168 lines
14 KiB
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168 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.205.3491" ID-GBIF-Dataset="16f08073-854b-40a2-aef9-72a97c85c8bd" ID-PMC="PMC3391731" ID-Pensoft-Pub="1313-2970-205-59" ID-PubMed="22792035" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-205-59" ModsDocOrigin="ZooKeys 205" ModsDocTitle="A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae, Gnaphosidae)" checkinTime="1451248915768" checkinUser="pensoft" docAuthor="Kovblyuk, Mykola M., Kastrygina, Zoya A. & Omelko, Mikhail M." docDate="2012" docId="A96D7DE7517C0FEA3550AAE99268F064" docLanguage="en" docName="ZooKeys 205: 59-89" docOrigin="ZooKeys 205" docSource="http://dx.doi.org/10.3897/zookeys.205.3491" docTitle="Haplodrassus pseudosignifer Marusik, Hippa & Koponen 1996" docType="treatment" docVersion="3" lastPageNumber="68" masterDocId="FFC2BD6FFFD62C69FFC3EC55350AFFA3" masterDocTitle="A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae, Gnaphosidae)" masterLastPageNumber="89" masterPageNumber="59" pageNumber="67" updateTime="1668154130483" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae, Gnaphosidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kovblyuk, Mykola M.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kastrygina, Zoya A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Omelko, Mikhail M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>205</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>59</mods:start>
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<mods:end>89</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.205.3491</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.205.3491</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-205-59</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152036246" LSID="urn:lsid:plazi:treatment:A96D7DE7517C0FEA3550AAE99268F064" httpUri="http://treatment.plazi.org/id/A96D7DE7517C0FEA3550AAE99268F064" lastPageId="9" lastPageNumber="68" pageId="8" pageNumber="67">
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<subSubSection pageId="8" pageNumber="67" type="nomenclature">
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<paragraph pageId="8" pageNumber="67">
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<taxonomicName LSID="http://species-id.net/wiki/Haplodrassus_pseudosignifer" authority="Marusik, Hippa & Koponen, 1996" authorityName="Marusik, Hippa & Koponen" authorityYear="1996" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pseudosignifer" order="Araneae" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="pseudosignifer">
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<pageBreakToken pageId="8" pageNumber="67" start="start">Haplodrassus</pageBreakToken>
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pseudosignifer Marusik, Hippa & Koponen, 1996
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</taxonomicName>
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Figs 68
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<normalizedToken originalValue="–7075–">-7075-</normalizedToken>
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76
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="67" type="reference_group">
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<paragraph pageId="8" pageNumber="67">
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pseudosignifer" order="Araneae" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="pseudosignifer">Haplodrassus pseudosignifer</taxonomicName>
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Marusik et al., 1996: 26, f. 63-65, 69 (♂♀).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="67" type="type material">
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<paragraph pageId="8" pageNumber="67">Type material.</paragraph>
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<paragraph pageId="8" pageNumber="67">RUSSIA. 2 ♂♂ paratypes (ZMT), SW Altai, 7 km W Katanda, Katun River valley, forest steppe, pitfall traps, 22.06.-26.07.1983, H. Hippa.</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="67" type="material">
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<paragraph pageId="8" pageNumber="67">Material.</paragraph>
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<paragraph pageId="8" pageNumber="67">UKRAINE, CRIMEA: Feodosiya Distr.:19 ♂♂, 6 ♀♀ (TNU), Karadag Nature Reserve, 15.10.2006-05.2011, M.M. Kovblyuk, O.V. Kukushkin, A.A. Nadolny. Simferopol Distr.: 2 ♂♂, 2 ♀♀ (TNU), Bayrakly Mt. (519 m), ~ 400 m, 14.05.-23.06.2000, M.M. Kovblyuk; 1 ♂ (TNU), Chatyr-Dag Mt., east slope, 10-28.06.2000, M.M. Kovblyuk; 1 ♀ (TNU), near Skvortsovo Vill., 9-30.06.2002, M.M. Kovblyuk. Sudak Distr.: 28 ♂♂, 7 ♀♀ (TNU), 10 km west from Sudak Town, Mezhdurechie Vill., 3.05.-3.06.2010, M.K. Yusufova. Yalta Distr.: 1 ♂ (TNU), Ay-Petri Yaila Mt., west part, 12-13.06.1999, O.V. Kukushkin; 15 ♂♂, 1 ♀ (TNU), Nikitskaya Yaila Mt. (=Skrinita), ~ 1200 m, 2.06.-24.07.2001, M.M. Kovblyuk.</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="67" type="additional material">
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<paragraph pageId="8" pageNumber="67">Additional material.</paragraph>
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<paragraph pageId="8" pageNumber="67">UKRAINE. Nikolaev Area: 1 ♀ (TNU), Pervomaysky Distr., Kuripchane Vil., 26.05-8.06.2006, N.Yu. Polchaninova.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="68" type="note">
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<paragraph pageId="9" pageNumber="68">
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<pageBreakToken pageId="9" pageNumber="68" start="start">Note</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="9" pageNumber="68">Identification of this species was based on comparison of our specimens with the male paratypes from ZMT. Specimens from Crimea and Altai differ only slightly in the shape of the tooth on embolus. In our opinion the specimens from Crimea, Nikolaev Area and Altai are conspecific.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="68" type="diagnosis">
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<paragraph pageId="9" pageNumber="68">Diagnosis.</paragraph>
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<paragraph pageId="9" pageNumber="68">
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pseudosignifer" order="Araneae" pageId="9" pageNumber="68" phylum="Arthropoda" rank="species" species="pseudosignifer">Haplodrassus pseudosignifer</taxonomicName>
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is very similar to
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus signifer" order="Araneae" pageId="9" pageNumber="68" phylum="Arthropoda" rank="species" species="signifer">Haplodrassus signifer</taxonomicName>
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. The two species have no distinct differences in coloration, size or leg spination, but
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pseudosignifer" order="Araneae" pageId="9" pageNumber="68" phylum="Arthropoda" rank="species" species="pseudosignifer">Haplodrassus pseudosignifer</taxonomicName>
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can be differentiated from
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus signifer" order="Araneae" pageId="9" pageNumber="68" phylum="Arthropoda" rank="species" species="signifer">Haplodrassus signifer</taxonomicName>
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by having an almost straight and shorter terminal apophysis and thinner embolus, and by the shape of the lateral pockets and fovea of the epigyne.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="68" type="description">
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<paragraph pageId="9" pageNumber="68">Description.</paragraph>
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<paragraph pageId="9" pageNumber="68">Males (n = 5) and females (n = 5). Measurements (♂ / ♀): total length 5.7-8.0 (6.79) / 6.6-10.5 (7.8); carapace 2.8-3.5 (3.1) / 2.6-3.6 (3.3) long, 2.2-2.6 (2.4) / 2.1-3.0 (2.6) wide; abdomen 3.1-4.8 (3.8) / 3.4-6.7 (4.7) long, 1.7-2.4 (2.0) / 2.1-4.2 (2.8) wide.</paragraph>
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<paragraph pageId="9" pageNumber="68">Length of leg segments:</paragraph>
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<paragraph pageId="9" pageNumber="68">
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<table pageId="9" pageNumber="68">
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<tr pageId="9" pageNumber="68">
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<th colspan="1" pageId="9" pageNumber="68" rowspan="1">Leg</th>
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<th colspan="1" pageId="9" pageNumber="68" rowspan="1">Femur</th>
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<th colspan="1" pageId="9" pageNumber="68" rowspan="1">Patella</th>
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<th colspan="1" pageId="9" pageNumber="68" rowspan="1">Tibia</th>
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<th colspan="1" pageId="9" pageNumber="68" rowspan="1">Metatarsus</th>
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<th colspan="1" pageId="9" pageNumber="68" rowspan="1">Tarsus</th>
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<th colspan="1" pageId="9" pageNumber="68" rowspan="1">Total</th>
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</tr>
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</table>
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</paragraph>
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<paragraph pageId="9" pageNumber="68">Length of palp segments (male / female): femur 1.0-1.3 (1.2) / 0.8-1.2 (1.1), patella 0.4-0.7 (0.5) / 0.5-0.6 (0.5), tibia 0.3-0.4 (0.38) / 0.4-0.6 (0.5), tarsus 1.0-1.2 (1.1) / 0.7-0.9 (0.8).</paragraph>
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<paragraph pageId="9" pageNumber="68">Chelicerae with 2-3 promarginal and 2 retromarginal teeth in males and females. Number of promarginal teeth varies from 2 (frequently) to 3 (rarely).</paragraph>
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<paragraph pageId="9" pageNumber="68">Coloration grey.</paragraph>
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<paragraph pageId="9" pageNumber="68">Male palp as in Figs 68-70. Terminal apophysis short (length/width ratio ~ 2) and straight, ridge poorly developed; embolus almost straight and with a tooth.</paragraph>
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<paragraph pageId="9" pageNumber="68">Epigyne as in Figs 75-76. Fovea elongated, rectangular (longer than wide) with narrow longitudinal groove.</paragraph>
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<caption pageId="9" pageNumber="68">
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<paragraph pageId="9" pageNumber="68">
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Figures 68-74. Males of
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pseudosignifer" order="Araneae" pageId="9" pageNumber="68" phylum="Arthropoda" rank="species" species="pseudosignifer">Haplodrassus pseudosignifer</taxonomicName>
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(68-70 from Crimea) and
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus signifer" order="Araneae" pageId="9" pageNumber="68" phylum="Arthropoda" rank="species" species="signifer">Haplodrassus signifer</taxonomicName>
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(71-74 from Crimea): 68, 71 RTA, retrolateral view 69, 72 palp, ventral view 70, 73-74 bulbus, apical view.
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</paragraph>
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</caption>
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<caption pageId="9" pageNumber="68">
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<paragraph pageId="9" pageNumber="68">
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Figures 75-78. Females of
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pseudosignifer" order="Araneae" pageId="9" pageNumber="68" phylum="Arthropoda" rank="species" species="pseudosignifer">Haplodrassus pseudosignifer</taxonomicName>
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(75-76 from Crimea) and
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus signifer" order="Araneae" pageId="9" pageNumber="68" phylum="Arthropoda" rank="species" species="signifer">Haplodrassus signifer</taxonomicName>
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(77-78 from Crimea): 75, 77 epigyne, ventral view 76, 78 epigyne, dorsal view. Abbreviations: Ah anterior hood; Gr groove of epigyne; Spspermatheca.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="68" type="distribution">
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<paragraph pageId="9" pageNumber="68">Distribution.</paragraph>
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<paragraph pageId="9" pageNumber="68">
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Steppe zone of Eurasia: from Crimea and Nikolaev Area of Ukraine east to Altai (
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<bibRefCitation author="Marusik, YM" journalOrPublisher="Acta Zoologica Fennica" pageId="13" pageNumber="72" pagination="11 - 45" title="Spiders (Araneae) from the Altai area, southern Siberia." volume="201" year="1996">Marusik et al. 1996</bibRefCitation>
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; present data).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="68" type="comments">
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<paragraph pageId="9" pageNumber="68">Comments.</paragraph>
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<paragraph pageId="9" pageNumber="68">
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pseudosignifer" order="Araneae" pageId="9" pageNumber="68" phylum="Arthropoda" rank="species" species="pseudosignifer">Haplodrassus pseudosignifer</taxonomicName>
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is a new species record for Crimea and Ukraine. Crimea is the westernmost point of the known distribution range.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="68" type="habitats">
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<paragraph pageId="9" pageNumber="68">Habitats.</paragraph>
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<paragraph pageId="9" pageNumber="68">
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Forests with
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<taxonomicName genus="Pistaca" lsidName="Pistaca mutica" pageId="9" pageNumber="68" rank="species" species="mutica">Pistaca mutica</taxonomicName>
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or
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<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Quercus pubescens" order="Fagales" pageId="9" pageNumber="68" phylum="Tracheophyta" rank="species" species="pubescens">Quercus pubescens</taxonomicName>
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, forest-steppes, forest stripes (=shelter belts), rocky steppes, grasslands.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="68" type="phenology">
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<paragraph pageId="9" pageNumber="68">Phenology.</paragraph>
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<paragraph pageId="9" pageNumber="68">♂♀ - V-VII, ♂♂ - IV, X. In Crimea the peak of activity in adults occurs in May.</paragraph>
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</subSubSection>
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</treatment>
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</document> |