281 lines
35 KiB
XML
281 lines
35 KiB
XML
<document id="562CEDECEAF67FF9541EA1A0CE20DF47" ID-DOI="10.5281/zenodo.156407" ID-GBIF-Dataset="f41d3d44-0e64-4ad1-841f-dea3d4a551e6" ID-ISSN="11755326" ID-Zenodo-Dep="156407" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459595625205" checkinUser="plazi" docAuthor="Gusarov, Vladimir I." docDate="2003" docId="E310B73CFF94FFEB1E2AFC7110D1D3CE" docLanguage="en" docName="zt00164.pdf" docOrigin="Zootaxa 164" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Goniusa alperti Kistner 1976" docType="treatment" docVersion="9" lastPageNumber="17" masterDocId="1F29CF44FF99FFFB1F22FFEA114FD248" masterDocTitle="A revision of the genus Goniusa Casey, 1906 (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="14" updateTime="1698202593239" updateUser="plazi">
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<mods:title id="D0E98276549CE4DFEC73A2964935DCD3">A revision of the genus Goniusa Casey, 1906 (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="43C62D68A20F63D422F2222990E1965D">Gusarov, Vladimir I.</mods:namePart>
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<mods:title id="18A7DACFA80A4614AB129EF422B7ABA6">Zootaxa</mods:title>
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<mods:part id="CCBE4A70EA85931A69ADCC933037CD55">
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<mods:date id="177F0E0E6E47C79387D54F7FAA368B28">2003</mods:date>
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<mods:classification id="288A1C209786F0D671E9232629BD3211">journal article</mods:classification>
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<mods:identifier id="A99FEAEF4CD4512E9776F59C2A30B8A2" type="DOI">10.5281/zenodo.156407</mods:identifier>
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<treatment id="E310B73CFF94FFEB1E2AFC7110D1D3CE" ID-DOI="http://doi.org/10.5281/zenodo.6273699" ID-Zenodo-Dep="6273699" LSID="urn:lsid:plazi:treatment:E310B73CFF94FFEB1E2AFC7110D1D3CE" httpUri="http://treatment.plazi.org/id/E310B73CFF94FFEB1E2AFC7110D1D3CE" lastPageId="16" lastPageNumber="17" pageId="13" pageNumber="14">
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<paragraph id="6B06062AFF94FFF61E2AFC71127FD1FD" blockId="13.[264,816,923,950]" box="[264,816,923,950]" pageId="13" pageNumber="14">
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<heading id="304EB146FF94FFF61E2AFC71127FD1FD" bold="true" box="[264,816,923,950]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
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<emphasis id="59CDDA38FF94FFF61E2AFC71127FD1FD" bold="true" box="[264,816,923,950]" pageId="13" pageNumber="14">
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<keyStep id="D04D1E8FFF94FFF61E2AFC71127FD1FD" box="[264,816,923,950]" pageId="13" pageNumber="14">
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<key id="1F28511BFF94FFF61E2AFC71127FD1FD" box="[264,816,923,950]" pageId="13" pageNumber="14">
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<keyLead id="D048A51FFF94FFF61E2AFC71127FD1FD" box="[264,816,923,950]" pageId="13" pageNumber="14">
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2.
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<taxonomicName id="ACB97DA9FF94FFF61E04FC7613C1D1FD" ID-CoL="3GY2D" authority="Kistner, 1976" authorityName="Kistner" authorityYear="1976" box="[294,654,923,950]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="alperti">
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||
<emphasis id="59CDDA38FF94FFF61E04FC7610ADD1FE" bold="true" box="[294,482,924,950]" italics="true" pageId="13" pageNumber="14">Goniusa alperti</emphasis>
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||
<bibRefCitation id="0F287BDBFF94FFF61ECBFC7113C1D1FD" author="Kistner" box="[489,654,923,949]" pageId="13" pageNumber="19" refString="Kistner, D. H. (1976) Revision and Reclassification of the Genus Goniusa Casey with a Larval Description and Ant Host Records (Coleoptera: Staphylinidae). Sociobiology, 2 (1), 83 - 95." type="journal article" year="1976">Kistner, 1976</bibRefCitation>
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||
</taxonomicName>
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||
(
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||
<figureCitation id="F3821AAFFF94FFF61DBCFC711267D1FD" box="[670,808,923,949]" captionStart-0="FIGURES 37 41" captionStart-1="FIGURES 42 47" captionStart-2="FIGURES 48 51" captionStart-3="FIGURES 52 56" captionStartId-0="12.[264,382,1692,1716]" captionStartId-1="14.[264,382,1769,1793]" captionStartId-2="15.[264,383,1288,1312]" captionStartId-3="16.[264,383,1747,1771]" captionTargetBox-0="[301,1285,783,1653]" captionTargetBox-1="[288,1298,540,1737]" captionTargetBox-2="[314,1270,697,1252]" captionTargetBox-3="[292,1298,498,1731]" captionTargetId-0="figure@12.[288,1299,769,1665]" captionTargetId-1="figure@14.[279,1307,529,1742]" captionTargetId-2="figure@15.[305,1283,689,1261]" captionTargetId-3="figure@16.[281,1307,489,1736]" captionTargetPageId-0="12" captionTargetPageId-1="14" captionTargetPageId-2="15" captionTargetPageId-3="16" captionText-0="FIGURES 37 41. Pronotum of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (37); and Stonewall, Manitoba (38 )) and G. alperti Kistner (holotype (39); female from San Bernardino Mts., California (40), paratype from 10 mi. E of Coville, Washington (41 )). 37, 39 – male pronotum; 38, 40 41 – female pronotum. Scale bar 0.5 mm." captionText-1="FIGURES 42 47. Abdominal segment 8 of Goniusa alperti Kistner (holotype (42 44); male from San Bernardino Mts., California (45); female from 16 km NW of Granby, Colorado (46 47 )). 42 – male tergum 8; 43 – male sternum 8; 44 45 – apex of male tergum 8; 46 – female tergum 8; 47 – female sternum 8. Scale bar 0.4 mm (42 43, 46 47), 0.2 mm (44 45)." captionText-2="FIGURES 48 51. Median lobe of aedeagus of Goniusa alperti Kistner (male from West Point, Nebraska). 48 – median lobe, parameral view; 49 – apex of median lobe, parameral view; 50 – median lobe, lateral view; 51 – apex of median lobe, lateral view. Scale bar 0.4 mm (48, 50), 0.2 mm (49, 51)." captionText-3="FIGURES 52 56. Genitalia of Goniusa alperti Kistner (paratype from 10 mi. E of Coville, Washington (52); female from Tahoe City, California (53); female from 16 km NW of Granby, Colorado (54); holotype (55 56 )). 52 54 – spermatheca; 55 – everted internal sac of aedeagus, lateral view; 56 – everted internal sac of aedeagus, parameral view. Scale bar 0.2 mm." httpUri-0="https://zenodo.org/record/156415/files/figure.png" httpUri-1="https://zenodo.org/record/156416/files/figure.png" httpUri-2="https://zenodo.org/record/156417/files/figure.png" httpUri-3="https://zenodo.org/record/156418/files/figure.png" pageId="13" pageNumber="14">Figs. 3956</figureCitation>
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||
)
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||
</keyLead>
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||
</key>
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||
</keyStep>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="23A355A1FF94FFF61E2AFC0D1331D656" pageId="13" pageNumber="14" type="reference_group">
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||
<paragraph id="6B06062AFF94FFF61E2AFC0D1331D656" blockId="13.[264,861,999,1054]" pageId="13" pageNumber="14">
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||
<treatmentCitationGroup id="4BA92104FF94FFF61E2AFC0D1331D656" pageId="13" pageNumber="14">
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||
<taxonomicName id="ACB97DA9FF94FFF61E2AFC0D1354D1B6" authority="Fenyes" authorityName="Fenyes" box="[264,539,999,1022]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="montana">
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<emphasis id="59CDDA38FF94FFF61E2AFC0D1089D1B6" box="[264,454,999,1022]" italics="true" pageId="13" pageNumber="14">Goniusa montana</emphasis>
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Fenyes
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</taxonomicName>
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,
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<emphasis id="59CDDA38FF94FFF61D04FC0D13C9D1B6" box="[550,646,999,1022]" italics="true" pageId="13" pageNumber="14">in litteris</emphasis>
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(manuscript name).
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<taxonomicName id="ACB97DA9FF94FFF61E2AFBED1335D656" ID-CoL="3GY2D" authority="Kistner, 1976: 89" authorityName="Kistner" authorityPageNumber="89" authorityYear="1976" box="[264,634,1031,1054]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="alperti">
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||
<emphasis id="59CDDA38FF94FFF61E2AFBED10FFD656" box="[264,432,1031,1054]" italics="true" pageId="13" pageNumber="14">Goniusa alperti</emphasis>
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||
<treatmentCitation id="EA18203BFF94FFF61E95FBED1335D656" author="Kistner" box="[439,634,1031,1054]" page="89" pageId="13" pageNumber="14" year="1976">
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||
<bibRefCitation id="0F287BDBFF94FFF61E95FBED1304D656" author="Kistner" box="[439,587,1031,1054]" pageId="13" pageNumber="19" refString="Kistner, D. H. (1976) Revision and Reclassification of the Genus Goniusa Casey with a Larval Description and Ant Host Records (Coleoptera: Staphylinidae). Sociobiology, 2 (1), 83 - 95." type="journal article" year="1976">Kistner, 1976</bibRefCitation>
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||
: 89
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||
</treatmentCitation>
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||
</taxonomicName>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="23A355A1FF94FFF61E2AFBA413A6D469" pageId="13" pageNumber="14" type="materials_examined">
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||
<paragraph id="6B06062AFF94FFF61E2AFBA41237D6D9" blockId="13.[264,1324,1102,1849]" pageId="13" pageNumber="14">
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||
<emphasis id="59CDDA38FF94FFF61E2AFBA410F3D620" bold="true" box="[264,444,1102,1128]" pageId="13" pageNumber="14">
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||
<typeStatus id="B402B888FF94FFF61E2AFBA4100CD620" box="[264,323,1102,1128]" pageId="13" pageNumber="14">Type</typeStatus>
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||
material.
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||
</emphasis>
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||
<typeStatus id="B402B888FF94FFF61EECFBA51374D621" box="[462,571,1103,1129]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
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||
of
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||
<taxonomicName id="ACB97DA9FF94FFF61D40FBA5139AD620" box="[610,725,1103,1128]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="alperti">
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||
<emphasis id="59CDDA38FF94FFF61D40FBA5139AD620" box="[610,725,1103,1128]" italics="true" pageId="13" pageNumber="14">G. alperti</emphasis>
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||
</taxonomicName>
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||
:,
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||
<emphasis id="59CDDA38FF94FFF61C2DFBA412BBD620" bold="true" box="[783,1012,1102,1128]" pageId="13" pageNumber="14">
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||
<collectingCountry id="13AE46BAFF94FFF61C2DFBA412BBD620" box="[783,1012,1102,1128]" name="United States of America" pageId="13" pageNumber="14">UNITED STATES</collectingCountry>
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||
</emphasis>
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||
:
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||
<emphasis id="59CDDA38FF94FFF61B29FBA415D0D620" bold="true" box="[1035,1183,1102,1128]" pageId="13" pageNumber="14">Washington</emphasis>
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||
: Whatcom Co.: Blaine, ant nest (G.Alpert),
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||
<date id="1F0720EAFF94FFF61DADFB9D124DD6D9" box="[655,770,1143,1169]" pageId="13" pageNumber="14" value="1971-05-10">10.v.1971</date>
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||
(
|
||
<collectionCode id="0DA89EEFFF94FFF61C30FB9D1223D6D9" box="[786,876,1143,1169]" pageId="13" pageNumber="14">DHKC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6B06062AFF94FFF61E1AFB751311D741" blockId="13.[264,1324,1102,1849]" pageId="13" pageNumber="14">
|
||
<typeStatus id="B402B888FF94FFF61E1AFB7510E5D6F1" box="[312,426,1183,1209]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
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||
of
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||
<taxonomicName id="ACB97DA9FF94FFF61EF2FB75130DD6F0" box="[464,578,1183,1208]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="alperti">
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||
<emphasis id="59CDDA38FF94FFF61EF2FB75130DD6F0" box="[464,578,1183,1208]" italics="true" pageId="13" pageNumber="14">G. alperti</emphasis>
|
||
</taxonomicName>
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||
:
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||
<emphasis id="59CDDA38FF94FFF61D7BFB741271D6F0" bold="true" box="[601,830,1182,1208]" pageId="13" pageNumber="14">
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||
<collectingCountry id="13AE46BAFF94FFF61D7BFB741271D6F0" box="[601,830,1182,1208]" name="United States of America" pageId="13" pageNumber="14">UNITED STATES</collectingCountry>
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||
</emphasis>
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||
:
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<emphasis id="59CDDA38FF94FFF61C76FB7412A6D6F0" bold="true" box="[852,1001,1182,1208]" pageId="13" pageNumber="14">Washington</emphasis>
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||
: Clallam Co.:,
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||
<quantity id="AC41ABCFFF94FFF61BE5FB751442D6F1" box="[1223,1293,1183,1209]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="13" pageNumber="14" unit="mi" value="10.0">10 mi</quantity>
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||
. E Coville, ant nest (G.Alpert),
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||
<date id="1F0720EAFF94FFF61D42FB2D1391D6A9" box="[608,734,1223,1249]" pageId="13" pageNumber="14" value="1971-04-11">11.iv.1971</date>
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||
; Whatcom Co.:, Bellingham, ant nest (G.Alpert),
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||
<date id="1F0720EAFF94FFF61E1DFB0510FDD741" box="[319,434,1263,1289]" pageId="13" pageNumber="14" value="1971-05-11">11.v.1971</date>
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||
(all
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||
<collectionCode id="0DA89EEFFF94FFF61EDBFB05131DD741" box="[505,594,1263,1289]" pageId="13" pageNumber="14">DHKC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6B06062AFF94FFF61E1AFAFC13A6D469" blockId="13.[264,1324,1102,1849]" pageId="13" pageNumber="14">
|
||
<emphasis id="59CDDA38FF94FFF61E1AFAFC127ED778" bold="true" box="[312,817,1302,1328]" pageId="13" pageNumber="14">
|
||
Additional material.
|
||
<collectingCountry id="13AE46BAFF94FFF61D68FAFC127ED778" box="[586,817,1302,1328]" name="United States of America" pageId="13" pageNumber="14">UNITED STATES</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="59CDDA38FF94FFF61C6EFAFC1283D778" bold="true" box="[844,972,1302,1328]" pageId="13" pageNumber="14">California</emphasis>
|
||
: San Bernardino Co.: 2,, San Bernardino Mts.,
|
||
<quantity id="AC41ABCFFF94FFF61D62FAD51332D711" box="[576,637,1343,1369]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="13" pageNumber="14" unit="mi" value="2.0">2 mi</quantity>
|
||
. NW Fawnskin, host:
|
||
<taxonomicName id="ACB97DA9FF94FFF61C80FAD51599D710" box="[930,1238,1343,1368]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
|
||
<emphasis id="59CDDA38FF94FFF61C80FAD51599D710" box="[930,1238,1343,1368]" italics="true" pageId="13" pageNumber="14">Formica haemorrhoidalis</emphasis>
|
||
</taxonomicName>
|
||
(B. & E.MacKay) (all
|
||
<collectionCode id="0DA89EEFFF94FFF61EF9FA8D137BD7C9" box="[475,564,1383,1409]" pageId="13" pageNumber="14">DHKC</collectionCode>
|
||
); Placer Co.:, Tahoe City;
|
||
<emphasis id="59CDDA38FF94FFF61CB7FA8C1545D7C8" bold="true" box="[917,1034,1382,1408]" pageId="13" pageNumber="14">Colorado</emphasis>
|
||
: Ouray Co.:, Ouray, 9000' (mounted with a
|
||
<taxonomicName id="ACB97DA9FF94FFF61D37FA651335D7E0" box="[533,634,1423,1448]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF94FFF61D37FA651335D7E0" box="[533,634,1423,1448]" italics="true" pageId="13" pageNumber="14">Formica</emphasis>
|
||
</taxonomicName>
|
||
specimen) (Wickham), vii (all
|
||
<collectionCode id="0DA89EEFFF94FFF61CD5FA651506D7E1" box="[1015,1097,1423,1449]" pageId="13" pageNumber="14">CASC</collectionCode>
|
||
); Grand Co.:,
|
||
<quantity id="AC41ABCFFF94FFF61A2FFA651062D799" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="13" pageNumber="14" unit="km" value="16.0">16 km</quantity>
|
||
NW Granby, Arapaho National Forest, Hwy. 125,
|
||
<geoCoordinate id="0E8D60EDFF94FFF61CA3FA5D1290D799" box="[897,991,1463,1489]" direction="north" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="40.2">40º12'N</geoCoordinate>
|
||
<geoCoordinate id="0E8D60EDFF94FFF61CC5FA5D1515D799" box="[999,1114,1463,1489]" direction="west" orientation="longitude" pageId="13" pageNumber="14" precision="925" value="-106.05">106º03'W</geoCoordinate>
|
||
,
|
||
<quantity id="AC41ABCFFF94FFF61B44FA5D15F5D799" box="[1126,1210,1463,1489]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="13" pageNumber="14" unit="m" value="2600.0">2600m</quantity>
|
||
,
|
||
<taxonomicName id="ACB97DA9FF94FFF61BE4FA5D1464D798" box="[1222,1323,1463,1488]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF94FFF61BE4FA5D1464D798" box="[1222,1323,1463,1488]" italics="true" pageId="13" pageNumber="14">Formica</emphasis>
|
||
</taxonomicName>
|
||
nest (mounted with a
|
||
<taxonomicName id="ACB97DA9FF94FFF61D24FA351324D7B0" box="[518,619,1503,1528]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF94FFF61D24FA351324D7B0" box="[518,619,1503,1528]" italics="true" pageId="13" pageNumber="14">Formica</emphasis>
|
||
</taxonomicName>
|
||
specimen) (V.I.Gusarov),
|
||
<date id="1F0720EAFF94FFF61C83FA351560D7B1" box="[929,1071,1503,1529]" pageId="13" pageNumber="14" value="2000-08-23">23.viii.2000</date>
|
||
(
|
||
<collectionCode id="0DA89EEFFF94FFF61B1CFA3515D9D7B1" box="[1086,1174,1503,1529]" pageId="13" pageNumber="14">KSEM</collectionCode>
|
||
);
|
||
<emphasis id="59CDDA38FF94FFF61B88FA34146DD7B0" bold="true" box="[1194,1314,1502,1528]" pageId="13" pageNumber="14">Nebraska</emphasis>
|
||
: Cuming Co.:,, West Point (
|
||
<collectionCode id="0DA89EEFFF94FFF61DA3F9ED1392D469" LSID="urn:lsid:biocol.org:col:34795" box="[641,733,1543,1569]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="13" pageNumber="14">FMNH</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23A355A1FF94FFF61E1AF9C4132BD489" pageId="13" pageNumber="14" type="diagnosis">
|
||
<paragraph id="6B06062AFF94FFF61E1AF9C4132BD489" blockId="13.[264,1324,1102,1849]" pageId="13" pageNumber="14">
|
||
<emphasis id="59CDDA38FF94FFF61E1AF9C410F8D400" bold="true" box="[312,439,1582,1608]" pageId="13" pageNumber="14">Diagnosis.</emphasis>
|
||
<taxonomicName id="ACB97DA9FF94FFF61EEAF9C513CDD400" box="[456,642,1583,1608]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="alperti">
|
||
<emphasis id="59CDDA38FF94FFF61EEAF9C513CDD400" box="[456,642,1583,1608]" italics="true" pageId="13" pageNumber="14">Goniusa alperti</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from
|
||
<taxonomicName id="ACB97DA9FF94FFF61CE7F9C5157DD400" box="[965,1074,1583,1608]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="caseyi">
|
||
<emphasis id="59CDDA38FF94FFF61CE7F9C5157DD400" box="[965,1074,1583,1608]" italics="true" pageId="13" pageNumber="14">G. caseyi</emphasis>
|
||
</taxonomicName>
|
||
by having pronotum with lateral sides converging anteriad, reaching maximum width behind the middle (
|
||
<figureCitation id="F3821AAFFF94FFF61BD2F9BD101FD4D1" captionStart="FIGURES 37 41" captionStartId="12.[264,382,1692,1716]" captionTargetBox="[301,1285,783,1653]" captionTargetId="figure@12.[288,1299,769,1665]" captionTargetPageId="12" captionText="FIGURES 37 41. Pronotum of Goniusa caseyi Gusarov, sp. n. (paratypes from Aweme, Manitoba (37); and Stonewall, Manitoba (38 )) and G. alperti Kistner (holotype (39); female from San Bernardino Mts., California (40), paratype from 10 mi. E of Coville, Washington (41 )). 37, 39 – male pronotum; 38, 40 41 – female pronotum. Scale bar 0.5 mm." httpUri="https://zenodo.org/record/156415/files/figure.png" pageId="13" pageNumber="14">Figs. 3941</figureCitation>
|
||
); antennal segments 510 slightly transverse; smaller body and short apical process of median lobe (
|
||
<figureCitation id="F3821AAFFF94FFF61EEAF94D1319D489" box="[456,598,1703,1729]" captionStart="FIGURES 48 51" captionStartId="15.[264,383,1288,1312]" captionTargetBox="[314,1270,697,1252]" captionTargetId="figure@15.[305,1283,689,1261]" captionTargetPageId="15" captionText="FIGURES 48 51. Median lobe of aedeagus of Goniusa alperti Kistner (male from West Point, Nebraska). 48 – median lobe, parameral view; 49 – apex of median lobe, parameral view; 50 – median lobe, lateral view; 51 – apex of median lobe, lateral view. Scale bar 0.4 mm (48, 50), 0.2 mm (49, 51)." httpUri="https://zenodo.org/record/156417/files/figure.png" pageId="13" pageNumber="14">Figs. 49, 51</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23A355A1FF94FFF41E1AF92413EDD575" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
|
||
<paragraph id="6B06062AFF94FFF61E1AF9241391D571" blockId="13.[264,1324,1102,1849]" pageId="13" pageNumber="14">
|
||
<emphasis id="59CDDA38FF94FFF61E1AF924109FD4A0" bold="true" box="[312,464,1742,1768]" pageId="13" pageNumber="14">Description.</emphasis>
|
||
Length 3.24.0 mm. Head brownish black, pronotum and abdomen dark brown, often with lighter segments 34, elytra reddish brown, legs and antennae brownish red, mouthparts yellow to brownish red.
|
||
</paragraph>
|
||
<paragraph id="6B06062AFF97FFF51E1AFEF61330D39E" blockId="14.[264,1324,284,470]" pageId="14" pageNumber="15">In females head surface glossy, with weak isodiametric microsculpture, with fine punctation, distance between punctures equals 2 times their diameter. In males head surface matte, with strong isodiametric microsculpture, with weak, sparse and poorly visible punctation. Temple length to eye length ratio 0.60.9. Antennal article 4 slightly elongate, articles 510 slightly transverse.</paragraph>
|
||
<caption id="3FC656A2FF97FFF51E2AF90312A1D520" httpUri="https://zenodo.org/record/156416/files/figure.png" pageId="14" pageNumber="15" targetBox="[288,1298,540,1737]" targetPageId="14">
|
||
<paragraph id="6B06062AFF97FFF51E2AF90312A1D520" blockId="14.[264,1323,1769,1896]" pageId="14" pageNumber="15">
|
||
<emphasis id="59CDDA38FF97FFF51E2AF9031085D549" bold="true" box="[264,458,1769,1793]" pageId="14" pageNumber="15">FIGURES 4247.</emphasis>
|
||
Abdominal segment 8 of
|
||
<taxonomicName id="ACB97DA9FF97FFF51DC8F90012A6D548" authority="Kistner" authorityName="Kistner" box="[746,1001,1769,1793]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="alperti">
|
||
<emphasis id="59CDDA38FF97FFF51DC8F90012DCD549" box="[746,915,1770,1793]" italics="true" pageId="14" pageNumber="15">Goniusa alperti</emphasis>
|
||
Kistner
|
||
</taxonomicName>
|
||
(holotype (4244); male from San Bernardino Mts., California (45); female from 16 km NW of Granby, Colorado (4647)). 42 – male tergum 8; 43 – male sternum 8; 4445 – apex of male tergum 8; 46 – female tergum 8; 47 – female sternum 8. Scale bar 0.4 mm (4243, 4647), 0.2 mm (4445).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6B06062AFF96FFF41E1AFEF612E7D006" blockId="15.[264,1324,284,590]" pageId="15" pageNumber="16">
|
||
Pronotum strongly transverse, 1.6 times as wide as head, width
|
||
<quantity id="AC41ABCFFF96FFF41B0FFEF61599D37E" box="[1069,1238,284,310]" metricMagnitude="-4" metricUnit="m" metricValue="9.25" metricValueMax="10.1" metricValueMin="8.4" pageId="15" pageNumber="16" unit="mm" value="0.925" valueMax="1.01" valueMin="0.84">0.841.01 mm</quantity>
|
||
, length
|
||
<quantity id="AC41ABCFFF96FFF41E2AFEAE10FED316" box="[264,433,324,350]" metricMagnitude="-4" metricUnit="m" metricValue="5.85" metricValueMax="6.6" metricValueMin="5.1" pageId="15" pageNumber="16" unit="mm" value="0.585" valueMax="0.66" valueMin="0.51">0.510.66 mm</quantity>
|
||
, width to length ratio 1.6; in females surface glossy, with weak isodiametric microsculpture; punctation as fine as on head, slightly asperate, distance between punctures equal to 2 times their diameter; in males surface matte, with strong isodiametric microsculpture, with fine and poorly visible punctation. Elytra slightly wider (
|
||
<quantity id="AC41ABCFFF96FFF41B9BFE561077D3B6" metricMagnitude="-3" metricUnit="m" metricValue="1.08" metricValueMax="1.17" metricValueMin="0.99" pageId="15" pageNumber="16" unit="mm" value="1.08" valueMax="1.17" valueMin="0.99">0.991.17 mm</quantity>
|
||
) and longer (
|
||
<quantity id="AC41ABCFFF96FFF41EFAFE0E13CBD3B6" box="[472,644,484,510]" metricMagnitude="-4" metricUnit="m" metricValue="7.15" metricValueMax="7.9" metricValueMin="6.4" pageId="15" pageNumber="16" unit="mm" value="0.715" valueMax="0.79" valueMin="0.64">0.640.79 mm</quantity>
|
||
, measured from humeral angle) than pronotum (pronotal length to elytral length ratio 0.8), 1.5 times as wide as long, surface glossy, with weak isodiametric microsculpture and punctation as on pronotum.
|
||
</paragraph>
|
||
<caption id="3FC656A2FF96FFF41E2AFAE210DCD7CF" httpUri="https://zenodo.org/record/156417/files/figure.png" pageId="15" pageNumber="16" targetBox="[314,1270,697,1252]" targetPageId="15">
|
||
<paragraph id="6B06062AFF96FFF41E2AFAE210DCD7CF" blockId="15.[264,1322,1288,1415]" pageId="15" pageNumber="16">
|
||
<emphasis id="59CDDA38FF96FFF41E2AFAE21082D768" bold="true" box="[264,461,1288,1312]" pageId="15" pageNumber="16">FIGURES 4851.</emphasis>
|
||
Median lobe of aedeagus of
|
||
<taxonomicName id="ACB97DA9FF96FFF41C3FFAE3156FD757" authority="Kistner" authorityName="Kistner" box="[797,1056,1288,1312]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="alperti">
|
||
<emphasis id="59CDDA38FF96FFF41C3FFAE31287D768" box="[797,968,1289,1312]" italics="true" pageId="15" pageNumber="16">Goniusa alperti</emphasis>
|
||
Kistner
|
||
</taxonomicName>
|
||
(male from West Point, Nebraska). 48 – median lobe, parameral view; 49 – apex of median lobe, parameral view; 50 – median lobe, lateral view; 51 – apex of median lobe, lateral view. Scale bar 0.4 mm (48, 50), 0.2 mm (49, 51).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="6B06062AFF96FFF41E1AFA5115B6D46D" blockId="15.[264,1324,1467,1893]" pageId="15" pageNumber="16">Abdominal terga glossy, with fine and often poorly visible transverse microsculpture, with fine punctation, distance between punctures equals 12 times their diameter on terga 35 and 36 times on terga 67. Apical margin of tergum 7 with white palisade fringe.</paragraph>
|
||
<paragraph id="6B06062AFF96FFF41E1AF9D913B2D4A5" blockId="15.[264,1324,1467,1893]" pageId="15" pageNumber="16">
|
||
In males head with broad medial impression; pronotum with broad median impression that is half as wide as pronotum, deeper in posterior half, posterolateral portions of the impression with less dense microsculpture, without punctation and pubescence. Male sternum 8 with wide apical emargination (
|
||
<figureCitation id="F3821AAFFF96FFF41DF6F9411263D48D" box="[724,812,1707,1733]" captionStart="FIGURES 42 47" captionStartId="14.[264,382,1769,1793]" captionTargetBox="[288,1298,540,1737]" captionTargetId="figure@14.[279,1307,529,1742]" captionTargetPageId="14" captionText="FIGURES 42 47. Abdominal segment 8 of Goniusa alperti Kistner (holotype (42 44); male from San Bernardino Mts., California (45); female from 16 km NW of Granby, Colorado (46 47 )). 42 – male tergum 8; 43 – male sternum 8; 44 45 – apex of male tergum 8; 46 – female tergum 8; 47 – female sternum 8. Scale bar 0.4 mm (42 43, 46 47), 0.2 mm (44 45)." httpUri="https://zenodo.org/record/156416/files/figure.png" pageId="15" pageNumber="16">Fig. 43</figureCitation>
|
||
). Male tergum 8 with slightly crenulate or straight posterior margin (
|
||
<figureCitation id="F3821AAFFF96FFF41D1BF93913BED4A5" box="[569,753,1747,1773]" captionStart="FIGURES 42 47" captionStartId="14.[264,382,1769,1793]" captionTargetBox="[288,1298,540,1737]" captionTargetId="figure@14.[279,1307,529,1742]" captionTargetPageId="14" captionText="FIGURES 42 47. Abdominal segment 8 of Goniusa alperti Kistner (holotype (42 44); male from San Bernardino Mts., California (45); female from 16 km NW of Granby, Colorado (46 47 )). 42 – male tergum 8; 43 – male sternum 8; 44 45 – apex of male tergum 8; 46 – female tergum 8; 47 – female sternum 8. Scale bar 0.4 mm (42 43, 46 47), 0.2 mm (44 45)." httpUri="https://zenodo.org/record/156416/files/figure.png" pageId="15" pageNumber="16">Figs. 42, 4445</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6B06062AFF96FFF41E1AF911139ED55D" blockId="15.[264,1324,1467,1893]" box="[312,721,1787,1813]" pageId="15" pageNumber="16">
|
||
Aedeagus as in
|
||
<figureCitation id="F3821AAFFF96FFF41ED3F9111336D55D" box="[497,633,1787,1813]" captionStart="FIGURES 48 51" captionStartId="15.[264,383,1288,1312]" captionTargetBox="[314,1270,697,1252]" captionTargetId="figure@15.[305,1283,689,1261]" captionTargetPageId="15" captionText="FIGURES 48 51. Median lobe of aedeagus of Goniusa alperti Kistner (male from West Point, Nebraska). 48 – median lobe, parameral view; 49 – apex of median lobe, parameral view; 50 – median lobe, lateral view; 51 – apex of median lobe, lateral view. Scale bar 0.4 mm (48, 50), 0.2 mm (49, 51)." httpUri="https://zenodo.org/record/156417/files/figure.png" pageId="15" pageNumber="16">Figs. 4851</figureCitation>
|
||
,
|
||
<figureCitation id="F3821AAFFF96FFF41DA7F9111382D55D" box="[645,717,1787,1813]" captionStart="FIGURES 52 56" captionStartId="16.[264,383,1747,1771]" captionTargetBox="[292,1298,498,1731]" captionTargetId="figure@16.[281,1307,489,1736]" captionTargetPageId="16" captionText="FIGURES 52 56. Genitalia of Goniusa alperti Kistner (paratype from 10 mi. E of Coville, Washington (52); female from Tahoe City, California (53); female from 16 km NW of Granby, Colorado (54); holotype (55 56 )). 52 54 – spermatheca; 55 – everted internal sac of aedeagus, lateral view; 56 – everted internal sac of aedeagus, parameral view. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156418/files/figure.png" pageId="15" pageNumber="16">5556</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6B06062AFF96FFF41E1AF8C913EDD575" blockId="15.[264,1324,1467,1893]" box="[312,674,1827,1853]" pageId="15" pageNumber="16">
|
||
Spermatheca as in
|
||
<figureCitation id="F3821AAFFF96FFF41D37F8C913D1D575" box="[533,670,1827,1853]" captionStart="FIGURES 52 56" captionStartId="16.[264,383,1747,1771]" captionTargetBox="[292,1298,498,1731]" captionTargetId="figure@16.[281,1307,489,1736]" captionTargetPageId="16" captionText="FIGURES 52 56. Genitalia of Goniusa alperti Kistner (paratype from 10 mi. E of Coville, Washington (52); female from Tahoe City, California (53); female from 16 km NW of Granby, Colorado (54); holotype (55 56 )). 52 54 – spermatheca; 55 – everted internal sac of aedeagus, lateral view; 56 – everted internal sac of aedeagus, parameral view. Scale bar 0.2 mm." httpUri="https://zenodo.org/record/156418/files/figure.png" pageId="15" pageNumber="16">Figs. 5254</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="23A355A1FF96FFEB1E1AF8A010D1D3CE" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="distribution">
|
||
<paragraph id="6B06062AFF96FFF41E1AF8A01463D52D" blockId="15.[264,1324,1467,1893]" box="[312,1324,1866,1893]" pageId="15" pageNumber="16">
|
||
<emphasis id="59CDDA38FF96FFF41E1AF8A01097D52C" bold="true" box="[312,472,1866,1892]" pageId="15" pageNumber="16">Distribution.</emphasis>
|
||
Known from Washington, California, Colorado and Nebraska (
|
||
<figureCitation id="F3821AAFFF96FFF41BE4F8A11451D52D" box="[1222,1310,1867,1893]" captionStart="FIGURE 57" captionStartId="17.[264,368,1343,1367]" captionTargetBox="[295,1292,323,1314]" captionTargetId="figure@17.[293,1294,320,1316]" captionTargetPageId="17" captionText="FIGURE 57. Geographical distribution of Goniusa caseyi Gusarov, sp. n. and G. alperti Kistner. Black circles and triangles indicate the localities of examined specimens. Open circles and triangles indicate the localities of additional specimens listed by Kistner (1976) but not examined by me. The exact locality of the Texas specimens of Goniusa caseyi is unknown." httpUri="https://zenodo.org/record/156419/files/figure.png" pageId="15" pageNumber="16">Fig. 57</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6B06062AFF89FFEB1E1AFEF110D1D3CE" blockId="16.[264,1322,283,390]" pageId="16" pageNumber="17">
|
||
<emphasis id="59CDDA38FF89FFEB1E1AFEF11349D37D" bold="true" box="[312,518,283,309]" pageId="16" pageNumber="17">Natural History.</emphasis>
|
||
<taxonomicName id="ACB97DA9FF89FFEB1D34FEF6139FD37D" box="[534,720,284,309]" class="Insecta" family="Staphylinidae" genus="Goniusa" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="alperti">
|
||
<emphasis id="59CDDA38FF89FFEB1D34FEF6139FD37D" box="[534,720,284,309]" italics="true" pageId="16" pageNumber="17">Goniusa alperti</emphasis>
|
||
</taxonomicName>
|
||
is associated with ants of the genus
|
||
<taxonomicName id="ACB97DA9FF89FFEB1BA0FEF615A8D37D" box="[1154,1255,284,309]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="59CDDA38FF89FFEB1BA0FEF615A8D37D" box="[1154,1255,284,309]" italics="true" pageId="16" pageNumber="17">Formica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="59CDDA38FF89FFEB1BDBFEF6146CD37D" box="[1273,1315,284,309]" italics="true" pageId="16" pageNumber="17">e.g.</emphasis>
|
||
,
|
||
<taxonomicName id="ACB97DA9FF89FFEB1E2AFEAE10BBD315" box="[264,500,324,349]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="obscuripes">
|
||
<emphasis id="59CDDA38FF89FFEB1E2AFEAE10BBD315" box="[264,500,324,349]" italics="true" pageId="16" pageNumber="17">Formica obscuripes</emphasis>
|
||
</taxonomicName>
|
||
(identified by Dr. D.R.Smith:
|
||
<bibRefCitation id="0F287BDBFF89FFEB1C4AFEAE1546D316" author="Kistner" box="[872,1033,324,350]" pageId="16" pageNumber="19" refString="Kistner, D. H. (1976) Revision and Reclassification of the Genus Goniusa Casey with a Larval Description and Ant Host Records (Coleoptera: Staphylinidae). Sociobiology, 2 (1), 83 - 95." type="journal article" year="1976">Kistner, 1976</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="ACB97DA9FF89FFEB1B68FEAE1465D315" box="[1098,1322,324,349]" class="Insecta" family="Formicidae" genus="Formica" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="haemorrhoidalis">
|
||
<emphasis id="59CDDA38FF89FFEB1B68FEAE1465D315" box="[1098,1322,324,349]" italics="true" pageId="16" pageNumber="17">F. haemorrhoidalis</emphasis>
|
||
</taxonomicName>
|
||
(label data)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |