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<document id="F8A5A1744B805A32C321B17D0CA7E5D3" ID-DOI="10.5281/zenodo.210542" ID-GBIF-Dataset="c9e5cc54-2a89-4b75-91aa-a4fce79f17fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="210542" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460500319311" checkinUser="plazi" docAuthor="Ramírez-Ponce, Andrés &amp; Morón, Miguel Angel" docDate="2012" docId="A567C22F585EFF98FF64A4A3534483EC" docLanguage="en" docName="zt03394p024.pdf" docOrigin="Zootaxa 3394" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pachystethus matzapatlecus Ramirez-Ponce &amp; Moron, new species" docType="treatment" docVersion="9" lastPageNumber="17" masterDocId="595EBA575850FF88FFF3A25B52478434" masterDocTitle="Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="15" updateTime="1698311598655" updateUser="plazi">
<mods:mods id="4C4F021EAF5A61880BBD7407B824A2FA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="030B17B928E73E2E51F3442F4D106489">
<mods:title id="AC99D62B5FC6EADFCDC44D5CB1E0C42D">Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)</mods:title>
</mods:titleInfo>
<mods:name id="E5CF273E3B52489150C3DEA013145935" type="personal">
<mods:role id="32E1DFFFEB84F95AEAE126B9B1C17E36">
<mods:roleTerm id="76067E2EAFBF45BBA0813819E9E81D9F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F65DBF1EED7C723EFAF5B62663D0AEC3">Ramírez-Ponce, Andrés</mods:namePart>
</mods:name>
<mods:name id="E8A24A97B2DA3F5AA47C3D4DE5219977" type="personal">
<mods:role id="4488AC825CABE7B16E655BF045D1346D">
<mods:roleTerm id="72E0A082DA789315B4EFFE59C42E18E6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="09DC42B9F689DEE811F2841C528B9463">Morón, Miguel Angel</mods:namePart>
</mods:name>
<mods:typeOfResource id="5E0C11581B12CECCB48851A5805A976F">text</mods:typeOfResource>
<mods:relatedItem id="27ECCAC948DE32B2812A94E44FC29EB6" type="host">
<mods:titleInfo id="DF221C83573D51B65302203B9FFBB9EC">
<mods:title id="B5C4C6D9200C7368546BB336D3EA024C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="4F3B525AEE07B36169D2C4AB03E688B0">
<mods:date id="67A8B3945ED26546177DE63B588A0178">2012</mods:date>
<mods:detail id="A1D54249ED58319E1FE9257024063AD5" type="volume">
<mods:number id="9B203962847DFE1E55BB62D9EE54B830">3394</mods:number>
</mods:detail>
<mods:extent id="35E88D47CD38A646C18873DDF034A3A5" unit="page">
<mods:start id="C45E46526690837A48D6084267F1A4EE">1</mods:start>
<mods:end id="5ED8B586D917C5F0792EB884CA371326">24</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="39F4C9D3E2DFAA1DF724568410856533">journal article</mods:classification>
<mods:identifier id="42328470ABFEA87F5692962E4E05BAD7" type="DOI">10.5281/zenodo.210542</mods:identifier>
<mods:identifier id="9BDA0B328F1FD7E5CE8C681A42DCD6AA" type="GBIF-Dataset">c9e5cc54-2a89-4b75-91aa-a4fce79f17fc</mods:identifier>
<mods:identifier id="E7421FDE64B9DC9A50B7EC4CDD2CE77E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="52D6D3D3AAE7E9150903B787765B0DF3" type="Zenodo-Dep">210542</mods:identifier>
</mods:mods>
<treatment id="A567C22F585EFF98FF64A4A3534483EC" ID-DOI="http://doi.org/10.5281/zenodo.6179202" ID-GBIF-Taxon="119521050" ID-Zenodo-Dep="6179202" LSID="urn:lsid:plazi:treatment:A567C22F585EFF98FF64A4A3534483EC" httpUri="http://treatment.plazi.org/id/A567C22F585EFF98FF64A4A3534483EC" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
<subSubSection id="65D420B2585EFF86FF64A4A353E88307" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="2D717339585EFF86FF64A4A3518C8326" blockId="14.[151,971,1784,1843]" box="[151,971,1784,1810]" pageId="14" pageNumber="15">
<heading id="7639C455585EFF86FF64A4A3518C8326" bold="true" box="[151,971,1784,1810]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<emphasis id="1FBAAF2B585EFF86FF64A4A3518C8326" bold="true" box="[151,971,1784,1810]" pageId="14" pageNumber="15">
<taxonomicName id="EACE08BA585EFF86FF64A4A351698326" authority="Ramirez-Ponce &amp; Moron" authorityName="Ramirez-Ponce &amp; Moron" box="[151,814,1784,1810]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="matzapatlecus" status="sp. nov.">
<emphasis id="1FBAAF2B585EFF86FF64A4A353AE8326" bold="true" box="[151,489,1784,1810]" italics="true" pageId="14" pageNumber="15">Pachystethus matzapatlecus</emphasis>
Ramírez-Ponce &amp; Morón
</taxonomicName>
,
<taxonomicNameLabel id="04891250585EFF86FCC8A4A3518C8326" box="[827,971,1784,1810]" pageId="14" pageNumber="15" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2D717339585EFF86FF6DA54053E88307" blockId="14.[151,971,1784,1843]" box="[158,431,1819,1843]" pageId="14" pageNumber="15">
(
<figureCitation id="B5F56FBC585EFF86FF55A54053698307" box="[166,302,1819,1843]" captionStart="FIGURES 26 – 35" captionStartId="15.[151,264,1938,1961]" captionTargetBox="[238,1394,484,1899]" captionTargetId="figure@15.[151,1436,445,1918]" captionTargetPageId="15" captionText="FIGURES 26 – 35. Diagnostic structures in P. matzapatlecus: 26) right proleg, 27) right antenna, 28) male genitalia in dorsal view, 29) male genitalia in lateral view, 30) spiculum gastrale, 31) female genital plates, 32) labium, 33) labrum, 34) right maxilla, 35) left mandible. Scale lines 1 mm." httpUri="https://zenodo.org/record/210548/files/figure.png" pageId="14" pageNumber="15">Figs. 26–35</figureCitation>
,
<figureCitation id="B5F56FBC585EFF86FECAA54053108307" box="[313,343,1819,1843]" captionStart="FIGURES 42 – 47" captionStartId="19.[151,264,1909,1932]" captionTargetBox="[214,1377,207,1832]" captionTargetId="figure@19.[198,1392,193,1847]" captionTargetPageId="19" captionText="FIGURES 42 – 47. Dorsal view of Pachystethus species: 42 a, b) P. viduus males; 42 c) P. viduus female; 42 d) P. viduus female, lectotype; 43 a) P. nectoctenus male; 43 b) P. nectoctenus female; 44 a) P. crassesculptus male; 44 b) P. crassesculptus female; 45 a, b) P. nutans males; 45 c) P. nu t an s female; 46 a, b) P. matzapatlecus males; 46 c) P. matzapatlecus female; 47 a) P. ixtacomitanus male; 47 b) P. ixtacomitanus female. Scale lines 1 mm." httpUri="https://zenodo.org/record/210550/files/figure.png" pageId="14" pageNumber="15">46</figureCitation>
a–c, 52)
</paragraph>
</subSubSection>
<subSubSection id="65D420B2585EFF87FF64A53F508D85B3" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="2D717339585EFF86FF64A53F57DB83D3" blockId="14.[151,1436,1891,2024]" pageId="14" pageNumber="15">
<emphasis id="1FBAAF2B585EFF86FF64A53F53068348" bold="true" box="[151,321,1891,1916]" pageId="14" pageNumber="15">
<typeStatus id="F275CD9B585EFF86FF64A53F52958348" box="[151,210,1892,1916]" pageId="14" pageNumber="15">Type</typeStatus>
material
</emphasis>
(
<specimenCount id="3BC8B8B0585EFF86FEA7A5385387834F" box="[340,448,1891,1915]" pageId="14" pageNumber="15" type="male">20 males</specimenCount>
,
<specimenCount id="3BC8B8B0585EFF86FE23A5385005834F" box="[464,578,1891,1915]" pageId="14" pageNumber="15" type="female">5 females</specimenCount>
).
<emphasis id="1FBAAF2B585EFF86FDA8A53F508F8348" bold="true" box="[603,712,1892,1916]" pageId="14" pageNumber="15">
<typeStatus id="F275CD9B585EFF86FDA8A53F508F8348" box="[603,712,1892,1916]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
</emphasis>
male housed at
<collectionCode id="4BDFEBFC585EFF86FC7DA53F51968348" box="[910,977,1892,1916]" pageId="14" pageNumber="15">IEXA</collectionCode>
with following label data: a) “
<collectingCountry id="55D933A9585EFF86FACEA53857DF834F" box="[1341,1432,1891,1915]" name="Mexico" pageId="14" pageNumber="15">México</collectingCountry>
, Oaxaca, Santiago Comaltepec, La Esperanza,
<date id="597055F9585EFF86FD5AA5DC516B83AB" box="[681,812,1927,1951]" pageId="14" pageNumber="15" value="1931-08-05">31-VIII-05.</date>
Mora, E. col.” (typeface), b) “Colecta directa en flor,
<quantity id="EA36DEDC585EFF86FF64A5F052B583F7" box="[151,242,1963,1987]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="14" pageNumber="15" unit="m" value="1550.0">1550 m</quantity>
, Bosque mesófilo de montaña-
<taxonomicName id="EACE08BA585EFF86FDADA5F7512283F7" box="[606,869,1964,1987]" class="Magnoliopsida" family="Juglandaceae" genus="Oreomunnea" kingdom="Plantae" order="Fagales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="mexicana">
<emphasis id="1FBAAF2B585EFF86FDADA5F7512283F7" box="[606,869,1964,1987]" italics="true" pageId="14" pageNumber="15">Oreomunnea mexicana</emphasis>
</taxonomicName>
” (typeface), c) our
<typeStatus id="F275CD9B585EFF86FBA1A5F056F383F7" box="[1106,1204,1963,1987]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
red label.
<emphasis id="1FBAAF2B585EFF86FAC4A5F057DB83F0" bold="true" box="[1335,1436,1963,1988]" pageId="14" pageNumber="15">Allotype</emphasis>
female housed at
<collectionCode id="4BDFEBFC585EFF86FE99A58B53EA83DC" box="[362,429,2000,2024]" pageId="14" pageNumber="15">IEXA</collectionCode>
with following label data: a, b) same data as
<typeStatus id="F275CD9B585EFF86FC2BA594567983D3" box="[984,1086,1999,2023]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
; c) our allotype yellow label.
</paragraph>
<paragraph id="2D717339585FFF87FF64A2C3508D85B3" blockId="15.[151,1436,151,391]" pageId="15" pageNumber="16">
<emphasis id="1FBAAF2B585FFF87FF64A2C353488484" bold="true" box="[151,271,152,176]" pageId="15" pageNumber="16">
<typeStatus id="F275CD9B585FFF87FF64A2C353488484" box="[151,271,152,176]" pageId="15" pageNumber="16" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="3BC8B8B0585FFF87FEECA2CC53C0849B" box="[287,391,151,175]" pageId="15" pageNumber="16" type="male">19 males</specimenCount>
,
<specimenCount id="3BC8B8B0585FFF87FE60A2CC5043849B" box="[403,516,151,175]" pageId="15" pageNumber="16" type="female">4 females</specimenCount>
;
<collectionCode id="4BDFEBFC585FFF87FDE3A2CC5010849B" box="[528,599,151,175]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="15" pageNumber="16">CNIN</collectionCode>
,
<collectionCode id="4BDFEBFC585FFF87FD90A2C350EE8484" box="[611,681,152,176]" pageId="15" pageNumber="16">IEXA</collectionCode>
,
<collectionCode id="4BDFEBFC585FFF87FD47A2CC514F849B" box="[692,776,151,175]" pageId="15" pageNumber="16">MXAL</collectionCode>
) with our
<typeStatus id="F275CD9B585FFF87FC8DA2C351998484" box="[894,990,152,176]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
yellow labels and following label data: “A. Ibarra.
<collectingCountry id="55D933A9585FFF87FEE6A2E753C084E0" box="[277,391,188,212]" name="Mexico" pageId="15" pageNumber="16">MEXICO</collectingCountry>
. Oaxaca, Puerto Eligio, S. de Juarez
<date id="597055F9585FFF87FCD2A2E751CC84E0" box="[801,907,188,212]" pageId="15" pageNumber="16" value="1982-09-12">12-IX-82</date>
” (handwritten) (
<specimenCount id="3BC8B8B0585FFF87FBB6A2E756D984E0" box="[1093,1182,188,212]" pageId="15" pageNumber="16" type="male">7 males</specimenCount>
,
<specimenCount id="3BC8B8B0585FFF87FB5AA2E7574A84E0" box="[1193,1293,188,212]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
;
<collectionCode id="4BDFEBFC585FFF87FAEBA2E7572684E0" box="[1304,1377,188,212]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="15" pageNumber="16">CNIN</collectionCode>
); “A. Ibarra.
<collectingCountry id="55D933A9585FFF87FF1FA284531984C3" box="[236,350,223,247]" name="Mexico" pageId="15" pageNumber="16">MEXICO</collectingCountry>
. Oaxaca, Metates, S. de Juárez.
<date id="597055F9585FFF87FD2CA284510F84C3" box="[735,840,223,247]" pageId="15" pageNumber="16" value="1982-09-16">16-IX-82</date>
” (handwritten) (
<specimenCount id="3BC8B8B0585FFF87FBF8A284562E84C3" box="[1035,1129,223,247]" pageId="15" pageNumber="16" type="male">8 males</specimenCount>
,
<specimenCount id="3BC8B8B0585FFF87FB8BA28456AB84C3" box="[1144,1260,223,247]" pageId="15" pageNumber="16" type="female">2 females</specimenCount>
;
<collectionCode id="4BDFEBFC585FFF87FB08A284570584C3" box="[1275,1346,223,247]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="15" pageNumber="16">CNIN</collectionCode>
,
<collectionCode id="4BDFEBFC585FFF87FAA1A2BB57DF84CC" box="[1362,1432,224,248]" pageId="15" pageNumber="16">IEXA</collectionCode>
,
<collectionCode id="4BDFEBFC585FFF87FF64A35F52A88528" box="[151,239,260,284]" pageId="15" pageNumber="16">MXAL</collectionCode>
); “
<collectingCountry id="55D933A9585FFF87FEFDA35F532D8528" box="[270,362,260,284]" name="Mexico" pageId="15" pageNumber="16">Mexico</collectingCountry>
: Oaxaca Metates Sierra de Juárez.
<date id="597055F9585FFF87FD09A35F51158528" box="[762,850,260,284]" pageId="15" pageNumber="16" value="1983-03-08">8-III-83</date>
J. Bueno” (handwritten) (
<specimenCount id="3BC8B8B0585FFF87FB73A35F56898528" box="[1152,1230,260,284]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<collectionCode id="4BDFEBFC585FFF87FB28A35F57638528" box="[1243,1316,260,284]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="15" pageNumber="16">CNIN</collectionCode>
); “Km. 65 carr. Tuxtepec Oaxaca,
<collectingCountry id="55D933A9585FFF87FE57A37C53BE850B" box="[420,505,295,319]" name="Mexico" pageId="15" pageNumber="16">México</collectingCountry>
<date id="597055F9585FFF87FDF1A37C502D850B" box="[514,618,295,319]" pageId="15" pageNumber="16" value="1982-09-18">18-IX-82</date>
A. Ibarra” (handwritten) (
<specimenCount id="3BC8B8B0585FFF87FC6AA37C51B3850B" box="[921,1012,295,319]" pageId="15" pageNumber="16" type="male">2 males</specimenCount>
,
<collectionCode id="4BDFEBFC585FFF87FBF1A37C560B850B" box="[1026,1100,295,319]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="15" pageNumber="16">CNIN</collectionCode>
); “A. Ibarra,
<collectingCountry id="55D933A9585FFF87FB10A37C577F850B" box="[1251,1336,295,319]" name="Mexico" pageId="15" pageNumber="16">México</collectingCountry>
Oaxaca. Km. 65 Carr. Tuxtepec. Oax.
<date id="597055F9585FFF87FE03A317501D8550" box="[496,602,332,356]" pageId="15" pageNumber="16" value="1982-09-18">18-IX-82</date>
” (handwritten) (
<specimenCount id="3BC8B8B0585FFF87FCEFA31751C68550" box="[796,897,332,356]" pageId="15" pageNumber="16" type="female">1 female</specimenCount>
,
<collectionCode id="4BDFEBFC585FFF87FC7CA317519E8550" box="[911,985,332,356]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="15" pageNumber="16">CNIN</collectionCode>
); “
<quantity id="EA36DEDC585FFF87FC08A31756708550" box="[1019,1079,332,356]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="16" unit="km" value="5.0">5 km</quantity>
W La Esperanza,
<date id="597055F9585FFF87FAE2A31757DF8550" box="[1297,1432,332,356]" pageId="15" pageNumber="16" value="1977-09-27">27-IX-1977</date>
,
<quantity id="EA36DEDC585FFF87FF64A33452B285B3" box="[151,245,367,391]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="15" pageNumber="16" unit="m" value="1500.0">1,500 m</quantity>
, J. Llorente” (typeface) (
<specimenCount id="3BC8B8B0585FFF87FDFCA334501C85B3" box="[527,603,367,391]" pageId="15" pageNumber="16" type="male">1 male</specimenCount>
,
<collectionCode id="4BDFEBFC585FFF87FD94A33450F885B3" box="[615,703,367,391]" pageId="15" pageNumber="16">MXAL</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="79B123B1585FFF87FF64A5C9507383D2" httpUri="https://zenodo.org/record/210548/files/figure.png" pageId="15" pageNumber="16" targetBox="[238,1394,484,1899]" targetPageId="15">
<paragraph id="2D717339585FFF87FF64A5C9507383D2" blockId="15.[151,1436,1938,2023]" pageId="15" pageNumber="16">
<emphasis id="1FBAAF2B585FFF87FF64A5C95309839C" bold="true" box="[151,334,1938,1961]" pageId="15" pageNumber="16">FIGURES 26–35</emphasis>
. Diagnostic structures in
<taxonomicName id="EACE08BA585FFF87FDABA5CF5143839D" box="[600,772,1939,1961]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="matzapatlecus">
<emphasis id="1FBAAF2B585FFF87FDABA5CF5143839D" box="[600,772,1939,1961]" italics="true" pageId="15" pageNumber="16">P. matzapatlecus</emphasis>
</taxonomicName>
:
<emphasis id="1FBAAF2B585FFF87FCE1A5C95173839D" bold="true" box="[786,820,1938,1961]" pageId="15" pageNumber="16">26)</emphasis>
right proleg,
<emphasis id="1FBAAF2B585FFF87FC31A5C951A3839D" bold="true" box="[962,996,1938,1961]" pageId="15" pageNumber="16">27)</emphasis>
right antenna,
<emphasis id="1FBAAF2B585FFF87FB72A5C956E5839D" bold="true" box="[1153,1186,1938,1961]" pageId="15" pageNumber="16">28)</emphasis>
male genitalia in dorsal view,
<emphasis id="1FBAAF2B585FFF87FF20A5EA52B383FC" bold="true" box="[211,244,1969,1992]" pageId="15" pageNumber="16">29)</emphasis>
male genitalia in lateral view,
<emphasis id="1FBAAF2B585FFF87FDD9A5EA500C83FC" bold="true" box="[554,587,1969,1992]" pageId="15" pageNumber="16">30)</emphasis>
spiculum gastrale,
<emphasis id="1FBAAF2B585FFF87FCFCA5EA517783FC" bold="true" box="[783,816,1969,1992]" pageId="15" pageNumber="16">31)</emphasis>
female genital plates,
<emphasis id="1FBAAF2B585FFF87FBE7A5EA567183FC" bold="true" box="[1044,1078,1969,1992]" pageId="15" pageNumber="16">32)</emphasis>
labium,
<emphasis id="1FBAAF2B585FFF87FB7DA5EA56F783FC" bold="true" box="[1166,1200,1969,1992]" pageId="15" pageNumber="16">33)</emphasis>
labrum,
<emphasis id="1FBAAF2B585FFF87FAF9A5EA576B83FC" bold="true" box="[1290,1324,1969,1992]" pageId="15" pageNumber="16">34)</emphasis>
right maxilla,
<emphasis id="1FBAAF2B585FFF87FF37A58B52A283D3" bold="true" box="[196,229,2000,2023]" pageId="15" pageNumber="16">35)</emphasis>
left mandible. Scale lines 1 mm.
</paragraph>
</caption>
<subSubSection id="65D420B25840FF98FF3AA2CC53B48197" pageId="16" pageNumber="17" type="description">
<paragraph id="2D7173395840FF98FF3AA2CC518780BD" blockId="16.[151,1437,151,2008]" pageId="16" pageNumber="17">
<emphasis id="1FBAAF2B5840FF98FF3AA2CC539B8484" bold="true" box="[201,476,151,176]" pageId="16" pageNumber="17">
Description.
<typeStatus id="F275CD9B5840FF98FE95A2C353908484" box="[358,471,152,176]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
:
</emphasis>
male. Length 9.0 mm, width 4.0 mm. Color shiny black on almost entire body; reddish dark brown on legs.
<emphasis id="1FBAAF2B5840FF98FE25A2E0505D84E7" box="[470,538,187,211]" italics="true" pageId="16" pageNumber="17">Head.</emphasis>
Clypeus subtrapezoidal, anterior border straight, briefly raised, and not directed forward; anterior angles widely rounded; clypeal surface rugose with abundant large punctures. Frontoclypeal suture straight, clearly defined. Frons slightly convex, glabrous, densely punctate; puctures rugose at center, more disperse on lateral and posterior margins. Punctures around eyes scattered. Antennal club slightly longer than the length of all preceding antennomeres combined; lamellar shape semiovate, nearly straight (
<figureCitation id="B5F56FBC5840FF98FB6AA31C56A9856B" box="[1177,1262,327,351]" captionStart="FIGURES 26 – 35" captionStartId="15.[151,264,1938,1961]" captionTargetBox="[238,1394,484,1899]" captionTargetId="figure@15.[151,1436,445,1918]" captionTargetPageId="15" captionText="FIGURES 26 – 35. Diagnostic structures in P. matzapatlecus: 26) right proleg, 27) right antenna, 28) male genitalia in dorsal view, 29) male genitalia in lateral view, 30) spiculum gastrale, 31) female genital plates, 32) labium, 33) labrum, 34) right maxilla, 35) left mandible. Scale lines 1 mm." httpUri="https://zenodo.org/record/210548/files/figure.png" pageId="16" pageNumber="17">Fig. 27</figureCitation>
). Eyes medium size, ovate; ocular margin dorsally noticeable and setose. Eye canthus thick with glabrous, rounded apex and flattened surface; surface with shallow minute punctures. Mouthparts (
<figureCitation id="B5F56FBC5840FF98FC45A3D556058592" box="[950,1090,398,422]" captionStart="FIGURES 26 – 35" captionStartId="15.[151,264,1938,1961]" captionTargetBox="[238,1394,484,1899]" captionTargetId="figure@15.[151,1436,445,1918]" captionTargetPageId="15" captionText="FIGURES 26 – 35. Diagnostic structures in P. matzapatlecus: 26) right proleg, 27) right antenna, 28) male genitalia in dorsal view, 29) male genitalia in lateral view, 30) spiculum gastrale, 31) female genital plates, 32) labium, 33) labrum, 34) right maxilla, 35) left mandible. Scale lines 1 mm." httpUri="https://zenodo.org/record/210548/files/figure.png" pageId="16" pageNumber="17">Figs. 32–35</figureCitation>
): maxillary basistipe flattened, with irregular punctation; labrum wide and sinuate; labium sinuate, as long as wide, with irregularly rugopunctate, setose prementum.
<emphasis id="1FBAAF2B5840FF98FE83A38D539885D9" box="[368,479,470,493]" italics="true" pageId="16" pageNumber="17">Pronotum</emphasis>
: wider than long, moderately convex; surface with scattered, small, round punctures; lateral borders angled with bulky sides and large, deep lateral pits; posterior angles raised. Mesoepimera briefly exposed over humeral calli. Space between mesocoxae wide; mesometasternal process long and stout, briefly projecting anteriorly past mesocoxa, with wide, rounded apex directed downward; metasternum covered with numerous wide punctures, sides moderately setiferous, mesial region glabrous with deep longitudinal sulcus.
<emphasis id="1FBAAF2B5840FF98FF64A0DE529A86A9" box="[151,221,645,669]" italics="true" pageId="16" pageNumber="17">Elytra</emphasis>
flattened and ovate, striae weakly impressed with disperse and shallow, small punctures; elytral costae well defined; elytral suture raised on posterior third; brief preapical slope; lateral margins nearly straight with scattered stout, short setae on epipleural area and other slender setae near margin. Epipleural fold narrow, briefly excavated in lateral view, abruptly narrowed after metacoxa.
<taxonomicName id="EACE08BA5840FF98FD15A0B551138732" box="[742,852,750,774]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="1FBAAF2B5840FF98FD15A0B551138732" box="[742,852,750,774]" italics="true" pageId="16" pageNumber="17">Scutellum</emphasis>
</taxonomicName>
: slightly longer than wide; lateral margins briefly angled, apex rounded; surface with irregular, large punctures.
<emphasis id="1FBAAF2B5840FF98FC91A14A518B871D" box="[866,972,785,809]" italics="true" pageId="16" pageNumber="17">Abdomen</emphasis>
: sternites with scattered setae on nearly continuous row of irregular, large punctures.
<emphasis id="1FBAAF2B5840FF98FD69A16D50888779" box="[666,719,822,845]" italics="true" pageId="16" pageNumber="17">Legs</emphasis>
: protarsomeres 1–4 short and wide, as wide as protarsomere 5 and equal in length. Protibia stout, progressively widened distally, lateral borders divergent towards apex; external border with proximal denticle short and rounded; distal denticle long, slightly curved (
<figureCitation id="B5F56FBC5840FF98FBA8A12056E887A7" box="[1115,1199,891,915]" captionStart="FIGURES 26 – 35" captionStartId="15.[151,264,1938,1961]" captionTargetBox="[238,1394,484,1899]" captionTargetId="figure@15.[151,1436,445,1918]" captionTargetPageId="15" captionText="FIGURES 26 – 35. Diagnostic structures in P. matzapatlecus: 26) right proleg, 27) right antenna, 28) male genitalia in dorsal view, 29) male genitalia in lateral view, 30) spiculum gastrale, 31) female genital plates, 32) labium, 33) labrum, 34) right maxilla, 35) left mandible. Scale lines 1 mm." httpUri="https://zenodo.org/record/210548/files/figure.png" pageId="16" pageNumber="17">Fig. 26</figureCitation>
). Mesofemora with a well-defined longitudinal keel and setiferous punctures. Metafemora without keels. Metatibia with 3 deep transverse keels and spine-like, short setae; apex of metatibia clearly swollen. Metatarsomeres 1–2 longer than tarsomeres 3–5.
<emphasis id="1FBAAF2B5840FF98FEA6A1BE53F887C9" box="[341,447,997,1021]" italics="true" pageId="16" pageNumber="17">Genitalia</emphasis>
(
<figureCitation id="B5F56FBC5840FF98FE22A1BF501C87C8" box="[465,603,996,1020]" captionStart="FIGURES 26 – 35" captionStartId="15.[151,264,1938,1961]" captionTargetBox="[238,1394,484,1899]" captionTargetId="figure@15.[151,1436,445,1918]" captionTargetPageId="15" captionText="FIGURES 26 – 35. Diagnostic structures in P. matzapatlecus: 26) right proleg, 27) right antenna, 28) male genitalia in dorsal view, 29) male genitalia in lateral view, 30) spiculum gastrale, 31) female genital plates, 32) labium, 33) labrum, 34) right maxilla, 35) left mandible. Scale lines 1 mm." httpUri="https://zenodo.org/record/210548/files/figure.png" pageId="16" pageNumber="17">Figs. 28–31</figureCitation>
) parameres complex; shorter than the length of tectum; with upper part irregular; apices narrowed, upturned; lower part of parameres briefly curved backwards. Basal half of tectum wider than distal half, with outstanding posterolateral calli. Basal piece with long and rounded, pointed apodema.
<emphasis id="1FBAAF2B5840FF98FF64A61553258052" box="[151,354,1102,1126]" italics="true" pageId="16" pageNumber="17">Spiculum gastrale</emphasis>
(
<figureCitation id="B5F56FBC5840FF98FE82A61553858052" box="[369,450,1102,1126]" captionStart="FIGURES 26 – 35" captionStartId="15.[151,264,1938,1961]" captionTargetBox="[238,1394,484,1899]" captionTargetId="figure@15.[151,1436,445,1918]" captionTargetPageId="15" captionText="FIGURES 26 – 35. Diagnostic structures in P. matzapatlecus: 26) right proleg, 27) right antenna, 28) male genitalia in dorsal view, 29) male genitalia in lateral view, 30) spiculum gastrale, 31) female genital plates, 32) labium, 33) labrum, 34) right maxilla, 35) left mandible. Scale lines 1 mm." httpUri="https://zenodo.org/record/210548/files/figure.png" pageId="16" pageNumber="17">Fig. 30</figureCitation>
) narrow; basal part straight with uniform width, shorter than lateral sclerites; associated plates large, subrounded, with numerous long setae near exterior border.
</paragraph>
<paragraph id="2D7173395840FF98FF3AA6CF53478122" blockId="16.[151,1437,151,2008]" pageId="16" pageNumber="17">
<emphasis id="1FBAAF2B5840FF98FF3AA6CF53CA8099" bold="true" box="[201,397,1172,1197]" pageId="16" pageNumber="17">Allotype female.</emphasis>
Length 11.0 mm, width
<quantity id="EA36DEDC5840FF98FD5FA6CF51408098" box="[684,775,1172,1196]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="16" pageNumber="17" unit="mm" value="4.2">4.2 mm</quantity>
. Color shiny black on most of body surface; reddish dark brown on all legs. Antennal club as long as the length of funicle. Epipleural fold swollen and enlarged. Protibiae and protarsi narrower than in male. Genital plates elongate, subtriangular, with angled apex and scattered setae (
<figureCitation id="B5F56FBC5840FF98FF6CA6A552B48122" box="[159,243,1278,1302]" captionStart="FIGURES 26 – 35" captionStartId="15.[151,264,1938,1961]" captionTargetBox="[238,1394,484,1899]" captionTargetId="figure@15.[151,1436,445,1918]" captionTargetPageId="15" captionText="FIGURES 26 – 35. Diagnostic structures in P. matzapatlecus: 26) right proleg, 27) right antenna, 28) male genitalia in dorsal view, 29) male genitalia in lateral view, 30) spiculum gastrale, 31) female genital plates, 32) labium, 33) labrum, 34) right maxilla, 35) left mandible. Scale lines 1 mm." httpUri="https://zenodo.org/record/210548/files/figure.png" pageId="16" pageNumber="17">Fig. 31</figureCitation>
).
</paragraph>
<paragraph id="2D7173395840FF98FF3AA77953B48197" blockId="16.[151,1437,151,2008]" pageId="16" pageNumber="17">
<emphasis id="1FBAAF2B5840FF98FF3AA7795306810E" bold="true" box="[201,321,1314,1338]" pageId="16" pageNumber="17">
<typeStatus id="F275CD9B5840FF98FF3AA7795306810E" box="[201,321,1314,1338]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<emphasis id="1FBAAF2B5840FF98FEA0A7795326810D" box="[339,353,1314,1337]" italics="true" pageId="16" pageNumber="17">n</emphasis>
= 27). Length: 9.0–
<quantity id="EA36DEDC5840FF98FDBAA77A50E8810D" box="[585,687,1313,1337]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="16" pageNumber="17" unit="mm" value="10.7">10.7 mm</quantity>
, width:
<quantity id="EA36DEDC5840FF98FCFEA77A51DE810D" box="[781,921,1313,1337]" metricMagnitude="-3" metricUnit="m" metricValue="3.9499999999999997" metricValueMax="4.1" metricValueMin="3.8" pageId="16" pageNumber="17" unit="mm" value="3.9499999999999997" valueMax="4.1" valueMin="3.8">3.8–4.1 mm</quantity>
. Fifteen specimens (
<specimenCount id="3BC8B8B05840FF98FB77A77A56AA810D" box="[1156,1261,1313,1337]" pageId="16" pageNumber="17" type="male">13 males</specimenCount>
,
<specimenCount id="3BC8B8B05840FF98FB09A77A572D810D" box="[1274,1386,1313,1337]" pageId="16" pageNumber="17" type="female">2 females</specimenCount>
) are completely shiny black, with reddish brown legs and
<specimenCount id="3BC8B8B05840FF98FD09A71F51D48168" box="[762,915,1348,1372]" pageId="16" pageNumber="17" type="generic">12 specimens</specimenCount>
(
<specimenCount id="3BC8B8B05840FF98FC56A71F56478168" box="[933,1024,1348,1372]" pageId="16" pageNumber="17" type="male">9 males</specimenCount>
,
<specimenCount id="3BC8B8B05840FF98FBFEA71F563B8168" box="[1037,1148,1348,1372]" pageId="16" pageNumber="17" type="female">3 females</specimenCount>
) have the pronotum with reddish yellow lateral margins and black disc. Mesometasternal process in bicolored specimens appears slightly larger than in black specimens.
</paragraph>
</subSubSection>
<subSubSection id="65D420B25840FF98FF3AA7F557258267" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="2D7173395840FF98FF3AA7F557258267" blockId="16.[151,1437,151,2008]" pageId="16" pageNumber="17">
<emphasis id="1FBAAF2B5840FF98FF3AA7F5530481F3" bold="true" box="[201,323,1454,1479]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
Clypeus subrectangular. Pronotum with scattered small, circular punctures (approximate diameter
<quantity id="EA36DEDC5840FF98FF64A78A534881DD" box="[151,271,1489,1513]" metricMagnitude="-5" metricUnit="m" metricValue="2.7" pageId="16" pageNumber="17" unit="mm" value="0.027">0.027 mm</quantity>
). Elytral surface nearly smooth, with striae weakly indicated by discontinuos rows of shallow, minute punctures; margins without bend. Protarsomeres 1–4 shortened, but slightly wider than protarsomere 5. Protibia stout, progressively widened distally; external border with proximal denticle short and rounded; distal denticle long, straight. Parameres short and complex with upper part narrowed, widely curved towards apex (Table 1).
</paragraph>
</subSubSection>
<subSubSection id="65D420B25840FF98FF3AA40551BD82AD" pageId="16" pageNumber="17" type="etymology">
<paragraph id="2D7173395840FF98FF3AA40551BD82AD" blockId="16.[151,1437,151,2008]" pageId="16" pageNumber="17">
<emphasis id="1FBAAF2B5840FF98FF3AA40553088242" bold="true" box="[201,335,1630,1654]" pageId="16" pageNumber="17">Etymology.</emphasis>
From the ancient indigenous Nahuatl language “
<emphasis id="1FBAAF2B5840FF98FC86A40556458242" box="[885,1026,1630,1654]" italics="true" pageId="16" pageNumber="17">Matzapatlan</emphasis>
” a mythical city in the mountains of northern Oaxaca where the first mazateco or “
<emphasis id="1FBAAF2B5840FF98FD6FA4D9511282AD" box="[668,853,1665,1689]" italics="true" pageId="16" pageNumber="17">Ha Shuta Enima</emphasis>
” people lived.
</paragraph>
</subSubSection>
<subSubSection id="65D420B25840FF98FF3AA4FF5361837D" pageId="16" pageNumber="17" type="distribution">
<paragraph id="2D7173395840FF98FF3AA4FF5361837D" blockId="16.[151,1437,151,2008]" pageId="16" pageNumber="17">
<emphasis id="1FBAAF2B5840FF98FF3AA4FF50A68289" bold="true" box="[201,737,1700,1725]" pageId="16" pageNumber="17">Distribution, phenology, and natural history</emphasis>
. This species is distributed in the humid slopes of Sierra de Juarez, northern State of Oaxaca in
<collectingCountry id="55D933A95840FF98FDD9A493503882D4" box="[554,639,1736,1760]" name="Mexico" pageId="16" pageNumber="17">Mexico</collectingCountry>
(
<figureCitation id="B5F56FBC5840FF98FD7CA49350A682D4" box="[655,737,1736,1760]" captionStart="FIGURE 54" captionStartId="21.[151,250,1234,1257]" captionTargetBox="[157,1436,382,1193]" captionTargetId="figure@21.[157,1442,373,1193]" captionTargetPageId="21" captionText="FIGURE 54. Distribution of species of Pachystethus." httpUri="https://zenodo.org/record/210552/files/figure.png" pageId="16" pageNumber="17">Fig. 54</figureCitation>
) between
<quantity id="EA36DEDC5840FF98FCA6A49351AE82D4" box="[853,1001,1736,1760]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.55" metricValueMin="0.65" pageId="16" pageNumber="17" unit="m" value="1100.0" valueMax="1550.0" valueMin="650.0">650–1,550 m</quantity>
of elevation where cloud forests overlaps with tropical rain forest and coffee plantations. Adults are active during March, August, and September. Flowers of an unidentified species of
<taxonomicName id="EACE08BA5840FF98FDD2A55450DB8313" box="[545,668,1807,1831]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
are recorded as host for adults, and the immature stages remain undescribed.
</paragraph>
</subSubSection>
<subSubSection id="65D420B25840FF98FF3AA50E534483EC" pageId="16" pageNumber="17" type="discussion">
<paragraph id="2D7173395840FF98FF3AA50E534483EC" blockId="16.[151,1437,151,2008]" pageId="16" pageNumber="17">
<emphasis id="1FBAAF2B5840FF98FF3AA50E537A8359" bold="true" box="[201,317,1877,1901]" pageId="16" pageNumber="17">Remarks.</emphasis>
<taxonomicName id="EACE08BA5840FF98FEB7A50E50C78359" box="[324,640,1877,1901]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="matzapatlecus">
<emphasis id="1FBAAF2B5840FF98FEB7A50E50C78359" box="[324,640,1877,1901]" italics="true" pageId="16" pageNumber="17">Pachystethus matzapatlecus</emphasis>
</taxonomicName>
has resemblance with
<taxonomicName id="EACE08BA5840FF98FC73A50D567E8359" box="[896,1081,1878,1901]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ixtacomitanus">
<emphasis id="1FBAAF2B5840FF98FC73A50D567E8359" box="[896,1081,1878,1901]" italics="true" pageId="16" pageNumber="17">P. ixtacomitanus</emphasis>
</taxonomicName>
and
<emphasis id="1FBAAF2B5840FF98FB83A50D574B8359" box="[1136,1292,1878,1901]" italics="true" pageId="16" pageNumber="17">P. n e c t o c t e n u s</emphasis>
, but may be distinguished from the former by the elytra with the posterior callus not prominent and a shallower punctation, and from the latter by the protibia thinner and protarsus thinner and elongate. The shape of parameres is also distinctive (Table 1).
</paragraph>
</subSubSection>
</treatment>
</document>
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