480 lines
67 KiB
XML
480 lines
67 KiB
XML
<document id="401D049DBA6FC7605375931FEBD444C0" ID-DOI="10.11646/zootaxa.3866.1.1" ID-GBIF-Dataset="56308a7b-8f25-42e3-9dfd-9ca5322aec2a" ID-ISSN="1175-5326" ID-Zenodo-Dep="287158" ID-ZooBank="64DBAA6D-1CFA-451E-9613-B4A6321D8C7D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460915585981" checkinUser="plazi" docAuthor="Jaschhof, Mathias & Jaschhof, Catrin" docDate="2014" docId="EC7D87B42A2A052FFF03FF2D94D8F813" docLanguage="en" docName="zt03866p029.pdf" docOrigin="Zootaxa 3866 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zadbimyia Jaschhof & Jaschhof, 2014, gen. n." docType="treatment" docVersion="6" lastPageNumber="6" masterDocId="1044FFCC2A28052AFF94FFF09746FFF4" masterDocTitle="Zadbimyia, a new genus of asynaptine Porricondylinae (Diptera: Cecidomyiidae) with twenty-two new species from the cloud forest of Costa Rica" masterLastPageNumber="29" masterPageNumber="1" pageNumber="3" updateTime="1698361648592" updateUser="plazi">
|
||
<mods:mods id="96B32A7940D78CF7094FDEB1994E619D" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="2769FDB4A8A26182E945684826C30A96">
|
||
<mods:title id="A2A1C2C69A643DF0236D48F2DEEA4E16">Zadbimyia, a new genus of asynaptine Porricondylinae (Diptera: Cecidomyiidae) with twenty-two new species from the cloud forest of Costa Rica</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="7FFBB41BA8BD6C9C54468FC85D0B6179" type="personal">
|
||
<mods:role id="434565FB8599102FA498F6AF9C59006E">
|
||
<mods:roleTerm id="EC96932AA4843F41970195170360F565">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C7E667B0B877DD4692245E6597A0D9C5">Jaschhof, Mathias</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D4DB8CDF9568B452F1EA7060DC540787" type="personal">
|
||
<mods:role id="9CB8AE1FFD27816B9B5DAE26927A8825">
|
||
<mods:roleTerm id="51C2EA2A86515B78280564856B92A6F7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8073A8A61B631B07BED2BCC9C3B9AF61">Jaschhof, Catrin</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="705A9854873B2F6B99FB934BA30E4974">text</mods:typeOfResource>
|
||
<mods:relatedItem id="8F7F2E4C238B9BF90A67DF5A998AD9E5" type="host">
|
||
<mods:titleInfo id="E8A0DAE6050AAE6182753A4C83AF447F">
|
||
<mods:title id="B1B44D53F0735D6BD3473BCCE9BF7264">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="756B8B17FA90259850DC51B0766807DB">
|
||
<mods:date id="37B054C3F4E5A75D02A471F4F4FEC769">2014</mods:date>
|
||
<mods:detail id="3A634DBFAFC441125F5A85A562C83A77" type="volume">
|
||
<mods:number id="B4BBA0FCA9D05CC25448B3C7E1ED59D0">3866</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="E51B68CDFC3730668142C3A0C8011AAF" type="issue">
|
||
<mods:number id="6244F45432373D50A404A5D7954FF722">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="7630213D41AFF1C126BDD45D5D58C303" unit="page">
|
||
<mods:start id="99F464AC09C96DEFA6E8A1180C1660D1">1</mods:start>
|
||
<mods:end id="DF88CA9A9403940312D0BBA9FF16341D">29</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="334C2712D33038C5AD532EEB9C66815D">journal article</mods:classification>
|
||
<mods:identifier id="E49CF0FBC3218104E0BDD6E8ECE74552" type="DOI">10.11646/zootaxa.3866.1.1</mods:identifier>
|
||
<mods:identifier id="D1E2C2669B1B35F24ADFD869C292705D" type="GBIF-Dataset">56308a7b-8f25-42e3-9dfd-9ca5322aec2a</mods:identifier>
|
||
<mods:identifier id="E9DE468D3C5120FED5ADF3689EAADC6F" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="712A71FF24F24DE80110A1F55F04C664" type="Zenodo-Dep">287158</mods:identifier>
|
||
<mods:identifier id="A0283D9BF7A16815BBB1D570B4BC0DFF" type="ZooBank">64DBAA6D-1CFA-451E-9613-B4A6321D8C7D</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="EC7D87B42A2A052FFF03FF2D94D8F813" ID-DOI="http://doi.org/10.5281/zenodo.6132596" ID-GBIF-Taxon="127670135" ID-Zenodo-Dep="6132596" LSID="urn:lsid:plazi:treatment:EC7D87B42A2A052FFF03FF2D94D8F813" httpUri="http://treatment.plazi.org/id/EC7D87B42A2A052FFF03FF2D94D8F813" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
|
||
<subSubSection id="2CCE65292A2A0528FF03FF2D968EFF03" box="[151,456,221,247]" pageId="2" pageNumber="3" type="nomenclature">
|
||
<paragraph id="646B36A22A2A0528FF03FF2D968EFF03" blockId="2.[151,456,221,247]" box="[151,456,221,247]" pageId="2" pageNumber="3">
|
||
<heading id="3F2381CE2A2A0528FF03FF2D968EFF03" bold="true" box="[151,456,221,247]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
|
||
<emphasis id="56A0EAB02A2A0528FF03FF2D968EFF03" bold="true" box="[151,456,221,247]" pageId="2" pageNumber="3">
|
||
Genus
|
||
<taxonomicName id="A3D44D212A2A0528FF7AFF2D9629FF03" ID-CoL="89RR5" box="[238,367,221,247]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56A0EAB02A2A0528FF7AFF2D9629FF03" bold="true" box="[238,367,221,247]" italics="true" pageId="2" pageNumber="3">Zadbimyia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="4D9357CB2A2A0528FEE2FF2D968EFF03" box="[374,456,221,247]" pageId="2" pageNumber="3" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2CCE65292A2A0528FF03FED5951FFECE" box="[151,601,292,314]" pageId="2" pageNumber="3" type="materials_examined">
|
||
<paragraph id="646B36A22A2A0528FF03FED5951FFECE" blockId="2.[151,601,292,314]" box="[151,601,292,314]" pageId="2" pageNumber="3">
|
||
<typeStatus id="BB6F88002A2A0528FF03FED5978FFECE" box="[151,201,293,314]" pageId="2" pageNumber="3">Type</typeStatus>
|
||
species,
|
||
<taxonomicName id="A3D44D212A2A0528FEB1FED4955AFECE" box="[293,540,292,314]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="costaricensis" status="sp. nov.">
|
||
<emphasis id="56A0EAB02A2A0528FEB1FED4955AFECE" box="[293,540,292,314]" italics="true" pageId="2" pageNumber="3">Zadbimyia costaricensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="4D9357CB2A2A0528FDB6FED5951FFECE" box="[546,601,293,314]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2CCE65292A2A0528FF03FE969361FC8E" pageId="2" pageNumber="3" type="diagnosis">
|
||
<paragraph id="646B36A22A2A0528FF03FE969361FC8E" blockId="2.[151,1437,358,2014]" pageId="2" pageNumber="3">
|
||
<emphasis id="56A0EAB02A2A0528FF03FE969657FE8B" bold="true" box="[151,273,358,383]" pageId="2" pageNumber="3">Diagnosis.</emphasis>
|
||
Based on males (
|
||
<figureCitation id="FCEF2A272A2A0528FE41FE969550FE8B" box="[469,534,358,383]" captionStart="FIGURE 1" captionStartId="3.[151,250,1868,1890]" captionTargetBox="[418,1163,564,1833]" captionTargetId="figure@3.[411,1175,555,1844]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of male adult Zadbimyia sp., most of wing setae omitted. Scale 1.0 mm." httpUri="https://zenodo.org/record/287159/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||
); females and immature stages are unknown.
|
||
<taxonomicName id="A3D44D212A2A0528FB82FE9693C8FE8A" box="[1046,1166,358,382]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56A0EAB02A2A0528FB82FE9693C8FE8A" box="[1046,1166,358,382]" italics="true" pageId="2" pageNumber="3">Zadbimyia</emphasis>
|
||
</taxonomicName>
|
||
can be recognized as an
|
||
<taxonomicName id="A3D44D212A2A0528FF03FE7A9654FE57" box="[151,274,394,419]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
|
||
using the key by
|
||
<bibRefCitation id="00454B532A2A0528FE41FE7A947CFE57" author="Jaschhof" box="[469,826,394,419]" pageId="2" pageNumber="3" refString="Jaschhof, M. & Jaschhof, C. (2013) The Porricondylinae (Diptera: Cecidomyiidae) of Sweden, with notes on extralimital species. Studia dipterologica Supplement 20." type="book" year="2013">Jaschhof & Jaschhof (2013: 46)</bibRefCitation>
|
||
.
|
||
<taxonomicName id="A3D44D212A2A0528FCD7FE7A94F8FE57" box="[835,958,394,419]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
|
||
are the only
|
||
<taxonomicName id="A3D44D212A2A0528FBDAFE7A9247FE57" box="[1102,1281,394,419]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="subFamily" subFamily="Porricondylinae">Porricondylinae</taxonomicName>
|
||
that regularly have more than 14 flagellomeres and the abdominal sclerites subdivided in two or four raised portions.
|
||
<taxonomicName id="A3D44D212A2A0528FAB0FE5E92DAFE32" box="[1316,1436,430,454]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56A0EAB02A2A0528FAB0FE5E92DAFE32" box="[1316,1436,430,454]" italics="true" pageId="2" pageNumber="3">Zadbimyia</emphasis>
|
||
</taxonomicName>
|
||
is distinct from other
|
||
<taxonomicName id="A3D44D212A2A0528FE05FE22954AFE1F" box="[401,524,466,491]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
|
||
based on the following combination of character states:
|
||
<emphasis id="56A0EAB02A2A0528FB0DFE2393FBFE1F" box="[1177,1213,467,491]" italics="true" pageId="2" pageNumber="3">[1]</emphasis>
|
||
flagellomere necks usually have microtrichia, at least basally (
|
||
<figureCitation id="FCEF2A272A2A0528FD33FE0695B5FDFB" box="[679,755,502,527]" captionStart="FIGURE 4" captionStartId="6.[151,250,1936,1958]" captionTargetBox="[338,1247,917,1910]" captionTargetId="figure@6.[334,1253,914,1915]" captionTargetPageId="6" captionText="FIGURE 4. Zadbimyia costaricensis, male. A: ejaculatory apodeme and parameres, ventral. B: ditto. C: terminalia, ventral. D: fourth flagellomere, lateral. Scale 0.05 mm. ej ap = ejaculatory apodeme, par = paramere, par ap = parameral apodeme, par pr = paramere process." httpUri="https://zenodo.org/record/287162/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
|
||
D);
|
||
<emphasis id="56A0EAB02A2A0528FCB2FE07940DFDFB" box="[806,843,503,527]" italics="true" pageId="2" pageNumber="3">[2]</emphasis>
|
||
circumfila, in the typical outline, form a single circumferential band with short, irregular loops (
|
||
<figureCitation id="FCEF2A272A2A0528FD47FDEA945DFDC7" box="[723,795,538,563]" captionStart="FIGURE 4" captionStartId="6.[151,250,1936,1958]" captionTargetBox="[338,1247,917,1910]" captionTargetId="figure@6.[334,1253,914,1915]" captionTargetPageId="6" captionText="FIGURE 4. Zadbimyia costaricensis, male. A: ejaculatory apodeme and parameres, ventral. B: ditto. C: terminalia, ventral. D: fourth flagellomere, lateral. Scale 0.05 mm. ej ap = ejaculatory apodeme, par = paramere, par ap = parameral apodeme, par pr = paramere process." httpUri="https://zenodo.org/record/287162/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
|
||
D); both
|
||
<emphasis id="56A0EAB02A2A0528FC13FDEB94EDFDC7" box="[903,939,539,563]" italics="true" pageId="2" pageNumber="3">[3]</emphasis>
|
||
labella and
|
||
<emphasis id="56A0EAB02A2A0528FBD4FDEB9323FDC7" box="[1088,1125,539,563]" italics="true" pageId="2" pageNumber="3">[4]</emphasis>
|
||
palpus are atrophied (
|
||
<figureCitation id="FCEF2A272A2A0528FAFAFDEA97E0FDA3" captionStart="FIGURE 2" captionStartId="4.[151,250,1551,1573]" captionTargetBox="[340,1249,204,1524]" captionTargetId="figure@4.[328,1259,193,1529]" captionTargetPageId="4" captionText="FIGURE 2. Morphology of male Zadbimyia spp. A: basitarsus of foreleg, dorsolateral, of Z. browni. B: head, lateral, of Z. artborkenti. C: ditto, of Z. anniae. D: ditto, of Z. spinapiscis. E: fifth tarsomere of foreleg, lateral, of Z. browni. F: ditto, of Z. aberrans. Scale for A, E – F, 0.05 mm; B – D, 0.1 mm. clp = clypeus, eye br = eye bridge, fd chnl = food channel, lbl = labellum, lbr = labrum, mx prtb = maxillary protuberance, pfr = postfrons, plp = palpus, prfr = prefrons." httpUri="https://zenodo.org/record/287160/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
B–D);
|
||
<emphasis id="56A0EAB02A2A0528FF62FDCF965DFDA2" box="[246,283,575,598]" italics="true" pageId="2" pageNumber="3">[5]</emphasis>
|
||
vein r-m+m-cu is almost straight to slightly curved or sinuous, i.e. the point where the media is supposed to branch off is obscured (
|
||
<figureCitation id="FCEF2A272A2A0528FDB8FD929536FD8F" box="[556,624,610,635]" captionStart="FIGURE 3" captionStartId="5.[151,250,1361,1383]" captionTargetBox="[346,1242,632,1335]" captionTargetId="figure@5.[340,1248,625,1340]" captionTargetPageId="5" captionText="FIGURE 3. Wings of male Zadbimyia spp., setae omitted. A: Z. membranacea. B: Z. lasalturas. Scale 1.0 mm." httpUri="https://zenodo.org/record/287161/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||
);
|
||
<emphasis id="56A0EAB02A2A0528FD13FD9395EDFD8F" box="[647,683,611,635]" italics="true" pageId="2" pageNumber="3">[6]</emphasis>
|
||
vein M is absent (
|
||
<figureCitation id="FCEF2A272A2A0528FCEEFD9294F8FD8F" box="[890,958,610,635]" captionStart="FIGURE 3" captionStartId="5.[151,250,1361,1383]" captionTargetBox="[346,1242,632,1335]" captionTargetId="figure@5.[340,1248,625,1340]" captionTargetPageId="5" captionText="FIGURE 3. Wings of male Zadbimyia spp., setae omitted. A: Z. membranacea. B: Z. lasalturas. Scale 1.0 mm." httpUri="https://zenodo.org/record/287161/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||
);
|
||
<emphasis id="56A0EAB02A2A0528FC41FD9394BFFD8E" box="[981,1017,611,634]" italics="true" pageId="2" pageNumber="3">[7]</emphasis>
|
||
vein CuA1 is absent (
|
||
<figureCitation id="FCEF2A272A2A0528FB78FD92926BFD8F" box="[1260,1325,610,635]" captionStart="FIGURE 3" captionStartId="5.[151,250,1361,1383]" captionTargetBox="[346,1242,632,1335]" captionTargetId="figure@5.[340,1248,625,1340]" captionTargetPageId="5" captionText="FIGURE 3. Wings of male Zadbimyia spp., setae omitted. A: Z. membranacea. B: Z. lasalturas. Scale 1.0 mm." httpUri="https://zenodo.org/record/287161/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||
A) except in one species (
|
||
<figureCitation id="FCEF2A272A2A0528FEDCFD7996CAFD56" box="[328,396,649,674]" captionStart="FIGURE 3" captionStartId="5.[151,250,1361,1383]" captionTargetBox="[346,1242,632,1335]" captionTargetId="figure@5.[340,1248,625,1340]" captionTargetPageId="5" captionText="FIGURE 3. Wings of male Zadbimyia spp., setae omitted. A: Z. membranacea. B: Z. lasalturas. Scale 1.0 mm." httpUri="https://zenodo.org/record/287161/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||
B);
|
||
<emphasis id="56A0EAB02A2A0528FE2DFD79969BFD55" box="[441,477,649,673]" italics="true" pageId="2" pageNumber="3">[8]</emphasis>
|
||
each basitarsus has a spine ventro-apically (
|
||
<figureCitation id="FCEF2A272A2A0528FC4EFD799359FD56" box="[986,1055,649,674]" captionStart="FIGURE 2" captionStartId="4.[151,250,1551,1573]" captionTargetBox="[340,1249,204,1524]" captionTargetId="figure@4.[328,1259,193,1529]" captionTargetPageId="4" captionText="FIGURE 2. Morphology of male Zadbimyia spp. A: basitarsus of foreleg, dorsolateral, of Z. browni. B: head, lateral, of Z. artborkenti. C: ditto, of Z. anniae. D: ditto, of Z. spinapiscis. E: fifth tarsomere of foreleg, lateral, of Z. browni. F: ditto, of Z. aberrans. Scale for A, E – F, 0.05 mm; B – D, 0.1 mm. clp = clypeus, eye br = eye bridge, fd chnl = food channel, lbl = labellum, lbr = labrum, mx prtb = maxillary protuberance, pfr = postfrons, plp = palpus, prfr = prefrons." httpUri="https://zenodo.org/record/287160/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
A);
|
||
<emphasis id="56A0EAB02A2A0528FBD8FD799336FD55" box="[1100,1136,649,673]" italics="true" pageId="2" pageNumber="3">[9]</emphasis>
|
||
claws are strongly curved (
|
||
<figureCitation id="FCEF2A272A2A0528FF0BFD5D97AEFD32" box="[159,232,685,710]" captionStart="FIGURE 2" captionStartId="4.[151,250,1551,1573]" captionTargetBox="[340,1249,204,1524]" captionTargetId="figure@4.[328,1259,193,1529]" captionTargetPageId="4" captionText="FIGURE 2. Morphology of male Zadbimyia spp. A: basitarsus of foreleg, dorsolateral, of Z. browni. B: head, lateral, of Z. artborkenti. C: ditto, of Z. anniae. D: ditto, of Z. spinapiscis. E: fifth tarsomere of foreleg, lateral, of Z. browni. F: ditto, of Z. aberrans. Scale for A, E – F, 0.05 mm; B – D, 0.1 mm. clp = clypeus, eye br = eye bridge, fd chnl = food channel, lbl = labellum, lbr = labrum, mx prtb = maxillary protuberance, pfr = postfrons, plp = palpus, prfr = prefrons." httpUri="https://zenodo.org/record/287160/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
E) except in one species (
|
||
<figureCitation id="FCEF2A272A2A0528FDB1FD5D9528FD32" box="[549,622,685,710]" captionStart="FIGURE 2" captionStartId="4.[151,250,1551,1573]" captionTargetBox="[340,1249,204,1524]" captionTargetId="figure@4.[328,1259,193,1529]" captionTargetPageId="4" captionText="FIGURE 2. Morphology of male Zadbimyia spp. A: basitarsus of foreleg, dorsolateral, of Z. browni. B: head, lateral, of Z. artborkenti. C: ditto, of Z. anniae. D: ditto, of Z. spinapiscis. E: fifth tarsomere of foreleg, lateral, of Z. browni. F: ditto, of Z. aberrans. Scale for A, E – F, 0.05 mm; B – D, 0.1 mm. clp = clypeus, eye br = eye bridge, fd chnl = food channel, lbl = labellum, lbr = labrum, mx prtb = maxillary protuberance, pfr = postfrons, plp = palpus, prfr = prefrons." httpUri="https://zenodo.org/record/287160/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
F);
|
||
<emphasis id="56A0EAB02A2A0528FD0FFD5D9588FD31" box="[667,718,685,709]" italics="true" pageId="2" pageNumber="3">[10]</emphasis>
|
||
abdominal tergites are present as a single narrow, medially interrupted band on the posterior margin of segments I–VI (rarely VII);
|
||
<emphasis id="56A0EAB02A2A0528FC2AFD2194A9FD1D" box="[958,1007,721,745]" italics="true" pageId="2" pageNumber="3">[11]</emphasis>
|
||
the anterior portions of the gonocoxal apodemes are separated medially, i.e. not merged to form a common shaft (
|
||
<figureCitation id="FCEF2A272A2A0528FB80FD05931CFCFA" box="[1044,1114,757,782]" captionStart="FIGURE 5" captionStartId="8.[151,250,1882,1904]" captionTargetBox="[347,1241,991,1854]" captionTargetId="figure@8.[340,1248,988,1860]" captionTargetPageId="8" captionText="FIGURE 5. Zadbimyia browni, male. A: gonostylus, ventral. B: ejaculatory apodeme and parameres, ventral. C: terminalia, ventral. D: fourth flagellomere, lateral. Scale 0.05 mm. acc gl dc = accessory gland duct, antGA = anterior portion of gonocoxal apodeme, lat shld = lateral shoulder, vtr em = ventral emargination." httpUri="https://zenodo.org/record/287163/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
|
||
C);
|
||
<emphasis id="56A0EAB02A2A0528FB1DFD0593FDFCF9" box="[1161,1211,757,781]" italics="true" pageId="2" pageNumber="3">[12]</emphasis>
|
||
gonostylus lacks a pectinate claw and is densely microtrichose apically [
|
||
<figureCitation id="FCEF2A272A2A0528FD7FFCE99476FCC6" box="[747,816,793,818]" captionStart="FIGURE 4" captionStartId="6.[151,250,1936,1958]" captionTargetBox="[338,1247,917,1910]" captionTargetId="figure@6.[334,1253,914,1915]" captionTargetPageId="6" captionText="FIGURE 4. Zadbimyia costaricensis, male. A: ejaculatory apodeme and parameres, ventral. B: ditto. C: terminalia, ventral. D: fourth flagellomere, lateral. Scale 0.05 mm. ej ap = ejaculatory apodeme, par = paramere, par ap = parameral apodeme, par pr = paramere process." httpUri="https://zenodo.org/record/287162/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
|
||
C];
|
||
<emphasis id="56A0EAB02A2A0528FCCCFCE994CDFCC5" box="[856,907,793,817]" italics="true" pageId="2" pageNumber="3">[13]</emphasis>
|
||
the ejaculatory apodeme is well developed and distinct from the parameres [
|
||
<figureCitation id="FCEF2A272A2A0528FE41FCCD955CFCA2" box="[469,538,829,854]" captionStart="FIGURE 4" captionStartId="6.[151,250,1936,1958]" captionTargetBox="[338,1247,917,1910]" captionTargetId="figure@6.[334,1253,914,1915]" captionTargetPageId="6" captionText="FIGURE 4. Zadbimyia costaricensis, male. A: ejaculatory apodeme and parameres, ventral. B: ditto. C: terminalia, ventral. D: fourth flagellomere, lateral. Scale 0.05 mm. ej ap = ejaculatory apodeme, par = paramere, par ap = parameral apodeme, par pr = paramere process." httpUri="https://zenodo.org/record/287162/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
|
||
A];
|
||
<emphasis id="56A0EAB02A2A0528FDD0FCCD9530FCA1" box="[580,630,829,853]" italics="true" pageId="2" pageNumber="3">[14]</emphasis>
|
||
parameres are merged medially to form a structural unit (‘tegmen’); and
|
||
<emphasis id="56A0EAB02A2A0528FF03FC91978FFC8D" box="[151,201,865,889]" italics="true" pageId="2" pageNumber="3">[15]</emphasis>
|
||
bear 1–3 pairs of sclerotized processes ventrally to ventrolaterally (
|
||
<figureCitation id="FCEF2A272A2A0528FC54FC919345FC8E" box="[960,1027,865,890]" captionStart="FIGURE 4" captionStartId="6.[151,250,1936,1958]" captionTargetBox="[338,1247,917,1910]" captionTargetId="figure@6.[334,1253,914,1915]" captionTargetPageId="6" captionText="FIGURE 4. Zadbimyia costaricensis, male. A: ejaculatory apodeme and parameres, ventral. B: ditto. C: terminalia, ventral. D: fourth flagellomere, lateral. Scale 0.05 mm. ej ap = ejaculatory apodeme, par = paramere, par ap = parameral apodeme, par pr = paramere process." httpUri="https://zenodo.org/record/287162/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2CCE65292A2A052FFF53FC7593BCF86B" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="646B36A22A2A0529FF53FC7594C3FE00" blockId="2.[151,1437,358,2014]" lastBlockId="3.[151,1436,151,501]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||
<emphasis id="56A0EAB02A2A0528FF53FC7596DCFC6A" bold="true" box="[199,410,901,926]" pageId="2" pageNumber="3">Male description.</emphasis>
|
||
Slender, delicate midges, body lengths 1.3–2.0 mm (
|
||
<figureCitation id="FCEF2A272A2A0528FB90FC75930FFC6A" box="[1028,1097,901,926]" captionStart="FIGURE 1" captionStartId="3.[151,250,1868,1890]" captionTargetBox="[418,1163,564,1833]" captionTargetId="figure@3.[411,1175,555,1844]" captionTargetPageId="3" captionText="FIGURE 1. Habitus of male adult Zadbimyia sp., most of wing setae omitted. Scale 1.0 mm." httpUri="https://zenodo.org/record/287159/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||
). Color: body light brown to yellow, with dark setae and scales, latter densely covering legs and halter (trap-collected specimens usually rubbed, vestiture often missing).
|
||
<emphasis id="56A0EAB02A2A0528FE27FC3D96B5FC12" bold="true" box="[435,499,973,998]" pageId="2" pageNumber="3">Head</emphasis>
|
||
(
|
||
<figureCitation id="FCEF2A272A2A0528FD91FC3D950FFC12" box="[517,585,973,998]" captionStart="FIGURE 2" captionStartId="4.[151,250,1551,1573]" captionTargetBox="[340,1249,204,1524]" captionTargetId="figure@4.[328,1259,193,1529]" captionTargetPageId="4" captionText="FIGURE 2. Morphology of male Zadbimyia spp. A: basitarsus of foreleg, dorsolateral, of Z. browni. B: head, lateral, of Z. artborkenti. C: ditto, of Z. anniae. D: ditto, of Z. spinapiscis. E: fifth tarsomere of foreleg, lateral, of Z. browni. F: ditto, of Z. aberrans. Scale for A, E – F, 0.05 mm; B – D, 0.1 mm. clp = clypeus, eye br = eye bridge, fd chnl = food channel, lbl = labellum, lbr = labrum, mx prtb = maxillary protuberance, pfr = postfrons, plp = palpus, prfr = prefrons." httpUri="https://zenodo.org/record/287160/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
B–D). Eyes covering most of head, in some species narrowed ventrally, at vertex connate, resulting eye bridge 6–11 ommatidia long. Occiput evenly convex, with various sized setae including pair of large dorsal setae. Postfrons asetose. Prefrons extensive, usually well sclerotized. Scape markedly tapered toward base, with setal tuft distomedially, 1.5–2.5 times longer than pedicel, in
|
||
<taxonomicName id="A3D44D212A2A0528FB3FFBCA9248FBA5" box="[1195,1294,1081,1105]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dubia">
|
||
<emphasis id="56A0EAB02A2A0528FB3FFBCA9248FBA5" box="[1195,1294,1081,1105]" italics="true" pageId="2" pageNumber="3">Z. dubia</emphasis>
|
||
</taxonomicName>
|
||
shorter and conspicuously broad. Number of flagellomeres 16–31 with some intraspecific variation; with distinct necks, barrelshaped to subglobular nodes; in many species necks partly or entirely covered with microtrichia; node, from base to apex, with irregular double whorl of setae, one circumfilum, few to many large sensory hairs arising from hooded alveoli, one whorl of sensory hairs arising from ordinary alveoli; circumfilum typically sinuous, appressed to somewhat bowed, forming irregular loops between adjoining alveoli (
|
||
<figureCitation id="FCEF2A272A2A0528FC41FB1D9377FAF2" box="[981,1073,1261,1286]" captionStart="FIGURE 16" captionStartId="18.[151,250,1270,1292]" captionTargetBox="[341,1248,416,1245]" captionTargetId="figure@18.[333,1254,409,1249]" captionTargetPageId="18" captionText="FIGURE 16. Zadbimyia anniae, male. A: gonostylus, ventral. B: fourth flagellomere, lateral, only basal pores of circumfilum shown. C: terminalia, gonostyli omitted, ventral. D: ejaculatory apodeme and parameres, ventral. Scale 0.05 mm." httpUri="https://zenodo.org/record/287174/files/figure.png" pageId="2" pageNumber="3">Figs 16</figureCitation>
|
||
B, 19A); ultimate flagellomere neckless, often with additional apical bud. Clypeus small, with large setae or asetose. Labrum asetose. Labellum vestigial, basal segment tiny, smooth, asetose, apical segment with 1–3 setae. Maxillary protuberance large, in some species elongate (
|
||
<figureCitation id="FCEF2A272A2A0528FE25FAA996BDFA86" box="[433,507,1369,1394]" captionStart="FIGURE 2" captionStartId="4.[151,250,1551,1573]" captionTargetBox="[340,1249,204,1524]" captionTargetId="figure@4.[328,1259,193,1529]" captionTargetPageId="4" captionText="FIGURE 2. Morphology of male Zadbimyia spp. A: basitarsus of foreleg, dorsolateral, of Z. browni. B: head, lateral, of Z. artborkenti. C: ditto, of Z. anniae. D: ditto, of Z. spinapiscis. E: fifth tarsomere of foreleg, lateral, of Z. browni. F: ditto, of Z. aberrans. Scale for A, E – F, 0.05 mm; B – D, 0.1 mm. clp = clypeus, eye br = eye bridge, fd chnl = food channel, lbl = labellum, lbr = labrum, mx prtb = maxillary protuberance, pfr = postfrons, plp = palpus, prfr = prefrons." httpUri="https://zenodo.org/record/287160/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
D). Palpus atrophied, usually shorter than length of head, with 1–4 segments.
|
||
<emphasis id="56A0EAB02A2A0528FF03FA8D97B0FA62" bold="true" box="[151,246,1405,1430]" pageId="2" pageNumber="3">Thorax.</emphasis>
|
||
Scutum with large lateral and dorsocentral setae. Anepisternum and anepimeron setose except in
|
||
<taxonomicName id="A3D44D212A2A0528FAAAFA8E92DAFA61" box="[1342,1436,1405,1429]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="dubia">
|
||
<emphasis id="56A0EAB02A2A0528FAAAFA8E92DAFA61" box="[1342,1436,1405,1429]" italics="true" pageId="2" pageNumber="3">Z. dubia</emphasis>
|
||
</taxonomicName>
|
||
with asetose anepisternum, anepimeral setae twice as long as anepisternal setae.
|
||
<emphasis id="56A0EAB02A2A0528FBB9FA519328FA4E" bold="true" box="[1069,1134,1441,1466]" pageId="2" pageNumber="3">Wing</emphasis>
|
||
(
|
||
<figureCitation id="FCEF2A272A2A0528FBEBFA519383FA4E" box="[1151,1221,1441,1466]" captionStart="FIGURE 3" captionStartId="5.[151,250,1361,1383]" captionTargetBox="[346,1242,632,1335]" captionTargetId="figure@5.[340,1248,625,1340]" captionTargetPageId="5" captionText="FIGURE 3. Wings of male Zadbimyia spp., setae omitted. A: Z. membranacea. B: Z. lasalturas. Scale 1.0 mm." httpUri="https://zenodo.org/record/287161/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||
). About as long as body. Membrane usually tinged slightly gray or gray-brown, costal cell often darker than rest of membrane. Veins: R1 long; Rs short, oblique, in line with R5; R5 slightly curved, progressively departing from C, joining C beyond apex of wing; r-m+m-cu straight to slightly curved or slightly sinuous; M absent; no wingfold anteriad of CuA; CuA1 typically absent (
|
||
<figureCitation id="FCEF2A272A2A0528FE0CF9C4969FF9B9" box="[408,473,1588,1613]" captionStart="FIGURE 3" captionStartId="5.[151,250,1361,1383]" captionTargetBox="[346,1242,632,1335]" captionTargetId="figure@5.[340,1248,625,1340]" captionTargetPageId="5" captionText="FIGURE 3. Wings of male Zadbimyia spp., setae omitted. A: Z. membranacea. B: Z. lasalturas. Scale 1.0 mm." httpUri="https://zenodo.org/record/287161/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||
A), present only in
|
||
<taxonomicName id="A3D44D212A2A0528FD24F9C5947AF9B8" box="[688,828,1588,1612]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="lasalturas">
|
||
<emphasis id="56A0EAB02A2A0528FD24F9C5947AF9B8" box="[688,828,1588,1612]" italics="true" pageId="2" pageNumber="3">Z. lasalturas</emphasis>
|
||
</taxonomicName>
|
||
, there forming fork with CuA2 (
|
||
<figureCitation id="FCEF2A272A2A0528FB35F9C493A4F9B9" box="[1185,1250,1588,1613]" captionStart="FIGURE 3" captionStartId="5.[151,250,1361,1383]" captionTargetBox="[346,1242,632,1335]" captionTargetId="figure@5.[340,1248,625,1340]" captionTargetPageId="5" captionText="FIGURE 3. Wings of male Zadbimyia spp., setae omitted. A: Z. membranacea. B: Z. lasalturas. Scale 1.0 mm." httpUri="https://zenodo.org/record/287161/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||
B); CuA2 mostly straight, only distally curved, usually evanescent apically; CuP long, reaching to bend of CuA2.
|
||
<emphasis id="56A0EAB02A2A0528FB4BF9AB9258F980" bold="true" box="[1247,1310,1627,1652]" pageId="2" pageNumber="3">Legs.</emphasis>
|
||
More than twice body length. Each basitarsus with long, thin spine ventro-apically (
|
||
<figureCitation id="FCEF2A272A2A0528FC50F9719341F96E" box="[964,1031,1665,1690]" captionStart="FIGURE 2" captionStartId="4.[151,250,1551,1573]" captionTargetBox="[340,1249,204,1524]" captionTargetId="figure@4.[328,1259,193,1529]" captionTargetPageId="4" captionText="FIGURE 2. Morphology of male Zadbimyia spp. A: basitarsus of foreleg, dorsolateral, of Z. browni. B: head, lateral, of Z. artborkenti. C: ditto, of Z. anniae. D: ditto, of Z. spinapiscis. E: fifth tarsomere of foreleg, lateral, of Z. browni. F: ditto, of Z. aberrans. Scale for A, E – F, 0.05 mm; B – D, 0.1 mm. clp = clypeus, eye br = eye bridge, fd chnl = food channel, lbl = labellum, lbr = labrum, mx prtb = maxillary protuberance, pfr = postfrons, plp = palpus, prfr = prefrons." httpUri="https://zenodo.org/record/287160/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
A). Claws typically strongly curved, a large curved tooth basally, rarely a tiny tooth subapically (
|
||
<figureCitation id="FCEF2A272A2A0528FCC4F95594D0F94A" box="[848,918,1701,1726]" captionStart="FIGURE 2" captionStartId="4.[151,250,1551,1573]" captionTargetBox="[340,1249,204,1524]" captionTargetId="figure@4.[328,1259,193,1529]" captionTargetPageId="4" captionText="FIGURE 2. Morphology of male Zadbimyia spp. A: basitarsus of foreleg, dorsolateral, of Z. browni. B: head, lateral, of Z. artborkenti. C: ditto, of Z. anniae. D: ditto, of Z. spinapiscis. E: fifth tarsomere of foreleg, lateral, of Z. browni. F: ditto, of Z. aberrans. Scale for A, E – F, 0.05 mm; B – D, 0.1 mm. clp = clypeus, eye br = eye bridge, fd chnl = food channel, lbl = labellum, lbr = labrum, mx prtb = maxillary protuberance, pfr = postfrons, plp = palpus, prfr = prefrons." httpUri="https://zenodo.org/record/287160/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
E);
|
||
<taxonomicName id="A3D44D212A2A0528FC54F9579303F94A" box="[960,1093,1702,1726]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="aberrans">
|
||
<emphasis id="56A0EAB02A2A0528FC54F9579303F94A" box="[960,1093,1702,1726]" italics="true" pageId="2" pageNumber="3">Z. aberrans</emphasis>
|
||
</taxonomicName>
|
||
exceptional with thin, almost straight claws with fine basal tooth (
|
||
<figureCitation id="FCEF2A272A2A0528FDDEF93995D4F916" box="[586,658,1737,1762]" captionStart="FIGURE 2" captionStartId="4.[151,250,1551,1573]" captionTargetBox="[340,1249,204,1524]" captionTargetId="figure@4.[328,1259,193,1529]" captionTargetPageId="4" captionText="FIGURE 2. Morphology of male Zadbimyia spp. A: basitarsus of foreleg, dorsolateral, of Z. browni. B: head, lateral, of Z. artborkenti. C: ditto, of Z. anniae. D: ditto, of Z. spinapiscis. E: fifth tarsomere of foreleg, lateral, of Z. browni. F: ditto, of Z. aberrans. Scale for A, E – F, 0.05 mm; B – D, 0.1 mm. clp = clypeus, eye br = eye bridge, fd chnl = food channel, lbl = labellum, lbr = labrum, mx prtb = maxillary protuberance, pfr = postfrons, plp = palpus, prfr = prefrons." httpUri="https://zenodo.org/record/287160/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
F). Empodia and pulvilli rudimentary.
|
||
<emphasis id="56A0EAB02A2A0528FBC3F93993B1F916" bold="true" box="[1111,1271,1737,1762]" pageId="2" pageNumber="3">Preabdomen.</emphasis>
|
||
Tergites I–VI (rarely VII) present as narrow, raised, setose, medially interrupted bands on posterior margin of segments, with strikingly large setae, terga VII–VIII largely unsclerotized, asetose. Pleural membrane at least of anterior segments setose, pleural setae 1/3 as long as tergal setae. Sternum I unsclerotized, asetose, sternites II–VIII evenly sclerotized, setose, sternal setae 1/3 to 1/2 as long as tergal setae.
|
||
<emphasis id="56A0EAB02A2A0528FC08F8A99359F886" bold="true" box="[924,1055,1881,1906]" pageId="2" pageNumber="3">Terminalia</emphasis>
|
||
(see
|
||
<figureCitation id="FCEF2A272A2A0528FBF4F8A993ECF886" box="[1120,1194,1881,1906]" captionStart="FIGURE 4" captionStartId="6.[151,250,1936,1958]" captionTargetBox="[338,1247,917,1910]" captionTargetId="figure@6.[334,1253,914,1915]" captionTargetPageId="6" captionText="FIGURE 4. Zadbimyia costaricensis, male. A: ejaculatory apodeme and parameres, ventral. B: ditto. C: terminalia, ventral. D: fourth flagellomere, lateral. Scale 0.05 mm. ej ap = ejaculatory apodeme, par = paramere, par ap = parameral apodeme, par pr = paramere process." httpUri="https://zenodo.org/record/287162/files/figure.png" pageId="2" pageNumber="3">Figs 4</figureCitation>
|
||
A, 5B–C for specific terms). Tergite IX subtrapezoid, with posterior or lateroposterior setae, posterior margin straight or medially concave, anterior margin membranous, obscured. Gonocoxites broadly united ventrobasally, with posteromedial concavity (= ventral emargination), ventrobasal portion narrowed to various extent, often subrectangular, gonocoxal apodemes with distinct anterior and posterior portions, anterior portions (antGA in species descriptions) maximally as long as distance separating them. Gonostylus robust, curved, tapered toward apex to various extent, with dense, often large microtrichia apically, without spines, subapical bristles usually absent, present only in
|
||
<taxonomicName id="A3D44D212A2B0529FA11FF1197BAFEE8" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="aberrans">
|
||
<emphasis id="56A0EAB02A2B0529FA11FF1197BAFEE8" italics="true" pageId="3" pageNumber="4">Z. aberrans</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A3D44D212A2B0529FEA0FEF596D4FEE8" box="[308,402,260,284]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="dubia">
|
||
<emphasis id="56A0EAB02A2B0529FEA0FEF596D4FEE8" box="[308,402,260,284]" italics="true" pageId="3" pageNumber="4">Z. dubia</emphasis>
|
||
</taxonomicName>
|
||
. Ejaculatory apodeme long, strongly sclerotized; ducts of accessory glands usually distinct, joining apodeme near apex. Parameres merged medially, typical outline (
|
||
<figureCitation id="FCEF2A272A2B0529FC44FED7935FFEB4" box="[976,1049,295,320]" captionStart="FIGURE 4" captionStartId="6.[151,250,1936,1958]" captionTargetBox="[338,1247,917,1910]" captionTargetId="figure@6.[334,1253,914,1915]" captionTargetPageId="6" captionText="FIGURE 4. Zadbimyia costaricensis, male. A: ejaculatory apodeme and parameres, ventral. B: ditto. C: terminalia, ventral. D: fourth flagellomere, lateral. Scale 0.05 mm. ej ap = ejaculatory apodeme, par = paramere, par ap = parameral apodeme, par pr = paramere process." httpUri="https://zenodo.org/record/287162/files/figure.png" pageId="3" pageNumber="4">Figs 4</figureCitation>
|
||
A, 5B) with broad portion basally, usually small, weakly sclerotized apodemes, and narrow, subcylindrical, often seemingly two-pointed portion apicomedially, typically 1–3 pairs of diversely shaped, sclerotized processes and, in some species, pronounced lateral shoulders at transition from basal to apical portions; atypical parameres may occur (see
|
||
<figureCitation id="FCEF2A272A2B0529FB77FE64927DFE58" box="[1251,1339,404,429]" captionStart="FIGURE 23" captionStartId="24.[151,250,1710,1732]" captionTargetBox="[340,1246,830,1689]" captionTargetId="figure@24.[334,1253,825,1694]" captionTargetPageId="24" captionText="FIGURE 23. Zadbimyia inornata, male. A: terminalia, ventral. B: fourth flagellomere, lateral. C: ejaculatory apodeme and parameres, ventral. Scale 0.05 mm." httpUri="https://zenodo.org/record/287181/files/figure.png" pageId="3" pageNumber="4">Figs 23</figureCitation>
|
||
C, 25C). Hypoproct deeply incised, resulting in two rounded lobes apically, each with a few apical setulae. Cerci similar to hypoproct but larger, usually extending beyond level of parameres.
|
||
</paragraph>
|
||
<caption id="30AB662A2A2B0529FF03F8BC9301F896" box="[151,1095,1868,1890]" httpUri="https://zenodo.org/record/287159/files/figure.png" pageId="3" pageNumber="4" targetBox="[418,1163,564,1833]" targetPageId="3">
|
||
<paragraph id="646B36A22A2B0529FF03F8BC9301F896" blockId="3.[151,1095,1868,1890]" box="[151,1095,1868,1890]" pageId="3" pageNumber="4">
|
||
<emphasis id="56A0EAB02A2B0529FF03F8BC9655F895" bold="true" box="[151,275,1868,1890]" pageId="3" pageNumber="4">FIGURE 1.</emphasis>
|
||
Habitus of male adult
|
||
<taxonomicName id="A3D44D212A2B0529FE6EF8BC9520F896" box="[506,614,1868,1890]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56A0EAB02A2B0529FE6EF8BC9520F896" box="[506,614,1868,1890]" italics="true" pageId="3" pageNumber="4">Zadbimyia</emphasis>
|
||
</taxonomicName>
|
||
sp., most of wing setae omitted. Scale 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="30AB662A2A2C052EFF03F9FF9309F975" httpUri="https://zenodo.org/record/287160/files/figure.png" pageId="4" pageNumber="5" targetBox="[340,1249,204,1524]" targetPageId="4">
|
||
<paragraph id="646B36A22A2C052EFF03F9FF9309F975" blockId="4.[151,1437,1551,1665]" pageId="4" pageNumber="5">
|
||
<emphasis id="56A0EAB02A2C052EFF03F9FF9653F9D0" bold="true" box="[151,277,1551,1573]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
|
||
Morphology of male
|
||
<taxonomicName id="A3D44D212A2C052EFE6FF9FF9521F9D1" box="[507,615,1551,1573]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56A0EAB02A2C052EFE6FF9FF9521F9D1" box="[507,615,1551,1573]" italics="true" pageId="4" pageNumber="5">Zadbimyia</emphasis>
|
||
</taxonomicName>
|
||
spp.
|
||
<emphasis id="56A0EAB02A2C052EFD36F9FF95FAF9D0" bold="true" box="[674,700,1551,1572]" pageId="4" pageNumber="5">A:</emphasis>
|
||
basitarsus of foreleg, dorsolateral, of
|
||
<taxonomicName id="A3D44D212A2C052EFBDDF9E093EAF9D1" box="[1097,1196,1551,1573]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="browni">
|
||
<emphasis id="56A0EAB02A2C052EFBDDF9E093EAF9D1" box="[1097,1196,1551,1573]" italics="true" pageId="4" pageNumber="5">Z. browni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="56A0EAB02A2C052EFB28F9FF9393F9D0" bold="true" box="[1212,1237,1551,1572]" pageId="4" pageNumber="5">B:</emphasis>
|
||
head, lateral, of
|
||
<taxonomicName id="A3D44D212A2C052EFA13F9E0964FF9B7" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="artborkenti">
|
||
<emphasis id="56A0EAB02A2C052EFA13F9E0964FF9B7" italics="true" pageId="4" pageNumber="5">Z. artborkenti</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="56A0EAB02A2C052EFE82F9DD9677F9B7" bold="true" box="[278,305,1581,1603]" pageId="4" pageNumber="5">C:</emphasis>
|
||
ditto, of
|
||
<taxonomicName id="A3D44D212A2C052EFE1AF9DF96A9F9B0" box="[398,495,1583,1604]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="anniae">
|
||
<emphasis id="56A0EAB02A2C052EFE1AF9DF96A9F9B0" box="[398,495,1583,1604]" italics="true" pageId="4" pageNumber="5">Z. anniae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="56A0EAB02A2C052EFE68F9DE9550F9B7" bold="true" box="[508,534,1582,1603]" pageId="4" pageNumber="5">D:</emphasis>
|
||
ditto, of
|
||
<taxonomicName id="A3D44D212A2C052EFDE0F9DF9446F9B0" box="[628,768,1583,1604]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="spinapiscis">
|
||
<emphasis id="56A0EAB02A2C052EFDE0F9DF9446F9B0" box="[628,768,1583,1604]" italics="true" pageId="4" pageNumber="5">Z. spinapiscis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="56A0EAB02A2C052EFC99F9DE9463F9B7" bold="true" box="[781,805,1582,1603]" pageId="4" pageNumber="5">E:</emphasis>
|
||
fifth tarsomere of foreleg, lateral, of
|
||
<taxonomicName id="A3D44D212A2C052EFB37F9DF9242F9B7" box="[1187,1284,1581,1604]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="browni">
|
||
<emphasis id="56A0EAB02A2C052EFB37F9DF9242F9B7" box="[1187,1284,1581,1604]" italics="true" pageId="4" pageNumber="5">Z. browni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="56A0EAB02A2C052EFA86F9DE926FF9B7" bold="true" box="[1298,1321,1582,1603]" pageId="4" pageNumber="5">F:</emphasis>
|
||
ditto, of
|
||
<taxonomicName id="A3D44D212A2C052EFA13F9DF97B5F996" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="aberrans">
|
||
<emphasis id="56A0EAB02A2C052EFA13F9DF97B5F996" italics="true" pageId="4" pageNumber="5">Z. aberrans</emphasis>
|
||
</taxonomicName>
|
||
. Scale for A, E–F, 0.05 mm; B–D, 0.1 mm. clp = clypeus, eye br = eye bridge, fd chnl = food channel, lbl = labellum, lbr = labrum, mx prtb = maxillary protuberance, pfr = postfrons, plp = palpus, prfr = prefrons.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="646B36A22A2C052FFF53F9589407FDC9" blockId="4.[151,1436,1704,2017]" lastBlockId="5.[151,1437,151,573]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
<emphasis id="56A0EAB02A2C052EFF53F95894EAF935" bold="true" box="[199,940,1704,1729]" pageId="4" pageNumber="5">Phylogenetic relationships as inferred from adult morphology.</emphasis>
|
||
The basic construction of the male genitalia in
|
||
<taxonomicName id="A3D44D212A2C052EFF27F93C966DF910" box="[179,299,1740,1764]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56A0EAB02A2C052EFF27F93C966DF910" box="[179,299,1740,1764]" italics="true" pageId="4" pageNumber="5">Zadbimyia</emphasis>
|
||
</taxonomicName>
|
||
is uniform (see character states
|
||
<emphasis id="56A0EAB02A2C052EFD06F93C9447F910" box="[658,769,1740,1764]" italics="true" pageId="4" pageNumber="5">[11]–[15]</emphasis>
|
||
specified in the generic diagnosis), and character state
|
||
<emphasis id="56A0EAB02A2C052EFAFEF93C92DAF910" box="[1386,1436,1740,1764]" italics="true" pageId="4" pageNumber="5">[15]</emphasis>
|
||
can be regarded as a synapomorphy of the included species. Also, the genus contains the only
|
||
<taxonomicName id="A3D44D212A2C052EFB26F900926BF8FD" box="[1202,1325,1776,1801]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
|
||
known to have the flagellomere necks completely covered with microtrichia.
|
||
<taxonomicName id="A3D44D212A2C052EFC00F8E4934AF8D8" box="[916,1036,1812,1836]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56A0EAB02A2C052EFC00F8E4934AF8D8" box="[916,1036,1812,1836]" italics="true" pageId="4" pageNumber="5">Zadbimyia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A3D44D212A2C052EFBD1F8E49245F8D9" authority="Spungis" authorityName="Spungis" box="[1093,1283,1812,1837]" class="Insecta" family="Cecidomyiidae" genus="Asycola" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56A0EAB02A2C052EFBD1F8E493D8F8D8" box="[1093,1182,1812,1836]" italics="true" pageId="4" pageNumber="5">Asycola</emphasis>
|
||
Spungis
|
||
</taxonomicName>
|
||
, a monotypic genus of
|
||
<taxonomicName id="A3D44D212A2C052EFE93F8C896C4F8A5" box="[263,386,1848,1873]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
|
||
from the Palearctic, are the only
|
||
<taxonomicName id="A3D44D212A2C052EFC83F8C8948CF8A5" box="[791,970,1848,1873]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Porricondylinae">Porricondylinae</taxonomicName>
|
||
with multi-looped circumfila, yet other characters indicate that the two genera are not closely related (see
|
||
<bibRefCitation id="00454B532A2C052EFCE8F8AC935FF881" author="Spungis" box="[892,1049,1884,1909]" pageId="4" pageNumber="5" refString="Spungis, V. (1991) [Review of European species of gall-midges of the subtribe Asynaptina (Diptera, Cecidomyiidae).] Entomologicheskoe Obozrenie 70, 196 - 202. Translated in Entomological Review, 70 (9), 44 - 52. [1992]" type="journal article" year="1991">Spungis 1991</bibRefCitation>
|
||
). Altogether, there is no doubt that
|
||
<taxonomicName id="A3D44D212A2C052EFF03F8709649F86C" box="[151,271,1920,1944]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56A0EAB02A2C052EFF03F8709649F86C" box="[151,271,1920,1944]" italics="true" pageId="4" pageNumber="5">Zadbimyia</emphasis>
|
||
</taxonomicName>
|
||
is a monophyletic group. In searching for its closest relatives one is reliant upon fragments of knowledge available for other New World and tropical
|
||
<taxonomicName id="A3D44D212A2C052EFC94F8549439F849" box="[768,895,1956,1981]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
|
||
. Based on this
|
||
<taxonomicName id="A3D44D212A2C052EFBB2F8549247F848" box="[1062,1281,1956,1980]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56A0EAB02A2C052EFBB2F8549247F848" box="[1062,1281,1956,1980]" italics="true" pageId="4" pageNumber="5">Pseudocamptomyia</emphasis>
|
||
</taxonomicName>
|
||
appears to be closer to
|
||
<taxonomicName id="A3D44D212A2C052EFF6AF8389630F814" box="[254,374,1992,2016]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56A0EAB02A2C052EFF6AF8389630F814" box="[254,374,1992,2016]" italics="true" pageId="4" pageNumber="5">Zadbimyia</emphasis>
|
||
</taxonomicName>
|
||
than any other
|
||
<taxonomicName id="A3D44D212A2C052EFDB1F83895E6F815" box="[549,672,1992,2017]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
|
||
described. The North American
|
||
<typeStatus id="BB6F88002A2C052EFB86F8399304F815" box="[1042,1090,1993,2017]" pageId="4" pageNumber="5">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="A3D44D212A2C052FFB55F83897AFFF44" authority="Parnell" authorityName="Parnell" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" lastPageId="5" lastPageNumber="6" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56A0EAB02A2C052EFB55F83892DAF814" box="[1217,1436,1992,2016]" italics="true" pageId="4" pageNumber="5">Pseudocamptomyia</emphasis>
|
||
Parnell
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A3D44D212A2D052FFF60FF699683FF44" authority="Felt" authorityName="Felt" box="[244,453,151,176]" class="Insecta" family="Cecidomyiidae" genus="Asynapta" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="phosphila">
|
||
<emphasis id="56A0EAB02A2D052FFF60FF69963BFF44" box="[244,381,152,176]" italics="true" pageId="5" pageNumber="6">P. phosphila</emphasis>
|
||
(Felt)
|
||
</taxonomicName>
|
||
, resembles
|
||
<taxonomicName id="A3D44D212A2D052FFDD3FF6895F9FF44" box="[583,703,152,176]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56A0EAB02A2D052FFDD3FF6895F9FF44" box="[583,703,152,176]" italics="true" pageId="5" pageNumber="6">Zadbimyia</emphasis>
|
||
</taxonomicName>
|
||
species with respect to character states
|
||
<emphasis id="56A0EAB02A2D052FFBEEFF6893D8FF44" box="[1146,1182,152,176]" italics="true" pageId="5" pageNumber="6">[3]</emphasis>
|
||
,
|
||
<emphasis id="56A0EAB02A2D052FFB38FF689396FF44" box="[1196,1232,152,176]" italics="true" pageId="5" pageNumber="6">[4]</emphasis>
|
||
,
|
||
<emphasis id="56A0EAB02A2D052FFB49FF689247FF44" box="[1245,1281,152,176]" italics="true" pageId="5" pageNumber="6">[6]</emphasis>
|
||
,
|
||
<emphasis id="56A0EAB02A2D052FFA9AFF689274FF5B" box="[1294,1330,152,175]" italics="true" pageId="5" pageNumber="6">[7]</emphasis>
|
||
,
|
||
<emphasis id="56A0EAB02A2D052FFAD4FF6892DBFF44" box="[1344,1437,152,176]" italics="true" pageId="5" pageNumber="6">[10–12]</emphasis>
|
||
and
|
||
<emphasis id="56A0EAB02A2D052FFF52FF4C97B1FF20" box="[198,247,188,212]" italics="true" pageId="5" pageNumber="6">[14]</emphasis>
|
||
, yet there are a number of differences. Most notably, in
|
||
<taxonomicName id="A3D44D212A2D052FFCFCFF4D94B4FF20" box="[872,1010,188,212]" class="Insecta" family="Cecidomyiidae" genus="Asynapta" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="phosphila">
|
||
<emphasis id="56A0EAB02A2D052FFCFCFF4D94B4FF20" box="[872,1010,188,212]" italics="true" pageId="5" pageNumber="6">P. phosphila</emphasis>
|
||
</taxonomicName>
|
||
circumfila are not looped but are only slightly sinuous; vein r-m+m-cu is strongly sinuous and enters the radius at an almost right angle; each basitarsus has a microtrichose projection rather than a spine ventro-apically; the gonostylus lacks microtrichia and possibly has a small spine apically (which cannot be accurately determined in the two specimens existing of this species); and the ejaculatory apodeme and parameres together form a complex structure, in which the apodeme bears two pairs of apical processes and the parameres one lateral pair. Two species from
|
||
<collectingCountry id="1CC376322A2D052FFB84FE9F9328FE7C" box="[1040,1134,367,392]" name="Somalia" pageId="5" pageNumber="6">Somalia</collectingCountry>
|
||
, unavailable for our study, were classified in
|
||
<taxonomicName id="A3D44D212A2D052FFEE7FE649509FE58" box="[371,591,404,428]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56A0EAB02A2D052FFEE7FE649509FE58" box="[371,591,404,428]" italics="true" pageId="5" pageNumber="6">Pseudocamptomyia</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="00454B532A2D052FFD10FE659490FE58" author="Mamaev" box="[644,982,404,429]" pageId="5" pageNumber="6" refString="Mamaev, B. M. & Zaitzev, A. I. (1997) New genus and species of free-developing gall midges of the subfamily Porricondylinae from Somalia (Diptera, Cecidomyiidae). Journal of the Ukrainian entomological Society, 3 (2), 5 - 13." type="journal article" year="1997">Mamaev and Zaitzev (1997)</bibRefCitation>
|
||
. Their genitalia are figured as having processes similar to that of
|
||
<taxonomicName id="A3D44D212A2D052FFE42FE499525FE24" box="[470,611,440,464]" class="Insecta" family="Cecidomyiidae" genus="Asynapta" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="phosphila">
|
||
<emphasis id="56A0EAB02A2D052FFE42FE499525FE24" box="[470,611,440,464]" italics="true" pageId="5" pageNumber="6">P. phosphila</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="00454B532A2D052FFDE0FE4894DCFE24" author="Mamaev" box="[628,922,439,464]" pageId="5" pageNumber="6" refString="Mamaev, B. M. & Zaitzev, A. I. (1997) New genus and species of free-developing gall midges of the subfamily Porricondylinae from Somalia (Diptera, Cecidomyiidae). Journal of the Ukrainian entomological Society, 3 (2), 5 - 13." type="journal article" year="1997">Mamaev & Zaitzev 1997</bibRefCitation>
|
||
: fig. 3b–c), but it is impossible to determine where exactly these processes are inserted. Other important characters were also not adequately described. Regardless of whether these two Afrotropical species are properly placed in the genus
|
||
<taxonomicName id="A3D44D212A2D052FFBF9FDF0920EFDEC" box="[1133,1352,512,536]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56A0EAB02A2D052FFBF9FDF0920EFDEC" box="[1133,1352,512,536]" italics="true" pageId="5" pageNumber="6">Pseudocamptomyia</emphasis>
|
||
</taxonomicName>
|
||
, which remains to be verified, they are certainly closely related to it.
|
||
</paragraph>
|
||
<caption id="30AB662A2A2D052FFF03FAA19243FA93" box="[151,1285,1361,1383]" httpUri="https://zenodo.org/record/287161/files/figure.png" pageId="5" pageNumber="6" targetBox="[346,1242,632,1335]" targetPageId="5">
|
||
<paragraph id="646B36A22A2D052FFF03FAA19243FA93" blockId="5.[151,1285,1361,1383]" box="[151,1285,1361,1383]" pageId="5" pageNumber="6">
|
||
<emphasis id="56A0EAB02A2D052FFF03FAA19654FA92" bold="true" box="[151,274,1361,1383]" pageId="5" pageNumber="6">FIGURE 3.</emphasis>
|
||
Wings of male
|
||
<taxonomicName id="A3D44D212A2D052FFE27FAA19559FA93" box="[435,543,1361,1383]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56A0EAB02A2D052FFE27FAA19559FA93" box="[435,543,1361,1383]" italics="true" pageId="5" pageNumber="6">Zadbimyia</emphasis>
|
||
</taxonomicName>
|
||
spp., setae omitted.
|
||
<emphasis id="56A0EAB02A2D052FFD7BFAA1944FFA92" bold="true" box="[751,777,1361,1382]" pageId="5" pageNumber="6">A:</emphasis>
|
||
<taxonomicName id="A3D44D212A2D052FFC84FAA294FCFA93" box="[784,954,1361,1383]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="membranacea">
|
||
<emphasis id="56A0EAB02A2D052FFC84FAA294FCFA93" box="[784,954,1361,1383]" italics="true" pageId="5" pageNumber="6">Z. membranacea</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="56A0EAB02A2D052FFC53FAA194A6FA92" bold="true" box="[967,992,1361,1382]" pageId="5" pageNumber="6">B:</emphasis>
|
||
<taxonomicName id="A3D44D212A2D052FFC72FAA29320FA93" box="[998,1126,1361,1383]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lasalturas">
|
||
<emphasis id="56A0EAB02A2D052FFC72FAA29320FA93" box="[998,1126,1361,1383]" italics="true" pageId="5" pageNumber="6">Z. lasalturas</emphasis>
|
||
</taxonomicName>
|
||
. Scale 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="646B36A22A2D052FFF53FA7F94ABF8FB" blockId="5.[151,1437,1422,2023]" pageId="5" pageNumber="6">
|
||
As regards the interspecific relationships of
|
||
<taxonomicName id="A3D44D212A2D052FFD50FA7E947AFA52" box="[708,828,1422,1446]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56A0EAB02A2D052FFD50FA7E947AFA52" box="[708,828,1422,1446]" italics="true" pageId="5" pageNumber="6">Zadbimyia</emphasis>
|
||
</taxonomicName>
|
||
, there is a large core group of species, including the
|
||
<typeStatus id="BB6F88002A2D052FFF03FA439781FA3F" box="[151,199,1459,1483]" pageId="5" pageNumber="6">type</typeStatus>
|
||
species, whose parameres have moderately sized, toothlike processes (see, for instance,
|
||
<figureCitation id="FCEF2A272A2D052FFB30FA4293A1FA3F" box="[1188,1255,1458,1483]" captionStart="FIGURE 4" captionStartId="6.[151,250,1936,1958]" captionTargetBox="[338,1247,917,1910]" captionTargetId="figure@6.[334,1253,914,1915]" captionTargetPageId="6" captionText="FIGURE 4. Zadbimyia costaricensis, male. A: ejaculatory apodeme and parameres, ventral. B: ditto. C: terminalia, ventral. D: fourth flagellomere, lateral. Scale 0.05 mm. ej ap = ejaculatory apodeme, par = paramere, par ap = parameral apodeme, par pr = paramere process." httpUri="https://zenodo.org/record/287162/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
|
||
B). It is possible that these species are more closely related to each other than to those exhibiting different
|
||
<typeStatus id="BB6F88002A2D052FFB05FA27938AFA1B" box="[1169,1228,1495,1519]" pageId="5" pageNumber="6">types</typeStatus>
|
||
of parameres (see
|
||
<figureCitation id="FCEF2A272A2D052FFF03FA0A97A8F9E7" box="[151,238,1530,1555]" captionStart="FIGURE 22" captionStartId="23.[151,250,1942,1964]" captionTargetBox="[338,1246,1026,1913]" captionTargetId="figure@23.[332,1256,1020,1921]" captionTargetPageId="23" captionText="FIGURE 22. Zadbimyia spinapiscis, male. A: terminalia, ventral. B: fourth flagellomere, lateral, only basal pores of circumfilum shown. C: ejaculatory apodeme and parameres, ventral. Scale 0.05 mm." httpUri="https://zenodo.org/record/287180/files/figure.png" pageId="5" pageNumber="6">Figs 22</figureCitation>
|
||
C, 24C, 25C). However, this assumption is not supported by additional evidence, so for the time being it remains uncertain whether or not this core group is monophyletic. Even subsets of the core group, such as those species having two pairs of paramere processes, are usually quite variable regarding other characters, such as the outline of the eye bridge, size of the maxillary protuberance, or coverage of flagellomere necks with microtrichiae. As an exception,
|
||
<taxonomicName id="A3D44D212A2D052FFEF1F97B955FF956" box="[357,537,1675,1698]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="costaricensis">
|
||
<emphasis id="56A0EAB02A2D052FFEF1F97B955FF956" box="[357,537,1675,1698]" italics="true" pageId="5" pageNumber="6">Z. costaricensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A3D44D212A2D052FFDBFF97B95DCF957" box="[555,666,1675,1699]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="browni">
|
||
<emphasis id="56A0EAB02A2D052FFDBFF97B95DCF957" box="[555,666,1675,1699]" italics="true" pageId="5" pageNumber="6">Z. browni</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="A3D44D212A2D052FFD74F97B94C6F957" box="[736,896,1675,1699]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="artborkenti">
|
||
<emphasis id="56A0EAB02A2D052FFD74F97B94C6F957" box="[736,896,1675,1699]" italics="true" pageId="5" pageNumber="6">Z. artborkenti</emphasis>
|
||
</taxonomicName>
|
||
correspond in having tarsal claws with a tiny subapical ancillary tooth in addition to parameres of the same outline, so this trio is certainly monophyletic. Various
|
||
<taxonomicName id="A3D44D212A2D052FFF60F923962AF91F" box="[244,364,1747,1771]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56A0EAB02A2D052FFF60F923962AF91F" box="[244,364,1747,1771]" italics="true" pageId="5" pageNumber="6">Zadbimyia</emphasis>
|
||
</taxonomicName>
|
||
with unusual parameres, namely
|
||
<taxonomicName id="A3D44D212A2D052FFD71F9239439F91E" box="[741,895,1747,1770]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="spinapiscis">
|
||
<emphasis id="56A0EAB02A2D052FFD71F9239439F91E" box="[741,895,1747,1770]" italics="true" pageId="5" pageNumber="6">Z. spinapiscis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A3D44D212A2D052FFC1AF923934AF91E" box="[910,1036,1747,1770]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="inornata">
|
||
<emphasis id="56A0EAB02A2D052FFC1AF923934AF91E" box="[910,1036,1747,1770]" italics="true" pageId="5" pageNumber="6">Z. inornata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A3D44D212A2D052FFB8FF92393D8F91F" box="[1051,1182,1747,1771]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="aberrans">
|
||
<emphasis id="56A0EAB02A2D052FFB8FF92393D8F91F" box="[1051,1182,1747,1771]" italics="true" pageId="5" pageNumber="6">Z. aberrans</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="A3D44D212A2D052FFB48F923927CF91F" box="[1244,1338,1747,1771]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="dubia">
|
||
<emphasis id="56A0EAB02A2D052FFB48F923927CF91F" box="[1244,1338,1747,1771]" italics="true" pageId="5" pageNumber="6">Z. dubia</emphasis>
|
||
</taxonomicName>
|
||
, suggest a degree of intrageneric complexity that we have only just begun to explore.
|
||
</paragraph>
|
||
<paragraph id="646B36A22A2D052FFF53F8EA93BCF86B" blockId="5.[151,1437,1422,2023]" pageId="5" pageNumber="6">
|
||
All in all, the present body of evidence indicates that
|
||
<taxonomicName id="A3D44D212A2D052FFC8FF8EB94D5F8C7" box="[795,915,1819,1843]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56A0EAB02A2D052FFC8FF8EB94D5F8C7" box="[795,915,1819,1843]" italics="true" pageId="5" pageNumber="6">Zadbimyia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A3D44D212A2D052FFC5DF8EB93E2F8C7" box="[969,1188,1819,1843]" class="Insecta" family="Cecidomyiidae" genus="Pseudocamptomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56A0EAB02A2D052FFC5DF8EB93E2F8C7" box="[969,1188,1819,1843]" italics="true" pageId="5" pageNumber="6">Pseudocamptomyia</emphasis>
|
||
</taxonomicName>
|
||
belong to one and the same clade of
|
||
<taxonomicName id="A3D44D212A2D052FFED1F8CE9686F8A3" box="[325,448,1854,1879]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
|
||
that is distributed in the New World and occurs also in the Afrotropics. This clade appears to be similarly complex and biodiverse as are both
|
||
<taxonomicName id="A3D44D212A2D052FFCC6F8939340F88E" authority="Loew" authorityName="Loew" box="[850,1030,1891,1914]" class="Insecta" family="Cecidomyiidae" genus="Asynapta" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56A0EAB02A2D052FFCC6F89394FCF88E" box="[850,954,1891,1914]" italics="true" pageId="5" pageNumber="6">Asynapta</emphasis>
|
||
Loew
|
||
</taxonomicName>
|
||
, with 44 species, and
|
||
<taxonomicName id="A3D44D212A2D052FFA99F89397AFF86B" authority="Kieffer" authorityName="Kieffer" class="Insecta" family="Cecidomyiidae" genus="Camptomyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="56A0EAB02A2D052FFA99F89392DAF88E" box="[1293,1436,1891,1914]" italics="true" pageId="5" pageNumber="6">Camptomyia</emphasis>
|
||
Kieffer
|
||
</taxonomicName>
|
||
, with 65 species, two genera that hitherto accounted for the greater part of world
|
||
<taxonomicName id="A3D44D212A2D052FFBECF87693B0F86B" box="[1144,1270,1926,1951]" class="Insecta" family="Cecidomyiidae" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Asynaptini">Asynaptini</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2CCE65292A2D052FFF53F85A94D8F813" pageId="5" pageNumber="6" type="etymology">
|
||
<paragraph id="646B36A22A2D052FFF53F85A94D8F813" blockId="5.[151,1437,1422,2023]" pageId="5" pageNumber="6">
|
||
<emphasis id="56A0EAB02A2D052FFF53F85A9608F837" bold="true" box="[199,334,1962,1987]" pageId="5" pageNumber="6">Etymology.</emphasis>
|
||
The generic name,
|
||
<taxonomicName id="A3D44D212A2D052FFDAFF85B95F5F837" box="[571,691,1963,1987]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="56A0EAB02A2D052FFDAFF85B95F5F837" box="[571,691,1963,1987]" italics="true" pageId="5" pageNumber="6">Zadbimyia</emphasis>
|
||
</taxonomicName>
|
||
, is composed of
|
||
<emphasis id="56A0EAB02A2D052FFC15F85B9484F837" box="[897,962,1963,1987]" italics="true" pageId="5" pageNumber="6">Zadbi</emphasis>
|
||
-, the abbreviation of
|
||
<taxonomicName id="A3D44D212A2D052FFB57F85B9249F837" box="[1219,1295,1963,1987]" class="Insecta" family="Cecidomyiidae" genus="Zadbimyia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="23" phylum="Arthropoda" rank="species" species="zurqui">Zurquí</taxonomicName>
|
||
All-Diptera Biodiversity Inventory, and -Μγία (lat. -
|
||
<emphasis id="56A0EAB02A2D052FFDDBF83F95C3F812" box="[591,645,1999,2022]" italics="true" pageId="5" pageNumber="6">myia</emphasis>
|
||
), the Greek word for fly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |