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<document id="2DE3F90ACA6BF4C0FB014F4B59758209" ID-CLB-Dataset="32590" ID-DOI="10.11646/zootaxa.4299.4.1" ID-GBIF-Dataset="e9fcac10-f2bf-4f24-b015-b8c0f18faa14" ID-ISSN="1175-5326" ID-Zenodo-Dep="836971" ID-ZooBank="DEB133FB-9D5A-4F17-A718-568ACA9BA9D7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1501579068187" checkinUser="plazi" docAuthor="Sirenko, Boris I. &amp; Saito, Hiroshi" docDate="2017" docId="2C2987BBDD40FFA0FF10335CFC3CD2DD" docLanguage="en" docName="zootaxa.4299.4.1.pdf" docOrigin="Zootaxa 4299 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptoplax richardi Sirenko &amp; Saito 2017, n. comb." docType="treatment" docVersion="8" lastPageNumber="469" masterDocId="D010FFC3DD52FFB2FF87362AFFCBD52F" masterDocTitle="New species of chitons of the superfamily Cryptoplacoidea (Mollusca: Polyplacophora) from Vietnamese waters" masterLastPageNumber="506" masterPageNumber="451" pageNumber="469" updateTime="1698466057706" updateUser="ExternalLinkService">
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<mods:title id="C382EA8B00CAFB9806BA9E00CBAD1876">New species of chitons of the superfamily Cryptoplacoidea (Mollusca: Polyplacophora) from Vietnamese waters</mods:title>
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<mods:namePart id="C8ACD0635AB9B143CD169F612066B5C3">Sirenko, Boris I.</mods:namePart>
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<treatment id="2C2987BBDD40FFA0FF10335CFC3CD2DD" ID-DOI="http://doi.org/10.5281/zenodo.5998841" ID-GBIF-Taxon="132617940" ID-Zenodo-Dep="5998841" LSID="urn:lsid:plazi:treatment:2C2987BBDD40FFA0FF10335CFC3CD2DD" httpUri="http://treatment.plazi.org/id/2C2987BBDD40FFA0FF10335CFC3CD2DD" lastPageNumber="469" pageId="18" pageNumber="469">
<subSubSection id="EC9A6526DD40FFA0FF10335CFF2DD09D" pageId="18" pageNumber="469" type="nomenclature">
<paragraph id="A43F36ADDD40FFA0FF10335CFD46D0BF" blockId="18.[151,653,1398,1458]" box="[151,653,1398,1424]" pageId="18" pageNumber="469">
<heading id="FF7781C1DD40FFA0FF10335CFD46D0BF" bold="true" box="[151,653,1398,1424]" fontSize="11" level="1" pageId="18" pageNumber="469" reason="1">
<emphasis id="96F4EABFDD40FFA0FF10335CFD46D0BF" bold="true" box="[151,653,1398,1424]" pageId="18" pageNumber="469">
<taxonomicName id="63804D2EDD40FFA0FF10335CFDD7D0BF" ID-CoL="3TTGM" authority="Kaas, 1990" authorityName="Sirenko &amp; Saito" authorityYear="2017" box="[151,540,1398,1424]" class="Polyplacophora" family="Acanthochitonidae" genus="Leptoplax" kingdom="Animalia" order="Chitonida" pageId="18" pageNumber="469" phylum="Mollusca" rank="species" species="richardi" status="comb. nov.">
<emphasis id="96F4EABFDD40FFA0FF10335CFEB3D0BF" bold="true" box="[151,376,1398,1424]" italics="true" pageId="18" pageNumber="469">Leptoplax richardi</emphasis>
(
<bibRefCitation id="C0114B5CDD40FFA0FE0E335CFDD9D0BF" author="Kaas" box="[393,530,1398,1424]" pageId="18" pageNumber="469" refString="Kaas, P. (1990) New species and further records of known species of Polyplacophora from the tropical western Pacific. Basteria, 54, 175 - 186." type="journal article" year="1990">Kaas, 1990</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="8DC757C4DD40FFA0FDA4335CFD46D0BF" box="[547,653,1398,1424]" pageId="18" pageNumber="469">n. comb.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="A43F36ADDD40FFA0FF1033B3FF2DD09D" blockId="18.[151,653,1398,1458]" box="[151,230,1433,1458]" pageId="18" pageNumber="469">
<figureCitation id="3CBB2A28DD40FFA0FF1033B3FF2DD09D" box="[151,230,1433,1458]" captionStart="FIGURE 16" captionStartId="19.[151,249,917,939]" captionTargetBox="[163,1409,195,886]" captionTargetId="figure@19.[151,1436,173,917]" captionTargetPageId="19" captionText="FIGURE 16. Leptoplax richardi, Southern Vietnam, Cau Id.: A. Valve VIII, dorsal view; B. Valve VIII, lateral view; C. Valve VIII, detail of tegmentum in antemucronal area." httpUri="https://zenodo.org/record/837003/files/figure.png" pageId="18" pageNumber="469">Fig. 16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="EC9A6526DD40FFA0FF1033F0FD1FD301" pageId="18" pageNumber="469" type="reference_group">
<paragraph id="A43F36ADDD40FFA0FF1033F0FD42D320" blockId="18.[151,784,1498,1582]" pageId="18" pageNumber="469">
<treatmentCitationGroup id="84901183DD40FFA0FF1033F0FD42D320" pageId="18" pageNumber="469">
<taxonomicName id="63804D2EDD40FFA0FF1033F0FDA0D0DF" ID-CoL="47XW3" authority="Kaas, 1990: 178" authorityName="Kaas" authorityPageNumber="178" authorityYear="1990" box="[151,619,1498,1520]" class="Polyplacophora" family="Acanthochitonidae" genus="Notoplax" kingdom="Animalia" order="Chitonida" pageId="18" pageNumber="469" phylum="Mollusca" rank="species" species="richardi" subGenus="Notoplax">
<emphasis id="96F4EABFDD40FFA0FF1033F0FF3FD0DF" box="[151,244,1498,1520]" italics="true" pageId="18" pageNumber="469">Notoplax</emphasis>
(
<emphasis id="96F4EABFDD40FFA0FE8533F0FE94D0DF" box="[258,351,1498,1520]" italics="true" pageId="18" pageNumber="469">Notoplax</emphasis>
)
<emphasis id="96F4EABFDD40FFA0FEE933F0FE0BD0DF" box="[366,448,1498,1520]" italics="true" pageId="18" pageNumber="469">richardi</emphasis>
<treatmentCitation id="252110BCDD40FFA0FE4133F1FDA0D0DF" author="Kaas" box="[454,619,1498,1520]" page="178" pageId="18" pageNumber="469" year="1990">
<bibRefCitation id="C0114B5CDD40FFA0FE4133F1FDF0D0DF" author="Kaas" box="[454,571,1498,1520]" pageId="18" pageNumber="469" refString="Kaas, P. (1990) New species and further records of known species of Polyplacophora from the tropical western Pacific. Basteria, 54, 175 - 186." type="journal article" year="1990">Kaas, 1990</bibRefCitation>
: 178
</treatmentCitation>
</taxonomicName>
179, figs 110.
<taxonomicName id="63804D2EDD40FFA0FF1033D3FD4ED320" ID-CoL="47XW3" authority="Kaas &amp; Van Belle, 1998: 158" authorityName="Kaas &amp; Van Belle" authorityPageNumber="158" authorityYear="1998" box="[151,645,1529,1551]" class="Polyplacophora" family="Acanthochitonidae" genus="Notoplax" kingdom="Animalia" order="Chitonida" pageId="18" pageNumber="469" phylum="Mollusca" rank="species" species="richardi">
<emphasis id="96F4EABFDD40FFA0FF1033D3FE87D320" box="[151,332,1529,1551]" italics="true" pageId="18" pageNumber="469">Notoplax richardi</emphasis>
:
<treatmentCitation id="252110BCDD40FFA0FEDE33D0FD4ED320" author="Kaas" box="[345,645,1529,1551]" page="158" pageId="18" pageNumber="469" year="1998">
<bibRefCitation id="C0114B5CDD40FFA0FEDE33D0FD99D320" author="Kaas" box="[345,594,1529,1551]" pageId="18" pageNumber="469" refString="Kaas, P. &amp; Van Belle, R. A. (1998) Catalogue of living chitons (Mollusca, Polyplacophora). 2 nd Revised Edition. Dr. W. Backhuys, Publisher, Leiden, 204 pp." type="book" year="1998">Kaas &amp; Van Belle, 1998</bibRefCitation>
: 158
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="A43F36ADDD40FFA0FF103032FD1FD301" blockId="18.[151,784,1498,1582]" box="[151,724,1560,1582]" pageId="18" pageNumber="469">
<treatmentCitationGroup id="84901183DD40FFA0FF103032FD1FD301" box="[151,724,1560,1582]" pageId="18" pageNumber="469">
<taxonomicName id="63804D2EDD40FFA0FF253032FF34D301" ID-CoL="7P9Z8" authorityName="H. Adams" authorityYear="1861" box="[162,255,1560,1582]" class="Polyplacophora" family="Acanthochitonidae" genus="Notoplax" kingdom="Animalia" order="Chitonida" pageId="18" pageNumber="469" phylum="Mollusca" rank="genus">
<emphasis id="96F4EABFDD40FFA0FF253032FF34D301" box="[162,255,1560,1582]" italics="true" pageId="18" pageNumber="469">Notoplax</emphasis>
</taxonomicName>
” sp.
<taxonomicName id="63804D2EDD40FFA0FEB43032FD91D301" ID-CoL="47XW3" authority="Saito, 1998: 158" authorityName="Saito" authorityPageNumber="158" authorityYear="1998" box="[307,602,1560,1582]" class="Polyplacophora" family="Acanthochitonidae" genus="Notoplax" kingdom="Animalia" order="Chitonida" pageId="18" pageNumber="469" phylum="Mollusca" rank="species" species="richardi">
cf.
<emphasis id="96F4EABFDD40FFA0FED33032FE6DD301" box="[340,422,1560,1582]" italics="true" pageId="18" pageNumber="469">richardi</emphasis>
:
<treatmentCitation id="252110BCDD40FFA0FE343032FD91D301" author="Saito" box="[435,602,1560,1582]" page="158" pageId="18" pageNumber="469" year="1998">
<bibRefCitation id="C0114B5CDD40FFA0FE343032FDE0D301" author="Saito" box="[435,555,1560,1582]" pageId="18" pageNumber="469" refString="Saito, H. (1998) The chitons (Mollusca: Polyplacophora) of the Goto Islands, west coast of Kyushu, Japan. Memoir of the National Science Museum, Tokyo, 31, 145 - 163." type="journal article" year="1998">Saito, 1998</bibRefCitation>
: 158
</treatmentCitation>
</taxonomicName>
160, fig. 8.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="EC9A6526DD40FFA0FF10307EFDD6D3F8" pageId="18" pageNumber="469" type="materials_examined">
<paragraph id="A43F36ADDD40FFA0FF10307EFD00D342" blockId="18.[151,1436,1620,2035]" box="[151,715,1620,1645]" pageId="18" pageNumber="469">
<emphasis id="96F4EABFDD40FFA0FF10307EFE8ED342" bold="true" box="[151,325,1620,1645]" pageId="18" pageNumber="469">Type material.</emphasis>
Holotype, MNHN-IM-2000-6111.
</paragraph>
<paragraph id="A43F36ADDD40FFA0FF40305DFC56D3A0" blockId="18.[151,1436,1620,2035]" box="[199,925,1655,1680]" pageId="18" pageNumber="469">
<emphasis id="96F4EABFDD40FFA0FF40305DFEAED3BF" bold="true" box="[199,357,1655,1680]" pageId="18" pageNumber="469">Type locality.</emphasis>
Capel Bank, Coral Sea, 25°20´S, 159°44´E,
<quantity id="63789B48DD40FFA0FCDA305DFC52D3A0" box="[861,921,1655,1680]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="18" pageNumber="469" unit="m" value="56.0">56 m</quantity>
.
</paragraph>
<paragraph id="A43F36ADDD40FFA0FF4030B1FDD6D3F8" blockId="18.[151,1436,1620,2035]" pageId="18" pageNumber="469">
<materialsCitation id="14E83CF0DD40FFA0FF4030B1FDD6D3F8" ID-GBIF-Occurrence="1585189403" collectingDate="2013-05-14" collectionCode="SCUBA" collectorName="Cau Id. &amp; B. Sirenko." country="Vietnam" elevation="13" latitude="11.2220335" location="Material" longLatPrecision="1" longitude="108.8311" pageId="18" pageNumber="469" specimenCount="1">
<emphasis id="96F4EABFDD40FFA0FF4030B1FE78D39B" bold="true" box="[199,435,1691,1716]" pageId="18" pageNumber="469">
<location id="A15F6076DD40FFA0FF4030B1FEE4D39B" LSID="urn:lsid:plazi:treatment:2C2987BBDD40FFA0FF10335CFC3CD2DD:A15F6076DD40FFA0FF4030B1FEE4D39B" box="[199,303,1691,1716]" country="Vietnam" latitude="11.2220335" longLatPrecision="1" longitude="108.8311" name="Material" pageId="18" pageNumber="469">Material</location>
examined.
</emphasis>
Southern
<collectingCountry id="DC97763DDD40FFA0FDAB30B1FD45D39B" box="[556,654,1691,1716]" name="Vietnam" pageId="18" pageNumber="469">Vietnam</collectingCountry>
,
<collectorName id="0975537BDD40FFA0FD1C30B1FD39D39B" box="[667,754,1691,1716]" pageId="18" pageNumber="469">Cau Id.</collectorName>
,
<geoCoordinate id="C1B4506ADD40FFA0FC8630B1FC5FD39C" box="[769,916,1691,1715]" direction="north" orientation="latitude" pageId="18" pageNumber="469" precision="1" value="11.2220335">11°13.322´N</geoCoordinate>
,
<geoCoordinate id="C1B4506ADD40FFA0FC2530B1FB8AD39B" box="[930,1089,1691,1716]" direction="east" orientation="longitude" pageId="18" pageNumber="469" precision="1" value="108.8311">108°49.866´E</geoCoordinate>
,
<elevation id="2FADD19EDD40FFA0FBD730B1FB7CD39C" box="[1104,1207,1691,1715]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="18" pageNumber="469" unit="m" value="12.5" valueMax="13.0" valueMin="12.0">
<quantity id="63789B48DD40FFA0FBD730B1FB7CD39C" box="[1104,1207,1691,1715]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="18" pageNumber="469" unit="m" value="12.5" valueMax="13.0" valueMin="12.0">1213 m</quantity>
</elevation>
,
<collectionCode id="C291AE68DD40FFA0FB4130B1FAEDD39C" box="[1222,1318,1691,1715]" pageId="18" pageNumber="469">SCUBA</collectionCode>
, sand, tail valve,
<collectingDate id="C07AE985DD40FFA0FF663094FEA9D3F8" box="[225,354,1726,1751]" pageId="18" pageNumber="469" value="2013-05-14">
<date id="D03E106DDD40FFA0FF663094FEA9D3F8" box="[225,354,1726,1751]" pageId="18" pageNumber="469" value="2013-05-14">14.05.2013</date>
</collectingDate>
, leg.
<collectorName id="0975537BDD40FFA0FE193095FDD6D3F8" box="[414,541,1726,1751]" pageId="18" pageNumber="469">B. Sirenko.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="EC9A6526DD40FFA0FF4030CBFBC8D3D6" box="[199,1027,1761,1786]" pageId="18" pageNumber="469" type="distribution">
<paragraph id="A43F36ADDD40FFA0FF4030CBFBC8D3D6" blockId="18.[151,1436,1620,2035]" box="[199,1027,1761,1786]" pageId="18" pageNumber="469">
<emphasis id="96F4EABFDD40FFA0FF4030CBFEABD3D5" bold="true" box="[199,352,1761,1786]" pageId="18" pageNumber="469">Distribution.</emphasis>
Coral Sea; Southern
<collectingCountry id="DC97763DDD40FFA0FDC830CBFD79D3D5" box="[591,690,1761,1786]" name="Vietnam" pageId="18" pageNumber="469">Vietnam</collectingCountry>
; Goto Islands,
<collectingCountry id="DC97763DDD40FFA0FCDE30C8FC57D3D6" box="[857,924,1762,1785]" name="Japan" pageId="18" pageNumber="469">Japan</collectingCountry>
,
<quantity id="63789B48DD40FFA0FC2030CBFC34D3D6" box="[935,1023,1761,1786]" metricMagnitude="1" metricUnit="m" metricValue="3.1" metricValueMax="5.6" metricValueMin="0.6" pageId="18" pageNumber="469" unit="m" value="31.0" valueMax="56.0" valueMin="6.0">
6
<specimenCount id="B286FD24DD40FFA0FC4430CBFC34D3D6" box="[963,1023,1761,1786]" pageId="18" pageNumber="469" type="generic">56 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="EC9A6526DD40FFA0FF40312FFC3CD2DD" pageId="18" pageNumber="469" type="discussion">
<paragraph id="A43F36ADDD40FFA0FF40312FFB2AD2A7" blockId="18.[151,1436,1620,2035]" pageId="18" pageNumber="469">
<emphasis id="96F4EABFDD40FFA0FF40312FFEF0D231" bold="true" box="[199,315,1797,1822]" pageId="18" pageNumber="469">Remarks.</emphasis>
The tail valve (
<figureCitation id="3CBB2A28DD40FFA0FE7D312FFD9BD231" box="[506,592,1797,1822]" captionStart="FIGURE 16" captionStartId="19.[151,249,917,939]" captionTargetBox="[163,1409,195,886]" captionTargetId="figure@19.[151,1436,173,917]" captionTargetPageId="19" captionText="FIGURE 16. Leptoplax richardi, Southern Vietnam, Cau Id.: A. Valve VIII, dorsal view; B. Valve VIII, lateral view; C. Valve VIII, detail of tegmentum in antemucronal area." httpUri="https://zenodo.org/record/837003/files/figure.png" pageId="18" pageNumber="469">Fig. 16</figureCitation>
) which is the only known specimen of this species collected during the survey of collected sand is identifiable to
<taxonomicName id="63804D2EDD40FFA0FD043102FC10D26E" authority="Kaas, 1990" authorityName="Kaas" authorityYear="1990" box="[643,987,1832,1857]" class="Polyplacophora" family="Acanthochitonidae" genus="Notoplax" kingdom="Animalia" order="Chitonida" pageId="18" pageNumber="469" phylum="Mollusca" rank="species" species="richardi">
<emphasis id="96F4EABFDD40FFA0FD043102FC84D26E" box="[643,847,1832,1857]" italics="true" pageId="18" pageNumber="469">Notoplax richardi</emphasis>
<bibRefCitation id="C0114B5CDD40FFA0FCDE3103FC10D26E" author="Kaas" box="[857,987,1833,1857]" pageId="18" pageNumber="469" refString="Kaas, P. (1990) New species and further records of known species of Polyplacophora from the tropical western Pacific. Basteria, 54, 175 - 186." type="journal article" year="1990">Kaas, 1990</bibRefCitation>
</taxonomicName>
[=
<taxonomicName id="63804D2EDD40FFA0FB8E3102FAB6D26E" authority="Kaas, 1990" authorityName="Sirenko &amp; Saito" authorityYear="2017" box="[1033,1405,1832,1857]" class="Polyplacophora" family="Acanthochitonidae" genus="Leptoplax" kingdom="Animalia" order="Chitonida" pageId="18" pageNumber="469" phylum="Mollusca" rank="species" species="richardi" status="comb. nov.">
<emphasis id="96F4EABFDD40FFA0FB8E3102FB15D26E" box="[1033,1246,1832,1857]" italics="true" pageId="18" pageNumber="469">Leptoplax richardi</emphasis>
(
<bibRefCitation id="C0114B5CDD40FFA0FB773103FABFD26E" author="Kaas" box="[1264,1396,1833,1857]" pageId="18" pageNumber="469" refString="Kaas, P. (1990) New species and further records of known species of Polyplacophora from the tropical western Pacific. Basteria, 54, 175 - 186." type="journal article" year="1990">Kaas, 1990</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="8DC757C4DD40FFA0FA003103FF14D24A" pageId="18" pageNumber="469">n. comb.</taxonomicNameLabel>
] described from Coral Sea, at the depth of
<quantity id="63789B48DD40FFA0FD4A3166FCC3D24B" box="[717,776,1868,1893]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="18" pageNumber="469" unit="m" value="56.0">56 m</quantity>
by the outline of tegmentum, wavy margins of the jugum which has oblique sulci and 27 micraesthete pores distributed in the proximal half of the pustule.
</paragraph>
<paragraph id="A43F36ADDD40FFA0FF4031B9FC3CD2DD" blockId="18.[151,1436,1620,2035]" pageId="18" pageNumber="469">
The generic assignment of this species is changed from
<taxonomicName id="63804D2EDD40FFA0FCBA31B9FC6FD283" authorityName="H. Adams" authorityYear="1861" box="[829,932,1939,1964]" class="Polyplacophora" family="Acanthochitonidae" genus="Notoplax" kingdom="Animalia" order="Chitonida" pageId="18" pageNumber="469" phylum="Mollusca" rank="genus">
<emphasis id="96F4EABFDD40FFA0FCBA31B9FC6FD283" box="[829,932,1939,1964]" italics="true" pageId="18" pageNumber="469">Notoplax</emphasis>
</taxonomicName>
to
<taxonomicName id="63804D2EDD40FFA0FC4F31B9FBF3D283" authorityName="Dall" authorityYear="1882" box="[968,1080,1939,1964]" class="Polyplacophora" family="Acanthochitonidae" genus="Leptoplax" kingdom="Animalia" order="Chitonida" pageId="18" pageNumber="469" phylum="Mollusca" rank="genus">
<emphasis id="96F4EABFDD40FFA0FC4F31B9FBF3D283" box="[968,1080,1939,1964]" italics="true" pageId="18" pageNumber="469">Leptoplax</emphasis>
</taxonomicName>
herein, based on the diagnostic features of the sutural laminae of the tail valve and the central part of the radula. The latter is shown in
<bibRefCitation id="C0114B5CDD40FFA0FA94319CFEFAD2DD" author="Saito" pageId="18" pageNumber="469" refString="Saito, H. (1998) The chitons (Mollusca: Polyplacophora) of the Goto Islands, west coast of Kyushu, Japan. Memoir of the National Science Museum, Tokyo, 31, 145 - 163." type="journal article" year="1998">
Saito (1998: 159,
<figureCitation id="3CBB2A28DD40FFA0FF4831F0FEDBD2DD" box="[207,272,2010,2035]" captionStart="FIGURE 8" captionStartId="10.[151,249,1700,1722]" captionTargetBox="[277,1283,203,1612]" captionTargetId="figure@10.[257,1330,173,1750]" captionTargetPageId="10" captionText="FIGURE 8. Notoplax odysseyi n. sp., Southern Vietnam, Off Phan Rang, holotype, BL 51 mm: A. Needle of tufts; B. Dorsal needles, C. Ventral spicules, D. Head of major lateral tooth of radula; E. Central and first lateral teeth of radula; F. Pustule of lateropleural areas of tegmentum. Scale bars: 100 µm." httpUri="https://zenodo.org/record/836987/files/figure.png" pageId="18" pageNumber="469">Fig. 8</figureCitation>
A)
</bibRefCitation>
. This is the first record of this species from Vietnamese waters.
</paragraph>
</subSubSection>
</treatment>
</document>