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<document id="9C38945A74425A46983CAE7F63692DC6" ID-DOI="10.5281/zenodo.210542" ID-GBIF-Dataset="c9e5cc54-2a89-4b75-91aa-a4fce79f17fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="210542" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460500319311" checkinUser="plazi" docAuthor="Ramírez-Ponce, Andrés &amp; Morón, Miguel Angel" docDate="2012" docId="A567C22F5856FF80FF64A7FC52A883E8" docLanguage="en" docName="zt03394p024.pdf" docOrigin="Zootaxa 3394" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pachystethus crassesculptus Bates 1888, new combination" docType="treatment" docVersion="9" lastPageNumber="9" masterDocId="595EBA575850FF88FFF3A25B52478434" masterDocTitle="Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)" masterLastPageNumber="24" masterPageNumber="1" pageNumber="7" updateTime="1698311598655" updateUser="plazi">
<mods:mods id="F97B65B153D153CF80654860C38C0790" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="359483C25A29E1CC2187C3FD334E1C87">
<mods:title id="4AF37A0700B2CB87A58D0AE6AF16022B">Revision of the genus Pachystethus Blanchard with description of three new species from Mexico (Coleoptera: Scarabaeidae: Rutelinae: Anomalini)</mods:title>
</mods:titleInfo>
<mods:name id="9791DA07E3BDF9D4A47C6FCB53BC7337" type="personal">
<mods:role id="088A7E474F432B8160364CCC00B52227">
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</mods:role>
<mods:namePart id="CAF2A0213B4ECA4498745953150280A7">Ramírez-Ponce, Andrés</mods:namePart>
</mods:name>
<mods:name id="4676B972CB778CAD1CFF08512A479C2A" type="personal">
<mods:role id="CC38A7ECE94AEFD6BEE96E5AD6345AF8">
<mods:roleTerm id="34525A1BE343C7FB07233BE354F5D8CA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="02FD278E1D963D942EDDA55DE600C8A8">Morón, Miguel Angel</mods:namePart>
</mods:name>
<mods:typeOfResource id="426C7CBD11CBBCB62B8B97BCF254DA7B">text</mods:typeOfResource>
<mods:relatedItem id="C206792A422DF56C85EC647D49366340" type="host">
<mods:titleInfo id="1BB8FD5EC41D21DD8C6A69ECDF22A8D4">
<mods:title id="991E06917FCEC1767E41608F6A0E90A4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B3B25E07E6FCFA5FC22762ECCD3A56C1">
<mods:date id="744D927FAF99DCF91870792586226112">2012</mods:date>
<mods:detail id="679DEA3D1292D72FFDE66E0B335138B8" type="volume">
<mods:number id="36647C8C2A19BFCF21D9F7A8DD2931CA">3394</mods:number>
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<mods:start id="4BDA38F6B8CFAD2C64A7FA6E355D8557">1</mods:start>
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<mods:classification id="F2FE75172F21C6CF0E5E14D28A95C27F">journal article</mods:classification>
<mods:identifier id="D7E8F1E862F6E7AB85B9DBE5CD6344BF" type="DOI">10.5281/zenodo.210542</mods:identifier>
<mods:identifier id="3683A1B66FCC858E19FD6D58FD19CAD1" type="GBIF-Dataset">c9e5cc54-2a89-4b75-91aa-a4fce79f17fc</mods:identifier>
<mods:identifier id="821BC9804A631DF6E2B2F2295D6A473F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D754DC0CE3C0B84FCABBE2CE552499E0" type="Zenodo-Dep">210542</mods:identifier>
</mods:mods>
<treatment id="A567C22F5856FF80FF64A7FC52A883E8" ID-DOI="http://doi.org/10.5281/zenodo.6179196" ID-GBIF-Taxon="119521053" ID-Zenodo-Dep="6179196" LSID="urn:lsid:plazi:treatment:A567C22F5856FF80FF64A7FC52A883E8" httpUri="http://treatment.plazi.org/id/A567C22F5856FF80FF64A7FC52A883E8" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="65D420B25856FF8EFF64A7FC53D281D6" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="2D7173395856FF8EFF64A7FC513681F5" blockId="6.[151,881,1447,1506]" box="[151,881,1447,1473]" pageId="6" pageNumber="7">
<heading id="7639C4555856FF8EFF64A7FC513681F5" bold="true" box="[151,881,1447,1473]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="1FBAAF2B5856FF8EFF64A7FC513681F5" bold="true" box="[151,881,1447,1473]" pageId="6" pageNumber="7">
<taxonomicName id="EACE08BA5856FF8EFF64A7FC50D381F5" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" box="[151,660,1447,1473]" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="crassesculptus" status="comb. nov.">
<emphasis id="1FBAAF2B5856FF8EFF64A7FC53AB81F5" bold="true" box="[151,492,1447,1473]" italics="true" pageId="6" pageNumber="7">Pachystethus crassesculptus</emphasis>
(
<bibRefCitation id="495F0EC85856FF8EFE08A7FC50CB81F5" author="Bates" box="[507,652,1447,1473]" pageId="6" pageNumber="7" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia. In: O. Salvin &amp; F. D. Godwin (eds.), Biologia Centrali-Americana. Insecta Coleoptera. Volume 2, part 2: 1 - 432." type="book chapter" year="1888">Bates, 1888</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="048912505856FF8EFD68A7FC513681F5" box="[667,881,1447,1473]" pageId="6" pageNumber="7">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="2D7173395856FF8EFF64A79153D281D6" blockId="6.[151,881,1447,1506]" box="[151,405,1482,1506]" pageId="6" pageNumber="7">
(
<figureCitation id="B5F56FBC5856FF8EFF6CA791535E81D6" box="[159,281,1482,1506]" captionStart="FIGURES 8 – 13" captionStartId="7.[151,264,1669,1692]" captionTargetBox="[185,1401,251,1668]" captionTargetId="figure@7.[151,1436,225,1677]" captionTargetPageId="7" captionText="FIGURES 8 – 13. Diagnostic structures in P. crassesculptus. 8) right proleg, 9) right antenna, 10) male genitalia in dorsal view, 11) male genitalia in lateral view, 12) spiculum gastrale, 13) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210545/files/figure.png" pageId="6" pageNumber="7">Figs. 8–13</figureCitation>
,
<figureCitation id="B5F56FBC5856FF8EFED7A791530481D6" box="[292,323,1482,1506]" captionStart="FIGURES 42 – 47" captionStartId="19.[151,264,1909,1932]" captionTargetBox="[214,1377,207,1832]" captionTargetId="figure@19.[198,1392,193,1847]" captionTargetPageId="19" captionText="FIGURES 42 – 47. Dorsal view of Pachystethus species: 42 a, b) P. viduus males; 42 c) P. viduus female; 42 d) P. viduus female, lectotype; 43 a) P. nectoctenus male; 43 b) P. nectoctenus female; 44 a) P. crassesculptus male; 44 b) P. crassesculptus female; 45 a, b) P. nutans males; 45 c) P. nu t an s female; 46 a, b) P. matzapatlecus males; 46 c) P. matzapatlecus female; 47 a) P. ixtacomitanus male; 47 b) P. ixtacomitanus female. Scale lines 1 mm." httpUri="https://zenodo.org/record/210550/files/figure.png" pageId="6" pageNumber="7">44</figureCitation>
a,b, 50)
</paragraph>
</subSubSection>
<subSubSection id="65D420B25856FF8EFF64A44950D9821C" box="[151,670,1554,1576]" pageId="6" pageNumber="7" type="reference_group">
<paragraph id="2D7173395856FF8EFF64A44950D9821C" blockId="6.[151,670,1554,1576]" box="[151,670,1554,1576]" pageId="6" pageNumber="7">
<treatmentCitationGroup id="0DDE54175856FF8EFF64A44950D9821C" box="[151,670,1554,1576]" pageId="6" pageNumber="7">
<taxonomicName id="EACE08BA5856FF8EFF64A4495376821C" box="[151,305,1554,1576]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="vidua">
<emphasis id="1FBAAF2B5856FF8EFF64A4495376821C" box="[151,305,1554,1576]" italics="true" pageId="6" pageNumber="7">Anomala vidua</emphasis>
</taxonomicName>
<taxonomicName id="EACE08BA5856FF8EFEC4A44850D9821C" authority="Bates, 1888: 406" authorityName="Bates" authorityPageNumber="406" authorityYear="1888" box="[311,670,1554,1576]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="variety" species="vidua" variety="crassesculpta">
var.
<emphasis id="1FBAAF2B5856FF8EFE91A44953AB821C" box="[354,492,1554,1576]" italics="true" pageId="6" pageNumber="7">crassesculpta</emphasis>
<treatmentCitation id="AC6F55285856FF8EFE00A44850D9821C" author="Bates" box="[499,670,1554,1576]" page="406" pageId="6" pageNumber="7" year="1888">
<bibRefCitation id="495F0EC85856FF8EFE00A4485029821C" author="Bates" box="[499,622,1555,1576]" pageId="6" pageNumber="7" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia. In: O. Salvin &amp; F. D. Godwin (eds.), Biologia Centrali-Americana. Insecta Coleoptera. Volume 2, part 2: 1 - 432." type="book chapter" year="1888">Bates, 1888</bibRefCitation>
: 406
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="65D420B25856FF8FFF64A40E572584E0" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="2D7173395856FF8EFF64A40E535A82A5" blockId="6.[151,1437,1621,2005]" pageId="6" pageNumber="7">
<emphasis id="1FBAAF2B5856FF8EFF64A40E5327825A" bold="true" box="[151,352,1621,1646]" pageId="6" pageNumber="7">
<typeStatus id="F275CD9B5856FF8EFF64A40E52958259" box="[151,210,1621,1645]" pageId="6" pageNumber="7">Type</typeStatus>
specimens.
</emphasis>
Male and female
<typeStatus id="F275CD9B5856FF8EFDB4A40D50EE825A" box="[583,681,1622,1646]" pageId="6" pageNumber="7" type="syntype">syntypes</typeStatus>
examined, deposited in
<collectionCode id="4BDFEBFC5856FF8EFC24A40D565B8259" box="[983,1052,1622,1645]" pageId="6" pageNumber="7">NHM</collectionCode>
.
<typeStatus id="F275CD9B5856FF8EFBDDA40D5621825A" box="[1070,1126,1622,1646]" pageId="6" pageNumber="7">Type</typeStatus>
locality: “Chimaltenango,
<collectingCountry id="55D933A95856FF8EFF64A422535482A5" box="[151,275,1657,1681]" name="Guatemala" pageId="6" pageNumber="7">Guatemala</collectingCountry>
”
</paragraph>
<paragraph id="2D7173395856FF8FFF3AA4C6572584E0" blockId="6.[151,1437,1621,2005]" lastBlockId="7.[151,1436,151,212]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="1FBAAF2B5856FF8EFF3AA4C6539A8282" bold="true" box="[201,477,1693,1718]" pageId="6" pageNumber="7">
<typeStatus id="F275CD9B5856FF8EFF3AA4C653438281" box="[201,260,1693,1717]" pageId="6" pageNumber="7">Type</typeStatus>
material revised
</emphasis>
(
<specimenCount id="3BC8B8B05856FF8EFE04A4C6500E8281" box="[503,585,1693,1717]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
lectoype,
<specimenCount id="3BC8B8B05856FF8EFD3CA4C6517F8281" box="[719,824,1693,1717]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
<typeStatus id="F275CD9B5856FF8EFCBBA4C651A48281" box="[840,995,1693,1717]" pageId="6" pageNumber="7" type="paralectotype">paralectotype</typeStatus>
;
<collectionCode id="4BDFEBFC5856FF8EFC05A4C6567E8281" box="[1014,1081,1693,1717]" pageId="6" pageNumber="7">NHM</collectionCode>
) labeled:
<emphasis id="1FBAAF2B5856FF8EFB4AA4C657318281" bold="true" box="[1209,1398,1693,1717]" pageId="6" pageNumber="7">
<typeStatus id="F275CD9B5856FF8EFB4AA4C657628281" box="[1209,1317,1693,1717]" pageId="6" pageNumber="7" type="lectotype">lectotype</typeStatus>
male:
</emphasis>
a) “
<collectingCountry id="55D933A95856FF8EFF52A49A531182ED" box="[161,342,1729,1753]" name="Guatemala" pageId="6" pageNumber="7">GUATEMALA</collectingCountry>
: Chimaltenango, Conradt, B.C.A.Col.II (2)” (typeface), b) “
<taxonomicName id="EACE08BA5856FF8EFBF7A49A529F82C9" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" class="Insecta" family="Rutelidae" genus="Pachystethus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="crassesculptus">
<emphasis id="1FBAAF2B5856FF8EFBF7A49A570382ED" box="[1028,1348,1729,1753]" italics="true" pageId="6" pageNumber="7">Pachystethus crassesculptus</emphasis>
(
<bibRefCitation id="495F0EC85856FF8EFAA6A49A529782C9" author="Bates" pageId="6" pageNumber="7" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia. In: O. Salvin &amp; F. D. Godwin (eds.), Biologia Centrali-Americana. Insecta Coleoptera. Volume 2, part 2: 1 - 432." type="book chapter" year="1888">Bates, 1888</bibRefCitation>
)
</taxonomicName>
<typeStatus id="F275CD9B5856FF8EFF10A4BE53C482C9" box="[227,387,1765,1789]" pageId="6" pageNumber="7" type="lectotype">LECTOTYPE</typeStatus>
male Ramírez-Ponce y Morón des. 2011” (typeface).
<emphasis id="1FBAAF2B5856FF8EFBF1A4BE570582CA" bold="true" box="[1026,1346,1765,1790]" pageId="6" pageNumber="7">
<typeStatus id="F275CD9B5856FF8EFBF1A4BE563F82C9" box="[1026,1144,1765,1789]" pageId="6" pageNumber="7" type="lectotype">Lectotype</typeStatus>
here designated
</emphasis>
for the taxonomic purpose to preserve nomenclatural stability by selecting a single specimen as the name-bearing
<typeStatus id="F275CD9B5856FF8EFABEA551573A8316" box="[1357,1405,1802,1826]" pageId="6" pageNumber="7">type</typeStatus>
of the name of this taxon.
<emphasis id="1FBAAF2B5856FF8EFE5DA57650F18372" bold="true" box="[430,694,1837,1862]" pageId="6" pageNumber="7">
<typeStatus id="F275CD9B5856FF8EFE5DA57650148371" box="[430,595,1837,1861]" pageId="6" pageNumber="7" type="paralectotype">Paralectotype</typeStatus>
female:
</emphasis>
a) “
<collectingCountry id="55D933A95856FF8EFD1AA57651D98371" box="[745,926,1837,1861]" name="Guatemala" pageId="6" pageNumber="7">GUATEMALA</collectingCountry>
: Chimaltenango, Conradt, B.C.A.Col.II (2)” (typeface), b) “
<taxonomicName id="EACE08BA5856FF8EFEBBA50A516A835D" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" box="[328,813,1873,1897]" class="Insecta" family="Scarabaeidae" genus="Pachyststhus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="crassesculptus">
<emphasis id="1FBAAF2B5856FF8EFEBBA50A50CE835D" box="[328,649,1873,1897]" italics="true" pageId="6" pageNumber="7">Pachyststhus crassesculptus</emphasis>
(
<bibRefCitation id="495F0EC85856FF8EFD68A50A5162835D" author="Bates" box="[667,805,1873,1897]" pageId="6" pageNumber="7" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia. In: O. Salvin &amp; F. D. Godwin (eds.), Biologia Centrali-Americana. Insecta Coleoptera. Volume 2, part 2: 1 - 432." type="book chapter" year="1888">Bates, 1888</bibRefCitation>
)
</taxonomicName>
<typeStatus id="F275CD9B5856FF8EFCCBA50A5667835D" box="[824,1056,1873,1897]" pageId="6" pageNumber="7" type="paralectotype">PARALECTOTYPE</typeStatus>
female Ramírez-Ponce y Morón des. 2011” (typeface).
<emphasis id="1FBAAF2B5856FF8EFE5BA52E50BA83BA" bold="true" box="[424,765,1909,1934]" pageId="6" pageNumber="7">Additional material revised</emphasis>
(
<specimenCount id="3BC8B8B05856FF8EFCE0A52E51C683B9" box="[787,897,1909,1933]" pageId="6" pageNumber="7" type="male">21 males</specimenCount>
,
<specimenCount id="3BC8B8B05856FF8EFC60A52E564D83B9" box="[915,1034,1909,1933]" pageId="6" pageNumber="7" type="female">9 females</specimenCount>
;
<collectionCode id="4BDFEBFC5856FF8EFBE8A52E562583B9" box="[1051,1122,1909,1933]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="6" pageNumber="7">CNIN</collectionCode>
,
<collectionCode id="4BDFEBFC5856FF8EFB86A52D568D83BA" box="[1141,1226,1910,1934]" pageId="6" pageNumber="7">MXAL</collectionCode>
,
<collectionCode id="4BDFEBFC5856FF8EFB2FA52E576883B9" box="[1244,1327,1909,1933]" pageId="6" pageNumber="7">ZMHB</collectionCode>
) labeled:
<collectingCountry id="55D933A95856FF8EFF64A5C253578385" box="[151,272,1945,1969]" name="Guatemala" pageId="6" pageNumber="7">Guatemala</collectingCountry>
/ L. Conradt S. (
<specimenCount id="3BC8B8B05856FF8EFE22A5C250708385" box="[465,567,1945,1969]" pageId="6" pageNumber="7" type="male">18 males</specimenCount>
and
<specimenCount id="3BC8B8B05856FF8EFD83A5C250A78385" box="[624,736,1945,1969]" pageId="6" pageNumber="7" type="female">4 females</specimenCount>
,
<collectionCode id="4BDFEBFC5856FF8EFD1FA5C251788385" box="[748,831,1945,1969]" pageId="6" pageNumber="7">ZMHB</collectionCode>
) (typeface);
<collectingCountry id="55D933A95856FF8EFC23A5C2560E8385" box="[976,1097,1945,1969]" name="Guatemala" pageId="6" pageNumber="7">Guatemala</collectingCountry>
/ Tumbador Riedel S. 4.12 (
<specimenCount id="3BC8B8B05856FF8EFA63A5C2528A83E1" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
and
<specimenCount id="3BC8B8B05856FF8EFEF9A5E6533A83E1" box="[266,381,1981,2005]" pageId="6" pageNumber="7" type="female">3 females</specimenCount>
,
<collectionCode id="4BDFEBFC5856FF8EFE78A5E6539983E1" box="[395,478,1981,2005]" pageId="6" pageNumber="7">ZMHB</collectionCode>
) (typeface);
<collectingCountry id="55D933A95856FF8EFD87A5E650AA83E1" box="[628,749,1981,2005]" name="Guatemala" pageId="6" pageNumber="7">Guatemala</collectingCountry>
(
<specimenCount id="3BC8B8B05856FF8EFCF1A5E6512F83E1" box="[770,872,1981,2005]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
,
<collectionCode id="4BDFEBFC5856FF8EFC84A5E6518A83E1" box="[887,973,1981,2005]" pageId="6" pageNumber="7">ZMHB</collectionCode>
);
<collectingCountry id="55D933A95856FF8EFC10A5E6561383E1" box="[995,1108,1981,2005]" name="Mexico" pageId="6" pageNumber="7">MEXICO</collectingCountry>
,
<emphasis id="1FBAAF2B5856FF8EFB90A5E6568783E1" box="[1123,1216,1981,2005]" italics="true" pageId="6" pageNumber="7">Chiapas</emphasis>
, Tapachula, Unión Roja, La Boquilla,
<date id="597055F95857FF8FFE98A2CC5391849B" box="[363,470,151,175]" pageId="7" pageNumber="8" value="1982-09-22">
22-IX-
<quantity id="EA36DEDC5857FF8FFE4AA2CC5391849B" box="[441,470,151,175]" metricMagnitude="4" metricUnit="m" metricValue="8.254999999999999" pageId="7" pageNumber="8" unit="m" value="82550.0">82</quantity>
</date>
, 550 m, planta silvestre, R. Terrón (
<specimenCount id="3BC8B8B05857FF8FFC97A2CC51FA849B" box="[868,957,151,175]" pageId="7" pageNumber="8" type="male">2 males</specimenCount>
,
<collectionCode id="4BDFEBFC5857FF8FFC3AA2CC565A849B" box="[969,1053,151,175]" pageId="7" pageNumber="8">MXAL</collectionCode>
) (typeface);
<collectingCountry id="55D933A95857FF8FFB59A2CC575C849B" box="[1194,1307,151,175]" name="Mexico" pageId="7" pageNumber="8">MEXICO</collectingCountry>
, Chiapas,
<quantity id="EA36DEDC5857FF8FFA7DA2CC52FC84E0" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="7" pageNumber="8" unit="km" value="9.0">9 km</quantity>
al sur de Union Juárez, Puente Shujukal.
<date id="597055F95857FF8FFD7CA2E750BF84E0" box="[655,760,188,212]" pageId="7" pageNumber="8" value="1983-04-21">21-IV-83</date>
. E. Barrera (handwritten) (
<specimenCount id="3BC8B8B05857FF8FFBD5A2E756CE84E0" box="[1062,1161,188,212]" pageId="7" pageNumber="8" type="female">1 female</specimenCount>
,
<collectionCode id="4BDFEBFC5857FF8FFB67A2E7569D84E0" box="[1172,1242,188,212]" httpUri="http://grbio.org/cool/qbc0-xx4g" name="Coleccion Nacional de Insectos, Universidad Nacional Autonoma de Mexico" pageId="7" pageNumber="8">CNIN</collectionCode>
) (typeface).
</paragraph>
</subSubSection>
<caption id="79B123B15857FF8FFF64A4DE56D7828E" httpUri="https://zenodo.org/record/210545/files/figure.png" pageId="7" pageNumber="8" targetBox="[185,1401,251,1668]" targetPageId="7">
<paragraph id="2D7173395857FF8FFF64A4DE56D7828E" blockId="7.[151,1436,1669,1723]" pageId="7" pageNumber="8">
<emphasis id="1FBAAF2B5857FF8FFF64A4DE530682AF" bold="true" box="[151,321,1669,1692]" pageId="7" pageNumber="8">FIGURES 8–13</emphasis>
. Diagnostic structures in
<taxonomicName id="EACE08BA5857FF8FFDB7A4DC50B682A8" box="[580,753,1670,1692]" class="Insecta" family="Scarabaeidae" genus="Pachyststhus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="crassesculptus">
<emphasis id="1FBAAF2B5857FF8FFDB7A4DC50B682A8" box="[580,753,1670,1692]" italics="true" pageId="7" pageNumber="8">P. crassesculptus</emphasis>
</taxonomicName>
.
<emphasis id="1FBAAF2B5857FF8FFD0EA4DE515482A8" bold="true" box="[765,787,1669,1692]" pageId="7" pageNumber="8">8)</emphasis>
right proleg,
<emphasis id="1FBAAF2B5857FF8FFC6EA4DE51F582A8" bold="true" box="[925,946,1669,1692]" pageId="7" pageNumber="8">9)</emphasis>
right antenna,
<emphasis id="1FBAAF2B5857FF8FFBB9A4DE562C82A8" bold="true" box="[1098,1131,1669,1692]" pageId="7" pageNumber="8">10)</emphasis>
male genitalia in dorsal view,
<emphasis id="1FBAAF2B5857FF8FFF64A4FE52F1828F" bold="true" box="[151,182,1700,1723]" pageId="7" pageNumber="8">11)</emphasis>
male genitalia in lateral view,
<emphasis id="1FBAAF2B5857FF8FFE1DA4FF5057828F" bold="true" box="[494,528,1700,1723]" pageId="7" pageNumber="8">12)</emphasis>
spiculum gastrale,
<emphasis id="1FBAAF2B5857FF8FFD26A4FF50B0828F" bold="true" box="[725,759,1700,1723]" pageId="7" pageNumber="8">13)</emphasis>
female genital plates. Scale lines 1 mm.
</paragraph>
</caption>
<subSubSection id="65D420B25857FF80FF3AA4BB53558160" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="2D7173395857FF80FF3AA4BB517C8094" blockId="7.[151,1437,1760,2037]" lastBlockId="8.[151,1436,151,2012]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="1FBAAF2B5857FF8FFF3AA4BB53FB82CD" bold="true" box="[201,444,1760,1785]" pageId="7" pageNumber="8">Redescription. Male</emphasis>
(
<emphasis id="1FBAAF2B5857FF8FFE3DA4B9539B82CD" box="[462,476,1762,1785]" italics="true" pageId="7" pageNumber="8">n</emphasis>
= 21). Length 10.7–11.0 mm, width
<quantity id="EA36DEDC5857FF8FFC7BA4BB565382CC" box="[904,1044,1760,1784]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.1" metricValueMin="3.9" pageId="7" pageNumber="8" unit="mm" value="4.0" valueMax="4.1" valueMin="3.9">3.9–4.1 mm</quantity>
. Color reddish yellow on clypeus, anterior half of frons, scape, pedicel, funicle, lateral margins of pronotum, abdomen, pygidium, femora, tibiae and tarsi; shiny black on antennal club, posterior half of frons, disc of pronotum,
<taxonomicName id="EACE08BA5857FF8FFC0CA57356288374" box="[1023,1135,1832,1856]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="7" pageNumber="18" phylum="Arthropoda" rank="genus">scutellum</taxonomicName>
, and elytra.
<emphasis id="1FBAAF2B5857FF8FFB09A57257708375" box="[1274,1335,1833,1857]" italics="true" pageId="7" pageNumber="8">Head</emphasis>
: clypeus subtrapezoidal, clearly wider than long; anterior border straight, slightly upturned and directed forward; anterior angles rounded; clypeal surface with irregular, dense, large punctures. Frontoclypeal suture well defined, straight. Frons nearly flattened, densely and irregularly rugopunctate at center, with scattered large punctures along lateral and posterior margins. Punctures around eyes are more disperse and smaller than in frons, and are combined with scarce, long setae. Antennal club nearly as long as all preceding antennomeres combined; lamellar shape ovate, straight (
<figureCitation id="B5F56FBC5858FF80FEF2A2CC530C849B" box="[257,331,151,175]" captionStart="FIGURES 8 – 13" captionStartId="7.[151,264,1669,1692]" captionTargetBox="[185,1401,251,1668]" captionTargetId="figure@7.[151,1436,225,1677]" captionTargetPageId="7" captionText="FIGURES 8 – 13. Diagnostic structures in P. crassesculptus. 8) right proleg, 9) right antenna, 10) male genitalia in dorsal view, 11) male genitalia in lateral view, 12) spiculum gastrale, 13) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210545/files/figure.png" pageId="8" pageNumber="9">Fig. 9</figureCitation>
). Eyes small, ovate, ocular margin noticeable and exposed dorsally. Eye canthus thick with glabrous, rounded apex and flattened surface; surface with irregular punctures. Maxillary basistipe flat with irregular, wide punctation; labrum wide and sinuate; labium sinuate, slightly longer than wide, with irregular rugopunctate; prementum setose.
<emphasis id="1FBAAF2B5858FF80FDE6A35E50C38528" box="[533,644,261,284]" italics="true" pageId="8" pageNumber="9">Pronotum</emphasis>
wider than long, slightly convex; surface with a moderate number of shallow, small sized, circular punctures; lateral borders nearly straight, weakly curved, with bulky sides and lateral pits; posterior angles raised. Mesoepimera scarcely exposed over humeral calli. Space between mesocoxae wide, meso-metasternal process long, projecting anteriorly past mesocoxa but not reaching procoxae; with wide, rounded apex directed downward; metasternum densely punctate, sides uniformly setiferous, mesial region glabrous and longitudinally sulcate.
<emphasis id="1FBAAF2B5858FF80FE6FA3E353A585E4" box="[412,482,440,464]" italics="true" pageId="8" pageNumber="9">Elytra</emphasis>
flattened, short, ovate; striae deeply impressed with coarse large punctures; elytral costae discrete, suture raised towards posterior half; brief preapical slope; lateral margins nearly straight (without wide bend at the end of epipleural fold) with sparse, short setae along apical third. Epipleural fold sharp in lateral view, gradually narrowed after metacoxa.
<taxonomicName id="EACE08BA5858FF80FD81A07F50A78608" box="[626,736,548,572]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="1FBAAF2B5858FF80FD81A07F50A78608" box="[626,736,548,572]" italics="true" pageId="8" pageNumber="9">Scutellum</emphasis>
</taxonomicName>
slightly wider than long; lateral margins rounded, apex acute, surface irregularly punctate.
<emphasis id="1FBAAF2B5858FF80FE2FA01350018654" box="[476,582,584,608]" italics="true" pageId="8" pageNumber="9">Abdomen</emphasis>
sternites scarcely setiferous, each with one discontinuous row of setiferous punctures near posterior border and wide, weakly punctate, glabrous central area.
<emphasis id="1FBAAF2B5858FF80FBCFA036563686B0" box="[1084,1137,621,644]" italics="true" pageId="8" pageNumber="9">Legs</emphasis>
: protarsomeres 1–4 short, slightly wider and longer than protarsomere 5. Protibia stout, slightly widened towards apex; lateral borders nearly parallel; external border with proximal denticle wide and acute; distal denticle long, oblique (
<figureCitation id="B5F56FBC5858FF80FB40A0EF56BF86F8" box="[1203,1272,692,716]" captionStart="FIGURES 8 – 13" captionStartId="7.[151,264,1669,1692]" captionTargetBox="[185,1401,251,1668]" captionTargetId="figure@7.[151,1436,225,1677]" captionTargetPageId="7" captionText="FIGURES 8 – 13. Diagnostic structures in P. crassesculptus. 8) right proleg, 9) right antenna, 10) male genitalia in dorsal view, 11) male genitalia in lateral view, 12) spiculum gastrale, 13) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210545/files/figure.png" pageId="8" pageNumber="9">Fig. 8</figureCitation>
). Mesofemora with a well-defined longitudinal keel, with long setae. Metafemora without longitudinal keel. Metatibia with scarce, setiferous, large punctures. Metatarsomeres 1–2 large, longer than tarsomeres 3–4.
<emphasis id="1FBAAF2B5858FF80FB7FA0A756B18720" box="[1164,1270,764,788]" italics="true" pageId="8" pageNumber="9">Genitalia</emphasis>
(
<figureCitation id="B5F56FBC5858FF80FAF6A0A757D78720" box="[1285,1424,764,788]" captionStart="FIGURES 8 – 13" captionStartId="7.[151,264,1669,1692]" captionTargetBox="[185,1401,251,1668]" captionTargetId="figure@7.[151,1436,225,1677]" captionTargetPageId="7" captionText="FIGURES 8 – 13. Diagnostic structures in P. crassesculptus. 8) right proleg, 9) right antenna, 10) male genitalia in dorsal view, 11) male genitalia in lateral view, 12) spiculum gastrale, 13) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210545/files/figure.png" pageId="8" pageNumber="9">Figs. 10–12</figureCitation>
): parameres shorter than the length of tectum, with upper part slightly concave and smooth, gradually narrowed and upturned towards apex; lower part of parameres slightly curved backwards. Basal half of tectum clearly wider than distal half, with outstanding posterolateral calli. Basal piece with long, acute apodema.
<emphasis id="1FBAAF2B5858FF80FB9BA133577487B4" box="[1128,1331,872,896]" italics="true" pageId="8" pageNumber="9">Spiculum gastrale</emphasis>
(
<figureCitation id="B5F56FBC5858FF80FAB1A13C57D3874B" box="[1346,1428,871,895]" captionStart="FIGURES 8 – 13" captionStartId="7.[151,264,1669,1692]" captionTargetBox="[185,1401,251,1668]" captionTargetId="figure@7.[151,1436,225,1677]" captionTargetPageId="7" captionText="FIGURES 8 – 13. Diagnostic structures in P. crassesculptus. 8) right proleg, 9) right antenna, 10) male genitalia in dorsal view, 11) male genitalia in lateral view, 12) spiculum gastrale, 13) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210545/files/figure.png" pageId="8" pageNumber="9">Fig. 12</figureCitation>
) narrow, basal part straight, slightly widened towards apex, slightly shorter than lateral sclerites; associated plates subtriangular with widely rounded apex, irregular upper inner margin and long setae on entire surface of exterior border.
<emphasis id="1FBAAF2B5858FF80FF1FA18F530487D8" bold="true" box="[236,323,980,1004]" pageId="8" pageNumber="9">Female</emphasis>
(
<emphasis id="1FBAAF2B5858FF80FEA0A18E532687D8" box="[339,353,981,1004]" italics="true" pageId="8" pageNumber="9">n</emphasis>
= 10). Length
<quantity id="EA36DEDC5858FF80FDF4A18F502A87D8" box="[519,621,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="8" pageNumber="9" unit="mm" value="11.2">11.2 mm</quantity>
, width
<quantity id="EA36DEDC5858FF80FD4DA18F515187D8" box="[702,790,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="8" pageNumber="9" unit="mm" value="4.2">4.2 mm</quantity>
. Color shiny reddish yellow or ochreous yellow on nearly entire body, except antennal club, two central-lateral spots on pronotal disc, and preapical elytral borders that are shiny black. Body stouter than in male; elytral striae marked with black, deep punctures; epipleural fold swollen; protibiae and protarsi not swollen; metatarsomeres wider than in male. Antennal club slightly longer than the length of the pedicel and funicle combined. Genital plates medium sized, elongate with round, convergent apex; regular inner borders with long setae on distal part (
<figureCitation id="B5F56FBC5858FF80FD29A6D351698094" box="[730,814,1160,1184]" captionStart="FIGURES 8 – 13" captionStartId="7.[151,264,1669,1692]" captionTargetBox="[185,1401,251,1668]" captionTargetId="figure@7.[151,1436,225,1677]" captionTargetPageId="7" captionText="FIGURES 8 – 13. Diagnostic structures in P. crassesculptus. 8) right proleg, 9) right antenna, 10) male genitalia in dorsal view, 11) male genitalia in lateral view, 12) spiculum gastrale, 13) female genital plates. Scale lines 1 mm." httpUri="https://zenodo.org/record/210545/files/figure.png" pageId="8" pageNumber="9">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="2D7173395858FF80FF3AA6F053558160" blockId="8.[151,1436,151,2012]" pageId="8" pageNumber="9">
<emphasis id="1FBAAF2B5858FF80FF3AA6F0537080F0" bold="true" box="[201,311,1195,1220]" pageId="8" pageNumber="9">Variation</emphasis>
(
<emphasis id="1FBAAF2B5858FF80FEBEA6F6531C80F0" box="[333,347,1197,1220]" italics="true" pageId="8" pageNumber="9">n</emphasis>
= 30). Extension of the black coloration in specimens with a contrasting bicolored pattern is variable mainly in the pronotum and head. Areas of the body with red color may be significantly more darkened (lateral borders on pronotum, frons, clypeus, abdomen, sternum, and legs). Although none were seen, some individuals are probably completely black in color. In one female, the distal part of the elytral interstriae was not prominent.
</paragraph>
</subSubSection>
<subSubSection id="65D420B25858FF80FF3AA70453A3823C" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="2D7173395858FF80FF3AA70453A3823C" blockId="8.[151,1436,151,2012]" pageId="8" pageNumber="9">
<emphasis id="1FBAAF2B5858FF80FF3AA7045304814C" bold="true" box="[201,323,1375,1400]" pageId="8" pageNumber="9">Diagnosis.</emphasis>
Clypeus subtrapezoidal. Pronotum with small size (approximate diameter
<quantity id="EA36DEDC5858FF80FB5CA73B576E814C" box="[1199,1321,1376,1400]" metricMagnitude="-5" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="9" unit="mm" value="0.025">0.025 mm</quantity>
), circular, shallow punctures. Elytral striae deeply indicated with punctures; posterior calli and distal part of interstriae prominent; lateral margins without bend. Protarsomeres 1–4 wider and shorter than distal protarsomere. Protibia slightly wider towards apex with lateral borders nearly parallel. Parameres large and complex, with dorsal surface smooth and convex (Table 1).
</paragraph>
</subSubSection>
<subSubSection id="65D420B25858FF80FF3AA44F53798240" pageId="8" pageNumber="9" type="etymology">
<paragraph id="2D7173395858FF80FF3AA44F53798240" blockId="8.[151,1436,151,2012]" pageId="8" pageNumber="9">
<emphasis id="1FBAAF2B5858FF80FF3AA44F53088218" bold="true" box="[201,335,1556,1580]" pageId="8" pageNumber="9">Etymology.</emphasis>
Although not cited in the original description, the name is derived from Latin terms
<emphasis id="1FBAAF2B5858FF80FABBA44E57DB8218" box="[1352,1436,1557,1580]" italics="true" pageId="8" pageNumber="9">crassus</emphasis>
(thick, heavy) and
<emphasis id="1FBAAF2B5858FF80FE94A463538B8264" box="[359,460,1592,1616]" italics="true" pageId="8" pageNumber="9">sculptum</emphasis>
(engraving, texture) (
<bibRefCitation id="495F0EC85858FF80FD4CA463510D8264" author="Jaeger" box="[703,842,1592,1616]" pageId="8" pageNumber="9" refString="Jaeger, E. C., (1978) A Source Book of Biological Names and Terms, 3 rd edition. Thomas Publisher, Springfield, Illinois." type="book" year="1978">Jaeger 1978</bibRefCitation>
), meaning heavy texture, probably in reference to the elytral surface.
</paragraph>
</subSubSection>
<subSubSection id="65D420B25858FF80FF3AA42453BF831C" pageId="8" pageNumber="9" type="distribution">
<paragraph id="2D7173395858FF80FF3AA42453BF831C" blockId="8.[151,1436,151,2012]" pageId="8" pageNumber="9">
<emphasis id="1FBAAF2B5858FF80FF3AA42450A782AC" bold="true" box="[201,736,1663,1688]" pageId="8" pageNumber="9">Distribution, phenology, and natural history</emphasis>
. This species is distributed along the humid slopes of Sierra Madre de Chiapas in
<collectingCountry id="55D933A95858FF80FE7CA4FF53A08288" box="[399,487,1700,1724]" name="Mexico" pageId="8" pageNumber="9">Mexico</collectingCountry>
, and Sierra Madre de
<collectingCountry id="55D933A95858FF80FD1BA4FF51268288" box="[744,865,1700,1724]" name="Guatemala" pageId="8" pageNumber="9">Guatemala</collectingCountry>
(
<figureCitation id="B5F56FBC5858FF80FC80A4FF518E8288" box="[883,969,1700,1724]" captionStart="FIGURE 54" captionStartId="21.[151,250,1234,1257]" captionTargetBox="[157,1436,382,1193]" captionTargetId="figure@21.[157,1442,373,1193]" captionTargetPageId="21" captionText="FIGURE 54. Distribution of species of Pachystethus." httpUri="https://zenodo.org/record/210552/files/figure.png" pageId="8" pageNumber="9">Fig. 54</figureCitation>
), between
<quantity id="EA36DEDC5858FF80FBB5A4FF56928288" box="[1094,1237,1700,1724]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.75" metricValueMin="0.55" pageId="8" pageNumber="9" unit="m" value="1150.0" valueMax="1750.0" valueMin="550.0">550–1750 m</quantity>
in elevation, and from the southeast of state of Chiapas,
<collectingCountry id="55D933A95858FF80FDBEA49350E482D4" box="[589,675,1736,1760]" name="Mexico" pageId="8" pageNumber="9">Mexico</collectingCountry>
(near Tapachula and Union Juárez) to south
<collectingCountry id="55D933A95858FF80FB69A493575482D4" box="[1178,1299,1736,1760]" name="Guatemala" pageId="8" pageNumber="9">Guatemala</collectingCountry>
(El Tumbador and Chimaltenango). Adults are active in September. No plants are recorded as hosts for adults, and the immature stages remain undescribed.
</paragraph>
</subSubSection>
<subSubSection id="65D420B25858FF80FF3AA56F52A883E8" pageId="8" pageNumber="9" type="discussion">
<paragraph id="2D7173395858FF80FF3AA56F52A883E8" blockId="8.[151,1436,151,2012]" pageId="8" pageNumber="9">
<emphasis id="1FBAAF2B5858FF80FF3AA56F537A8378" bold="true" box="[201,317,1844,1868]" pageId="8" pageNumber="9">Remarks.</emphasis>
In the supplement for Lamellicornia and Pectinicornia
<bibRefCitation id="495F0EC85858FF80FC41A56E56398378" author="Bates" box="[946,1150,1844,1868]" pageId="8" pageNumber="9" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia. In: O. Salvin &amp; F. D. Godwin (eds.), Biologia Centrali-Americana. Insecta Coleoptera. Volume 2, part 2: 1 - 432." type="book chapter" year="1888">Bates (1888: 406)</bibRefCitation>
described this taxon as a variety of
<taxonomicName id="EACE08BA5858FF80FEFAA50353F58344" box="[265,434,1880,1904]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="vidua">
<emphasis id="1FBAAF2B5858FF80FEFAA50353F58344" box="[265,434,1880,1904]" italics="true" pageId="8" pageNumber="9">Anomala vidua</emphasis>
</taxonomicName>
, arguing that it is a local modification and pointed out that the most distinct characters are the deep sculpture on dorsal surfaces and pale color in the tarsi of both sexes. Additional important differences include body shape, legs, ventral vestiture, body color, and in the form of the genitalia that support this as a valid species.
</paragraph>
</subSubSection>
</treatment>
</document>
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