451 lines
35 KiB
XML
451 lines
35 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e83035" ID-Pensoft-Pub="1864-8312-80-439" ID-Pensoft-UUID="3F4E21B524465471831D09738C2304A6" ID-ZooBank="62FFB94CEBF441639F22881435EFC37C" ModsDocID="1864-8312-80-439" checkinTime="1663021027595" checkinUser="pensoft" docAuthor="Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha" docDate="2022" docId="4D00CB1DCD52500BBA33B4C2221D493B" docLanguage="en" docName="ArthropodSystPhyl 80: 439-494" docOrigin="Arthropod Systematics & amp; Phylogeny 80" docPubDate="2022-09-12" docSource="http://dx.doi.org/10.3897/asp.80.e83035" docTitle="Amphineurus (Rhamphoneurus) nothofagetorum Alexander 1929" docType="treatment" docVersion="1" id="3F4E21B524465471831D09738C2304A6" lastPageNumber="439" masterDocId="3F4E21B524465471831D09738C2304A6" masterDocTitle="Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)" masterLastPageNumber="494" masterPageNumber="439" pageNumber="439" updateTime="1663021027595" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Santos, Daubian</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1220-1267</mods:nameIdentifier>
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<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">daubians@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Santos, Rodrigo dos Reis</mods:namePart>
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<mods:affiliation>Pos-Graduacao em Entomologia, Departamento de Zoologia, Universidade Federal do Parana, Curitiba, Parana, Brazil</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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||
</mods:role>
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<mods:namePart>Ribeiro, Guilherme Cunha</mods:namePart>
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<mods:affiliation>Universidade Federal do ABC, Centro de Ciencias Naturais e Humanas, Santo Andre, Sao Paulo, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-09-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>80</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>439</mods:start>
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<mods:end>494</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.80.e83035</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e83035</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-80-439</mods:identifier>
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<mods:identifier type="ZooBank">62FFB94CEBF441639F22881435EFC37C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">3F4E21B524465471831D09738C2304A6</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:4D00CB1DCD52500BBA33B4C2221D493B" httpUri="http://treatment.plazi.org/id/4D00CB1DCD52500BBA33B4C2221D493B" lastPageNumber="439" pageId="0" pageNumber="439">
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<subSubSection pageId="0" pageNumber="439" type="nomenclature">
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<paragraph pageId="0" pageNumber="439">
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3.3.18.
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||
<taxonomicName LSID="4D00CB1D-CD52-500B-BA33-B4C2221D493B" authority="Alexander, 1929 a" authorityName="Alexander" authorityYear="1929" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) nothofagetorum" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="nothofagetorum" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) nothofagetorum Alexander, 1929a</taxonomicName>
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</paragraph>
|
||
</subSubSection>
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<subSubSection pageId="0" pageNumber="439" type="reference_group">
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||
<paragraph pageId="0" pageNumber="439">
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||
<taxonomicName authorityName="Alexander" authorityYear="1929" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) nothofagetorum" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="nothofagetorum" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) nothofagetorum</taxonomicName>
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Alexander, 1929a: 187 (description, fig. 90: wing, fig. 223: male terminalia).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="439" type="materials_examined">
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||
<paragraph pageId="0" pageNumber="439">Material examined.</paragraph>
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||
<paragraph pageId="0" pageNumber="439">
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||
<emphasis bold="true" pageId="0" pageNumber="439">
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||
<typeStatus>Paratypes</typeStatus>
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||
</emphasis>
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||
:
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||
<specimenCount type="male">1 ♂</specimenCount>
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||
, Argentina, Nahuel Huapi, Lago Correntoso [
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<geoCoordinate degrees="40" direction="south" minutes="42" orientation="latitude" precision="925" value="-40.7">40°42′S</geoCoordinate>
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||
<geoCoordinate degrees="71" direction="west" minutes="39" orientation="longitude" precision="925" value="-71.65">71°39′W</geoCoordinate>
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||
],
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||
<date value="1926-11-02">2-XI-1926</date>
|
||
, R.C. Shannon (USNM)*;
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||
<specimenCount type="male">1 ♂</specimenCount>
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||
, Chile, Puerto Varas [
|
||
<geoCoordinate degrees="41" direction="south" minutes="19" orientation="latitude" precision="925" value="-41.316666">41°19′S</geoCoordinate>
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||
<geoCoordinate degrees="72" direction="west" minutes="59" orientation="longitude" precision="925" value="-72.98333">72°59′W</geoCoordinate>
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||
],
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||
<date value="1926-12-16">16-XII-1926</date>
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||
, F. & M. Edwards (USNM)*. -
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||
<emphasis bold="true" pageId="0" pageNumber="439">Additional material</emphasis>
|
||
:
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||
<materialsCitation collectingDate="1953-02-10" collectingDateMax="1965-01-30" collectingDateMin="1953-02-10" collectorName="Nahuelbuta, Pena, Cautin, Villarica, Km. NE" country="CHILE" latitude="-39.45" location="Arauco" longLatPrecision="1169" longitude="-71.9" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Arauco">
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||
<emphasis bold="true" pageId="0" pageNumber="439">
|
||
<collectingCountry name="Chile">CHILE</collectingCountry>
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||
.
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||
</emphasis>
|
||
<specimenCount type="male">1 ♂</specimenCount>
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||
,
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||
<specimenCount type="female">1 ♀</specimenCount>
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||
and 2 [sex unknown],
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||
<collectingRegion country="Chile" name="Araucania">Arauco</collectingRegion>
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||
,
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||
<collectorName>Nahuelbuta</collectorName>
|
||
[
|
||
<geoCoordinate degrees="37" direction="south" minutes="46" orientation="latitude" precision="925" value="-37.766666">37°46′S</geoCoordinate>
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||
<geoCoordinate degrees="72" direction="west" minutes="59" orientation="longitude" precision="925" value="-72.98333">72°59′W</geoCoordinate>
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||
], 5/
|
||
<collectingDate value="1953-02-10">10-II-1953</collectingDate>
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||
,
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||
<normalizedToken originalValue="Peña">
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||
<collectorName>Pena</collectorName>
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||
</normalizedToken>
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||
(USNM)*; 2 [sex unknown],
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||
<collectorName>Cautin</collectorName>
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||
,
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||
<collectorName>Villarica</collectorName>
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||
[
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||
<geoCoordinate degrees="39" direction="south" minutes="27" orientation="latitude" precision="925" value="-39.45">39°27′S</geoCoordinate>
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||
<geoCoordinate degrees="71" direction="west" minutes="54" orientation="longitude" precision="925" value="-71.9">71°54′W</geoCoordinate>
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||
], 30
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||
<collectorName>Km. NE</collectorName>
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||
, 1/
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||
<collectingDate value="1965-01-30">30-I-1965</collectingDate>
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||
,
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||
<normalizedToken originalValue="Peña">
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||
<collectorName>Pena</collectorName>
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||
</normalizedToken>
|
||
(USNM)
|
||
</materialsCitation>
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||
*;
|
||
<specimenCount type="male">2 ♂</specimenCount>
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||
and 1 [sex unknown],
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||
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
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||
Is. Ancud [
|
||
<geoCoordinate degrees="41" direction="south" minutes="52" orientation="latitude" precision="925" value="-41.866665">41°52′S</geoCoordinate>
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||
<geoCoordinate degrees="73" direction="west" minutes="48" orientation="longitude" precision="925" value="-73.8">73°48′W</geoCoordinate>
|
||
],
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||
<date value="1952-01-23">23-I-1952</date>
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||
, L.P. Guzman (USNM)*;
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||
<materialsCitation collectingDate="1954-01-31" collectingDateMax="1967-12-28" collectingDateMin="1954-01-31" collectorName="Chiloe Is., Chaiten, Pena, Nahuelbuta, Butamalal, Cordillera, Raices" country="Chile" elevation="1250" latitude="-38.516666" location="West of Lonquimay" longLatPrecision="1175" longitude="-71.46667" specimenCount="1" specimenCount-female="1" stateProvince="Arauco">
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||
<specimenCount type="female">1 ♀</specimenCount>
|
||
and 1 [sex unknown],
|
||
<collectorName>
|
||
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
|
||
Is.
|
||
</collectorName>
|
||
,
|
||
<collectorName>Chaiten</collectorName>
|
||
[
|
||
<geoCoordinate degrees="42" direction="south" minutes="58" orientation="latitude" precision="925" value="-42.966667">42°58′S</geoCoordinate>
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||
<geoCoordinate degrees="72" direction="west" minutes="32" orientation="longitude" precision="925" value="-72.53333">72°32′W</geoCoordinate>
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||
], 5/
|
||
<collectingDate value="1954-02-08">8-II-1954</collectingDate>
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||
,
|
||
<normalizedToken originalValue="Peña">
|
||
<collectorName>Pena</collectorName>
|
||
</normalizedToken>
|
||
(USNM)*; 1 [sex unknown],
|
||
<collectingRegion country="Chile" name="Araucania">Arauco</collectingRegion>
|
||
,
|
||
<collectorName>Nahuelbuta</collectorName>
|
||
,
|
||
<collectorName>Butamalal</collectorName>
|
||
[
|
||
<geoCoordinate degrees="37" direction="south" minutes="49" orientation="latitude" precision="925" value="-37.816666">37°49′S</geoCoordinate>
|
||
<geoCoordinate degrees="73" direction="west" minutes="14" orientation="longitude" precision="925" value="-73.23333">73°14′W</geoCoordinate>
|
||
],
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" unit="m" value="1250.0" valueMax="1400.0" valueMin="1100.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" unit="m" value="1250.0" valueMax="1400.0" valueMin="1100.0">1100-1400 m</elevation>
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||
</quantity>
|
||
, 23/
|
||
<collectingDate value="1954-01-31">31-I-1954</collectingDate>
|
||
,
|
||
<normalizedToken originalValue="Peña">
|
||
<collectorName>Pena</collectorName>
|
||
</normalizedToken>
|
||
(USNM)*; 1 [sex unknown],
|
||
<collectorName>Cordillera</collectorName>
|
||
las
|
||
<collectorName>Raices</collectorName>
|
||
[
|
||
<geoCoordinate degrees="38" direction="south" minutes="31" orientation="latitude" precision="925" value="-38.516666">38°31′S</geoCoordinate>
|
||
<geoCoordinate degrees="71" direction="west" minutes="28" orientation="longitude" precision="925" value="-71.46667">71°28′W</geoCoordinate>
|
||
],
|
||
<location LSID="urn:lsid:plazi:treatment:4D00CB1DCD52500BBA33B4C2221D493B:820B6001663405D078885E401F917EB8" country="Chile" latitude="-38.516666" longLatPrecision="1175" longitude="-71.46667" name="West of Lonquimay" stateProvince="Arauco">West of Lonquimay</location>
|
||
,
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">1500 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingDate value="1967-12-28">28-XII-1967</collectingDate>
|
||
,
|
||
<normalizedToken originalValue="Peña">
|
||
<collectorName>Pena</collectorName>
|
||
</normalizedToken>
|
||
(USNM)
|
||
</materialsCitation>
|
||
*;
|
||
<specimenCount type="male">2 ♂</specimenCount>
|
||
,
|
||
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
|
||
Is. Ancud [
|
||
<geoCoordinate degrees="41" direction="south" minutes="52" orientation="latitude" precision="925" value="-41.866665">41°52′S</geoCoordinate>
|
||
<geoCoordinate degrees="73" direction="west" minutes="48" orientation="longitude" precision="925" value="-73.8">73°48′W</geoCoordinate>
|
||
],
|
||
<date value="1952-01-23">23-I-1952</date>
|
||
,
|
||
<normalizedToken originalValue="Peña">Pena</normalizedToken>
|
||
(USNM);
|
||
<specimenCount type="male">2 ♂</specimenCount>
|
||
, Casa Pangue [
|
||
<geoCoordinate degrees="41" direction="south" minutes="03" orientation="latitude" precision="925" value="-41.05">41°03′S</geoCoordinate>
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||
<geoCoordinate degrees="71" direction="west" minutes="51" orientation="longitude" precision="925" value="-71.85">71°51′W</geoCoordinate>
|
||
], 6/
|
||
<date value="1926-12-10">10-XII-1926</date>
|
||
, F. & M. Edwards (USNM);
|
||
<specimenCount type="male">2 ♂</specimenCount>
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||
, Cautin, Villarica [
|
||
<geoCoordinate degrees="39" direction="south" minutes="27" orientation="latitude" precision="925" value="-39.45">39°27′S</geoCoordinate>
|
||
<geoCoordinate degrees="71" direction="west" minutes="54" orientation="longitude" precision="925" value="-71.9">71°54′W</geoCoordinate>
|
||
], 30 Km. NE, 1/
|
||
<date value="1965-01-30">30-I-1965</date>
|
||
,
|
||
<normalizedToken originalValue="Peña">Pena</normalizedToken>
|
||
(USNM);
|
||
<specimenCount type="male">1 ♂</specimenCount>
|
||
,
|
||
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
|
||
Is., Chaiten [
|
||
<geoCoordinate degrees="42" direction="south" minutes="58" orientation="latitude" precision="925" value="-42.966667">42°58′S</geoCoordinate>
|
||
<geoCoordinate degrees="72" direction="west" minutes="32" orientation="longitude" precision="925" value="-72.53333">72°32′W</geoCoordinate>
|
||
], 5/
|
||
<date value="1954-02-08">8-II-1954</date>
|
||
,
|
||
<normalizedToken originalValue="Peña">Pena</normalizedToken>
|
||
(USNM);
|
||
<materialsCitation collectingDate="1968-03" collectorName="Llanquihue, Hornohuinca, Pena" country="Chile" latitude="-41.416668" location="South of Volcan Calbuco" longLatPrecision="1156" longitude="-72.61667" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectorName>Llanquihue</collectorName>
|
||
,
|
||
<collectorName>Hornohuinca</collectorName>
|
||
[
|
||
<geoCoordinate degrees="41" direction="south" minutes="25" orientation="latitude" precision="925" value="-41.416668">41°25′S</geoCoordinate>
|
||
<geoCoordinate degrees="72" direction="west" minutes="37" orientation="longitude" precision="925" value="-72.61667">72°37′W</geoCoordinate>
|
||
],
|
||
<location LSID="urn:lsid:plazi:treatment:4D00CB1DCD52500BBA33B4C2221D493B:C80F02340DF1F6B3517C6A3486CAAFF1" country="Chile" latitude="-41.416668" longLatPrecision="1156" longitude="-72.61667" name="South of Volcan Calbuco">South of Volcan Calbuco</location>
|
||
,
|
||
<collectingDate value="1968-03">III-1968</collectingDate>
|
||
,
|
||
<normalizedToken originalValue="Peña">
|
||
<collectorName>Pena</collectorName>
|
||
</normalizedToken>
|
||
(USNM)
|
||
</materialsCitation>
|
||
;
|
||
<specimenCount type="male">1 ♂</specimenCount>
|
||
,
|
||
<normalizedToken originalValue="Chiloé">Chiloe</normalizedToken>
|
||
Is., Chepu [
|
||
<geoCoordinate degrees="42" direction="south" minutes="2" orientation="latitude" precision="925" value="-42.033333">42°2′S</geoCoordinate>
|
||
<geoCoordinate degrees="74" direction="west" minutes="1" orientation="longitude" precision="925" value="-74.01667">74°1′W</geoCoordinate>
|
||
], 4/
|
||
<date value="1968-04-06">6-IV-1968</date>
|
||
,
|
||
<normalizedToken originalValue="Peña">Pena</normalizedToken>
|
||
(USNM);
|
||
<specimenCount type="male">1 ♂</specimenCount>
|
||
, Osorno, Pucotrihue [
|
||
<geoCoordinate degrees="40" direction="south" minutes="32" orientation="latitude" precision="925" value="-40.533333">40°32′S</geoCoordinate>
|
||
<geoCoordinate degrees="73" direction="west" minutes="42" orientation="longitude" precision="925" value="-73.7">73°42′W</geoCoordinate>
|
||
],
|
||
<quantity metricMagnitude="1" metricUnit="m" metricValue="6.0" unit="m" value="60.0">60 m</quantity>
|
||
, 1/
|
||
<date value="1955-03-13">13-III-1955</date>
|
||
,
|
||
<normalizedToken originalValue="Peña">Pena</normalizedToken>
|
||
(USNM);
|
||
<specimenCount type="male">1 ♂</specimenCount>
|
||
, Caramavida [
|
||
<geoCoordinate degrees="37" direction="south" minutes="40" orientation="latitude" precision="925" value="-37.666668">37°40′S</geoCoordinate>
|
||
<geoCoordinate degrees="73" direction="west" minutes="20" orientation="longitude" precision="925" value="-73.333336">73°20′W</geoCoordinate>
|
||
],
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1000 m</quantity>
|
||
, 5/
|
||
<date value="1953-02-10">10-II-1953</date>
|
||
,
|
||
<normalizedToken originalValue="Peña">Pena</normalizedToken>
|
||
(USNM);
|
||
<materialsCitation collectingDate="1967-12-28" collectorName="Cordillera, Raices, Pena" country="Chile" elevation="1500" latitude="-38.516666" location="West of Lonquimay" longLatPrecision="1175" longitude="-71.46667" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectorName>Cordillera</collectorName>
|
||
las
|
||
<collectorName>Raices</collectorName>
|
||
[
|
||
<geoCoordinate degrees="38" direction="south" minutes="31" orientation="latitude" precision="925" value="-38.516666">38°31′S</geoCoordinate>
|
||
<geoCoordinate degrees="71" direction="west" minutes="28" orientation="longitude" precision="925" value="-71.46667">71°28′W</geoCoordinate>
|
||
],
|
||
<location LSID="urn:lsid:plazi:treatment:4D00CB1DCD52500BBA33B4C2221D493B:85ABDE8DDCC4FCCBB557E63748688212" country="Chile" latitude="-38.516666" longLatPrecision="1175" longitude="-71.46667" name="West of Lonquimay">West of Lonquimay</location>
|
||
,
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.5" unit="m" value="1500.0">1500 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingDate value="1967-12-28">28-XII-1967</collectingDate>
|
||
,
|
||
<normalizedToken originalValue="Peña">
|
||
<collectorName>Pena</collectorName>
|
||
</normalizedToken>
|
||
(USNM)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="439" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="439">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="439">The species is distinguished by the darkened distal quarter of the wings and M1+2 longer than half the length of the basal deflection of M1. Furthermore, the species has a distinctive clasper of gonostylus with setae on medial branch and straightened lateral branch.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="439" type="redescription">
|
||
<paragraph pageId="0" pageNumber="439">Redescription.</paragraph>
|
||
<paragraph pageId="0" pageNumber="439">
|
||
Wing length 6.10 mm, width 1.92 mm. -
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Coloration</emphasis>
|
||
: General coloration brown. Rostrum and palpus dark brown. First segments of antenna brownish yellow, remaining segments brown. Head dark brown. Eyes black. Thorax reddish-brown, pleura brown. Three brown stripes on scutum. Halter yellowish brown. Coxae and legs brownish yellow. Abdomen brown. -
|
||
<emphasis italics="true" pageId="0" pageNumber="439">Head</emphasis>
|
||
(ventral view Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphineurus (Rhamphoneurus) nothofagetorum Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure23" httpUri="https://binary.pensoft.net/fig/740576" pageId="0" pageNumber="439">23A</figureCitation>
|
||
, dorsal view Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphineurus (Rhamphoneurus) nothofagetorum Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure23" httpUri="https://binary.pensoft.net/fig/740576" pageId="0" pageNumber="439">23B</figureCitation>
|
||
): Rostrum bifurcated with short appendices; first palpal segment shorter than terminal segment; scape narrower than pedicel, terminal flagellomere as long as penultimate; antenna with several long setae. -
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Thorax</emphasis>
|
||
(lateral view Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphineurus (Rhamphoneurus) nothofagetorum Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure23" httpUri="https://binary.pensoft.net/fig/740576" pageId="0" pageNumber="439">23E</figureCitation>
|
||
, dorsal view Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphineurus (Rhamphoneurus) nothofagetorum Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure23" httpUri="https://binary.pensoft.net/fig/740576" pageId="0" pageNumber="439">23F</figureCitation>
|
||
): Anatergite as tall as katatergite. Halter with dilated knob. Wing (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphineurus (Rhamphoneurus) nothofagetorum Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure23" httpUri="https://binary.pensoft.net/fig/740576" pageId="0" pageNumber="439">23D</figureCitation>
|
||
) with several markings, including faint markings on A1 and beginning of Rs; markings highlighting m-cu and R2+3+4; stronger markings on R2 and near tip of R1; distal quarter of wing darkened; veins near fork of bM (Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphineurus (Rhamphoneurus) nothofagetorum Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure23" httpUri="https://binary.pensoft.net/fig/740576" pageId="0" pageNumber="439">23C</figureCitation>
|
||
) faded; M1+2 longer in length than half-length of basal deflection of M1; R2+3+4 about same length as R2+3. -
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Female terminalia</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphineurus (Rhamphoneurus) nothofagetorum Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure23" httpUri="https://binary.pensoft.net/fig/740576" pageId="0" pageNumber="439">23H</figureCitation>
|
||
): Female tergite IX longer and wider than tergite X. Hypogynial valve slender, longer than cercus. Cercus strongly curved dorsad. -
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="439">Male terminalia</emphasis>
|
||
(Fig.
|
||
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Amphineurus (Rhamphoneurus) nothofagetorum Alexander. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX." figureDoi="10.3897/asp.80.e83035.figure23" httpUri="https://binary.pensoft.net/fig/740576" pageId="0" pageNumber="439">23G</figureCitation>
|
||
): Male tergite IX unpigmented, with shallow curved median U-shaped notch. Dorsal branch of gonocoxite long, ca. 0.75 times length of ventral branch. Lobe of gonostylus with rounded lobule, lobule as long as wide, shorter than half-length of lobe of gonostylus, narrower than stem. Clasper of gonostylus with differently shaped branches; lateral branch straight, medial branch curved, club-shaped, with concentration of setae. Mesal lobes of gonocoxite asymmetrical, both with posterior margin rounded with bump on side; right mesal lobe wider. Sheath of aedeagus angulated, darkened through half its length.
|
||
</paragraph>
|
||
<caption doi="10.3897/asp.80.e83035.figure23" httpUri="https://binary.pensoft.net/fig/740576" pageId="0" pageNumber="439" start="Figure 23" startId="F23">
|
||
<paragraph pageId="0" pageNumber="439">
|
||
<emphasis bold="true" pageId="0" pageNumber="439">Figure 23.</emphasis>
|
||
<taxonomicName authorityName="Alexander" authorityYear="1929" class="Insecta" family="Limoniidae" genus="Amphineurus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Amphineurus (Rhamphoneurus) nothofagetorum" order="Diptera" pageId="0" pageNumber="439" phylum="Arthropoda" rank="species" species="nothofagetorum" subGenus="Rhamphoneurus">Amphineurus (Rhamphoneurus) nothofagetorum</taxonomicName>
|
||
Alexander.
|
||
<emphasis bold="true" pageId="0" pageNumber="439">A</emphasis>
|
||
head (ventral view);
|
||
<emphasis bold="true" pageId="0" pageNumber="439">B</emphasis>
|
||
head (dorsal view);
|
||
<emphasis bold="true" pageId="0" pageNumber="439">C</emphasis>
|
||
detail of wing;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">D</emphasis>
|
||
wing;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">E</emphasis>
|
||
thorax (lateral view);
|
||
<emphasis bold="true" pageId="0" pageNumber="439">F</emphasis>
|
||
thorax (dorsal view);
|
||
<emphasis bold="true" pageId="0" pageNumber="439">G</emphasis>
|
||
male tergite IX (dorsal view) and remainder of male terminalia (lateral view);
|
||
<emphasis bold="true" pageId="0" pageNumber="439">H</emphasis>
|
||
female terminalia (dorsal view). - Abbreviations:
|
||
<emphasis bold="true" pageId="0" pageNumber="439">aed sh</emphasis>
|
||
, sheath of aedeagus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">anatg</emphasis>
|
||
, anatergite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">anepm</emphasis>
|
||
, anepimeron;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">anepst</emphasis>
|
||
, anepisternum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">aprn</emphasis>
|
||
, anteropronotum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">cer</emphasis>
|
||
, cercus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">cgonst</emphasis>
|
||
, clasper of gonostylus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">cx</emphasis>
|
||
, coxa;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">goncx</emphasis>
|
||
, gonocoxite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">hlt</emphasis>
|
||
, halter;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">hyp val</emphasis>
|
||
, hypogynial valve;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">k</emphasis>
|
||
, knob of halter;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">kepm</emphasis>
|
||
, katepimeron;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">kepst</emphasis>
|
||
, katepisternum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">ktg</emphasis>
|
||
, katatergite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">l ms</emphasis>
|
||
, left mesal lobe of gonocoxite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">lgonst</emphasis>
|
||
; lobe of gonostylus;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mr</emphasis>
|
||
, meron;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mtanepst</emphasis>
|
||
, metanepisternum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mtepm</emphasis>
|
||
, metepimeron;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mtg</emphasis>
|
||
, mediotergite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mtkepst</emphasis>
|
||
, metakatepisternum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">mtn</emphasis>
|
||
, metanotum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">p</emphasis>
|
||
, posterior basalare;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">patg</emphasis>
|
||
, paratergite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">pprn</emphasis>
|
||
, postpronotum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">presct</emphasis>
|
||
, prescutum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">r ms</emphasis>
|
||
, right mesal lobe of gonocoxite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">sct</emphasis>
|
||
, scutum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">sctl</emphasis>
|
||
, scutellum;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">st</emphasis>
|
||
, sternite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">tg</emphasis>
|
||
, tergite;
|
||
<emphasis bold="true" pageId="0" pageNumber="439">t9</emphasis>
|
||
, male tergite IX.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="439" type="remarks">
|
||
<paragraph pageId="0" pageNumber="439">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="439">
|
||
We could not observe the holotype, but the paratypes were examined. This species resembles
|
||
<taxonomicName family="Limoniidae" kingdom="Animalia" lsidName="A. (R.) deceptus" order="Diptera" pageId="0" pageNumber="439" rank="species" species="deceptus" subGenus="R.">A. (R.) deceptus</taxonomicName>
|
||
sp. nov. but differs mainly in shape of branches of the clasper, male tergite IX, mesal lobes and the sheath of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |