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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.123.31738" ID-GBIF-Dataset="3fc0de51-2536-45b8-8be2-d8b860355539" ID-PMC="PMC6554266" ID-Pensoft-Pub="1314-2003-123-1" ID-Pensoft-UUID="8347FFA73705456BFF950322FFBBFFDB" ID-PubMed="31198402" ID-Zenodo-Dep="3240750" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1314-2003-123-1" ModsDocOrigin="PhytoKeys 123" ModsDocTitle="A revision of the Morelloid Clade of Solanum L. (Solanaceae) in North and Central America and the Caribbean" checkinTime="1559283366142" checkinUser="pensoft" docAuthor="Knapp, Sandra, Barboza, Gloria E., Bohs, Lynn &amp; Saerkinen, Tiina" docDate="2019" docId="271A31962CD6B2F79800D3331FB1BCAC" docLanguage="en" docName="PhytoKeys 123: 1-144" docOrigin="PhytoKeys 123" docSource="http://dx.doi.org/10.3897/phytokeys.123.31738" docTitle="Solanum pruinosum Dunal, Prodr. [A. P. de Candolle] 13 (1): 58. 1852" docType="treatment" docVersion="8" id="8347FFA73705456BFF950322FFBBFFDB" lastPageNumber="83" masterDocId="8347FFA73705456BFF950322FFBBFFDB" masterDocTitle="A revision of the Morelloid Clade of Solanum L. (Solanaceae) in North and Central America and the Caribbean" masterLastPageNumber="144" masterPageNumber="1" pageNumber="79" updateTime="1668139316858" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the Morelloid Clade of Solanum L. (Solanaceae) in North and Central America and the Caribbean</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Knapp, Sandra</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Barboza, Gloria E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bohs, Lynn</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Saerkinen, Tiina</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>123</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>144</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.123.31738</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.123.31738</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-123-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">8347FFA73705456BFF950322FFBBFFDB</mods:identifier>
<mods:identifier type="Zenodo-Dep">3240750</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="157330221" LSID="urn:lsid:plazi:treatment:271A31962CD6B2F79800D3331FB1BCAC" httpUri="http://treatment.plazi.org/id/271A31962CD6B2F79800D3331FB1BCAC" lastPageId="82" lastPageNumber="83" pageId="78" pageNumber="79">
<subSubSection pageId="78" pageNumber="79" type="nomenclature">
<paragraph pageId="78" pageNumber="79">
12.
<taxonomicName LSID="6A32C25E-1D5D-5673-AC15-341E2FE913C2" authority="Dunal, Prodr. [A. P. de Candolle] 13 (1): 58. 1852" authorityName="Dunal, Prodr. [A. P. de Candolle] 13 (1): 58." authorityYear="1852" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pruinosum" order="Solanales" pageId="78" pageNumber="79" phylum="Tracheophyta" rank="species" species="pruinosum">Solanum pruinosum Dunal, Prodr. [A. P. de Candolle] 13(1): 58. 1852</taxonomicName>
<figureCitation captionStart="Figure 36" captionStartId="F36" captionText="Figure 36. Solanum pruinosum Dunal. A Habit B flower C dissected flower D calyx E infructescence F elongate glandular trichome G sessile glandular trichome (A-E Amith JDA- 30248; F-G Ventura A. 2588). Drawing by L. Ribulgo." figureDoi="10.3897/phytokeys.123.31738.figure36" httpUri="https://binary.pensoft.net/fig/305335" pageId="78" pageNumber="79">Figures 36</figureCitation>
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Solanum pruinosum Dunal. A Leaves B young branch with flowering buds C inflorescence with flowers at full anthesis D developing fruits (A-D Amith JDA- 30248). Photos by M. Gorostiza Salazar." figureDoi="10.3897/phytokeys.123.31738.figure37" httpUri="https://binary.pensoft.net/fig/305336" pageId="78" pageNumber="79">, 37</figureCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80" type="reference_group">
<paragraph pageId="79" pageNumber="80">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum dasyadenium" order="Solanales" pageId="79" pageNumber="80" phylum="Tracheophyta" rank="species" species="dasyadenium">
<pageBreakToken pageId="79" pageNumber="80" start="start">Solanum</pageBreakToken>
dasyadenium
</taxonomicName>
Bitter, Repert. Spec. Nov. Regni Veg. 11: 8. 1912. Type. Mexico. Sin.loc.,
<emphasis italics="true" pageId="79" pageNumber="80">J. Schaffner 655</emphasis>
(syntype, B destroyed);
<emphasis italics="true" pageId="79" pageNumber="80">C.A. Uhde 80</emphasis>
(syntype, B destroyed; dups maybe at Halle?).
</paragraph>
<paragraph pageId="79" pageNumber="80">
<taxonomicName authority="subsp. uberius" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" infraspecific-rank="subsp." kingdom="Plantae" lsidName="Solanum dasyadenium" order="Solanales" pageId="79" pageNumber="80" phylum="Tracheophyta" rank="species" species="dasyadenium" subspecies="uberius">Solanum dasyadenium subsp. uberius</taxonomicName>
Bitter, Repert. Spec. Nov. Regni Veg. 11: 9. 1912. Type. Mexico. Sin.loc.,
<emphasis italics="true" pageId="79" pageNumber="80">A. Aschenborn 412</emphasis>
(syntype B, destroyed);
<emphasis italics="true" pageId="79" pageNumber="80">A. Aschenborn 413</emphasis>
(syntype B, destroyed).
</paragraph>
<paragraph lastPageId="80" lastPageNumber="81" pageId="79" pageNumber="80">
<taxonomicName authority="subsp. potosanum" class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" infraspecific-rank="subsp." kingdom="Plantae" lsidName="Solanum dasyadenium" order="Solanales" pageId="79" pageNumber="80" phylum="Tracheophyta" rank="species" species="dasyadenium" subspecies="potosanum">Solanum dasyadenium subsp. potosanum</taxonomicName>
Bitter, Repert. Spec. Nov. Regni Veg. 11: 9. 1912.
<pageBreakToken pageId="80" pageNumber="81" start="start">Type</pageBreakToken>
. Mexico. San Luis
<normalizedToken originalValue="Potosí">Potosi</normalizedToken>
: San Luis
<normalizedToken originalValue="Potosí">Potosi</normalizedToken>
,
<emphasis italics="true" pageId="80" pageNumber="81">J. Schaffner 408</emphasis>
(holotype: B, destroyed; lectotype, designated here: GOET [GOET003496]; isolectotypes: BM [BM000579277], M [M-0183327], NY [NY00751028], P [P00366754], US [US00027536, acc. # 939130]).
</paragraph>
</subSubSection>
<subSubSection pageId="80" pageNumber="81" type="materials_examined">
<paragraph pageId="80" pageNumber="81">Type.</paragraph>
<paragraph pageId="80" pageNumber="81">
Mexico. &quot;Circa Mexico&quot;,
<emphasis italics="true" pageId="80" pageNumber="81">J. Berlandier 751</emphasis>
(holotype: G [G00418346]).
</paragraph>
</subSubSection>
<subSubSection lastPageId="81" lastPageNumber="82" pageId="80" pageNumber="81" type="description">
<paragraph pageId="80" pageNumber="81">Description.</paragraph>
<paragraph lastPageId="81" lastPageNumber="82" pageId="80" pageNumber="81">
Perennial herb, 0.7-1 m tall, perhaps occasionally annual or only persisting for a few years. Stems angled to winged, lacking spinescent processes, usually erect, but occasionally lax and somewhat scrambling; young stems densely to sparsely pubescent with glandular, simple uniseriate trichomes 0.5-2 mm long, the trichomes (2
<normalizedToken originalValue=")4">-)4-</normalizedToken>
15 celled, the basal cells larger, the trichomes drying translucent; new growth densely glandular pubescent and sticky; bark of older stems greenish brown. Sympodial units difoliate, the leaves not geminate. Leaves simple, occasionally shallowly toothed, 2.5-6.5 cm long, 1.2-2.8 cm wide, elliptic to ovate, widest in the lower half, membranous; adaxial and abaxial surfaces evenly and densely glandular-pubescent with simple uniseriate trichomes to 2 mm long, these denser abaxially and along the veins; principal veins 4-6 pairs, drying paler than the lamina; base attenuate onto the petiole; margins entire to shallowly and irregularly toothed, the teeth mostly in the basal third of the blade, usually with minute glandular papillae with 2-celled glandular tips that dry dark brown; apex acute to acuminate; petiole 0.5-2 cm, narrowly winged from the attenuate leaf base. Inflorescences 0.8-2.5 cm long, unbranched, internodal, with 3-6 flowers (usually ca. 4) clustered in the distal third or quarter (sub-umbelliform), densely glandular-pubescent like the stems and leaves; peduncle 0.8-2.5 cm long; pedicels 0.7-0.9 cm long at anthesis, ca. 0.5 mm in diameter at the base, ca. 1 mm in diameter at the apex, slender and tapering, densely glandular-pubescent with short uniseriate trichomes and glandular papillae, with only a few trichomes to 2 mm long present, spreading at anthesis, articulated at the base; pedicels scars closely packed in the distal part of the inflorescence, with the lowermost ca. 1 mm distant from the rest. Buds globose to broadly ellipsoid, the corolla strongly exserted from the calyx tube before anthesis. Flowers 5-merous, all perfect. Calyx tube 1.5-2 mm long, conical to cylindrical, the lobes 0.5-1 mm long, 0.8-1 mm wide, deltate to triangular, the tips obtuse or rounded, densely glandular-pubescent like the pedicels with uniseriate trichomes and papillae. Corolla 10-15 mm in diameter, white or pale purple with a darker brownish purple central star, stellate, lobed 1/2 to 2/3 of the way to the base, the lobes 3.5-5 mm long, 2-3 mm wide, triangular, reflexed to spreading at anthesis, the abaxial surfaces densely papillate, the trichomes not glandular. Stamens equal; filament tube to 0.5 mm; free portion of the filaments 0.5-1 mm long, glabrous or with a few weak tangled simple uniseriate trichomes adaxially; anthers 2.5-3.5 mm long, 0.5-1 mm wide, ellipsoid, bright yellow, poricidal at the tips, the pores elongating to slits with age. Ovary conical, glabrous; style 4.5-6 mm long, sparsely pubescent with weak tangled trichomes to densely papillate in the lower part where included in the anther cone, only slightly (ca. 0.5 mm) exserted from the anther cone; stigma capitate, densely papillate. Fruit a globose berry, 0.5-1 cm in diameter, green to deep purple (red when ripe?
<emphasis italics="true" pageId="80" pageNumber="81">
<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
1211
</emphasis>
); opaque (mature fruits not seen on live plants but not markedly translucent when dry), the pericarp thin, matte; fruiting pedicels
<pageBreakToken pageId="81" pageNumber="82" start="start">6</pageBreakToken>
-9 mm long, enlarging from a base 0.6-1 mm in diameter to an apex 1-1.5 mm in diameter, not distinctly woody, spreading; fruiting calyx not accrescent, the tube less than 1 mm long, the lobes 1.5-2 mm long, appressed to the berry, venation very apparent and thickened. Seeds 10-30 per berry, 1-1.5 mm long, 1-1.2 mm wide, tear-drop shaped, reddish gold, the surfaces minutely putted, the testal cells pentagonal. Stone cells 2-4 (-6) per berry, 0.5-0.7 mm in diameter, pale cream. Chromosome number: not known.
</paragraph>
<caption ID-Zenodo-Dep="3240818" doi="10.3897/phytokeys.123.31738.figure36" httpUri="https://binary.pensoft.net/fig/305335" pageId="81" pageNumber="82" start="Figure 36" startId="F36">
<paragraph pageId="81" pageNumber="82">
<emphasis bold="true" pageId="81" pageNumber="82">Figure 36.</emphasis>
<taxonomicName class="Magnoliopsida" family="Verbenaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pruinosum" order="Solanales" pageId="81" pageNumber="82" phylum="Tracheophyta" rank="species" species="pruinosum">
<emphasis italics="true" pageId="81" pageNumber="82">Solanum pruinosum</emphasis>
</taxonomicName>
Dunal.
<emphasis bold="true" pageId="81" pageNumber="82">A</emphasis>
Habit
<emphasis bold="true" pageId="81" pageNumber="82">B</emphasis>
flower
<emphasis bold="true" pageId="81" pageNumber="82">C</emphasis>
dissected flower
<emphasis bold="true" pageId="81" pageNumber="82">D</emphasis>
calyx
<emphasis bold="true" pageId="81" pageNumber="82">E</emphasis>
infructescence
<emphasis bold="true" pageId="81" pageNumber="82">F</emphasis>
elongate glandular trichome
<emphasis bold="true" pageId="81" pageNumber="82">G</emphasis>
sessile glandular trichome (
<emphasis bold="true" pageId="81" pageNumber="82">A-E</emphasis>
<emphasis italics="true" pageId="81" pageNumber="82">Amith JDA-30248</emphasis>
;
<emphasis bold="true" pageId="81" pageNumber="82">F-G</emphasis>
<emphasis italics="true" pageId="81" pageNumber="82">Ventura A. 2588</emphasis>
). Drawing by L. Ribulgo.
</paragraph>
</caption>
<caption ID-Zenodo-Dep="3240820" doi="10.3897/phytokeys.123.31738.figure37" httpUri="https://binary.pensoft.net/fig/305336" pageId="81" pageNumber="82" start="Figure 37" startId="F37">
<paragraph pageId="81" pageNumber="82">
<emphasis bold="true" pageId="81" pageNumber="82">Figure 37.</emphasis>
<taxonomicName class="Magnoliopsida" family="Verbenaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pruinosum" order="Solanales" pageId="81" pageNumber="82" phylum="Tracheophyta" rank="species" species="pruinosum">
<emphasis italics="true" pageId="81" pageNumber="82">Solanum pruinosum</emphasis>
</taxonomicName>
Dunal.
<emphasis bold="true" pageId="81" pageNumber="82">A</emphasis>
Leaves
<emphasis bold="true" pageId="81" pageNumber="82">B</emphasis>
young branch with flowering buds
<emphasis bold="true" pageId="81" pageNumber="82">C</emphasis>
inflorescence with flowers at full anthesis
<emphasis bold="true" pageId="81" pageNumber="82">D</emphasis>
developing fruits (
<emphasis bold="true" pageId="81" pageNumber="82">A-D</emphasis>
<emphasis italics="true" pageId="81" pageNumber="82">Amith JDA-30248</emphasis>
). Photos by M. Gorostiza Salazar.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="81" pageNumber="82" type="distribution">
<paragraph pageId="81" pageNumber="82">Distribution.</paragraph>
<paragraph pageId="81" pageNumber="82">
(Figure
<figureCitation captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Distribution of Solanum pruinosum Dunal." figureDoi="10.3897/phytokeys.123.31738.figure38" httpUri="https://binary.pensoft.net/fig/305337" pageId="81" pageNumber="82">38</figureCitation>
)
<taxonomicName class="Magnoliopsida" family="Verbenaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pruinosum" order="Solanales" pageId="81" pageNumber="82" phylum="Tracheophyta" rank="species" species="pruinosum">
<emphasis italics="true" pageId="81" pageNumber="82">Solanum pruinosum</emphasis>
</taxonomicName>
occurs from the state of Nuevo
<normalizedToken originalValue="León">Leon</normalizedToken>
in Mexico south to the state of Oaxaca, across the central Volcanic Belt (
<bibRefCitation author="Rzedowski, J" journalOrPublisher="Bulletin of the Torrey Botanical Club" pageId="116" pageNumber="117" refId="B199" refString="Rzedowski, J, 1978. Vegetacion de Mexico. Editorial Limosa, Mexico." title="Vegetacion de Mexico. Editorial Limosa, Mexico." year="1978">Rzedowski 1978</bibRefCitation>
).
</paragraph>
<caption ID-Zenodo-Dep="3240822" doi="10.3897/phytokeys.123.31738.figure38" httpUri="https://binary.pensoft.net/fig/305337" pageId="81" pageNumber="82" start="Figure 38" startId="F38">
<paragraph pageId="81" pageNumber="82">
<emphasis bold="true" pageId="81" pageNumber="82">Figure 38.</emphasis>
Distribution of
<taxonomicName class="Magnoliopsida" family="Verbenaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pruinosum" order="Solanales" pageId="81" pageNumber="82" phylum="Tracheophyta" rank="species" species="pruinosum">
<emphasis italics="true" pageId="81" pageNumber="82">Solanum pruinosum</emphasis>
</taxonomicName>
Dunal.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="81" pageNumber="82" type="ecology">
<paragraph pageId="81" pageNumber="82">Ecology.</paragraph>
<paragraph pageId="81" pageNumber="82">
<taxonomicName class="Magnoliopsida" family="Verbenaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pruinosum" order="Solanales" pageId="81" pageNumber="82" phylum="Tracheophyta" rank="species" species="pruinosum">
<emphasis italics="true" pageId="81" pageNumber="82">Solanum pruinosum</emphasis>
</taxonomicName>
occurs in pine-oak forests, mesophyll forests and open areas from 1,000 to 2,500 m elevation.
</paragraph>
</subSubSection>
<subSubSection pageId="81" pageNumber="82" type="common names">
<paragraph pageId="81" pageNumber="82">Common names.</paragraph>
<paragraph pageId="81" pageNumber="82">
Mexico [Puebla]. Tomakilit (Nahuat,
<emphasis italics="true" pageId="81" pageNumber="82">Amith JDA-2084</emphasis>
), Tomakilit (silvestre) (Nahuat,
<emphasis italics="true" pageId="81" pageNumber="82">
<normalizedToken originalValue="Jiménez">Jimenez</normalizedToken>
Chimil JDA-30248
</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection pageId="81" pageNumber="82" type="uses">
<paragraph pageId="81" pageNumber="82">Uses.</paragraph>
<paragraph pageId="81" pageNumber="82">None recorded.</paragraph>
</subSubSection>
<subSubSection pageId="81" pageNumber="82" type="preliminary conservation status (iucn 2017)">
<paragraph pageId="81" pageNumber="82">
Preliminary conservation status (
<bibRefCitation author="IUCN" journalOrPublisher="Alaska Park Science" pageId="113" pageNumber="114" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B115" refString="IUCN, 2017. Guidelines for Using the IUCN Red List Categories and Criteria. Version 13. Prepared by the Standards and Petitions Subcommittee. http://www.iucnredlist.org/documents/RedListGuidelines.pdf" title="Guidelines for Using the IUCN Red List Categories and Criteria. Version 13. Prepared by the Standards and Petitions Subcommittee." url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2017">IUCN 2017</bibRefCitation>
).
</paragraph>
<paragraph pageId="81" pageNumber="82">
Least Concern (LC).
<taxonomicName class="Magnoliopsida" family="Verbenaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pruinosum" order="Solanales" pageId="81" pageNumber="82" phylum="Tracheophyta" rank="species" species="pruinosum">
<emphasis italics="true" pageId="81" pageNumber="82">Solanum pruinosum</emphasis>
</taxonomicName>
is widespread in Mexico but is not as common as
<taxonomicName family="Verbenaceae" kingdom="Plantae" lsidName="S. nigrescens" order="Solanales" pageId="81" pageNumber="82" rank="species" species="nigrescens">
<emphasis italics="true" pageId="81" pageNumber="82">S. nigrescens</emphasis>
</taxonomicName>
. For EOO see Table
<tableCitation captionStart="Table 6" captionStartId="T6" captionText="Table 6. Preliminary conservation assessments for morelloid species from the Caribbean and North and Central America. For details see Materials and Methods and individual species treatments. Preliminary assessments are based on EOO only (see Materials and Methods) and have been calculated for worldwide ranges for each species. The EOO and conservation status of species known to be solely cultivated, introduced or adventive in the region has been assessed in Saerkinen et al. (2018)." httpUri="http://table.plazi.org/id/C3922CEBD29B29D2373D9CB546F06573" pageId="81" pageNumber="82" tableUuid="C3922CEBD29B29D2373D9CB546F06573">6</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82" type="discussion">
<paragraph pageId="81" pageNumber="82">Discussion.</paragraph>
<paragraph lastPageId="82" lastPageNumber="83" pageId="81" pageNumber="82">
<taxonomicName class="Magnoliopsida" family="Verbenaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum pruinosum" order="Solanales" pageId="81" pageNumber="82" phylum="Tracheophyta" rank="species" species="pruinosum">
<emphasis italics="true" pageId="81" pageNumber="82">Solanum pruinosum</emphasis>
</taxonomicName>
differs from
<taxonomicName family="Verbenaceae" kingdom="Plantae" lsidName="S. nigrescens" order="Solanales" pageId="81" pageNumber="82" rank="species" species="nigrescens">
<emphasis italics="true" pageId="81" pageNumber="82">S. nigrescens</emphasis>
</taxonomicName>
and
<taxonomicName family="Verbenaceae" kingdom="Plantae" lsidName="S. douglasii" order="Solanales" pageId="81" pageNumber="82" rank="species" species="douglasii">
<emphasis italics="true" pageId="81" pageNumber="82">S. douglasii</emphasis>
</taxonomicName>
, with which it is sympatric, in its glandular pubescence. It is possible that these specimens represent isolated glandular populations of those two taxa, but in the absence of data showing this we elect to recognise these populations at the species level until further
<pageBreakToken pageId="82" pageNumber="83" start="start">work</pageBreakToken>
across the range in Mexico is done. The three taxa share numerous stone cells (ca. 5-6, more in
<taxonomicName family="Verbenaceae" kingdom="Plantae" lsidName="S. nigrescens" order="Solanales" pageId="82" pageNumber="83" rank="species" species="nigrescens">
<emphasis italics="true" pageId="82" pageNumber="83">S. nigrescens</emphasis>
</taxonomicName>
) in the ripe berries, and subumbellate to somewhat
<normalizedToken originalValue="“racemose”">&quot;racemose&quot;</normalizedToken>
inflorescences. The flowers of
<taxonomicName family="Verbenaceae" kingdom="Plantae" lsidName="S. pruinosum" order="Solanales" pageId="82" pageNumber="83" rank="species" species="pruinosum">
<emphasis italics="true" pageId="82" pageNumber="83">S. pruinosum</emphasis>
</taxonomicName>
are intermediate in size between
<taxonomicName family="Verbenaceae" kingdom="Plantae" lsidName="S. douglasii" order="Solanales" pageId="82" pageNumber="83" rank="species" species="douglasii">
<emphasis italics="true" pageId="82" pageNumber="83">S. douglasii</emphasis>
</taxonomicName>
(15-20 mm in diameter) and
<taxonomicName family="Verbenaceae" kingdom="Plantae" lsidName="S. nigrescens" order="Solanales" pageId="82" pageNumber="83" rank="species" species="nigrescens">
<emphasis italics="true" pageId="82" pageNumber="83">S. nigrescens</emphasis>
</taxonomicName>
(8-10 mm).
</paragraph>
<paragraph pageId="82" pageNumber="83">
Label data from
<emphasis italics="true" pageId="82" pageNumber="83">
<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
1211
</emphasis>
(MO) note the berries as
<normalizedToken originalValue="“rojo”">&quot;rojo&quot;</normalizedToken>
(red), but no other specimens have this data, so we suspect it is either a mistake, or an interpretation of purplish red.
</paragraph>
<paragraph pageId="82" pageNumber="83">
The locality on the type specimen of
<taxonomicName family="Verbenaceae" kingdom="Plantae" lsidName="S. pruinosum" order="Solanales" pageId="82" pageNumber="83" rank="species" species="pruinosum">
<emphasis italics="true" pageId="82" pageNumber="83">S. pruinosum</emphasis>
</taxonomicName>
is only given as &quot;circa Mexico&quot;, but is likely to have been collected in central Mexico; Berlandier was in Mexico City and vicinity from the time of his arrival in Mexico in 1826 until his journey north to the Tamaulipas-Texas borderlands in late 1827 (
<bibRefCitation author="Lawson, RM" journalOrPublisher="Taxon" pageId="113" pageNumber="114" refId="B142" refString="Lawson, RM, 2012. Frontier naturalist: Jean Louis Berlandier and the exploration of northern Mexico and Texas. UNM Press, Albuquerque." title="Frontier naturalist: Jean Louis Berlandier and the exploration of northern Mexico and Texas. UNM Press, Albuquerque." year="2012">Lawson 2012</bibRefCitation>
).
</paragraph>
<paragraph pageId="82" pageNumber="83">
We have not been able to trace any duplicates of the type specimens of
<taxonomicName family="Verbenaceae" kingdom="Plantae" lsidName="S. dasyadenium" order="Solanales" pageId="82" pageNumber="83" rank="species" species="dasyadenium">S. dasyadenium</taxonomicName>
and var.
<taxonomicName family="Verbenaceae" kingdom="Plantae" lsidName="var. uberius" order="Solanales" pageId="82" pageNumber="83" rank="variety" species="dasyadenium" variety="uberius">uberius</taxonomicName>
, both described from material held in Berlin (
<bibRefCitation DOI="https://doi.org/10.1002/fedr.19120110103" author="Bitter, G" journalOrPublisher="Botanische Jahrbuecher fuer Systematik, Pflanzengeschichte und Pflanzengeographie" pageId="107" pageNumber="108" refId="B20" refString="Bitter, G, 1912a. Solana nova vel minus cognita II. VIII. Weltere sudamerikanische Morellae. Repertorium Species Novarum Regni Vegetabilium 11(1-3): 1-9. https://doi.org/10.1002/fedr.19120110103" title="Solana nova vel minus cognita II. VIII. Weltere sudamerikanische Morellae. Repertorium Species Novarum Regni Vegetabilium 11 (1 - 3): 1 - 9." url="https://doi.org/10.1002/fedr.19120110103" year="1912 a">Bitter 1912a</bibRefCitation>
) and subsequently destroyed. Until we better understand the range of variation in these taxa, and their relationship to
<taxonomicName family="Verbenaceae" kingdom="Plantae" lsidName="S. douglasii" order="Solanales" pageId="82" pageNumber="83" rank="species" species="douglasii">
<emphasis italics="true" pageId="82" pageNumber="83">S. douglasii</emphasis>
</taxonomicName>
and
<taxonomicName family="Verbenaceae" kingdom="Plantae" lsidName="S. nigrescens" order="Solanales" pageId="82" pageNumber="83" rank="species" species="nigrescens">
<emphasis italics="true" pageId="82" pageNumber="83">S. nigrescens</emphasis>
</taxonomicName>
, we prefer not to neotypify these names at present.
</paragraph>
</subSubSection>
<subSubSection pageId="82" pageNumber="83" type="materials_examined">
<paragraph pageId="82" pageNumber="83">Specimens examined.</paragraph>
<paragraph pageId="82" pageNumber="83">See Suppl. materials 1 and 3.</paragraph>
</subSubSection>
</treatment>
</document>