointed, triangular, with 2 setae flanking base; with 18 setae arranged s4–5s: 4–5: 4: 2: 2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with two pairs of lyrifissures, first pair situated middle to the setae of ocular row, the second situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 13–15 terminally indented coxal spines on each side, set as an oblique row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for about half their length (
<figureCitation id="EDE62A22FF1EFF52FD795E373952FEF2" box="[718,834,295,321]" captionStart="FIGURE 114" captionStartId="139.[151,250,1943,1967]" captionTargetBox="[179,1383,181,1919]" captionTargetId="figure-18@139.[179,1408,181,1919]" captionTargetPageId="139" captionText="FIGURE 114. Tyrannochthonius quattuor sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795431" httpUri="https://zenodo.org/record/7795431/files/figure.png" pageId="136" pageNumber="137">Fig. 114C</figureCitation>
); intercoxal tubercle absent; without sub-oral seta.
</paragraph>
<caption id="21A2662FFF1EFF52FF205BD63AEAFAB0" ID-DOI="http://doi.org/10.5281/zenodo.7795423" ID-Zenodo-Dep="7795423" httpUri="https://zenodo.org/record/7795423/files/figure.png" pageId="136" pageNumber="137" startId="136.[151,250,1222,1246]" targetBox="[172,1414,353,1197]" targetPageId="136">
<paragraph id="756236A7FF1EFF52FF205BD63AEAFAB0" blockId="136.[151,1436,1222,1283]" pageId="136" pageNumber="137">
<emphasis id="47A9EAB5FF1EFF52FF205BD63B3BFB6C" bold="true" box="[151,299,1222,1247]" pageId="136" pageNumber="137">FIGURE 112.</emphasis>
<taxonomicName id="B2DD4D24FF1EFF52FE865BD73855FB6D" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[305,581,1223,1247]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="quattuor" status="sp. nov.">
<emphasis id="47A9EAB5FF1EFF52FE865BD73855FB6D" box="[305,581,1223,1247]" italics="true" pageId="136" pageNumber="137">Tyrannochthonius quattuor</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FF1EFF52FDFB5BD7388CFB6C" bold="true" box="[588,668,1223,1247]" pageId="136" pageNumber="137">
<taxonomicNameLabel id="5C9A57CEFF1EFF52FDFB5BD7388CFB6C" box="[588,668,1223,1247]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view).
</paragraph>
</caption>
<paragraph id="756236A7FF1EFF52FF705A213866F9F4" blockId="136.[151,1437,1329,2039]" pageId="136" pageNumber="137">
Chelicera (
<figureCitation id="EDE62A22FF1EFF52FEF65A213BA8FAF8" box="[321,440,1329,1355]" captionStart="FIGURE 113" captionStartId="138.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-20@138.[179,1408,181,1920]" captionTargetPageId="138" captionText="FIGURE 113. Tyrannochthonius quattuor sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795427" httpUri="https://zenodo.org/record/7795427/files/figure.png" pageId="136" pageNumber="137">Figs 113D</figureCitation>
,
<figureCitation id="EDE62A22FF1EFF52FE745A213813FAF8" box="[451,515,1329,1355]" captionStart="FIGURE 114" captionStartId="139.[151,250,1943,1967]" captionTargetBox="[179,1383,181,1919]" captionTargetId="figure-18@139.[179,1408,181,1919]" captionTargetPageId="139" captionText="FIGURE 114. Tyrannochthonius quattuor sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795431" httpUri="https://zenodo.org/record/7795431/files/figure.png" pageId="136" pageNumber="137">114B</figureCitation>
): large, about as long as carapace, 2.36–2.48 times longer than broad; 5 setae and 2 lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 18–21 teeth, distal one largest; movable finger with 17–18 retrorse contiguous small teeth; galea completely vestigial (
<figureCitation id="EDE62A22FF1EFF52FBD05AD13ECBFA68" box="[1127,1243,1473,1499]" captionStart="FIGURE 114" captionStartId="139.[151,250,1943,1967]" captionTargetBox="[179,1383,181,1919]" captionTargetId="figure-18@139.[179,1408,181,1919]" captionTargetPageId="139" captionText="FIGURE 114. Tyrannochthonius quattuor sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795431" httpUri="https://zenodo.org/record/7795431/files/figure.png" pageId="136" pageNumber="137">Fig. 114B</figureCitation>
). Serrula exterior with 22–24 and serrula interior with 14–15 blades. Rallum with 8 blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (
<figureCitation id="EDE62A22FF1EFF52FE41593D387AF9F4" box="[502,618,1581,1607]" captionStart="FIGURE 114" captionStartId="139.[151,250,1943,1967]" captionTargetBox="[179,1383,181,1919]" captionTargetId="figure-18@139.[179,1408,181,1919]" captionTargetPageId="139" captionText="FIGURE 114. Tyrannochthonius quattuor sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795431" httpUri="https://zenodo.org/record/7795431/files/figure.png" pageId="136" pageNumber="137">Fig. 114E</figureCitation>
).
</paragraph>
<paragraph id="756236A7FF1EFF53FF7059413B3DFEAE" blockId="136.[151,1437,1329,2039]" lastBlockId="137.[151,1437,150,1869]" lastPageId="137" lastPageNumber="138" pageId="136" pageNumber="137">
Pedipalp (
<figureCitation id="EDE62A22FF1EFF52FE8E59413BE4F9D8" box="[313,500,1617,1643]" captionStart="FIGURE 113" captionStartId="138.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-20@138.[179,1408,181,1920]" captionTargetPageId="138" captionText="FIGURE 113. Tyrannochthonius quattuor sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795427" httpUri="https://zenodo.org/record/7795427/files/figure.png" pageId="136" pageNumber="137">Figs 113A, B, E</figureCitation>
,
<figureCitation id="EDE62A22FF1EFF52FDB659413851F9D8" box="[513,577,1617,1643]" captionStart="FIGURE 114" captionStartId="139.[151,250,1943,1967]" captionTargetBox="[179,1383,181,1919]" captionTargetId="figure-18@139.[179,1408,181,1919]" captionTargetPageId="139" captionText="FIGURE 114. Tyrannochthonius quattuor sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795431" httpUri="https://zenodo.org/record/7795431/files/figure.png" pageId="136" pageNumber="137">114D</figureCitation>
,
<figureCitation id="EDE62A22FF1EFF52FDFA594138A0F9D8" box="[589,688,1617,1644]" captionStart="FIGURE 115" captionStartId="140.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@140.[179,1408,181,1919]" captionTargetPageId="140" captionText="FIGURE 115. Tyrannochthonius quattuor sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795433" httpUri="https://zenodo.org/record/7795433/files/figure.png" pageId="136" pageNumber="137">115A, B</figureCitation>
): long and slender, trochanter 1.18–1.28, femur 7.80–8.40, patella 2.44–2.50, chela 7.91–8.25, hand 2.91–2.95 times longer than broad; femur 2.66–2.80 times longer than patella; movable chelal finger 1.64–1.69 times longer than hand and 0.60–0.61 times longer than chela. Setae generally long and acuminate; 1 distal lyrifissure present on patella (
<figureCitation id="EDE62A22FF1EFF52FC9A59AD39B4F964" box="[813,932,1725,1751]" captionStart="FIGURE 113" captionStartId="138.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-20@138.[179,1408,181,1920]" captionTargetPageId="138" captionText="FIGURE 113. Tyrannochthonius quattuor sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795427" httpUri="https://zenodo.org/record/7795427/files/figure.png" pageId="136" pageNumber="137">Figs 113E</figureCitation>
,
<figureCitation id="EDE62A22FF1EFF52FC0759AD39E2F964" box="[944,1010,1725,1751]" captionStart="FIGURE 114" captionStartId="139.[151,250,1943,1967]" captionTargetBox="[179,1383,181,1919]" captionTargetId="figure-18@139.[179,1408,181,1919]" captionTargetPageId="139" captionText="FIGURE 114. Tyrannochthonius quattuor sp. nov., holotype male: A. Carapace (dorsal view); B. Left chelicera (dorsal view), with details of teeth; C. Coxal spines on coxae II (ventral view); D. Left pedipalp (minus chela, dorsal view); E. Rallum. Scale bars: 0.20 mm (A–B, D); 0.10 mm (C, E)." figureDoi="http://doi.org/10.5281/zenodo.7795431" httpUri="https://zenodo.org/record/7795431/files/figure.png" pageId="136" pageNumber="137">114D</figureCitation>
). Chelal palm not constricted towards fingers, apodeme complex of movable chelal finger only slightly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with 8 trichobothria, movable chelal finger with 4 trichobothria,
<emphasis id="47A9EAB5FF1EFF52FF2058393ABDF8F0" box="[151,173,1833,1859]" italics="true" pageId="136" pageNumber="137">ib</emphasis>
and
<emphasis id="47A9EAB5FF1EFF52FF5258393B16F8F0" box="[229,262,1833,1859]" italics="true" pageId="136" pageNumber="137">isb</emphasis>
situated close together, submedially on dorsum of chelal hand;
<emphasis id="47A9EAB5FF1EFF52FC6D583939E4F8F0" box="[986,1012,1833,1859]" italics="true" pageId="136" pageNumber="137">eb</emphasis>
,
<emphasis id="47A9EAB5FF1EFF52FBB458393E38F8F0" box="[1027,1064,1833,1859]" italics="true" pageId="136" pageNumber="137">esb</emphasis>
and
<emphasis id="47A9EAB5FF1EFF52FBD6583A3E6BF8F0" box="[1121,1147,1834,1859]" italics="true" pageId="136" pageNumber="137">ist</emphasis>
forming a nearly straight oblique row at base of fixed chelal finger;
<emphasis id="47A9EAB5FF1EFF52FDC5585E3892F8D4" box="[626,642,1870,1895]" italics="true" pageId="136" pageNumber="137">it</emphasis>
slightly distal to
<emphasis id="47A9EAB5FF1EFF52FCF4585E3972F8D4" box="[835,866,1870,1895]" italics="true" pageId="136" pageNumber="137">est</emphasis>
, situated subdistally;
<emphasis id="47A9EAB5FF1EFF52FBE1585E3E7AF8D4" box="[1110,1130,1870,1895]" italics="true" pageId="136" pageNumber="137">et</emphasis>
slightly near to tip of fixed chelal finger, very close to chelal teeth;
<emphasis id="47A9EAB5FF1EFF52FDE058613861F838" box="[599,625,1905,1931]" italics="true" pageId="136" pageNumber="137">dx</emphasis>
situated distal to
<emphasis id="47A9EAB5FF1EFF52FC805862395BF838" box="[823,843,1906,1931]" italics="true" pageId="136" pageNumber="137">et</emphasis>
;
<emphasis id="47A9EAB5FF1EFF52FCED58613963F838" box="[858,883,1905,1931]" italics="true" pageId="136" pageNumber="137">sb</emphasis>
closer to
<emphasis id="47A9EAB5FF1EFF52FC56586239E4F838" box="[993,1012,1906,1931]" italics="true" pageId="136" pageNumber="137">st</emphasis>
than to
<emphasis id="47A9EAB5FF1EFF52FBF858613E4DF838" box="[1103,1117,1905,1931]" italics="true" pageId="136" pageNumber="137">b</emphasis>
;
<emphasis id="47A9EAB5FF1EFF52FBDB58613E6AF838" box="[1132,1146,1905,1931]" italics="true" pageId="136" pageNumber="137">b</emphasis>
and
<emphasis id="47A9EAB5FF1EFF52FB0658623EA9F838" box="[1201,1209,1906,1931]" italics="true" pageId="136" pageNumber="137">t</emphasis>
situated subdistally,
<emphasis id="47A9EAB5FF1EFF52FF2058863A8FF81C" box="[151,159,1942,1967]" italics="true" pageId="136" pageNumber="137">t</emphasis>
situated distal to
<emphasis id="47A9EAB5FF1EFF52FED058853B65F81C" box="[359,373,1941,1967]" italics="true" pageId="136" pageNumber="137">b</emphasis>
and proximal to
<emphasis id="47A9EAB5FF1EFF52FD8158863845F81C" box="[566,597,1942,1967]" italics="true" pageId="136" pageNumber="137">est</emphasis>
(
<figureCitation id="EDE62A22FF1EFF52FDD2588538CBF803" box="[613,731,1941,1968]" captionStart="FIGURE 115" captionStartId="140.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@140.[179,1408,181,1919]" captionTargetPageId="140" captionText="FIGURE 115. Tyrannochthonius quattuor sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795433" httpUri="https://zenodo.org/record/7795433/files/figure.png" pageId="136" pageNumber="137">Fig. 115A</figureCitation>
). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to
<emphasis id="47A9EAB5FF1EFF52FE3D58AA3BB4F860" box="[394,420,1978,2003]" italics="true" pageId="136" pageNumber="137">ist</emphasis>
). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 31–35 macrodenticles, slightly retrorse, long and pointed; movable chelal finger with 49–51 macrodenticles (markedly smaller than teeth on fixed chelal finger), continuous and markedly retrorse (
<figureCitation id="EDE62A22FF1FFF53FAAC5FAB3F80FF66" box="[1307,1424,186,213]" captionStart="FIGURE 115" captionStartId="140.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@140.[179,1408,181,1919]" captionTargetPageId="140" captionText="FIGURE 115. Tyrannochthonius quattuor sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795433" httpUri="https://zenodo.org/record/7795433/files/figure.png" pageId="137" pageNumber="138">Fig. 115A</figureCitation>
). Chelal fingers straight in dorsal view; microsetae (chemosensory setae) present on dorsum of chelal hand (
<figureCitation id="EDE62A22FF1FFF53FADD5FCF3AC5FEAE" captionStart="FIGURE 113" captionStartId="138.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-20@138.[179,1408,181,1920]" captionTargetPageId="138" captionText="FIGURE 113. Tyrannochthonius quattuor sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795427" httpUri="https://zenodo.org/record/7795427/files/figure.png" pageId="137" pageNumber="138">Figs 113B</figureCitation>
,
<figureCitation id="EDE62A22FF1FFF53FF565E123B30FEAE" box="[225,288,258,285]" captionStart="FIGURE 115" captionStartId="140.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@140.[179,1408,181,1919]" captionTargetPageId="140" captionText="FIGURE 115. Tyrannochthonius quattuor sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795433" httpUri="https://zenodo.org/record/7795433/files/figure.png" pageId="137" pageNumber="138">115B</figureCitation>
).
</paragraph>
<paragraph id="756236A7FF1FFF53FF705E373997FE1E" blockId="137.[151,1437,150,1869]" pageId="137" pageNumber="138">
Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 4: 4: 4: 4: T2T: 0. Sternal chaetotaxy III–XII: 14–15: 13: 9–10: 8–9: 7–9: 8–9: 9: 9: 0: 2. Anterior genital operculum with 9–11 setae, genital opening slit-like, with 14–16 marginal setae on each side,
<quantity id="B2259B42FF1FFF53FDE95E8338D1FE1E" box="[606,705,403,429]" metricMagnitude="0" metricUnit="m" metricValue="1.016" metricValueMax="1.0668" metricValueMin="0.9652" pageId="137" pageNumber="138" unit="in" value="40.0" valueMax="42.0" valueMin="38.0">38–42 in</quantity>
total (
<figureCitation id="EDE62A22FF1FFF53FCBE5E83396AFE1E" box="[777,890,403,429]" captionStart="FIGURE 113" captionStartId="138.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-20@138.[179,1408,181,1920]" captionTargetPageId="138" captionText="FIGURE 113. Tyrannochthonius quattuor sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795427" httpUri="https://zenodo.org/record/7795427/files/figure.png" pageId="137" pageNumber="138">Fig. 113F</figureCitation>
).
</paragraph>
<paragraph id="756236A7FF1FFF53FF705EA73E17FD36" blockId="137.[151,1437,150,1869]" pageId="137" pageNumber="138">
Legs (
<figureCitation id="EDE62A22FF1FFF53FEB95EA73BB5FE62" box="[270,421,438,465]" captionStart="FIGURE 115" captionStartId="140.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1919]" captionTargetId="figure-20@140.[179,1408,181,1919]" captionTargetPageId="140" captionText="FIGURE 115. Tyrannochthonius quattuor sp. nov., holotype male: A. Left chela (lateral view), with details of teeth and with trichobothrial pattern; B. Left chela (dorsal view); C. Leg I (lateral view); D. Leg IV (lateral view). Scale bars: 0.20 mm." figureDoi="http://doi.org/10.5281/zenodo.7795433" httpUri="https://zenodo.org/record/7795433/files/figure.png" pageId="137" pageNumber="138">Fig. 115C, D</figureCitation>
): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 2.06–2.13 times longer than patella and with 1 lyrifissure at the base of femur; tarsus 2.23–2.35 times longer than tibia. Femoropatella of leg IV 4.17–4.79 times longer than deep; tibia 6.10–6.30 times longer than deep; with basal tactile setae on both tarsal segments: metatarsus 3.44–3.50 times longer than deep (TS= 0.36–0.39), tarsus 12.33–13.67 times longer than deep and 2.64–2.65 times longer than metatarsus (TS= 0.26–0.32). Arolium slightly shorter than the claws, not divided; claws simple.
</paragraph>
<paragraph id="756236A7FF1FFF53FF705D9F3B08FCEE" blockId="137.[151,1437,150,1869]" pageId="137" pageNumber="138">
Adult females (
<typeStatus id="AA668805FF1FFF53FEC35D9F3BF3FD1A" box="[372,483,655,681]" pageId="137" pageNumber="138" type="paratype">paratypes</typeStatus>
) (
<figureCitation id="EDE62A22FF1FFF53FE4E5D9F3860FD1A" box="[505,624,655,681]" captionStart="FIGURE 112" captionStartId="136.[151,250,1222,1246]" captionTargetBox="[172,1414,353,1197]" captionTargetId="figure-120@136.[161,1427,345,1199]" captionTargetPageId="136" captionText="FIGURE 112. Tyrannochthonius quattuor sp. nov., A. Holotype male, habitus (dorsal view); B. Paratype female, habitus (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.7795423" httpUri="https://zenodo.org/record/7795423/files/figure.png" pageId="137" pageNumber="138">Figs 112B</figureCitation>
,
<figureCitation id="EDE62A22FF1FFF53FDCB5D9F38ADFD1A" box="[636,701,655,681]" captionStart="FIGURE 113" captionStartId="138.[151,250,1943,1967]" captionTargetBox="[179,1408,181,1920]" captionTargetId="figure-20@138.[179,1408,181,1920]" captionTargetPageId="138" captionText="FIGURE 113. Tyrannochthonius quattuor sp. nov., holotype male (A–F), paratype female (G): A. Left chela (lateral view); B. Left chela (dorsal view); C. Carapace (dorsal view); D. Left chelicera (dorsal view); E. Left pedipalp (minus chela, dorsal view); F. Male genital area (ventral view); G. Female genital area (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.7795427" httpUri="https://zenodo.org/record/7795427/files/figure.png" pageId="137" pageNumber="138">113G</figureCitation>
). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5–6; tergal chaetotaxy I–XII: 3–4: 4: 4: 4: 4: 4: 4: 4: 4–5: 4: T2T: 0; sternal chaetotaxy IV–XII: 12–14: 9: 7–8: 9: 8–9: 9–10: 9: 0: 2; anterior genital operculum with 10 setae, posterior margin with 14–16 marginal setae,
<quantity id="B2259B42FF1FFF53FF6A5DEB3B2FFCA6" box="[221,319,763,789]" metricMagnitude="-1" metricUnit="m" metricValue="6.35" metricValueMax="6.604" metricValueMin="6.096" pageId="137" pageNumber="138" unit="in" value="25.0" valueMax="26.0" valueMin="24.0">24–26 in</quantity>
total; leg IV with a long tactile seta on both tarsal segments: metatarsus 3.20–3.33 times longer than deep (TS= 0.27–0.41), tarsus 13.00–13.50 times longer than deep and 2.53–2.60 times longer than metatarsus (TS= 0.33–0.37).
</paragraph>
<paragraph id="756236A7FF1FFF53FF705C773F64FBCE" blockId="137.[151,1437,150,1869]" pageId="137" pageNumber="138">Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses). Males: body length 2.28–2.48. Pedipalps: trochanter 0.20–0.23/0.17–0.18 (1.18–1.28), femur 1.17–1.26/0.15 (7.80–8.40), patella 0.44–0.45/0.18 (2.44–2.50), chela 1.65–1.74/0.20–0.22 (7.91–8.25), hand 0.59–0.64/0.20–0.22 (2.91–2.95), movable chelal finger length 1.00–1.05. Chelicera 0.59–0.62/0.25 (2.36–2.48), movable finger length 0.33–0.34. Carapace 0.56–0.59/0.55–0.57 (1.02–1.04). Leg I: trochanter 0.16/0.15–0.16 (1.00–1.07), femur 0.64–0.68/0.08–0.09 (7.56–8.00), patella 0.30–0.33/0.07 (4.29–4.71), tibia 0.30–0.31/0.06 (5.00–5.17), tarsus 0.67–0.73/0.06 (11.17– 12.17). Leg IV: trochanter 0.26–0.28/0.16–0.17 (1.63–1.65), femoropatella 0.91–0.96/0.19–0.23 (4.17–4.79), tibia 0.61–0.63/0.10 (6.10–6.30), metatarsus 0.28–0.31/0.08–0.09 (3.44–3.50), tarsus 0.74–0.82/0.06 (12.33–13.67).</paragraph>
<paragraph id="756236A7FF1FFF53FF705B9739A5FA2E" blockId="137.[151,1437,150,1869]" pageId="137" pageNumber="138">Females: body length 2.48–2.79. Pedipalps: trochanter 0.23–0.24/0.18–0.19 (1.26–1.28), femur 1.24– 1.32/0.16–0.18 (7.33–7.75), patella 0.46–0.50/0.18–0.21 (2.38–2.56), chela 1.74–1.86/0.24–0.27 (6.89–7.25), hand 0.63–0.70/0.24–0.27 (2.59–2.63), movable chelal finger length 1.05–1.02. Chelicera 0.62–0.71/0.27–0.29 (2.30–2.45), movable finger length 0.35–0.38. Carapace 0.58–0.63/0.58–0.62 (1.00–1.02). Leg I: trochanter 0.17– 0.19/0.16–0.17 (1.06–1.12), femur 0.66–0.75/0.09–0.10 (7.33–7.50), patella 0.35–0.38/0.07–0.09 (4.22–5.00), tibia 0.31–0.34/0.06–0.07 (4.86–5.17), tarsus 0.72–0.75/0.06 (12.00–12.50). Leg IV: trochanter 0.28–0.30/0.17–0.18 (1.65–1.67), femoropatella 0.94–1.01/0.22–0.25 (4.04–4.27), tibia 0.60–0.63/0.10–0.11 (5.45–6.30), metatarsus 0.30–0.32/0.09–0.10 (3.20–3.33), tarsus 0.78–0.81/0.06 (13.00–13.50).</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFF1FFF53FF705AB73F89F89A" pageId="137" pageNumber="138" type="discussion">
<paragraph id="756236A7FF1FFF53FF705AB739D2F90E" blockId="137.[151,1437,150,1869]" pageId="137" pageNumber="138">
<emphasis id="47A9EAB5FF1FFF53FF705AB73B2BFA72" bold="true" box="[199,315,1447,1473]" pageId="137" pageNumber="138">Remarks.</emphasis>
<taxonomicName id="B2DD4D24FF1FFF53FEF65AB73861FA73" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[321,625,1447,1473]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="quattuor" status="sp. nov.">
<emphasis id="47A9EAB5FF1FFF53FEF65AB73861FA73" box="[321,625,1447,1473]" italics="true" pageId="137" pageNumber="138">Tyrannochthonius quattuor</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FF1FFF53FDCF5AB738C0FA72" bold="true" box="[632,720,1447,1473]" pageId="137" pageNumber="138">
<taxonomicNameLabel id="5C9A57CEFF1FFF53FDCF5AB738C0FA72" box="[632,720,1447,1473]" pageId="137" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="B2DD4D24FF1FFF53FCE85AB739A0FA72" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[863,944,1447,1473]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="latus" status="sp. nov.">
<emphasis id="47A9EAB5FF1FFF53FCE85AB739A0FA72" box="[863,944,1447,1473]" italics="true" pageId="137" pageNumber="138">T. latus</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FF1FFF53FC015AB73E1FFA72" bold="true" box="[950,1039,1447,1473]" pageId="137" pageNumber="138">
<taxonomicNameLabel id="5C9A57CEFF1FFF53FC015AB73E1FFA72" box="[950,1039,1447,1473]" pageId="137" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in having the same number of setae on tergites I–VII (4) and lacking intercalary teeth on both chelal fingers, but differs by having a smaller body size (body length 2.28–2.45 (♂), 2.48–2.79 (
<specimenCount id="63DBFD2EFF1FFF53FDD75AFF3862F9BA" box="[608,626,1519,1545]" pageId="137" pageNumber="138" type="female">♀</specimenCount>
)
<emphasis id="47A9EAB5FF1FFF53FD335AFF38B3F9BB" box="[644,675,1519,1544]" italics="true" pageId="137" pageNumber="138">vs.</emphasis>
2.59–2.80 (♂), 2.91–3.11 (
<specimenCount id="63DBFD2EFF1FFF53FC6C5AFF39FDF9BA" box="[987,1005,1519,1545]" pageId="137" pageNumber="138" type="female">♀</specimenCount>
) mm; chela 7.91–8.25 (♂), 6.89–7.25 (
<specimenCount id="63DBFD2EFF1FFF53FF1559033AA4F99E" box="[162,180,1555,1581]" pageId="137" pageNumber="138" type="female">♀</specimenCount>
) times
<emphasis id="47A9EAB5FF1FFF53FEBD59033B38F99F" box="[266,296,1555,1580]" italics="true" pageId="137" pageNumber="138">vs.</emphasis>
6.69–7.13 (♂), 6.15–6.58 (
<specimenCount id="63DBFD2EFF1FFF53FDEA5903387FF99E" box="[605,623,1555,1581]" pageId="137" pageNumber="138" type="female">♀</specimenCount>
) times longer than board, length 1.65–1.74 (♂), 1.74–1.86 (
<specimenCount id="63DBFD2EFF1FFF53FAA359033F36F99E" box="[1300,1318,1555,1581]" pageId="137" pageNumber="138" type="female">♀</specimenCount>
)
<emphasis id="47A9EAB5FF1FFF53FA8F59033F46F99F" box="[1336,1366,1555,1580]" italics="true" pageId="137" pageNumber="138">vs.</emphasis>
2.14– 2.34 (♂), 2.37–2.46 (
<specimenCount id="63DBFD2EFF1FFF53FE3B59273B8EF9E2" box="[396,414,1591,1617]" pageId="137" pageNumber="138" type="female">♀</specimenCount>
) mm; palpal femur 7.80–8.40 (♂), 7.33–7.75 (
<specimenCount id="63DBFD2EFF1FFF53FC0D592739DCF9E2" box="[954,972,1591,1617]" pageId="137" pageNumber="138" type="female">♀</specimenCount>
) times
<emphasis id="47A9EAB5FF1FFF53FB9259273E53F9E3" box="[1061,1091,1591,1616]" italics="true" pageId="137" pageNumber="138">vs.</emphasis>
9.29–9.33 (♂), 8.75–8.79 (
<specimenCount id="63DBFD2EFF1FFF53FAC859273F81F9E2" box="[1407,1425,1591,1617]" pageId="137" pageNumber="138" type="female">♀</specimenCount>
) times longer than board, length 1.17–1.26 (♂), 1.24–1.32 (
<specimenCount id="63DBFD2EFF1FFF53FC8E594B395BF9C6" box="[825,843,1627,1653]" pageId="137" pageNumber="138" type="female">♀</specimenCount>
)
<emphasis id="47A9EAB5FF1FFF53FCE9594B396DF9C7" box="[862,893,1627,1652]" italics="true" pageId="137" pageNumber="138">vs.</emphasis>
1.58–1.68 (♂), 1.67–1.75 (
<specimenCount id="63DBFD2EFF1FFF53FB0E594B3EDBF9C6" box="[1209,1227,1627,1653]" pageId="137" pageNumber="138" type="female">♀</specimenCount>
) mm), the trait of teeth on the movable chelal finger (continuous and markedly retrorse
<emphasis id="47A9EAB5FF1FFF53FC10596F39D5F92B" box="[935,965,1663,1688]" italics="true" pageId="137" pageNumber="138">vs.</emphasis>
well-spaced and slightly retrorse) and the number of blades of rallum (8
<emphasis id="47A9EAB5FF1FFF53FE5A59B3381CF90F" box="[493,524,1699,1724]" italics="true" pageId="137" pageNumber="138">vs.</emphasis>
7) and coxal spines (13–15
<emphasis id="47A9EAB5FF1FFF53FCF059B33976F90F" box="[839,870,1699,1724]" italics="true" pageId="137" pageNumber="138">vs.</emphasis>
11–12).
</paragraph>
<paragraph id="756236A7FF1FFF53FF7059D73F89F89A" blockId="137.[151,1437,150,1869]" pageId="137" pageNumber="138">
<taxonomicName id="B2DD4D24FF1FFF53FF7059D73BE7F953" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" box="[199,503,1735,1761]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="quattuor" status="sp. nov.">
<emphasis id="47A9EAB5FF1FFF53FF7059D73BE7F953" box="[199,503,1735,1761]" italics="true" pageId="137" pageNumber="138">Tyrannochthonius quattuor</emphasis>
</taxonomicName>
<emphasis id="47A9EAB5FF1FFF53FE4A59D73845F952" bold="true" box="[509,597,1735,1761]" pageId="137" pageNumber="138">
<taxonomicNameLabel id="5C9A57CEFF1FFF53FE4A59D73845F952" box="[509,597,1735,1761]" pageId="137" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="B2DD4D24FF1FFF53FC3759D739E0F952" authorityName="Mahnert" authorityYear="2009" box="[896,1008,1735,1761]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="akaelus">
<emphasis id="47A9EAB5FF1FFF53FC3759D739E0F952" box="[896,1008,1735,1761]" italics="true" pageId="137" pageNumber="138">T. akaelus</emphasis>
</taxonomicName>
,
<taxonomicName id="B2DD4D24FF1FFF53FC4B59D73EBFF952" authorityName="Mahnert" authorityYear="2009" box="[1020,1199,1735,1761]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="antridraconis">
<emphasis id="47A9EAB5FF1FFF53FC4B59D73EBFF952" box="[1020,1199,1735,1761]" italics="true" pageId="137" pageNumber="138">T. antridraconis</emphasis>
</taxonomicName>
,
<taxonomicName id="B2DD4D24FF1FFF53FB0B59D73F38F952" authorityName="Gao, Wynne &amp; Fe.Zhang" authorityYear="2018" box="[1212,1320,1735,1761]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="chixing">
<emphasis id="47A9EAB5FF1FFF53FB0B59D73F38F952" box="[1212,1320,1735,1761]" italics="true" pageId="137" pageNumber="138">T. chixing</emphasis>
</taxonomicName>
,
<taxonomicName id="B2DD4D24FF1FFF53FA8159D73AE9F8B7" authorityName="Mahnert" authorityYear="2009" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="ganshuanensis">
<emphasis id="47A9EAB5FF1FFF53FA8159D73AE9F8B7" italics="true" pageId="137" pageNumber="138">T. ganshuanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="B2DD4D24FF1FFF53FE9959FB3B68F8B6" authorityName="Gao, Zhang &amp; Chen" authorityYear="2020" box="[302,376,1771,1797]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="zhai">
<emphasis id="47A9EAB5FF1FFF53FE9959FB3B68F8B6" box="[302,376,1771,1797]" italics="true" pageId="137" pageNumber="138">T. zhai</emphasis>
</taxonomicName>
by lacking intercalary teeth on the fixed chelal finger, from
<taxonomicName id="B2DD4D24FF1FFF53FBA659FB3E90F8B6" authorityName="Gao, Zhang &amp; Chen" authorityYear="2020" box="[1041,1152,1771,1797]" class="Arachnida" family="Chthoniidae" genus="Tyrannochthonius" kingdom="Animalia" order="Pseudoscorpiones" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="harveyi">
<emphasis id="47A9EAB5FF1FFF53FBA659FB3E90F8B6" box="[1041,1152,1771,1797]" italics="true" pageId="137" pageNumber="138">T. harveyi</emphasis>
</taxonomicName>
by the number of setae at the anterior margin of the carapace (6
<emphasis id="47A9EAB5FF1FFF53FDF4581F3871F89B" box="[579,609,1807,1832]" italics="true" pageId="137" pageNumber="138">vs.</emphasis>
4) and on tergites I–III (4
<emphasis id="47A9EAB5FF1FFF53FC3E581F39B8F89B" box="[905,936,1807,1832]" italics="true" pageId="137" pageNumber="138">vs.</emphasis>
2) (
<bibRefCitation id="114C4B56FF1FFF53FC62581F3E90F89A" author="Mahnert, V." box="[981,1152,1807,1833]" pageId="137" pageNumber="138" pagination="185 - 201" refId="ref77141" refString="Mahnert, V. (2009) New species of pseudoscorpions (Arachnida, Pseudoscorpiones, Chthoniidae, Chernetidae) from caves in China. Revue Suisse de Zoologie, 116 (2), 185 - 201. https: // doi. org / 10.5962 / bhl. part. 79492" type="journal article" year="2009">Mahnert 2009</bibRefCitation>
;
<bibRefCitation id="114C4B56FF1FFF53FB3C581F3F51F89A" author="Gao, Z. Z. &amp; Wynne, J. J. &amp; Zhang, F." box="[1163,1345,1807,1833]" pageId="137" pageNumber="138" pagination="345 - 354" refId="ref76312" refString="Gao, Z. Z., Wynne, J. J. &amp; Zhang, F. (2018) Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones: Neobisiidae, Chthoniidae) from Guangxi, China. Journal of Arachnology, 46, 345 - 354. https: // doi. org / 10.1636 / JoA-S- 17 - 063.1" type="journal article" year="2018">
Gao
<emphasis id="47A9EAB5FF1FFF53FB74581F3EEDF89A" box="[1219,1277,1807,1833]" italics="true" pageId="137" pageNumber="138">et al.</emphasis>
2018
</bibRefCitation>
,
<bibRefCitation id="114C4B56FF1FFF53FAFA581F3F9AF89A" author="Gao, Z. Z. &amp; Zhang, F. &amp; Chen, H. M." box="[1357,1418,1807,1833]" pageId="137" pageNumber="138" pagination="572 - 580" refId="ref76374" refString="Gao, Z. Z., Zhang, F. &amp; Chen, H. M. (2020) Two new cave-dwelling species of Tyrannochthonius Chamberlin 1929 (Pseudoscorpiones: Chthoniidae) from the Guizhou karst, China. Zootaxa, 4853 (4), 572 - 580. https: // doi. org / 10.11646 / zootaxa. 4853.4.6" type="journal article" year="2020">2020</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3DC7652CFF1FFF53FF70582238E1F8FE" box="[199,753,1842,1869]" pageId="137" pageNumber="138" type="distribution">
<paragraph id="756236A7FF1FFF53FF70582238E1F8FE" blockId="137.[151,1437,150,1869]" box="[199,753,1842,1869]" pageId="137" pageNumber="138">
<emphasis id="47A9EAB5FF1FFF53FF7058223B71F8FF" bold="true" box="[199,353,1842,1868]" pageId="137" pageNumber="138">Distribution.</emphasis>
Known only from the
<typeStatus id="AA668805FF1FFF53FDD758233880F8FE" box="[608,656,1843,1869]" pageId="137" pageNumber="138">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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