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<mods:title id="CD53AB66C9D950A2CF0B0C05246DEAE2">A revision of the genus Goniusa Casey, 1906 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<taxonomicName id="ACB97DA9FF98FFFA1E2AF9FC1346D478" ID-CoL="8VRPV" authority="Casey, 1906" authorityName="Casey" authorityYear="1906" box="[264,521,1558,1584]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF98FFFA1E2AF9FC1020D478" bold="true" box="[264,367,1558,1584]" italics="true" pageId="1" pageNumber="2">Goniusa</emphasis>
<bibRefCitation id="0F287BDBFF98FFFA1E54F9FC1346D478" author="Casey" box="[374,521,1558,1584]" pageId="1" pageNumber="19" refString="Casey, T. L. (1906) Observations of the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis, 16 (6), 125 - 434." type="journal article" year="1906">Casey, 1906</bibRefCitation>
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<figureCitation id="F3821AAFFF98FFFA1D3BF9FC13DCD478" box="[537,659,1558,1584]" captionStart-0="FIGURES 1 ­ 5" captionStart-1="FIGURES 6 ­ 11" captionStart-10="FIGURES 52 ­ 56" captionStart-2="FIGURES 12 ­ 15" captionStart-3="FIGURES 16 ­ 19" captionStart-4="FIGURES 20 ­ 24" captionStart-5="FIGURES 25 ­ 29" captionStart-6="FIGURES 30 ­ 36" captionStart-7="FIGURES 37 ­ 41" captionStart-8="FIGURES 42 ­ 47" captionStart-9="FIGURES 48 ­ 51" captionStartId-0="2.[264,383,1522,1546]" captionStartId-1="4.[264,383,1469,1493]" captionStartId-10="16.[264,383,1747,1771]" captionStartId-2="5.[264,383,1787,1811]" captionStartId-3="7.[264,383,1700,1724]" captionStartId-4="9.[264,383,1550,1574]" captionStartId-5="10.[264,383,1768,1792]" captionStartId-6="11.[264,382,1632,1656]" captionStartId-7="12.[264,382,1692,1716]" captionStartId-8="14.[264,382,1769,1793]" captionStartId-9="15.[264,383,1288,1312]" captionTargetBox-0="[290,1296,439,1505]" captionTargetBox-1="[298,1287,339,1429]" captionTargetBox-10="[292,1298,498,1731]" captionTargetBox-2="[289,1304,581,1774]" captionTargetBox-3="[281,1304,573,1663]" captionTargetBox-4="[290,1295,579,1513]" captionTargetBox-5="[280,1310,701,1744]" captionTargetBox-6="[288,1303,337,1596]" captionTargetBox-7="[301,1285,783,1653]" captionTargetBox-8="[288,1298,540,1737]" captionTargetBox-9="[314,1270,697,1252]" captionTargetId-0="figure@2.[280,1306,425,1512]" captionTargetId-1="figure@4.[288,1299,329,1443]" captionTargetId-10="figure@16.[281,1307,489,1736]" captionTargetId-2="figure@5.[272,1314,569,1776]" captionTargetId-3="figure@7.[271,1316,561,1676]" captionTargetId-4="figure@9.[282,1306,569,1524]" captionTargetId-5="figure@10.[263,1323,689,1755]" captionTargetId-6="figure@11.[273,1314,326,1608]" captionTargetId-7="figure@12.[288,1299,769,1665]" captionTargetId-8="figure@14.[279,1307,529,1742]" captionTargetId-9="figure@15.[305,1283,689,1261]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-10="16" captionTargetPageId-2="5" captionTargetPageId-3="7" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionTargetPageId-8="14" captionTargetPageId-9="15" captionText-0="FIGURES 1 ­ 5. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." captionText-1="FIGURES 6 ­ 11. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 6 – prementum; 7 – hypopharynx; 8 – right maxilla, ventral view; 9 – right lacinia, dorsal view; 10 – right lacinia, ventral view; 11 – right galea, dorsal view. Scale bar 0.1 mm (6 ­ 7, 9 ­ 11), 0.2 mm (8)." captionText-10="FIGURES 52 ­ 56. Genitalia of Goniusa alperti Kistner (paratype from 10 mi. E of Coville, Washington (52); female from Tahoe City, California (53); female from 16 km NW of Granby, Colorado (54); holotype (55 ­ 56 )). 52 ­ 54 – spermatheca; 55 – everted internal sac of aedeagus, lateral view; 56 – everted internal sac of aedeagus, parameral view. Scale bar 0.2 mm." captionText-2="FIGURES 12 ­ 15. Details of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 12 – pronotum; 13 – meso­ and metathorax, ventral view; 14 – right antenna; 15 – mentum. Scale bar 0.1 mm (15), 0.2 mm (12), 0.4 mm (13 ­ 14)." captionText-3="FIGURES 16 ­ 19. Details of Drusilla canaliculata (Fabricius) (female from Kiev, Ukraine (16); male from 15 km N of Magadan, Russia (17 )) and Zyras humeralis (Gravenhorst) (female from Briançon, France). 16, 19 – right maxilla, ventral view; 17 – meso­ and metathorax, ventral view; 18 – meso­ and metathorax, ventral view (setation not shown). Scale bar 0.2 mm (16), 0.4 mm (17, 19), 1.0 mm (18)." captionText-4="FIGURES 20 ­ 24. Abdominal segment 8 of Goniusa caseyi Gusarov, sp. n. (paratypes from Washington, D. C. (20 ­ 21); Aweme, Manitoba (22); and Stonewall, Manitoba (23 ­ 24 )). 20 – male tergum 8; 21 – male sternum 8; 22 – apex of male tergum 8; 23 – female tergum 8; 24 – female sternum 8. Scale bar 0.4 mm (20 ­ 21, 23 ­ 24), 0.2 mm (22)." captionText-5="FIGURES 25 ­ 29. Median lobe of aedeagus of Goniusa caseyi Gusarov, sp. n. (paratypes from Washington, D. C. (25 ­ 26, 28 ­ 29); and Sherborn, Massachusetts (27 )). 25 – median lobe, parameral view; 26 ­ 27 – apex of median lobe, parameral view; 28 – median lobe, lateral view; 29 – apex of median lobe, lateral view. Scale bar 0.4 mm (25, 28), 0.2 mm (26 ­ 27, 29)." captionText-6="FIGURES 30 ­ 36. Genitalia of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (30 ­ 31); Texas (32 ­ 34); Sherborn, Massachusetts (35); and Stonewall, Manitoba (36 )). 30 – everted internal sac of aedeagus, parameral view; 31 – everted internal sac of aedeagus, lateral view; 32 – copulatory piece, apical view; 33 – copulatory piece, lateral view; 34 – copulatory piece, abparameral view; 35 ­ 36 – spermatheca. Scale bar 0.1 mm (32 ­ 34), 0.2 mm (30 ­ 31, 35 ­ 36)." captionText-7="FIGURES 37 ­ 41. Pronotum of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (37); and Stonewall, Manitoba (38 )) and G. alperti Kistner (holotype (39); female from San Bernardino Mts., California (40), paratype from 10 mi. E of Coville, Washington (41 )). 37, 39 – male pronotum; 38, 40 ­ 41 – female pronotum. Scale bar 0.5 mm." captionText-8="FIGURES 42 ­ 47. Abdominal segment 8 of Goniusa alperti Kistner (holotype (42 ­ 44); male from San Bernardino Mts., California (45); female from 16 km NW of Granby, Colorado (46 ­ 47 )). 42 – male tergum 8; 43 – male sternum 8; 44 ­ 45 – apex of male tergum 8; 46 – female tergum 8; 47 – female sternum 8. Scale bar 0.4 mm (42 ­ 43, 46 ­ 47), 0.2 mm (44 ­ 45)." captionText-9="FIGURES 48 ­ 51. Median lobe of aedeagus of Goniusa alperti Kistner (male from West Point, Nebraska). 48 – median lobe, parameral view; 49 – apex of median lobe, parameral view; 50 – median lobe, lateral view; 51 – apex of median lobe, lateral view. Scale bar 0.4 mm (48, 50), 0.2 mm (49, 51)." httpUri-0="https://zenodo.org/record/156408/files/figure.png" httpUri-1="https://zenodo.org/record/156409/files/figure.png" httpUri-10="https://zenodo.org/record/156418/files/figure.png" httpUri-2="https://zenodo.org/record/156410/files/figure.png" httpUri-3="https://zenodo.org/record/156411/files/figure.png" httpUri-4="https://zenodo.org/record/156412/files/figure.png" httpUri-5="https://zenodo.org/record/156413/files/figure.png" httpUri-6="https://zenodo.org/record/156414/files/figure.png" httpUri-7="https://zenodo.org/record/156415/files/figure.png" httpUri-8="https://zenodo.org/record/156416/files/figure.png" httpUri-9="https://zenodo.org/record/156417/files/figure.png" pageId="1" pageNumber="2">Figs. 1­56</figureCitation>
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<taxonomicName id="ACB97DA9FF98FFFA1E2AF98B1367D430" ID-CoL="8VRPV" authority="Casey, 1906: 348" authorityName="Casey" authorityPageNumber="348" authorityYear="1906" box="[264,552,1633,1656]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF98FFFA1E2AF98B102DD430" box="[264,354,1633,1656]" italics="true" pageId="1" pageNumber="2">Goniusa</emphasis>
<treatmentCitation id="EA18203BFF98FFFA1E4BF98B1367D430" author="Casey" box="[361,552,1633,1656]" page="348" pageId="1" pageNumber="2" year="1906">
<bibRefCitation id="0F287BDBFF98FFFA1E4BF98B10A0D430" author="Casey" box="[361,495,1633,1656]" pageId="1" pageNumber="19" refString="Casey, T. L. (1906) Observations of the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis, 16 (6), 125 - 434." type="journal article" year="1906">Casey, 1906</bibRefCitation>
: 348
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(in tribe
<taxonomicName id="ACB97DA9FF98FFFA1DA8F98B1285D430" ID-CoL="8VWQ5" authority="Thomson, 1859" authorityName="Thomson" authorityYear="1859" box="[650,970,1633,1656]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Bolitocharini">
Bolitocharini
<bibRefCitation id="0F287BDBFF98FFFA1C3CF98B1285D430" author="Thomson" box="[798,970,1633,1656]" pageId="1" pageNumber="20" refString="Thomson, C. G. (1859) Skandinaviens Coleoptera, synoptiskt bearbetade. Tom 1. Berlingska Boktryckeriet, Lund, v + 290 pp." type="book" year="1859">Thomson, 1859</bibRefCitation>
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<emphasis id="59CDDA38FF98FFFA1E2AF96B102DD4D0" box="[264,354,1665,1688]" italics="true" pageId="1" pageNumber="2">Goniusa</emphasis>
:
<treatmentCitation id="EA18203BFF98FFFA1E55F96B137FD4D0" author="Casey" box="[375,560,1665,1688]" page="208" pageId="1" pageNumber="2" year="1911">
<bibRefCitation id="0F287BDBFF98FFFA1E55F96B10B8D4D0" author="Casey" box="[375,503,1665,1688]" pageId="1" pageNumber="19" refString="Casey, T. L. (1911) New American species of Aleocharinae and Myllaeninae. Memoirs on the Coleoptera II. The New Era Printing Company, Lancaster, pp. 1 - 245." type="book chapter" year="1911">Casey 1911</bibRefCitation>
: 208
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(in tribe
<taxonomicName id="ACB97DA9FF98FFFA1DB1F96B126AD4D0" ID-CoL="8VWQ5" box="[659,805,1665,1688]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Bolitocharini">Bolitocharini</taxonomicName>
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<taxonomicName id="ACB97DA9FF98FFFA1E2AF94B130ED4F0" ID-CoL="8VRPV" authority="Fenyes, 1918: 19" authorityName="Fenyes" authorityPageNumber="19" authorityYear="1918" box="[264,577,1697,1720]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF98FFFA1E2AF94B102DD4F0" box="[264,354,1697,1720]" italics="true" pageId="1" pageNumber="2">Goniusa</emphasis>
:
<treatmentCitation id="EA18203BFF98FFFA1E5EF94B130ED4F0" author="Fenyes" box="[380,577,1697,1720]" page="19" pageId="1" pageNumber="2" year="1918">
<bibRefCitation id="0F287BDBFF98FFFA1E5EF94B135FD4F0" author="Fenyes" box="[380,528,1697,1720]" pageId="1" pageNumber="19" refString="Fenyes, A. (1918) Coleoptera: Fam Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 A. L. Desmet-Verteneuil, Bruxelles, pp. 1 - 110." type="book chapter" year="1918">Fenyes, 1918</bibRefCitation>
: 19
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(in subtribe
<taxonomicName id="ACB97DA9FF98FFFA1DF2F94B12F6D4F0" ID-CoL="8VRC4" authority="Casey, 1910" authorityName="Casey" authorityYear="1910" box="[720,953,1697,1720]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="subTribe" subTribe="Athetina">
Athetina
<bibRefCitation id="0F287BDBFF98FFFA1C17F94B12F6D4F0" author="Casey" box="[821,953,1697,1720]" pageId="1" pageNumber="19" refString="Casey, T. L. (1910) New Species of the Staphylinid Tribe Myrmedoniini. Memoirs on the Coleoptera I. The New Era Printing Company, Lancaster, pp. 1 - 183." type="book chapter" year="1910">Casey, 1910</bibRefCitation>
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of tribe
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Myrmedoniini
<bibRefCitation id="0F287BDBFF98FFFA1B9DF94B103ED490" author="Thomson" pageId="1" pageNumber="20" refString="Thomson, C. G. (1867) Skandinaviens Coleoptera, synoptiskt bearbetade, Tom 9. Lundbergska Boktryckeriet, Lund, 408 pp." type="book" year="1867">Thomson, 1867</bibRefCitation>
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).
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<taxonomicName id="ACB97DA9FF98FFFA1E2AF90B130AD4B0" ID-CoL="8VRPV" authority="Fenyes, 1920: 235" authorityName="Fenyes" authorityPageNumber="235" authorityYear="1920" box="[264,581,1761,1784]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF98FFFA1E2AF90B102DD4B0" box="[264,354,1761,1784]" italics="true" pageId="1" pageNumber="2">Goniusa</emphasis>
:
<treatmentCitation id="EA18203BFF98FFFA1E55F90B130AD4B0" author="Fenyes" box="[375,581,1761,1784]" page="235" pageId="1" pageNumber="2" year="1920">
<bibRefCitation id="0F287BDBFF98FFFA1E55F90B1346D4B0" author="Fenyes" box="[375,521,1761,1784]" pageId="1" pageNumber="19" refString="Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles. Pp. 111 - 414." type="book chapter" year="1920">Fenyes, 1920</bibRefCitation>
: 235
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<paragraph id="6B06062AFF98FFFA1E2AF8EB12B7D550" blockId="1.[264,1323,1633,1880]" box="[264,1016,1793,1816]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="4BA92104FF98FFFA1E2AF8EB12B7D550" box="[264,1016,1793,1816]" pageId="1" pageNumber="2">
<taxonomicName id="ACB97DA9FF98FFFA1E2AF8EB124BD550" ID-CoL="8VRPV" authority="Bernhauer &amp; Scheerpeltz, 1926: 597" authorityName="Bernhauer &amp; Scheerpeltz" authorityPageNumber="597" authorityYear="1926" box="[264,772,1793,1816]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF98FFFA1E2AF8EB102DD550" box="[264,354,1793,1816]" italics="true" pageId="1" pageNumber="2">Goniusa</emphasis>
:
<treatmentCitation id="EA18203BFF98FFFA1E55F8EB124BD550" author="Bernhauer" box="[375,772,1793,1816]" page="597" pageId="1" pageNumber="2" year="1926">
<bibRefCitation id="0F287BDBFF98FFFA1E55F8EB1384D550" author="Bernhauer" box="[375,715,1793,1816]" pageId="1" pageNumber="19" refString="Bernhauer, M. &amp; Scheerpeltz, O. (1926) Staphylinidae VI. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus, Pars 82. W. Junk, Berlin, pp. 499 - 988." type="journal article" year="1926">Bernhauer &amp; Scheerpeltz, 1926</bibRefCitation>
: 597
</treatmentCitation>
</taxonomicName>
(in subtribe
<taxonomicName id="ACB97DA9FF98FFFA1CA9F8EB12A3D550" ID-CoL="8VRC4" box="[907,1004,1793,1816]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="subTribe" subTribe="Athetina">Athetina</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="6B06062AFF98FFFA1E2AF8CB13CBD570" blockId="1.[264,1323,1633,1880]" box="[264,644,1825,1848]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="4BA92104FF98FFFA1E2AF8CB13CBD570" box="[264,644,1825,1848]" pageId="1" pageNumber="2">
<taxonomicName id="ACB97DA9FF98FFFA1E2AF8CB13CFD570" ID-CoL="8VRPV" authority="Blackwelder, 1952: 174" authorityName="Blackwelder" authorityPageNumber="174" authorityYear="1952" box="[264,640,1825,1848]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF98FFFA1E2AF8CB102DD570" box="[264,354,1825,1848]" italics="true" pageId="1" pageNumber="2">Goniusa</emphasis>
:
<treatmentCitation id="EA18203BFF98FFFA1E55F8CB13CFD570" author="Blackwelder" box="[375,640,1825,1848]" page="174" pageId="1" pageNumber="2" year="1952">
<bibRefCitation id="0F287BDBFF98FFFA1E55F8CB130BD570" author="Blackwelder" box="[375,580,1825,1848]" pageId="1" pageNumber="19" refString="Blackwelder, R. E. (1952) The generic names of the beetle family Staphylinidae, with an essay on genotypy. U. S. National Museum Bulletin, 200, 1 - 483." type="journal article" year="1952">Blackwelder, 1952</bibRefCitation>
: 174
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6B06062AFF98FFFA1E2AF8AB12C7D510" blockId="1.[264,1323,1633,1880]" box="[264,904,1857,1880]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="4BA92104FF98FFFA1E2AF8AB12C7D510" box="[264,904,1857,1880]" pageId="1" pageNumber="2">
<taxonomicName id="ACB97DA9FF98FFFA1E2AF8AB1378D510" ID-CoL="8VRPV" authority="Kistner, 1976: 84" authorityName="Kistner" authorityPageNumber="84" authorityYear="1976" box="[264,567,1857,1880]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF98FFFA1E2AF8AB102DD510" box="[264,354,1857,1880]" italics="true" pageId="1" pageNumber="2">Goniusa</emphasis>
:
<treatmentCitation id="EA18203BFF98FFFA1E55F8AB1378D510" author="Kistner" box="[375,567,1857,1880]" page="84" pageId="1" pageNumber="2" year="1976">
<bibRefCitation id="0F287BDBFF98FFFA1E55F8AB1344D510" author="Kistner" box="[375,523,1857,1880]" pageId="1" pageNumber="19" refString="Kistner, D. H. (1976) Revision and Reclassification of the Genus Goniusa Casey with a Larval Description and Ant Host Records (Coleoptera: Staphylinidae). Sociobiology, 2 (1), 83 - 95." type="journal article" year="1976">Kistner, 1976</bibRefCitation>
: 84
</treatmentCitation>
</taxonomicName>
(in tribe Zyrini
<bibRefCitation id="0F287BDBFF98FFFA1DC1F8AB1233D510" author="Bradley" box="[739,892,1857,1880]" pageId="1" pageNumber="19" refString="Bradley, J. C. (1930) A manual of the genera of beetles of America north of Mexico. Daw, Illston &amp; Co., Ithaca, 360 pp." type="book" year="1930">Bradley, 1930</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="6B06062AFF9BFFF91E2AFEF610D1D33B" blockId="2.[264,1323,284,371]" pageId="2" pageNumber="3">
<treatmentCitationGroup id="4BA92104FF9BFFF91E2AFEF610D1D33B" pageId="2" pageNumber="3">
<taxonomicName id="ACB97DA9FF9BFFF91E2AFEF61305D37B" ID-CoL="8VRPV" authority="Seevers, 1978: 133" authorityName="Seevers" authorityPageNumber="133" authorityYear="1978" box="[264,586,284,307]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9BFFF91E2AFEF6102DD37B" box="[264,354,284,307]" italics="true" pageId="2" pageNumber="3">Goniusa</emphasis>
:
<treatmentCitation id="EA18203BFF9BFFF91E55FEF61305D37B" author="Seevers" box="[375,586,284,307]" page="133" pageId="2" pageNumber="3" year="1978">
<bibRefCitation id="0F287BDBFF9BFFF91E55FEF6135ED37B" author="Seevers" box="[375,529,284,307]" pageId="2" pageNumber="20" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">Seevers, 1978</bibRefCitation>
: 133
</treatmentCitation>
</taxonomicName>
(in
<taxonomicName id="ACB97DA9FF9BFFF91D57FEF61380D37B" ID-CoL="8VRPV" box="[629,719,284,307]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9BFFF91D57FEF61380D37B" box="[629,719,284,307]" italics="true" pageId="2" pageNumber="3">Goniusa</emphasis>
</taxonomicName>
group of tribe Athetini).
<taxonomicName id="ACB97DA9FF9BFFF91E2AFED610E1D31B" ID-CoL="8VRPV" authority="Ashe" authorityName="Ashe" box="[264,430,316,339]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9BFFF91E2AFED6102DD31B" box="[264,354,316,339]" italics="true" pageId="2" pageNumber="3">Goniusa</emphasis>
: Ashe
</taxonomicName>
<emphasis id="59CDDA38FF9BFFF91E97FED61085D31B" box="[437,458,316,339]" italics="true" pageId="2" pageNumber="3">in</emphasis>
<treatmentCitation id="EA18203BFF9BFFF91EF2FED6128CD31B" author="Newton" box="[464,963,316,339]" page="371" pageId="2" pageNumber="3" year="2000">
<bibRefCitation id="0F287BDBFF9BFFF91EF2FED612C5D31B" author="Newton" box="[464,906,316,339]" pageId="2" pageNumber="19" refString="Newton, A. F., Thayer, M. K., Ashe, J. S. &amp; Chandler, D. S. (2000) Staphylinidae Latreille, 1802. In: Arnett, R. H., Thomas, M. C. (Eds.), American Beetles. Vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press, Boca Raton, pp. 272 - 418." type="book chapter" year="2000">Newton, Thayer, Ashe &amp; Chandler, 2000</bibRefCitation>
: 371
</treatmentCitation>
(in tribe Athetini, not assigned to subtribe).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="3FC656A2FF9BFFF91E2AFA181387D407" httpUri="https://zenodo.org/record/156408/files/figure.png" pageId="2" pageNumber="3" targetBox="[290,1296,439,1505]" targetPageId="2">
<paragraph id="6B06062AFF9BFFF91E2AFA181387D407" blockId="2.[264,1324,1522,1615]" pageId="2" pageNumber="3">
<emphasis id="59CDDA38FF9BFFF91E2AFA1810FFD442" bold="true" box="[264,432,1522,1546]" pageId="2" pageNumber="3">FIGURES 1­5.</emphasis>
Mouthparts of
<taxonomicName id="ACB97DA9FF9BFFF91D7FFA19122DD442" authority="Gusarov" authorityName="Gusarov" box="[605,866,1523,1546]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="caseyi" status="sp. nov.">
<emphasis id="59CDDA38FF9BFFF91D7FFA19124ED442" box="[605,769,1523,1546]" italics="true" pageId="2" pageNumber="3">Goniusa caseyi</emphasis>
Gusarov
</taxonomicName>
,
<taxonomicNameLabel id="42FE6743FF9BFFF91C4CFA1812E6D442" box="[878,937,1522,1546]" pageId="2" pageNumber="3" rank="species">
<emphasis id="59CDDA38FF9BFFF91C4CFA1812E6D442" bold="true" box="[878,937,1522,1546]" pageId="2" pageNumber="3">sp. n.</emphasis>
</taxonomicNameLabel>
(paratype from Aweme, Manitoba). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm.
</paragraph>
</caption>
<subSubSection id="23A355A1FF9BFFF81E2AF940157AD0A6" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="6B06062AFF9BFFF81E2AF940157AD0A6" blockId="2.[264,1324,1706,1893]" lastBlockId="3.[264,1324,284,1790]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis id="59CDDA38FF9BFFF91E2AF94010C7D48C" bold="true" box="[264,392,1706,1732]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
<taxonomicName id="ACB97DA9FF9BFFF91EBAF94110B3D48C" box="[408,508,1707,1732]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9BFFF91EBAF94110B3D48C" box="[408,508,1707,1732]" italics="true" pageId="2" pageNumber="3">Goniusa</emphasis>
</taxonomicName>
can be distinguished from the other athetine genera by the combination of the following characters: body broad; antennal articles 5­10 slightly elongate (
<figureCitation id="F3821AAFFF9BFFF91BD9F9391069D55D" captionStart="FIGURES 12 ­ 15" captionStartId="5.[264,383,1787,1811]" captionTargetBox="[289,1304,581,1774]" captionTargetId="figure@5.[272,1314,569,1776]" captionTargetPageId="5" captionText="FIGURES 12 ­ 15. Details of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 12 – pronotum; 13 – meso­ and metathorax, ventral view; 14 – right antenna; 15 – mentum. Scale bar 0.1 mm (15), 0.2 mm (12), 0.4 mm (13 ­ 14)." httpUri="https://zenodo.org/record/156410/files/figure.png" pageId="2" pageNumber="3">Fig. 14</figureCitation>
) or slightly transverse; in dry specimens gaps between antennal articles inconspicuous; ligula split apically (
<figureCitation id="F3821AAFFF9BFFF91D25F8C9131BD575" box="[519,596,1827,1853]" captionStart="FIGURES 6 ­ 11" captionStartId="4.[264,383,1469,1493]" captionTargetBox="[298,1287,339,1429]" captionTargetId="figure@4.[288,1299,329,1443]" captionTargetPageId="4" captionText="FIGURES 6 ­ 11. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 6 – prementum; 7 – hypopharynx; 8 – right maxilla, ventral view; 9 – right lacinia, dorsal view; 10 – right lacinia, ventral view; 11 – right galea, dorsal view. Scale bar 0.1 mm (6 ­ 7, 9 ­ 11), 0.2 mm (8)." httpUri="https://zenodo.org/record/156409/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
); labial palpus with setae,, and present; pronotum strongly transverse, 1.5­1.6 times as wide as long, with microsetae directed posteriorly along the midline; in lateral portions of the disc microsetae directed posteriorly and obliquely laterally (
<typeStatus id="B402B888FF9AFFF81ED0FEAE1361D316" box="[498,558,324,350]" pageId="3" pageNumber="4">Type</typeStatus>
V,
<bibRefCitation id="0F287BDBFF9AFFF81D7BFEAE122BD316" author="Benick" box="[601,868,324,350]" pageId="3" pageNumber="19" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
) (
<figureCitation id="F3821AAFFF9AFFF81C5CFEAE1298D316" box="[894,983,324,350]" captionStart="FIGURES 12 ­ 15" captionStartId="5.[264,383,1787,1811]" captionTargetBox="[289,1304,581,1774]" captionTargetId="figure@5.[272,1314,569,1776]" captionTargetPageId="5" captionText="FIGURES 12 ­ 15. Details of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 12 – pronotum; 13 – meso­ and metathorax, ventral view; 14 – right antenna; 15 – mentum. Scale bar 0.1 mm (15), 0.2 mm (12), 0.4 mm (13 ­ 14)." httpUri="https://zenodo.org/record/156410/files/figure.png" pageId="3" pageNumber="4">Fig. 12</figureCitation>
); pronotal macrosetae short; pronotal hypomera fully visible in lateral view; medial macroseta of mesotibia inconspicuous, shorter than tibial width; tarsal formula 4­5­5; metatarsal segment 1 slightly shorter than segment 2; one empodial seta; abdominal sterna with numerous semierect macrosetae, sternum 8 with 30­60 macrosetae (
<figureCitation id="F3821AAFFF9AFFF81DC0FE0E1238D3B6" box="[738,887,484,510]" captionStart="FIGURES 20 ­ 24" captionStartId="9.[264,383,1550,1574]" captionTargetBox="[290,1295,579,1513]" captionTargetId="figure@9.[282,1306,569,1524]" captionTargetPageId="9" captionText="FIGURES 20 ­ 24. Abdominal segment 8 of Goniusa caseyi Gusarov, sp. n. (paratypes from Washington, D. C. (20 ­ 21); Aweme, Manitoba (22); and Stonewall, Manitoba (23 ­ 24 )). 20 – male tergum 8; 21 – male sternum 8; 22 – apex of male tergum 8; 23 – female tergum 8; 24 – female sternum 8. Scale bar 0.4 mm (20 ­ 21, 23 ­ 24), 0.2 mm (22)." httpUri="https://zenodo.org/record/156412/files/figure.png" pageId="3" pageNumber="4">Figs. 21, 24</figureCitation>
); male pronotum with broad medial impression that is half as wide as pronotum, deeper in the posterior half, postero­lateral portions of the impression with less dense microsculpture, without punctation and pubescence; male sternum 8 with broad apical emargination, apical portion of the sternum membranous (
<figureCitation id="F3821AAFFF9AFFF81E57FD6E1082D0D6" box="[373,461,644,670]" captionStart="FIGURES 20 ­ 24" captionStartId="9.[264,383,1550,1574]" captionTargetBox="[290,1295,579,1513]" captionTargetId="figure@9.[282,1306,569,1524]" captionTargetPageId="9" captionText="FIGURES 20 ­ 24. Abdominal segment 8 of Goniusa caseyi Gusarov, sp. n. (paratypes from Washington, D. C. (20 ­ 21); Aweme, Manitoba (22); and Stonewall, Manitoba (23 ­ 24 )). 20 – male tergum 8; 21 – male sternum 8; 22 – apex of male tergum 8; 23 – female tergum 8; 24 – female sternum 8. Scale bar 0.4 mm (20 ­ 21, 23 ­ 24), 0.2 mm (22)." httpUri="https://zenodo.org/record/156412/files/figure.png" pageId="3" pageNumber="4">Fig. 21</figureCitation>
); aedeagus with narrow but blunt apex (
<figureCitation id="F3821AAFFF9AFFF81C8BFD6E1544D0D6" box="[937,1035,644,670]" captionStart="FIGURES 25 ­ 29" captionStartId="10.[264,383,1768,1792]" captionTargetBox="[280,1310,701,1744]" captionTargetId="figure@10.[263,1323,689,1755]" captionTargetPageId="10" captionText="FIGURES 25 ­ 29. Median lobe of aedeagus of Goniusa caseyi Gusarov, sp. n. (paratypes from Washington, D. C. (25 ­ 26, 28 ­ 29); and Sherborn, Massachusetts (27 )). 25 – median lobe, parameral view; 26 ­ 27 – apex of median lobe, parameral view; 28 – median lobe, lateral view; 29 – apex of median lobe, lateral view. Scale bar 0.4 mm (25, 28), 0.2 mm (26 ­ 27, 29)." httpUri="https://zenodo.org/record/156413/files/figure.png" pageId="3" pageNumber="4">Figs. 25</figureCitation>
,
<figureCitation id="F3821AAFFF9AFFF81B3AFD6E1576D0D6" box="[1048,1081,644,670]" captionStart="FIGURES 48 ­ 51" captionStartId="15.[264,383,1288,1312]" captionTargetBox="[314,1270,697,1252]" captionTargetId="figure@15.[305,1283,689,1261]" captionTargetPageId="15" captionText="FIGURES 48 ­ 51. Median lobe of aedeagus of Goniusa alperti Kistner (male from West Point, Nebraska). 48 – median lobe, parameral view; 49 – apex of median lobe, parameral view; 50 – median lobe, lateral view; 51 – apex of median lobe, lateral view. Scale bar 0.4 mm (48, 50), 0.2 mm (49, 51)." httpUri="https://zenodo.org/record/156417/files/figure.png" pageId="3" pageNumber="4">48</figureCitation>
); medial lamellae of internal sac absent (
<figureCitation id="F3821AAFFF9AFFF81ECDFD46131FD08E" box="[495,592,684,710]" captionStart="FIGURES 30 ­ 36" captionStartId="11.[264,382,1632,1656]" captionTargetBox="[288,1303,337,1596]" captionTargetId="figure@11.[273,1314,326,1608]" captionTargetPageId="11" captionText="FIGURES 30 ­ 36. Genitalia of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (30 ­ 31); Texas (32 ­ 34); Sherborn, Massachusetts (35); and Stonewall, Manitoba (36 )). 30 – everted internal sac of aedeagus, parameral view; 31 – everted internal sac of aedeagus, lateral view; 32 – copulatory piece, apical view; 33 – copulatory piece, lateral view; 34 – copulatory piece, abparameral view; 35 ­ 36 – spermatheca. Scale bar 0.1 mm (32 ­ 34), 0.2 mm (30 ­ 31, 35 ­ 36)." httpUri="https://zenodo.org/record/156414/files/figure.png" pageId="3" pageNumber="4">Figs. 30</figureCitation>
,
<figureCitation id="F3821AAFFF9AFFF81D79FD461333D08E" box="[603,636,684,710]" captionStart="FIGURES 52 ­ 56" captionStartId="16.[264,383,1747,1771]" captionTargetBox="[292,1298,498,1731]" captionTargetId="figure@16.[281,1307,489,1736]" captionTargetPageId="16" captionText="FIGURES 52 ­ 56. Genitalia of Goniusa alperti Kistner (paratype from 10 mi. E of Coville, Washington (52); female from Tahoe City, California (53); female from 16 km NW of Granby, Colorado (54); holotype (55 ­ 56 )). 52 ­ 54 – spermatheca; 55 – everted internal sac of aedeagus, lateral view; 56 – everted internal sac of aedeagus, parameral view. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156418/files/figure.png" pageId="3" pageNumber="4">56</figureCitation>
); copulatory piece trough­shaped, with pointed apex (
<figureCitation id="F3821AAFFF9AFFF81BD2FD46101ED0A6" captionStart="FIGURES 30 ­ 36" captionStartId="11.[264,382,1632,1656]" captionTargetBox="[288,1303,337,1596]" captionTargetId="figure@11.[273,1314,326,1608]" captionTargetPageId="11" captionText="FIGURES 30 ­ 36. Genitalia of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (30 ­ 31); Texas (32 ­ 34); Sherborn, Massachusetts (35); and Stonewall, Manitoba (36 )). 30 – everted internal sac of aedeagus, parameral view; 31 – everted internal sac of aedeagus, lateral view; 32 – copulatory piece, apical view; 33 – copulatory piece, lateral view; 34 – copulatory piece, abparameral view; 35 ­ 36 – spermatheca. Scale bar 0.1 mm (32 ­ 34), 0.2 mm (30 ­ 31, 35 ­ 36)." httpUri="https://zenodo.org/record/156414/files/figure.png" pageId="3" pageNumber="4">Figs. 30­34</figureCitation>
); proximal portion of spermatheca with 2­3 coils (
<figureCitation id="F3821AAFFF9AFFF81CBCFD3E1566D0A6" box="[926,1065,724,750]" captionStart="FIGURES 30 ­ 36" captionStartId="11.[264,382,1632,1656]" captionTargetBox="[288,1303,337,1596]" captionTargetId="figure@11.[273,1314,326,1608]" captionTargetPageId="11" captionText="FIGURES 30 ­ 36. Genitalia of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (30 ­ 31); Texas (32 ­ 34); Sherborn, Massachusetts (35); and Stonewall, Manitoba (36 )). 30 – everted internal sac of aedeagus, parameral view; 31 – everted internal sac of aedeagus, lateral view; 32 – copulatory piece, apical view; 33 – copulatory piece, lateral view; 34 – copulatory piece, abparameral view; 35 ­ 36 – spermatheca. Scale bar 0.1 mm (32 ­ 34), 0.2 mm (30 ­ 31, 35 ­ 36)." httpUri="https://zenodo.org/record/156414/files/figure.png" pageId="3" pageNumber="4">Figs. 35­36</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="23A355A1FF9AFFF81E1AFD161216D666" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="6B06062AFF9AFFF81E1AFD16109BD1C6" blockId="3.[264,1324,284,1790]" pageId="3" pageNumber="4">
<taxonomicName id="ACB97DA9FF9AFFF81E1AFD1610D4D15D" ID-CoL="8VRPV" box="[312,411,764,789]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9AFFF81E1AFD1610D4D15D" box="[312,411,764,789]" italics="true" pageId="3" pageNumber="4">Goniusa</emphasis>
</taxonomicName>
differs from
<taxonomicName id="ACB97DA9FF9AFFF81D15FD161236D15E" ID-CoL="8VZSC" authority="Thomson, 1858" authorityName="Thomson" authorityYear="1858" box="[567,889,764,790]" class="Insecta" family="Staphylinidae" genus="Notothecta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9AFFF81D15FD1613F8D15D" box="[567,695,764,789]" italics="true" pageId="3" pageNumber="4">Notothecta</emphasis>
<bibRefCitation id="0F287BDBFF9AFFF81D9DFD161236D15E" author="Thomson" box="[703,889,764,790]" pageId="3" pageNumber="20" refString="Thomson, C. G. (1858) Forsok till uppstallning af Sveriges Staphyliner. Ofversigt af Kongl. Vetenskaps-Akademiens Forkhandlingar, 15, 27 - 40." type="journal article" year="1858">
Thomson,
<quantity id="AC41ABCFFF9AFFF81C1CFD161236D15E" box="[830,889,764,790]" metricMagnitude="1" metricUnit="m" metricValue="4.71932" pageId="3" pageNumber="4" unit="in" value="1858.0">1858</quantity>
</bibRefCitation>
</taxonomicName>
in having abdominal sternum 8 with numerous (30­60) macrosetae (
<figureCitation id="F3821AAFFF9AFFF81D5DFCCE125BD176" box="[639,788,804,830]" captionStart="FIGURES 20 ­ 24" captionStartId="9.[264,383,1550,1574]" captionTargetBox="[290,1295,579,1513]" captionTargetId="figure@9.[282,1306,569,1524]" captionTargetPageId="9" captionText="FIGURES 20 ­ 24. Abdominal segment 8 of Goniusa caseyi Gusarov, sp. n. (paratypes from Washington, D. C. (20 ­ 21); Aweme, Manitoba (22); and Stonewall, Manitoba (23 ­ 24 )). 20 – male tergum 8; 21 – male sternum 8; 22 – apex of male tergum 8; 23 – female tergum 8; 24 – female sternum 8. Scale bar 0.4 mm (20 ­ 21, 23 ­ 24), 0.2 mm (22)." httpUri="https://zenodo.org/record/156412/files/figure.png" pageId="3" pageNumber="4">Figs. 21, 24</figureCitation>
), male pronotum with broad medial impression, male sternum 8 with broad apical emargination, and different shape of the aedeagus and spermatheca.
</paragraph>
<paragraph id="6B06062AFF9AFFF81E1AFC761216D666" blockId="3.[264,1324,284,1790]" pageId="3" pageNumber="4">
<taxonomicName id="ACB97DA9FF9AFFF81E1AFC7610D4D1FD" ID-CoL="8VRPV" box="[312,411,924,949]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9AFFF81E1AFC7610D4D1FD" box="[312,411,924,949]" italics="true" pageId="3" pageNumber="4">Goniusa</emphasis>
</taxonomicName>
can be distinguished from similar­looking genera of the tribe
<taxonomicName id="ACB97DA9FF9AFFF81B4EFC76144AD1FE" ID-CoL="8VR8Z" box="[1132,1285,924,950]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Lomechusini">Lomechusini</taxonomicName>
by having longer and narrower mesothoracic process (
<figureCitation id="F3821AAFFF9AFFF81C46FC2E1288D196" box="[868,967,964,990]" captionStart="FIGURES 12 ­ 15" captionStartId="5.[264,383,1787,1811]" captionTargetBox="[289,1304,581,1774]" captionTargetId="figure@5.[272,1314,569,1776]" captionTargetPageId="5" captionText="FIGURES 12 ­ 15. Details of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 12 – pronotum; 13 – meso­ and metathorax, ventral view; 14 – right antenna; 15 – mentum. Scale bar 0.1 mm (15), 0.2 mm (12), 0.4 mm (13 ­ 14)." httpUri="https://zenodo.org/record/156410/files/figure.png" pageId="3" pageNumber="4">Figs. 13</figureCitation>
; 17­18); by shorter galea with its apical lobe only slightly projecting beyond the apex of lacinia (
<figureCitation id="F3821AAFFF9AFFF81B1AFC0615C0D64E" box="[1080,1167,1004,1030]" captionStart="FIGURES 6 ­ 11" captionStartId="4.[264,383,1469,1493]" captionTargetBox="[298,1287,339,1429]" captionTargetId="figure@4.[288,1299,329,1443]" captionTargetPageId="4" captionText="FIGURES 6 ­ 11. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 6 – prementum; 7 – hypopharynx; 8 – right maxilla, ventral view; 9 – right lacinia, dorsal view; 10 – right lacinia, ventral view; 11 – right galea, dorsal view. Scale bar 0.1 mm (6 ­ 7, 9 ­ 11), 0.2 mm (8)." httpUri="https://zenodo.org/record/156409/files/figure.png" pageId="3" pageNumber="4">Figs. 8</figureCitation>
; 16, 19) and stronger setae on its internal margin (
<figureCitation id="F3821AAFFF9AFFF81D9EFBFE1252D666" box="[700,797,1044,1070]" captionStart="FIGURES 6 ­ 11" captionStartId="4.[264,383,1469,1493]" captionTargetBox="[298,1287,339,1429]" captionTargetId="figure@4.[288,1299,329,1443]" captionTargetPageId="4" captionText="FIGURES 6 ­ 11. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 6 – prementum; 7 – hypopharynx; 8 – right maxilla, ventral view; 9 – right lacinia, dorsal view; 10 – right lacinia, ventral view; 11 – right galea, dorsal view. Scale bar 0.1 mm (6 ­ 7, 9 ­ 11), 0.2 mm (8)." httpUri="https://zenodo.org/record/156409/files/figure.png" pageId="3" pageNumber="4">Figs. 11</figureCitation>
; 16).
</paragraph>
</subSubSection>
<subSubSection id="23A355A1FF9AFFFD1E1AFBD1131ED0A6" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4" type="description">
<paragraph id="6B06062AFF9AFFF81E1AFBD1157BD636" blockId="3.[264,1324,284,1790]" pageId="3" pageNumber="4">
<emphasis id="59CDDA38FF9AFFF81E1AFBD1109FD61D" bold="true" box="[312,464,1083,1109]" pageId="3" pageNumber="4">Description.</emphasis>
Length
<quantity id="AC41ABCFFF9AFFF81D1AFBD6138CD61E" box="[568,707,1084,1110]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="4.2" metricValueMin="3.2" pageId="3" pageNumber="4" unit="mm" value="3.7" valueMax="4.2" valueMin="3.2">3.2­4.2 mm</quantity>
, pronotal width
<quantity id="AC41ABCFFF9AFFF81CA0FBD61565D61E" box="[898,1066,1084,1110]" metricMagnitude="-4" metricUnit="m" metricValue="9.55" metricValueMax="10.7" metricValueMin="8.4" pageId="3" pageNumber="4" unit="mm" value="0.955" valueMax="1.07" valueMin="0.84">0.84­1.07 mm</quantity>
. Body broad, reddish brown to dark brown with darker head and brownish red appendages.
</paragraph>
<paragraph id="6B06062AFF9AFFF81E1AFB66102DD4B6" blockId="3.[264,1324,284,1790]" pageId="3" pageNumber="4">
Head transverse; eyes large, temple length to eye length ratio 0.6­1.0; infraorbital carina very weak, complete, incomplete or absent altogether. Antennal article 2 as long as article 3, article 4 elongate, 5­10 slightly elongate or slightly transverse, apical article without coeloconic sensilla, longer than articles 9 and 10 combined (
<figureCitation id="F3821AAFFF9AFFF81B0EFAEE15CBD756" box="[1068,1156,1284,1310]" captionStart="FIGURES 12 ­ 15" captionStartId="5.[264,383,1787,1811]" captionTargetBox="[289,1304,581,1774]" captionTargetId="figure@5.[272,1314,569,1776]" captionTargetPageId="5" captionText="FIGURES 12 ­ 15. Details of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 12 – pronotum; 13 – meso­ and metathorax, ventral view; 14 – right antenna; 15 – mentum. Scale bar 0.1 mm (15), 0.2 mm (12), 0.4 mm (13 ­ 14)." httpUri="https://zenodo.org/record/156410/files/figure.png" pageId="3" pageNumber="4">Fig. 14</figureCitation>
). In dry specimens gaps between antennal articles inconspicuous. Labrum (
<figureCitation id="F3821AAFFF9AFFF81B2CFAC61519D70E" box="[1038,1110,1324,1350]" captionStart="FIGURES 1 ­ 5" captionStartId="2.[264,383,1522,1546]" captionTargetBox="[290,1296,439,1505]" captionTargetId="figure@2.[280,1306,425,1512]" captionTargetPageId="2" captionText="FIGURES 1 ­ 5. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/156408/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
) transverse, with straight anterior margin. Adoral surface of labrum (epipharynx) as in
<figureCitation id="F3821AAFFF9AFFF81B72FABE15D6D726" box="[1104,1177,1364,1390]" captionStart="FIGURES 1 ­ 5" captionStartId="2.[264,383,1522,1546]" captionTargetBox="[290,1296,439,1505]" captionTargetId="figure@2.[280,1306,425,1512]" captionTargetPageId="2" captionText="FIGURES 1 ­ 5. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/156408/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
. Mandibles (
<figureCitation id="F3821AAFFF9AFFF81E33FA961034D7DE" box="[273,379,1404,1430]" captionStart="FIGURES 1 ­ 5" captionStartId="2.[264,383,1522,1546]" captionTargetBox="[290,1296,439,1505]" captionTargetId="figure@2.[280,1306,425,1512]" captionTargetPageId="2" captionText="FIGURES 1 ­ 5. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 1 – labrum; 2 – epipharynx; 3 – left mandible, dorsal view; 4 – left mandible, ventral view; 5 – right mandible, dorsal view. Scale bar 0.1 mm." httpUri="https://zenodo.org/record/156408/files/figure.png" pageId="3" pageNumber="4">Figs. 3­5</figureCitation>
) broad, right mandible with small medial tooth; dorsal molar area with velvety patch consisting of very small denticles (poorly visible at 400x). Maxilla (
<figureCitation id="F3821AAFFF9AFFF81B4DFA4E15A9D7F6" box="[1135,1254,1444,1470]" captionStart="FIGURES 6 ­ 11" captionStartId="4.[264,383,1469,1493]" captionTargetBox="[298,1287,339,1429]" captionTargetId="figure@4.[288,1299,329,1443]" captionTargetPageId="4" captionText="FIGURES 6 ­ 11. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 6 – prementum; 7 – hypopharynx; 8 – right maxilla, ventral view; 9 – right lacinia, dorsal view; 10 – right lacinia, ventral view; 11 – right galea, dorsal view. Scale bar 0.1 mm (6 ­ 7, 9 ­ 11), 0.2 mm (8)." httpUri="https://zenodo.org/record/156409/files/figure.png" pageId="3" pageNumber="4">Figs. 8­11</figureCitation>
) with galea projecting slightly beyond apex of lacinia; apical lobe of galea covered with numerous fine and short setae; internal margin of galea with long subapical setae (
<figureCitation id="F3821AAFFF9AFFF81B58FA1E1580D446" box="[1146,1231,1524,1550]" captionStart="FIGURES 6 ­ 11" captionStartId="4.[264,383,1469,1493]" captionTargetBox="[298,1287,339,1429]" captionTargetId="figure@4.[288,1299,329,1443]" captionTargetPageId="4" captionText="FIGURES 6 ­ 11. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 6 – prementum; 7 – hypopharynx; 8 – right maxilla, ventral view; 9 – right lacinia, dorsal view; 10 – right lacinia, ventral view; 11 – right galea, dorsal view. Scale bar 0.1 mm (6 ­ 7, 9 ­ 11), 0.2 mm (8)." httpUri="https://zenodo.org/record/156409/files/figure.png" pageId="3" pageNumber="4">Fig. 11</figureCitation>
); apical 1/7 of lacinia with row of closely spaced spines, middle portion produced medially and covered with numerous setae (
<figureCitation id="F3821AAFFF9AFFF81D49F9AE13A9D416" box="[619,742,1604,1630]" captionStart="FIGURES 6 ­ 11" captionStartId="4.[264,383,1469,1493]" captionTargetBox="[298,1287,339,1429]" captionTargetId="figure@4.[288,1299,329,1443]" captionTargetPageId="4" captionText="FIGURES 6 ­ 11. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 6 – prementum; 7 – hypopharynx; 8 – right maxilla, ventral view; 9 – right lacinia, dorsal view; 10 – right lacinia, ventral view; 11 – right galea, dorsal view. Scale bar 0.1 mm (6 ­ 7, 9 ­ 11), 0.2 mm (8)." httpUri="https://zenodo.org/record/156409/files/figure.png" pageId="3" pageNumber="4">Figs. 9­10</figureCitation>
). Labium as in
<figureCitation id="F3821AAFFF9AFFF81C83F9AE1544D416" box="[929,1035,1604,1630]" captionStart="FIGURES 6 ­ 11" captionStartId="4.[264,383,1469,1493]" captionTargetBox="[298,1287,339,1429]" captionTargetId="figure@4.[288,1299,329,1443]" captionTargetPageId="4" captionText="FIGURES 6 ­ 11. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 6 – prementum; 7 – hypopharynx; 8 – right maxilla, ventral view; 9 – right lacinia, dorsal view; 10 – right lacinia, ventral view; 11 – right galea, dorsal view. Scale bar 0.1 mm (6 ­ 7, 9 ­ 11), 0.2 mm (8)." httpUri="https://zenodo.org/record/156409/files/figure.png" pageId="3" pageNumber="4">Figs. 6­7</figureCitation>
,
<figureCitation id="F3821AAFFF9AFFF81B35F9AE1578D416" box="[1047,1079,1604,1630]" captionStart="FIGURES 12 ­ 15" captionStartId="5.[264,383,1787,1811]" captionTargetBox="[289,1304,581,1774]" captionTargetId="figure@5.[272,1314,569,1776]" captionTargetPageId="5" captionText="FIGURES 12 ­ 15. Details of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 12 – pronotum; 13 – meso­ and metathorax, ventral view; 14 – right antenna; 15 – mentum. Scale bar 0.1 mm (15), 0.2 mm (12), 0.4 mm (13 ­ 14)." httpUri="https://zenodo.org/record/156410/files/figure.png" pageId="3" pageNumber="4">15</figureCitation>
; ligula split apically; medial area of prementum without pores but with 5 pseudopores, lateral areas with 2 pores, single setose pore and 12­16 pseudopores (
<figureCitation id="F3821AAFFF9AFFF81C6EF97E12DCD4E6" box="[844,915,1684,1710]" captionStart="FIGURES 6 ­ 11" captionStartId="4.[264,383,1469,1493]" captionTargetBox="[298,1287,339,1429]" captionTargetId="figure@4.[288,1299,329,1443]" captionTargetPageId="4" captionText="FIGURES 6 ­ 11. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 6 – prementum; 7 – hypopharynx; 8 – right maxilla, ventral view; 9 – right lacinia, dorsal view; 10 – right lacinia, ventral view; 11 – right galea, dorsal view. Scale bar 0.1 mm (6 ­ 7, 9 ­ 11), 0.2 mm (8)." httpUri="https://zenodo.org/record/156409/files/figure.png" pageId="3" pageNumber="4">Fig. 6</figureCitation>
). Hypopharyngeal lobes as in
<figureCitation id="F3821AAFFF9AFFF81BDFF97E1056D49E" captionStart="FIGURES 6 ­ 11" captionStartId="4.[264,383,1469,1493]" captionTargetBox="[298,1287,339,1429]" captionTargetId="figure@4.[288,1299,329,1443]" captionTargetPageId="4" captionText="FIGURES 6 ­ 11. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 6 – prementum; 7 – hypopharynx; 8 – right maxilla, ventral view; 9 – right lacinia, dorsal view; 10 – right lacinia, ventral view; 11 – right galea, dorsal view. Scale bar 0.1 mm (6 ­ 7, 9 ­ 11), 0.2 mm (8)." httpUri="https://zenodo.org/record/156409/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
. Labial palpus with setae,, and present. Mentum (
<figureCitation id="F3821AAFFF9AFFF81CEAF9561552D49E" box="[968,1053,1724,1750]" captionStart="FIGURES 12 ­ 15" captionStartId="5.[264,383,1787,1811]" captionTargetBox="[289,1304,581,1774]" captionTargetId="figure@5.[272,1314,569,1776]" captionTargetPageId="5" captionText="FIGURES 12 ­ 15. Details of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 12 – pronotum; 13 – meso­ and metathorax, ventral view; 14 – right antenna; 15 – mentum. Scale bar 0.1 mm (15), 0.2 mm (12), 0.4 mm (13 ­ 14)." httpUri="https://zenodo.org/record/156410/files/figure.png" pageId="3" pageNumber="4">Fig. 15</figureCitation>
) with concave anterior margin.
</paragraph>
<caption id="3FC656A2FF9DFFFF1E2AFA5710C9D475" httpUri="https://zenodo.org/record/156409/files/figure.png" pageId="4" pageNumber="5" targetBox="[298,1287,339,1429]" targetPageId="4">
<paragraph id="6B06062AFF9DFFFF1E2AFA5710C9D475" blockId="4.[264,1322,1469,1597]" pageId="4" pageNumber="5">
<emphasis id="59CDDA38FF9DFFFF1E2AFA5710F1D79D" bold="true" box="[264,446,1469,1493]" pageId="4" pageNumber="5">FIGURES 6­11.</emphasis>
Mouthparts of
<taxonomicName id="ACB97DA9FF9DFFFF1D5AFA5412CAD79D" authority="Gusarov" authorityName="Gusarov" box="[632,901,1470,1493]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="caseyi" status="sp. nov.">
<emphasis id="59CDDA38FF9DFFFF1D5AFA54126FD79D" box="[632,800,1470,1493]" italics="true" pageId="4" pageNumber="5">Goniusa caseyi</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="59CDDA38FF9DFFFF1CB0FA57129ED79D" bold="true" box="[914,977,1469,1493]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="42FE6743FF9DFFFF1CB0FA57129ED79D" box="[914,977,1469,1493]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratype from Aweme, Manitoba). 6 – prementum; 7 – hypopharynx; 8 – right maxilla, ventral view; 9 – right lacinia, dorsal view; 10 – right lacinia, ventral view; 11 – right galea, dorsal view. Scale bar 0.1 mm (6­7, 9­11), 0.2 mm (8).
</paragraph>
</caption>
<paragraph id="6B06062AFF9DFFFE1E1AF979126FD3B6" blockId="4.[264,1324,1683,1869]" lastBlockId="5.[264,1324,284,510]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
Pronotum (
<figureCitation id="F3821AAFFF9DFFFF1E99F9791353D4E5" box="[443,540,1683,1709]" captionStart="FIGURES 12 ­ 15" captionStartId="5.[264,383,1787,1811]" captionTargetBox="[289,1304,581,1774]" captionTargetId="figure@5.[272,1314,569,1776]" captionTargetPageId="5" captionText="FIGURES 12 ­ 15. Details of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 12 – pronotum; 13 – meso­ and metathorax, ventral view; 14 – right antenna; 15 – mentum. Scale bar 0.1 mm (15), 0.2 mm (12), 0.4 mm (13 ­ 14)." httpUri="https://zenodo.org/record/156410/files/figure.png" pageId="4" pageNumber="5">Figs. 12</figureCitation>
,
<figureCitation id="F3821AAFFF9DFFFF1D0AF9791322D4E5" box="[552,621,1683,1709]" captionStart="FIGURES 37 ­ 41" captionStartId="12.[264,382,1692,1716]" captionTargetBox="[301,1285,783,1653]" captionTargetId="figure@12.[288,1299,769,1665]" captionTargetPageId="12" captionText="FIGURES 37 ­ 41. Pronotum of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (37); and Stonewall, Manitoba (38 )) and G. alperti Kistner (holotype (39); female from San Bernardino Mts., California (40), paratype from 10 mi. E of Coville, Washington (41 )). 37, 39 – male pronotum; 38, 40 ­ 41 – female pronotum. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/156415/files/figure.png" pageId="4" pageNumber="5">37­41</figureCitation>
) strongly transverse, with microsetae directed posteriorly in midline; in lateral portions of disc microsetae directed posteriorly and obliquely laterally (
<typeStatus id="B402B888FF9DFFFF1E32F9091003D4B5" box="[272,332,1763,1789]" pageId="4" pageNumber="5">Type</typeStatus>
V,
<bibRefCitation id="0F287BDBFF9DFFFF1E57F9091332D4B5" author="Benick" box="[373,637,1763,1789]" pageId="4" pageNumber="19" refString="Benick, G. &amp; Lohse, G. A. (1974) 14. Tribus: Callicerini (Athetae). In: Freude, H., Harde, K. W. &amp; Lohse, G. A. (Eds.), Die Kafer Mitteleuropas. Band 5, Staphylinidae II (Hypocyphtinae und Aleocharinae). Pselaphidae. Goecke &amp; Evers Verlag, Krefeld, pp. 72 - 220." type="book chapter" year="1974">Benick &amp; Lohse 1974</bibRefCitation>
); macrosetae short; hypomera fully visible in lateral view. Meso­ and metasternum as in
<figureCitation id="F3821AAFFF9DFFFF1D5DF8E11396D56D" box="[639,729,1803,1829]" captionStart="FIGURES 12 ­ 15" captionStartId="5.[264,383,1787,1811]" captionTargetBox="[289,1304,581,1774]" captionTargetId="figure@5.[272,1314,569,1776]" captionTargetPageId="5" captionText="FIGURES 12 ­ 15. Details of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 12 – pronotum; 13 – meso­ and metathorax, ventral view; 14 – right antenna; 15 – mentum. Scale bar 0.1 mm (15), 0.2 mm (12), 0.4 mm (13 ­ 14)." httpUri="https://zenodo.org/record/156410/files/figure.png" pageId="4" pageNumber="5">Fig. 13</figureCitation>
, mesosternal process narrow, extending about ½ length of mesocoxal cavities, metasternal process short, mesosternum and mesosternal process not carinate medially; relative lengths of mesosternal process: isthmus: metasternal process in ratio of about 2:1:1; mesocoxal cavities margined posteriorly; mesocoxae narrowly separated. Medial macroseta of mesotibia inconspicuous, shorter than tibial width. Tarsal segmentation 4­5­5, metatarsal segment 1 slightly shorter than segment 2. One empodial seta, half as long as claws. Posterior margin of elytra slightly concave near postero­lateral angle. Wings fully developed.
</paragraph>
<caption id="3FC656A2FF9CFFFE1E2AF911124CD510" httpUri="https://zenodo.org/record/156410/files/figure.png" pageId="5" pageNumber="6" targetBox="[289,1304,581,1774]" targetPageId="5">
<paragraph id="6B06062AFF9CFFFE1E2AF911124CD510" blockId="5.[264,1323,1787,1880]" pageId="5" pageNumber="6">
<emphasis id="59CDDA38FF9CFFFE1E2AF9111083D55B" bold="true" box="[264,460,1787,1811]" pageId="5" pageNumber="6">FIGURES 12­15.</emphasis>
Details of
<taxonomicName id="ACB97DA9FF9CFFFE1D72F9161216D55A" authority="Gusarov" authorityName="Gusarov" box="[592,857,1787,1811]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="caseyi" status="sp. nov.">
<emphasis id="59CDDA38FF9CFFFE1D72F91613B9D55B" box="[592,758,1788,1811]" italics="true" pageId="5" pageNumber="6">Goniusa caseyi</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="59CDDA38FF9CFFFE1C44F91112ECD55B" bold="true" box="[870,931,1787,1811]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="42FE6743FF9CFFFE1C44F91112ECD55B" box="[870,931,1787,1811]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(paratype from Aweme, Manitoba). 12 – pronotum; 13 – meso­ and metathorax, ventral view; 14 – right antenna; 15 – mentum. Scale bar 0.1 mm (15), 0.2 mm (12), 0.4 mm (13­14).
</paragraph>
</caption>
<paragraph id="6B06062AFF9FFFFD1E1AFEF610AFD3E6" blockId="6.[264,1325,284,1870]" pageId="6" pageNumber="7">Abdominal terga 3­5 with moderate basal impressions. Tergum 7 1.1­1.2 times as long as tergum 6. Punctation on terga 6­7 sparser than on terga 3­5. Tergum 7 with wide white palisade fringe. Abdominal sterna with numerous semierect macrosetae, sternum 8 with 30­60 macrosetae.</paragraph>
<paragraph id="6B06062AFF9FFFFD1E1AFE56131ED0A6" blockId="6.[264,1325,284,1870]" pageId="6" pageNumber="7">
Male pronotum with broad medial impression that is half as wide as pronotum, deeper in posterior half, postero­lateral portions of the impression with less dense microsculpture, without punctation and pubescence. Compared to female, male pronotum matte, with stronger microsculpture and weaker punctation. Male sternum 8 with broad apical emargination, apical portion of the sternum membranous (
<figureCitation id="F3821AAFFF9FFFFD1CA4FDB61290D03E" box="[902,991,604,630]" captionStart="FIGURES 20 ­ 24" captionStartId="9.[264,383,1550,1574]" captionTargetBox="[290,1295,579,1513]" captionTargetId="figure@9.[282,1306,569,1524]" captionTargetPageId="9" captionText="FIGURES 20 ­ 24. Abdominal segment 8 of Goniusa caseyi Gusarov, sp. n. (paratypes from Washington, D. C. (20 ­ 21); Aweme, Manitoba (22); and Stonewall, Manitoba (23 ­ 24 )). 20 – male tergum 8; 21 – male sternum 8; 22 – apex of male tergum 8; 23 – female tergum 8; 24 – female sternum 8. Scale bar 0.4 mm (20 ­ 21, 23 ­ 24), 0.2 mm (22)." httpUri="https://zenodo.org/record/156412/files/figure.png" pageId="6" pageNumber="7">Fig. 21</figureCitation>
); aedeagus with narrow but blunt apex (
<figureCitation id="F3821AAFFF9FFFFD1EB6FD6E10B8D0D6" box="[404,503,644,670]" captionStart="FIGURES 25 ­ 29" captionStartId="10.[264,383,1768,1792]" captionTargetBox="[280,1310,701,1744]" captionTargetId="figure@10.[263,1323,689,1755]" captionTargetPageId="10" captionText="FIGURES 25 ­ 29. Median lobe of aedeagus of Goniusa caseyi Gusarov, sp. n. (paratypes from Washington, D. C. (25 ­ 26, 28 ­ 29); and Sherborn, Massachusetts (27 )). 25 – median lobe, parameral view; 26 ­ 27 – apex of median lobe, parameral view; 28 – median lobe, lateral view; 29 – apex of median lobe, lateral view. Scale bar 0.4 mm (25, 28), 0.2 mm (26 ­ 27, 29)." httpUri="https://zenodo.org/record/156413/files/figure.png" pageId="6" pageNumber="7">Figs. 25</figureCitation>
,
<figureCitation id="F3821AAFFF9FFFFD1D21FD6E136BD0D6" box="[515,548,644,670]" captionStart="FIGURES 48 ­ 51" captionStartId="15.[264,383,1288,1312]" captionTargetBox="[314,1270,697,1252]" captionTargetId="figure@15.[305,1283,689,1261]" captionTargetPageId="15" captionText="FIGURES 48 ­ 51. Median lobe of aedeagus of Goniusa alperti Kistner (male from West Point, Nebraska). 48 – median lobe, parameral view; 49 – apex of median lobe, parameral view; 50 – median lobe, lateral view; 51 – apex of median lobe, lateral view. Scale bar 0.4 mm (48, 50), 0.2 mm (49, 51)." httpUri="https://zenodo.org/record/156417/files/figure.png" pageId="6" pageNumber="7">48</figureCitation>
); medial lamellae of internal sac absent (
<figureCitation id="F3821AAFFF9FFFFD1B25FD6E1525D0D6" box="[1031,1130,644,670]" captionStart="FIGURES 30 ­ 36" captionStartId="11.[264,382,1632,1656]" captionTargetBox="[288,1303,337,1596]" captionTargetId="figure@11.[273,1314,326,1608]" captionTargetPageId="11" captionText="FIGURES 30 ­ 36. Genitalia of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (30 ­ 31); Texas (32 ­ 34); Sherborn, Massachusetts (35); and Stonewall, Manitoba (36 )). 30 – everted internal sac of aedeagus, parameral view; 31 – everted internal sac of aedeagus, lateral view; 32 – copulatory piece, apical view; 33 – copulatory piece, lateral view; 34 – copulatory piece, abparameral view; 35 ­ 36 – spermatheca. Scale bar 0.1 mm (32 ­ 34), 0.2 mm (30 ­ 31, 35 ­ 36)." httpUri="https://zenodo.org/record/156414/files/figure.png" pageId="6" pageNumber="7">Figs. 30</figureCitation>
,
<figureCitation id="F3821AAFFF9FFFFD1B55FD6E15D7D0D6" box="[1143,1176,644,670]" captionStart="FIGURES 52 ­ 56" captionStartId="16.[264,383,1747,1771]" captionTargetBox="[292,1298,498,1731]" captionTargetId="figure@16.[281,1307,489,1736]" captionTargetPageId="16" captionText="FIGURES 52 ­ 56. Genitalia of Goniusa alperti Kistner (paratype from 10 mi. E of Coville, Washington (52); female from Tahoe City, California (53); female from 16 km NW of Granby, Colorado (54); holotype (55 ­ 56 )). 52 ­ 54 – spermatheca; 55 – everted internal sac of aedeagus, lateral view; 56 – everted internal sac of aedeagus, parameral view. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156418/files/figure.png" pageId="6" pageNumber="7">56</figureCitation>
); copulatory piece trough­shaped, with pointed apex (
<figureCitation id="F3821AAFFF9FFFFD1DDBFD4612C8D08E" box="[761,903,684,710]" captionStart="FIGURES 30 ­ 36" captionStartId="11.[264,382,1632,1656]" captionTargetBox="[288,1303,337,1596]" captionTargetId="figure@11.[273,1314,326,1608]" captionTargetPageId="11" captionText="FIGURES 30 ­ 36. Genitalia of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (30 ­ 31); Texas (32 ­ 34); Sherborn, Massachusetts (35); and Stonewall, Manitoba (36 )). 30 – everted internal sac of aedeagus, parameral view; 31 – everted internal sac of aedeagus, lateral view; 32 – copulatory piece, apical view; 33 – copulatory piece, lateral view; 34 – copulatory piece, abparameral view; 35 ­ 36 – spermatheca. Scale bar 0.1 mm (32 ­ 34), 0.2 mm (30 ­ 31, 35 ­ 36)." httpUri="https://zenodo.org/record/156414/files/figure.png" pageId="6" pageNumber="7">Figs. 30­34</figureCitation>
); proximal portion of spermatheca with 2­3 coils (
<figureCitation id="F3821AAFFF9FFFFD1E9BFD3E130AD0A6" box="[441,581,724,750]" captionStart="FIGURES 30 ­ 36" captionStartId="11.[264,382,1632,1656]" captionTargetBox="[288,1303,337,1596]" captionTargetId="figure@11.[273,1314,326,1608]" captionTargetPageId="11" captionText="FIGURES 30 ­ 36. Genitalia of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (30 ­ 31); Texas (32 ­ 34); Sherborn, Massachusetts (35); and Stonewall, Manitoba (36 )). 30 – everted internal sac of aedeagus, parameral view; 31 – everted internal sac of aedeagus, lateral view; 32 – copulatory piece, apical view; 33 – copulatory piece, lateral view; 34 – copulatory piece, abparameral view; 35 ­ 36 – spermatheca. Scale bar 0.1 mm (32 ­ 34), 0.2 mm (30 ­ 31, 35 ­ 36)." httpUri="https://zenodo.org/record/156414/files/figure.png" pageId="6" pageNumber="7">Figs. 35­36</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="23A355A1FF9FFFFD1E1AFD11122CD176" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="6B06062AFF9FFFFD1E1AFD11122CD176" blockId="6.[264,1325,284,1870]" pageId="6" pageNumber="7">
<emphasis id="59CDDA38FF9FFFFD1E1AFD111096D15D" bold="true" box="[312,473,763,789]" pageId="6" pageNumber="7">
<typeStatus id="B402B888FF9FFFFD1E1AFD11103AD15D" box="[312,373,763,789]" pageId="6" pageNumber="7">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="ACB97DA9FF9FFFFD1EC5FD16124AD15E" authority="Gusarov" authorityName="Gusarov" box="[487,773,764,790]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="caseyi" status="sp. nov.">
<emphasis id="59CDDA38FF9FFFFD1EC5FD1613D5D15D" box="[487,666,764,789]" italics="true" pageId="6" pageNumber="7">Goniusa caseyi</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="59CDDA38FF9FFFFD1C32FD11121DD15D" bold="true" box="[784,850,763,789]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="42FE6743FF9FFFFD1C32FD11121DD15D" box="[784,850,763,789]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, by subsequent designation (see Discussion below), fixed under provisions of Article 70.3.
</paragraph>
</subSubSection>
<subSubSection id="23A355A1FF9FFFFC1E1AFCA11392D39E" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="discussion">
<paragraph id="6B06062AFF9FFFFD1E1AFCA110E2D7F6" blockId="6.[264,1325,284,1870]" pageId="6" pageNumber="7">
<emphasis id="59CDDA38FF9FFFFD1E1AFCA1108CD12D" bold="true" box="[312,451,843,869]" pageId="6" pageNumber="7">Discussion.</emphasis>
When proposing the new generic name
<taxonomicName id="ACB97DA9FF9FFFFD1C8EFCA615F2D12E" authority="Casey (1906)" authorityName="Casey" authorityYear="1906" box="[940,1213,844,870]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9FFFFD1C8EFCA61540D12D" box="[940,1039,844,869]" italics="true" pageId="6" pageNumber="7">Goniusa</emphasis>
,
<bibRefCitation id="0F287BDBFF9FFFFD1B3DFCA615F2D12E" author="Casey" box="[1055,1213,844,870]" pageId="6" pageNumber="19" refString="Casey, T. L. (1906) Observations of the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis, 16 (6), 125 - 434." type="journal article" year="1906">Casey (1906)</bibRefCitation>
</taxonomicName>
included in a single species,
<taxonomicName id="ACB97DA9FF9FFFFD1ED2FC9E122BD1C6" authority="LeConte, 1866" authorityName="LeConte" authorityYear="1866" box="[496,868,884,910]" class="Insecta" family="Staphylinidae" genus="Euryusa" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="obtusa">
<emphasis id="59CDDA38FF9FFFFD1ED2FC9E13E6D1C5" box="[496,681,884,909]" italics="true" pageId="6" pageNumber="7">Euryusa obtusa</emphasis>
<bibRefCitation id="0F287BDBFF9FFFFD1D90FC9E122BD1C6" author="LeConte" box="[690,868,884,910]" pageId="6" pageNumber="19" refString="LeConte, J. L. (1866) Additions to the coleopterous fauna of the United States. No. 1. Proceedings of the Academy of Sciences of Philadelphia, 18, 361 - 394." type="journal article" year="1866">LeConte, 1866</bibRefCitation>
</taxonomicName>
, and this species would be the
<typeStatus id="B402B888FF9FFFFD1BFAFC9E1445D1C6" box="[1240,1290,884,910]" pageId="6" pageNumber="7">type</typeStatus>
of the genus by monotypy. However, my examination of the
<typeStatus id="B402B888FF9FFFFD1CFEFC76150CD1FE" box="[988,1091,924,950]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
of
<taxonomicName id="ACB97DA9FF9FFFFD1B52FC761464D1FD" box="[1136,1323,924,949]" class="Insecta" family="Staphylinidae" genus="Euryusa" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="obtusa">
<emphasis id="59CDDA38FF9FFFFD1B52FC761464D1FD" box="[1136,1323,924,949]" italics="true" pageId="6" pageNumber="7">Euryusa obtusa</emphasis>
</taxonomicName>
(MCZ) and the specimens labeled as “
<taxonomicName id="ACB97DA9FF9FFFFD1DEFFC2E12C9D195" box="[717,902,964,989]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="obtusa">
<emphasis id="59CDDA38FF9FFFFD1DEFFC2E12C9D195" box="[717,902,964,989]" italics="true" pageId="6" pageNumber="7">Goniusa obtusa</emphasis>
</taxonomicName>
” in the Casey collection, as well as my analysis of characters mentioned by
<bibRefCitation id="0F287BDBFF9FFFFD1DCBFC0612C8D64E" author="Casey" box="[745,903,1004,1030]" pageId="6" pageNumber="19" refString="Casey, T. L. (1906) Observations of the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis, 16 (6), 125 - 434." type="journal article" year="1906">Casey (1906)</bibRefCitation>
demonstrated that Casey’s concept of the species does not agree with that of
<bibRefCitation id="0F287BDBFF9FFFFD1DCFFBFE12E3D666" author="LeConte" box="[749,940,1044,1070]" pageId="6" pageNumber="19" refString="LeConte, J. L. (1866) Additions to the coleopterous fauna of the United States. No. 1. Proceedings of the Academy of Sciences of Philadelphia, 18, 361 - 394." type="journal article" year="1866">LeConte (1866)</bibRefCitation>
. In fact the
<typeStatus id="B402B888FF9FFFFD1B1EFBFE15EDD666" box="[1084,1186,1044,1070]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
of
<taxonomicName id="ACB97DA9FF9FFFFD1BEBFBFE1019D61D" class="Insecta" family="Staphylinidae" genus="Euryusa" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="obtusa">
<emphasis id="59CDDA38FF9FFFFD1BEBFBFE1019D61D" italics="true" pageId="6" pageNumber="7">Euryusa obtusa</emphasis>
</taxonomicName>
belongs to the genus known as
<taxonomicName id="ACB97DA9FF9FFFFD1DF6FBD612B1D61E" authority="Fenyes, 1918" authorityName="Fenyes" authorityYear="1918" box="[724,1022,1084,1110]" class="Insecta" family="Staphylinidae" genus="Lypoglossa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9FFFFD1DF6FBD61216D61D" box="[724,857,1084,1109]" italics="true" pageId="6" pageNumber="7">Lypoglossa</emphasis>
<bibRefCitation id="0F287BDBFF9FFFFD1C42FBD612B1D61E" author="Fenyes" box="[864,1022,1084,1110]" pageId="6" pageNumber="19" refString="Fenyes, A. (1918) Coleoptera: Fam Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 A. L. Desmet-Verteneuil, Bruxelles, pp. 1 - 110." type="book chapter" year="1918">Fenyes, 1918</bibRefCitation>
</taxonomicName>
while Casey's specimens agree with current usage of the name &quot;
<taxonomicName id="ACB97DA9FF9FFFFD1DF3FB8E12C5D635" box="[721,906,1124,1149]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="obtusa">
<emphasis id="59CDDA38FF9FFFFD1DF3FB8E12C5D635" box="[721,906,1124,1149]" italics="true" pageId="6" pageNumber="7">Goniusa obtusa</emphasis>
</taxonomicName>
&quot;. The status of
<taxonomicName id="ACB97DA9FF9FFFFD1B6CFB8E1464D636" authority="LeConte" authorityName="LeConte" box="[1102,1323,1124,1150]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="obtusa">
<emphasis id="59CDDA38FF9FFFFD1B6CFB8E15F2D635" box="[1102,1213,1124,1149]" italics="true" pageId="6" pageNumber="7">E. obtusa</emphasis>
LeConte
</taxonomicName>
(nec Casey) will be discussed in a separate paper on the genus
<emphasis id="59CDDA38FF9FFFFD1CCBFB661539D6ED" box="[1001,1142,1164,1189]" italics="true" pageId="6" pageNumber="7">
<taxonomicName id="ACB97DA9FF9FFFFD1CCBFB66153DD6ED" box="[1001,1138,1164,1189]" class="Insecta" family="Staphylinidae" genus="Lypoglossa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">Lypoglossa</taxonomicName>
.
</emphasis>
Since the
<typeStatus id="B402B888FF9FFFFD1BDAFB661464D6EE" box="[1272,1323,1164,1190]" pageId="6" pageNumber="7">type</typeStatus>
species of the genus
<taxonomicName id="ACB97DA9FF9FFFFD1D21FB5E124DD686" authority="Casey, 1906" authorityName="Casey" authorityYear="1906" box="[515,770,1204,1230]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9FFFFD1D21FB5E1329D685" box="[515,614,1204,1229]" italics="true" pageId="6" pageNumber="7">Goniusa</emphasis>
<bibRefCitation id="0F287BDBFF9FFFFD1D52FB5E124DD686" author="Casey" box="[624,770,1204,1230]" pageId="6" pageNumber="19" refString="Casey, T. L. (1906) Observations of the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis, 16 (6), 125 - 434." type="journal article" year="1906">Casey, 1906</bibRefCitation>
</taxonomicName>
was misidentified by
<bibRefCitation id="0F287BDBFF9FFFFD1B2EFB5E15E4D686" author="Casey" box="[1036,1195,1204,1230]" pageId="6" pageNumber="19" refString="Casey, T. L. (1906) Observations of the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis, 16 (6), 125 - 434." type="journal article" year="1906">Casey (1906)</bibRefCitation>
the provisions of the Article 70.3 apply to it and the
<typeStatus id="B402B888FF9FFFFD1C30FB36120BD6BE" box="[786,836,1244,1270]" pageId="6" pageNumber="7">type</typeStatus>
species needs to be fixed (
<bibRefCitation id="0F287BDBFF9FFFFD1BA5FB361453D6BE" author="ICZN" box="[1159,1308,1244,1270]" pageId="6" pageNumber="19" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition. The International Trust for Zoological Nomenclature, London, xxix + 306 pp." type="book" year="1999">ICZN, 1999</bibRefCitation>
). To best serve stability and universality of nomenclature I select to fix as the
<typeStatus id="B402B888FF9FFFFD1B5CFAEE15FED756" box="[1150,1201,1284,1310]" pageId="6" pageNumber="7">type</typeStatus>
species of
<taxonomicName id="ACB97DA9FF9FFFFD1E2AFAC61024D70D" box="[264,363,1324,1349]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9FFFFD1E2AFAC61024D70D" box="[264,363,1324,1349]" italics="true" pageId="6" pageNumber="7">Goniusa</emphasis>
</taxonomicName>
the taxonomic species actually involved in the misidentification (Article 70.3.2). The
<typeStatus id="B402B888FF9FFFFD1E1FFABE103FD726" box="[317,368,1364,1390]" pageId="6" pageNumber="7">type</typeStatus>
species of the genus
<taxonomicName id="ACB97DA9FF9FFFFD1D4EFABE1228D726" authority="Casey, 1906" authorityName="Casey" authorityYear="1906" box="[620,871,1364,1390]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9FFFFD1D4EFABE139FD725" box="[620,720,1364,1389]" italics="true" pageId="6" pageNumber="7">Goniusa</emphasis>
<bibRefCitation id="0F287BDBFF9FFFFD1DF5FABE1228D726" author="Casey" box="[727,871,1364,1390]" pageId="6" pageNumber="19" refString="Casey, T. L. (1906) Observations of the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis, 16 (6), 125 - 434." type="journal article" year="1906">Casey, 1906</bibRefCitation>
</taxonomicName>
is now fixed as
<taxonomicName id="ACB97DA9FF9FFFFD1B0AFABE107FD7DE" authority="Gusarov" authorityName="Gusarov" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="caseyi" status="sp. nov.">
<emphasis id="59CDDA38FF9FFFFD1B0AFABE1592D725" box="[1064,1245,1364,1389]" italics="true" pageId="6" pageNumber="7">Goniusa caseyi</emphasis>
Gusarov
</taxonomicName>
,
<emphasis id="59CDDA38FF9FFFFD1E1CFA9110CED7DD" bold="true" box="[318,385,1403,1429]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="42FE6743FF9FFFFD1E1CFA9110CED7DD" box="[318,385,1403,1429]" pageId="6" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, misidentified as
<taxonomicName id="ACB97DA9FF9FFFFD1D75FA961286D7DE" authority="LeConte, 1866" authorityName="LeConte" authorityYear="1866" box="[599,969,1404,1430]" class="Insecta" family="Staphylinidae" genus="Euryusa" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="obtusa">
<emphasis id="59CDDA38FF9FFFFD1D75FA961240D7DD" box="[599,783,1404,1429]" italics="true" pageId="6" pageNumber="7">Euryusa obtusa</emphasis>
<bibRefCitation id="0F287BDBFF9FFFFD1C3AFA961286D7DE" author="LeConte" box="[792,969,1404,1430]" pageId="6" pageNumber="19" refString="LeConte, J. L. (1866) Additions to the coleopterous fauna of the United States. No. 1. Proceedings of the Academy of Sciences of Philadelphia, 18, 361 - 394." type="journal article" year="1866">
LeConte,
<quantity id="AC41ABCFFF9FFFFD1CACFA961286D7DE" box="[910,969,1404,1430]" metricMagnitude="1" metricUnit="m" metricValue="4.73964" pageId="6" pageNumber="7" unit="in" value="1866.0">1866</quantity>
</bibRefCitation>
</taxonomicName>
in the original description by
<bibRefCitation id="0F287BDBFF9FFFFD1E2AFA4E10E6D7F6" author="Casey" box="[264,425,1444,1470]" pageId="6" pageNumber="19" refString="Casey, T. L. (1906) Observations of the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis, 16 (6), 125 - 434." type="journal article" year="1906">Casey (1906)</bibRefCitation>
.
</paragraph>
<paragraph id="6B06062AFF9FFFFC1E1AFA2612FAD3CE" blockId="6.[264,1325,284,1870]" lastBlockId="7.[264,1323,284,470]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<bibRefCitation id="0F287BDBFF9FFFFD1E1AFA2610A7D7AE" author="Fenyes" box="[312,488,1484,1510]" pageId="6" pageNumber="19" refString="Fenyes, A. (1918) Coleoptera: Fam Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 A. L. Desmet-Verteneuil, Bruxelles, pp. 1 - 110." type="book chapter" year="1918">Fenyes (1918)</bibRefCitation>
placed
<taxonomicName id="ACB97DA9FF9FFFFD1D70FA2613F9D7AD" box="[594,694,1484,1509]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9FFFFD1D70FA2613F9D7AD" box="[594,694,1484,1509]" italics="true" pageId="6" pageNumber="7">Goniusa</emphasis>
</taxonomicName>
in the subtribe
<taxonomicName id="ACB97DA9FF9FFFFD1CA9FA2612BFD7AE" box="[907,1008,1484,1510]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subTribe" subTribe="Athetina">Athetina</taxonomicName>
and noted the similarity between
<taxonomicName id="ACB97DA9FF9FFFFD1E55FA1E1094D445" box="[375,475,1524,1549]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9FFFFD1E55FA1E1094D445" box="[375,475,1524,1549]" italics="true" pageId="6" pageNumber="7">Goniusa</emphasis>
</taxonomicName>
and the myrmecophilous genus
<taxonomicName id="ACB97DA9FF9FFFFD1C57FA1E101FD47E" authority="Thomson, 1858 (Fenyes 1920)" authorityName="Thomson, 1858 (Fenyes" authorityYear="1920" class="Insecta" family="Staphylinidae" genus="Notothecta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9FFFFD1C57FA1E12B9D445" box="[885,1014,1524,1549]" italics="true" pageId="6" pageNumber="7">Notothecta</emphasis>
<bibRefCitation id="0F287BDBFF9FFFFD1B21FA1E158ED446" author="Thomson" box="[1027,1217,1524,1550]" pageId="6" pageNumber="20" refString="Thomson, C. G. (1858) Forsok till uppstallning af Sveriges Staphyliner. Ofversigt af Kongl. Vetenskaps-Akademiens Forkhandlingar, 15, 27 - 40." type="journal article" year="1858">Thomson, 1858</bibRefCitation>
(
<bibRefCitation id="0F287BDBFF9FFFFD1BF4FA1E1008D47E" author="Fenyes" pageId="6" pageNumber="19" refString="Fenyes, A. (1920) Coleoptera. Fam. Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 B. L. Desmet-Verteneuil, Bruxelles. Pp. 111 - 414." type="book chapter" year="1920">Fenyes 1920</bibRefCitation>
)
</taxonomicName>
.
<bibRefCitation id="0F287BDBFF9FFFFD1E43F9F61343D47E" author="Kistner" box="[353,524,1564,1590]" pageId="6" pageNumber="19" refString="Kistner, D. H. (1976) Revision and Reclassification of the Genus Goniusa Casey with a Larval Description and Ant Host Records (Coleoptera: Staphylinidae). Sociobiology, 2 (1), 83 - 95." type="journal article" year="1976">Kistner (1976)</bibRefCitation>
disagreed with Fenyes and argued that based on &quot;the structure of the maxillae with their setigerous lacinia and galea&quot;
<taxonomicName id="ACB97DA9FF9FFFFD1C61F9AE12E8D415" box="[835,935,1604,1629]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9FFFFD1C61F9AE12E8D415" box="[835,935,1604,1629]" italics="true" pageId="6" pageNumber="7">Goniusa</emphasis>
</taxonomicName>
should be placed in Zyrini
<bibRefCitation id="0F287BDBFF9FFFFD1BCAF9AE103BD4CE" author="Bradley" pageId="6" pageNumber="19" refString="Bradley, J. C. (1930) A manual of the genera of beetles of America north of Mexico. Daw, Illston &amp; Co., Ithaca, 360 pp." type="book" year="1930">Bradley, 1930</bibRefCitation>
(spelled as
<taxonomicName id="ACB97DA9FF9FFFFD1D22F9861329D4CE" box="[512,614,1644,1670]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="tribe" tribe="Zyrasini">Zyrasini</taxonomicName>
), a junior synonym of
<taxonomicName id="ACB97DA9FF9FFFFD1C53F98615F2D4CE" authority="Fleming, 1821" authorityName="Fleming" authorityYear="1821" box="[881,1213,1644,1670]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="tribe" tribe="Lomechusini">
Lomechusini
<bibRefCitation id="0F287BDBFF9FFFFD1B33F98615F2D4CE" author="Fleming" box="[1041,1213,1644,1670]" pageId="6" pageNumber="19" refString="Fleming, J. (1821) Insecta. In: Supplement to the fourth, fifth and sixth editions of the Encyclopaedia Britannica, with preliminary dissertations on the history of the sciences. Vol. 5. Archibald Constable and Company, Edinburgh, pp. 41 - 56." type="book" year="1821">Fleming, 1821</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="0F287BDBFF9FFFFD1BEEF986108CD4E6" author="Newton" pageId="6" pageNumber="19" refString="Newton, A. F. &amp; Thayer, M. K. (1992) Current Classification and Family-Group Names in Staphyliniformia (Coleoptera). Fieldiana: Zoology, 67, 1 - 92." type="journal article" year="1992">Newton &amp; Thayer 1992</bibRefCitation>
). However, in
<taxonomicName id="ACB97DA9FF9FFFFD1D5BF97E1393D4E5" box="[633,732,1684,1709]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9FFFFD1D5BF97E1393D4E5" box="[633,732,1684,1709]" italics="true" pageId="6" pageNumber="7">Goniusa</emphasis>
</taxonomicName>
the galea is relatively short and it is only slightly projecting beyond the apex of lacinia (
<figureCitation id="F3821AAFFF9FFFFD1DC2F9561265D49E" box="[736,810,1724,1750]" captionStart="FIGURES 6 ­ 11" captionStartId="4.[264,383,1469,1493]" captionTargetBox="[298,1287,339,1429]" captionTargetId="figure@4.[288,1299,329,1443]" captionTargetPageId="4" captionText="FIGURES 6 ­ 11. Mouthparts of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 6 – prementum; 7 – hypopharynx; 8 – right maxilla, ventral view; 9 – right lacinia, dorsal view; 10 – right lacinia, ventral view; 11 – right galea, dorsal view. Scale bar 0.1 mm (6 ­ 7, 9 ­ 11), 0.2 mm (8)." httpUri="https://zenodo.org/record/156409/files/figure.png" pageId="6" pageNumber="7">Fig. 8</figureCitation>
) in comparison to the members of
<taxonomicName id="ACB97DA9FF9FFFFD1BFFF9561012D4B6" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="tribe" tribe="Lomechusini">Lomechusini</taxonomicName>
(
<emphasis id="59CDDA38FF9FFFFD1E4CF90E10D0D4B5" box="[366,415,1764,1789]" italics="true" pageId="6" pageNumber="7">e. g.</emphasis>
,
<taxonomicName id="ACB97DA9FF9FFFFD1E8CF90E1311D4B6" authority="Leach" authorityName="Leach" box="[430,606,1764,1790]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9FFFFD1E8CF90E1340D4B5" box="[430,527,1764,1789]" italics="true" pageId="6" pageNumber="7">Drusilla</emphasis>
Leach
</taxonomicName>
<emphasis id="59CDDA38FF9FFFFD1D44F90E1332D4B5" box="[614,637,1764,1789]" italics="true" pageId="6" pageNumber="7">in</emphasis>
<bibRefCitation id="0F287BDBFF9FFFFD1DA6F90E1205D4B6" author="Samouelle" box="[644,842,1764,1790]" lastPageId="6" lastPageNumber="20" pageId="6" pageNumber="19" refString="Samouelle, G. (1819) The Entomologist's useful compendium; or an introduction to the knowledge of British insects, comprising the best means of obtaining and preserving them, and a description of the apparatus generally used; together with the genera of Linne, and the modern method of arranging the classes Crustacea, Myriapoda, spiders, mites, and insects from their affinities and structure, according to the views of Dr. Leach. Also an explanation of the terms used in entomology; a calendar of the times of appearance, and usual situations of near 3000 species of British insects; with instructions for collecting and fitting up objects for the microscope. Thomas Boys, London, 496 pp." type="book" year="1819">Samouelle, 1819</bibRefCitation>
and
<taxonomicName id="ACB97DA9FF9FFFFD1CA0F90E15CCD4B6" authority="Stephens, 1835" authorityName="Stephens" authorityYear="1835" box="[898,1155,1764,1790]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9FFFFD1CA0F90E128AD4B5" box="[898,965,1764,1789]" italics="true" pageId="6" pageNumber="7">Zyras</emphasis>
<bibRefCitation id="0F287BDBFF9FFFFD1CEFF90E15CCD4B6" author="Stephens" box="[973,1155,1764,1790]" pageId="6" pageNumber="20" refString="Stephens, J. F. (1835) Illustrations of British entomology; or, a synopsis of indigenous insects; containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandibulata. 5. Baldwin and Cradock, London, pp. 369 - 448." type="journal article" year="1835">Stephens, 1835</bibRefCitation>
</taxonomicName>
;
<figureCitation id="F3821AAFFF9FFFFD1BADF90E1453D4B6" box="[1167,1308,1764,1790]" captionStart="FIGURES 16 ­ 19" captionStartId="7.[264,383,1700,1724]" captionTargetBox="[281,1304,573,1663]" captionTargetId="figure@7.[271,1316,561,1676]" captionTargetPageId="7" captionText="FIGURES 16 ­ 19. Details of Drusilla canaliculata (Fabricius) (female from Kiev, Ukraine (16); male from 15 km N of Magadan, Russia (17 )) and Zyras humeralis (Gravenhorst) (female from Briançon, France). 16, 19 – right maxilla, ventral view; 17 – meso­ and metathorax, ventral view; 18 – meso­ and metathorax, ventral view (setation not shown). Scale bar 0.2 mm (16), 0.4 mm (17, 19), 1.0 mm (18)." httpUri="https://zenodo.org/record/156411/files/figure.png" pageId="6" pageNumber="7">Figs. 16, 19</figureCitation>
). In
<taxonomicName id="ACB97DA9FF9FFFFD1E08F8E610C2D56D" box="[298,397,1804,1829]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9FFFFD1E08F8E610C2D56D" box="[298,397,1804,1829]" italics="true" pageId="6" pageNumber="7">Goniusa</emphasis>
</taxonomicName>
subapical setae of the internal margin of the galea are strong, and both galea and lacinia are very much like in other members of Athetini. Additionally, in
<taxonomicName id="ACB97DA9FF9FFFFD1BBEF8DE144FD505" box="[1180,1280,1844,1869]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9FFFFD1BBEF8DE144FD505" box="[1180,1280,1844,1869]" italics="true" pageId="6" pageNumber="7">Goniusa</emphasis>
</taxonomicName>
the mesosternal process is narrow and long (
<figureCitation id="F3821AAFFF9EFFFC1C23FEF61215D37E" box="[769,858,284,310]" captionStart="FIGURES 12 ­ 15" captionStartId="5.[264,383,1787,1811]" captionTargetBox="[289,1304,581,1774]" captionTargetId="figure@5.[272,1314,569,1776]" captionTargetPageId="5" captionText="FIGURES 12 ­ 15. Details of Goniusa caseyi Gusarov, sp. n. (paratype from Aweme, Manitoba). 12 – pronotum; 13 – meso­ and metathorax, ventral view; 14 – right antenna; 15 – mentum. Scale bar 0.1 mm (15), 0.2 mm (12), 0.4 mm (13 ­ 14)." httpUri="https://zenodo.org/record/156410/files/figure.png" pageId="7" pageNumber="8">Fig. 13</figureCitation>
) compared to the members of
<taxonomicName id="ACB97DA9FF9EFFFC1BFFFEF61012D316" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="tribe" tribe="Lomechusini">Lomechusini</taxonomicName>
(
<figureCitation id="F3821AAFFF9EFFFC1E4CFEAE10B7D316" box="[366,504,324,350]" captionStart="FIGURES 16 ­ 19" captionStartId="7.[264,383,1700,1724]" captionTargetBox="[281,1304,573,1663]" captionTargetId="figure@7.[271,1316,561,1676]" captionTargetPageId="7" captionText="FIGURES 16 ­ 19. Details of Drusilla canaliculata (Fabricius) (female from Kiev, Ukraine (16); male from 15 km N of Magadan, Russia (17 )) and Zyras humeralis (Gravenhorst) (female from Briançon, France). 16, 19 – right maxilla, ventral view; 17 – meso­ and metathorax, ventral view; 18 – meso­ and metathorax, ventral view (setation not shown). Scale bar 0.2 mm (16), 0.4 mm (17, 19), 1.0 mm (18)." httpUri="https://zenodo.org/record/156411/files/figure.png" pageId="7" pageNumber="8">Figs. 17­18</figureCitation>
). Presented arguments confirm the view of
<bibRefCitation id="0F287BDBFF9EFFFC1B24FEAE15FCD316" author="Fenyes" box="[1030,1203,324,350]" pageId="7" pageNumber="19" refString="Fenyes, A. (1918) Coleoptera: Fam Staphylinidae, subfam. Aleocharinae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 173 A. L. Desmet-Verteneuil, Bruxelles, pp. 1 - 110." type="book chapter" year="1918">Fenyes (1918)</bibRefCitation>
:
<taxonomicName id="ACB97DA9FF9EFFFC1BE5FEAE1464D315" box="[1223,1323,324,349]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9EFFFC1BE5FEAE1464D315" box="[1223,1323,324,349]" italics="true" pageId="7" pageNumber="8">Goniusa</emphasis>
</taxonomicName>
is not related to
<taxonomicName id="ACB97DA9FF9EFFFC1EE7FE861311D3CE" box="[453,606,364,390]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="tribe" tribe="Lomechusini">Lomechusini</taxonomicName>
and is a member of Athetini.
</paragraph>
<paragraph id="6B06062AFF9EFFFC1E1AFE7E1392D39E" blockId="7.[264,1323,284,470]" pageId="7" pageNumber="8">
Both known species of
<taxonomicName id="ACB97DA9FF9EFFFC1D69FE7E13E1D3E5" box="[587,686,404,429]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9EFFFC1D69FE7E13E1D3E5" box="[587,686,404,429]" italics="true" pageId="7" pageNumber="8">Goniusa</emphasis>
</taxonomicName>
are associated with the ants of the genus
<taxonomicName id="ACB97DA9FF9EFFFC1BB6FE7E15B6D3E5" box="[1172,1273,404,429]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="59CDDA38FF9EFFFC1BB6FE7E15B6D3E5" box="[1172,1273,404,429]" italics="true" pageId="7" pageNumber="8">Formica</emphasis>
</taxonomicName>
and have been collected inside the ant nests.
</paragraph>
</subSubSection>
</treatment>
</document>
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