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<document ID-DOI="10.25221/fee.370.1" ID-ISSN="2713-2196" ID-Zenodo-Dep="7164560" approvalRequired="86" approvalRequired_for_textStreams="86" checkinTime="1665243909279" checkinUser="felipe" docAuthor="Tanansathaporn, S., Zettel, H., Pinkaew, N., Kongmee, M., Waengsothorn, S. &amp; Jaitrong, W." docDate="2018" docId="A9508788742AFFBB07D5FC43FC6C1238" docLanguage="en" docName="FarEastEntomol.370.1-11.pdf" docOrigin="Far Eastern Entomologist 370" docSource="http://dx.doi.org/10.25221/fee.370.1" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.1:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="1" docTitle="Echinopla jeenthongi Tanansathaporn &amp; Zettel &amp; Pinkaew &amp; Kongmee &amp; Waengsothorn &amp; Jaitrong 2018, sp. n." docType="treatment" docVersion="2" lastPageNumber="10" masterDocId="5569FFF0742FFFB20736FFA5FFD01757" masterDocTitle="Two new species of the genus Echinopla Smith, 1857 (Hymenoptera: Formicidae: Formicinae) from Thailand" masterLastPageNumber="11" masterPageNumber="1" pageNumber="6" updateTime="1665256199426" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new species of the genus Echinopla Smith, 1857 (Hymenoptera: Formicidae: Formicinae) from Thailand</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tanansathaporn, S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zettel, H.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pinkaew, N.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kongmee, M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Waengsothorn, S.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jaitrong, W.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Far Eastern Entomologist</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-11-06</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>370</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>11</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.25221/fee.370.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.25221/fee.370.1</mods:identifier>
<mods:identifier type="ISSN">2713-2196</mods:identifier>
<mods:identifier type="Zenodo-Dep">7164560</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:A9508788742AFFBB07D5FC43FC6C1238" httpUri="http://treatment.plazi.org/id/A9508788742AFFBB07D5FC43FC6C1238" lastPageId="9" lastPageNumber="10" pageId="5" pageNumber="6">
<subSubSection box="[227,865,998,1023]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph blockId="5.[227,865,998,1053]" box="[227,865,998,1023]" pageId="5" pageNumber="6">
<heading box="[227,865,998,1023]" centered="true" fontSize="9" level="2" pageId="5" pageNumber="6" reason="2">
<emphasis bold="true" box="[227,865,998,1023]" pageId="5" pageNumber="6">
<taxonomicName authority="Tanansathaporn et Jaitrong" authorityName="Tanansathaporn &amp; Zettel &amp; Pinkaew &amp; Kongmee &amp; Waengsothorn &amp; Jaitrong" authorityYear="2018" box="[227,793,998,1023]" class="Insecta" family="Formicidae" genus="Echinopla" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="jeenthongi" status="sp. n.">
<emphasis bold="true" box="[227,465,998,1022]" italics="true" pageId="5" pageNumber="6">Echinopla jeenthongi</emphasis>
Tanansathaporn et Jaitrong
</taxonomicName>
,
<taxonomicNameLabel box="[805,865,999,1023]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[227,319,1029,1053]" pageId="5" pageNumber="6" type="description">
<paragraph blockId="5.[227,865,998,1053]" box="[227,319,1029,1053]" pageId="5" pageNumber="6">
<figureCitation box="[227,319,1029,1053]" captionStart-0="Figs 46" captionStart-1="Figs 79" captionStartId-0="6.[265,306,1258,1280]" captionStartId-1="8.[265,306,1476,1498]" captionTargetBox-0="[227,1122,256,1227]" captionTargetBox-1="[227,1122,472,1443]" captionTargetId-0="figure-2@6.[227,1122,256,1227]" captionTargetId-1="figure-71@8.[227,1122,472,1443]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionText-0="Figs 46. Echinopla jeenthongi, holotype worker. 4 head in full-face view; 5 body" captionText-1="Figs 79. Echinopla jeenthongi, paratype gyne. 7 head in full-face view; 8 body from" figureDoi-0="http://doi.org/10.5281/zenodo.7164564" figureDoi-1="http://doi.org/10.5281/zenodo.7164566" httpUri-0="https://zenodo.org/record/7164564/files/figure.png" httpUri-1="https://zenodo.org/record/7164566/files/figure.png" pageId="5" pageNumber="6">Figs 49</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="materials_examined">
<paragraph blockId="5.[227,1128,1090,1605]" pageId="5" pageNumber="6">
<materialsCitation collectionCode="TYPE, MATERIAL" country="Thailand" county="Sichon district" location="Yod Nam" municipality="Khao Noi" pageId="5" pageNumber="6" specimenCount="1" stateProvince="Nakhon Si Thammarat" typeStatus="holotype">
<collectionCode box="[265,483,1091,1115]" pageId="5" pageNumber="6">TYPE MATERIAL</collectionCode>
.
<typeStatus box="[496,595,1090,1114]" pageId="5" pageNumber="6">Holotype</typeStatus>
worker (THNHM-I-01368),
<emphasis bold="true" box="[940,1069,1090,1114]" pageId="5" pageNumber="6">
S
<collectingCountry box="[965,1069,1090,1114]" name="Thailand" pageId="5" pageNumber="6">Thailand</collectingCountry>
</emphasis>
:
<collectingRegion country="Thailand" name="Nakhon Si Thammarat" pageId="5" pageNumber="6">Nakhon Si Thammarat province</collectingRegion>
,
<collectingCounty box="[557,716,1121,1145]" pageId="5" pageNumber="6">Sichon district</collectingCounty>
,
<collectingMunicipality box="[728,834,1121,1145]" pageId="5" pageNumber="6">Khao Noi</collectingMunicipality>
subdistrict, near
<location LSID="urn:lsid:plazi:treatment:A9508788742AFFBB07D5FC43FC6C1238:24266045742AFFB704CBFBC4FBB6132E" box="[1021,1126,1121,1145]" country="Thailand" county="Sichon district" municipality="Khao Noi" name="Yod Nam" pageId="5" pageNumber="6" stateProvince="Nakhon Si Thammarat">Yod Nam</location>
</materialsCitation>
</paragraph>
<paragraph blockId="5.[227,1128,1090,1605]" box="[227,1128,1152,1176]" pageId="5" pageNumber="6">
<materialsCitation box="[227,766,1152,1176]" collectingDate="2008-11-01" collectionCode="T" latitude="8.9091" longLatPrecision="7" longitude="99.7319" pageId="5" pageNumber="6" specimenCount="1">
Waterfall,
<geoCoordinate box="[339,443,1152,1176]" degrees="8.9091" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="5" value="8.9091">8.9091°N</geoCoordinate>
,
<geoCoordinate box="[455,568,1152,1176]" degrees="99.7319" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="5" value="99.7319">99.7319°E</geoCoordinate>
,
<date box="[580,690,1152,1176]" pageId="5" pageNumber="6" value="2008-11-01">
<collectingDate box="[580,690,1152,1176]" pageId="5" pageNumber="6" value="2008-11-01">1.XI 2008</collectingDate>
</date>
, leg.
<collectionCode box="[747,766,1152,1175]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="6">T</collectionCode>
</materialsCitation>
.
<materialsCitation box="[777,1124,1152,1176]" collectionCode="THNHM" pageId="5" pageNumber="6" specimenCount="1" typeStatus="paratype">
Jeenthong (
<collectionCode box="[898,1001,1152,1176]" pageId="5" pageNumber="6">THNHM</collectionCode>
).
<typeStatus box="[1018,1124,1152,1176]" pageId="5" pageNumber="6">Paratypes</typeStatus>
</materialsCitation>
:
</paragraph>
<paragraph blockId="5.[227,1128,1090,1605]" box="[227,1127,1182,1207]" pageId="5" pageNumber="6">
<materialsCitation box="[227,1123,1182,1207]" collectionCode="NW" country="Thailand" location="Ban Nasarn district" pageId="5" pageNumber="6" specimenCode="NW310705-1" specimenCount="1" specimenCount-worker="1" stateProvince="Surat Thani">
<specimenCount box="[227,349,1182,1206]" pageId="5" pageNumber="6" type="worker">one worker</specimenCount>
(
<specimenCode box="[365,516,1182,1206]" collectionCode="NW" pageId="5" pageNumber="6">NW310705-1</specimenCode>
), S
<collectingCountry box="[559,657,1182,1206]" name="Thailand" pageId="5" pageNumber="6">Thailand</collectingCountry>
,
<collectingRegion box="[669,899,1182,1207]" country="Thailand" name="Surat Thani" pageId="5" pageNumber="6">Surat Thani province</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:A9508788742AFFBB07D5FC43FC6C1238:24266045742AFFB704B9FB3AFBB313E1" box="[911,1123,1182,1206]" country="Thailand" name="Ban Nasarn district" pageId="5" pageNumber="6" stateProvince="Surat Thani">Ban Nasarn district</location>
</materialsCitation>
,
</paragraph>
<paragraph blockId="5.[227,1128,1090,1605]" box="[227,1127,1213,1238]" pageId="5" pageNumber="6">
Julaporn,
<geoCoordinate box="[338,443,1213,1237]" degrees="8.8613" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="5" value="8.8613">8.8613°N</geoCoordinate>
,
<geoCoordinate box="[460,577,1213,1237]" degrees="99.4847" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="5" value="99.4847">99.4847°E</geoCoordinate>
, evergreen forest,
<quantity box="[790,919,1213,1237]" metricMagnitude="2" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="5" pageNumber="6" unit="m" value="475.0" valueMax="550.0" valueMin="400.0">400550 m</quantity>
, lower vegetation,
</paragraph>
<paragraph blockId="5.[227,1128,1090,1605]" box="[227,1127,1244,1268]" pageId="5" pageNumber="6">
<materialsCitation box="[227,1123,1244,1268]" collectingDate="2005-07-31" collectionCode="PSU" collectorName="N. Noon-anant" country="Thailand" location="Thailand" pageId="5" pageNumber="6" specimenCode="NW150405-2" specimenCount="1" specimenCount-worker="1">
<date box="[227,363,1244,1268]" pageId="5" pageNumber="6" value="2005-07-31">
<collectingDate box="[227,363,1244,1268]" pageId="5" pageNumber="6" value="2005-07-31">31.VII 2005</collectingDate>
</date>
, leg.
<collectorName box="[427,589,1244,1267]" pageId="5" pageNumber="6">N. Noon-anant</collectorName>
(
<collectionCode box="[607,661,1244,1268]" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="5" pageNumber="6" type="Museum">PSU</collectionCode>
);
<specimenCount box="[684,808,1244,1268]" pageId="5" pageNumber="6" type="worker">one worker</specimenCount>
(
<specimenCode box="[826,978,1244,1268]" collectionCode="NW" pageId="5" pageNumber="6">NW150405-2</specimenCode>
), S
<collectingCountry box="[1025,1123,1244,1268]" name="Thailand" pageId="5" pageNumber="6">Thailand</collectingCountry>
</materialsCitation>
,
</paragraph>
<paragraph blockId="5.[227,1128,1090,1605]" pageId="5" pageNumber="6">
<materialsCitation county="Noppitam district" elevation="200" latitude="8.6572" location="Krung Ching" longLatPrecision="7" longitude="99.6363" municipality="Noppitum" pageId="5" pageNumber="6" specimenCount="1" stateProvince="Nakhon Si Thammarat">
<collectingRegion box="[227,591,1274,1298]" country="Thailand" name="Nakhon Si Thammarat" pageId="5" pageNumber="6">Nakhon Si Thammarat province</collectingRegion>
,
<collectingCounty box="[609,805,1274,1298]" pageId="5" pageNumber="6">Noppitam district</collectingCounty>
,
<collectingMunicipality box="[822,930,1274,1298]" pageId="5" pageNumber="6">Noppitum</collectingMunicipality>
[Noppitam] subdistrict,
<location LSID="urn:lsid:plazi:treatment:A9508788742AFFBB07D5FC43FC6C1238:24266045742AFFB7060BFABFFE1F1266" box="[317,463,1305,1330]" county="Noppitam district" latitude="8.6572" longLatPrecision="7" longitude="99.6363" municipality="Noppitum" name="Krung Ching" pageId="5" pageNumber="6" stateProvince="Nakhon Si Thammarat">Krung Ching</location>
,
<geoCoordinate box="[477,583,1305,1329]" degrees="8.6572" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="5" value="8.6572">8.6572°N</geoCoordinate>
,
<geoCoordinate box="[596,713,1305,1329]" degrees="99.6363" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="5" value="99.6363">99.6363°E</geoCoordinate>
,
<quantity box="[727,853,1305,1329]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="5" pageNumber="6" unit="m" value="200.0" valueMax="300.0" valueMin="100.0">
<elevation box="[727,853,1305,1329]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="5" pageNumber="6" unit="m" value="200.0" valueMax="300.0" valueMin="100.0">100300 m</elevation>
</quantity>
, lower vegetation, 15.IV
</materialsCitation>
</paragraph>
<paragraph blockId="5.[227,1128,1090,1605]" box="[227,1127,1336,1360]" pageId="5" pageNumber="6">
2005, leg. N. Noon-anant (
<collectionCode box="[534,589,1336,1360]" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="5" pageNumber="6" type="Museum">PSU</collectionCode>
); one dealate gyne (
<accessionNumber box="[823,956,1336,1360]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/NW091105" pageId="5" pageNumber="6">
<collectionCode box="[823,871,1336,1360]" pageId="5" pageNumber="6">NW</collectionCode>
091105
</accessionNumber>
-3), S
<collectingCountry box="[1025,1123,1336,1360]" name="Thailand" pageId="5" pageNumber="6">Thailand</collectingCountry>
,
</paragraph>
<paragraph blockId="5.[227,1128,1090,1605]" box="[227,1126,1366,1391]" pageId="5" pageNumber="6">
<collectingRegion box="[227,454,1366,1390]" country="Thailand" name="Phangnga" pageId="5" pageNumber="6">Phangnga province</collectingRegion>
, Muang
<materialsCitation box="[571,1121,1366,1391]" latitude="8.6111" location="Song Prang" pageId="5" pageNumber="6" specimenCount="1" stateProvince="Phangnga">
<collectingRegion box="[571,683,1366,1390]" country="Thailand" name="Phangnga" pageId="5" pageNumber="6">Phangnga</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:A9508788742AFFBB07D5FC43FC6C1238:24266045742AFFB705F7FAF3FC9B1238" box="[705,843,1366,1391]" latitude="8.6111" name="Song Prang" pageId="5" pageNumber="6" stateProvince="Phangnga">Song Prang</location>
sub-district,
<geoCoordinate box="[1012,1121,1366,1390]" degrees="8.6111" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="5" value="8.6111">8.6111°N</geoCoordinate>
</materialsCitation>
,
</paragraph>
<paragraph blockId="5.[227,1128,1090,1605]" box="[227,1099,1397,1421]" pageId="5" pageNumber="6">
<materialsCitation box="[227,1096,1397,1421]" collectingDate="2005-11-09" collectionCode="PSU" collectorName="N. Noon-anant" elevation="250" longitude="98.5505" pageId="5" pageNumber="6" specimenCount="1">
<geoCoordinate box="[227,343,1397,1421]" degrees="98.5505" direction="east" orientation="longitude" pageId="5" pageNumber="6" precision="5" value="98.5505">98.5505°E</geoCoordinate>
,
<quantity box="[354,477,1397,1421]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="6" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">
<elevation box="[354,477,1397,1421]" metricMagnitude="2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="5" pageNumber="6" unit="m" value="250.0" valueMax="300.0" valueMin="200.0">200300 m</elevation>
</quantity>
, lower vegetation,
<date box="[680,804,1397,1421]" pageId="5" pageNumber="6" value="2005-11-09">
<collectingDate box="[680,804,1397,1421]" pageId="5" pageNumber="6" value="2005-11-09">09.XI 2005</collectingDate>
</date>
, leg.
<collectorName box="[860,1019,1398,1421]" pageId="5" pageNumber="6">N. Noon-anant</collectorName>
(
<collectionCode box="[1033,1088,1397,1421]" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="5" pageNumber="6" type="Museum">PSU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
<paragraph blockId="5.[227,1128,1090,1605]" box="[265,1127,1428,1452]" pageId="5" pageNumber="6">
MEASUREMENTS.
<emphasis bold="true" box="[497,601,1428,1452]" pageId="5" pageNumber="6">
<typeStatus box="[497,601,1428,1452]" pageId="5" pageNumber="6">Holotype</typeStatus>
</emphasis>
: TL 3.80, HL 1.12, HW 1.04, SL 1.12, ED 0.24,
</paragraph>
<paragraph blockId="5.[227,1128,1090,1605]" box="[227,1126,1458,1482]" pageId="5" pageNumber="6">
PW 0.92, MSL 1.62, PL 0.53,
<collectingCountry box="[558,592,1459,1482]" name="Philippines" pageId="5" pageNumber="6">PH</collectingCountry>
0.50, DPW 0.96, GW 1.25, CI 93, EI 23, SI 108,
</paragraph>
<paragraph blockId="5.[227,1128,1090,1605]" box="[227,1127,1489,1513]" pageId="5" pageNumber="6">
LPI 94, DPI 181.
<emphasis bold="true" box="[422,536,1489,1513]" pageId="5" pageNumber="6">
<typeStatus box="[422,536,1489,1513]" pageId="5" pageNumber="6">Paratypes</typeStatus>
</emphasis>
(n = 2): TL 4.404.50, HL 0.891.15, HW 0.961.00,
</paragraph>
<paragraph blockId="5.[227,1128,1090,1605]" box="[227,1126,1520,1544]" pageId="5" pageNumber="6">
SL 1.061.15, ED 0.230.25, PW 0.850.86, MSL 1.722.35, PL 0.650.66,
<collectingCountry box="[1092,1126,1520,1543]" name="Philippines" pageId="5" pageNumber="6">PH</collectingCountry>
</paragraph>
<paragraph blockId="5.[227,1128,1090,1605]" lastBlockId="6.[227,1091,1258,1307]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">0.530.60, DPW 0.730.80, GW 1.301.58, CI 87108, EI 2425, SI 110115, LPI from dorsal view; 6 body in profile.</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7164564" ID-Zenodo-Dep="7164564" box="[265,1091,1258,1280]" httpUri="https://zenodo.org/record/7164564/files/figure.png" pageId="6" pageNumber="7" startId="6.[265,306,1258,1280]" targetBox="[227,1122,256,1227]" targetPageId="6">
<paragraph blockId="6.[227,1091,1258,1307]" box="[265,1091,1258,1280]" pageId="6" pageNumber="7">
Figs 46.
<taxonomicName authorityName="Tanansathaporn &amp; Zettel &amp; Pinkaew &amp; Kongmee &amp; Waengsothorn &amp; Jaitrong" authorityYear="2018" box="[360,566,1258,1279]" class="Insecta" family="Formicidae" genus="Echinopla" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="jeenthongi">
<emphasis box="[360,566,1258,1279]" italics="true" pageId="6" pageNumber="7">Echinopla jeenthongi</emphasis>
</taxonomicName>
, holotype worker. 4 head in full-face view; 5 body
</paragraph>
</caption>
<subSubSection lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="diagnosis">
<paragraph blockId="6.[227,1127,1342,1630]" pageId="6" pageNumber="7">
DIAGNOSIS. Medium-sized, HW
<quantity box="[671,841,1342,1366]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.04" metricValueMin="0.96" pageId="6" pageNumber="7" unit="mm" value="1.0" valueMax="1.04" valueMin="0.96">0.961.04 mm</quantity>
, head in full-face view elliptical, clearly longer than broad; eye relatively large, weakly convex; mesosoma very elongated and subcylindrical; pronotum with tooth-like angles; in dorsal view mesosoma with waist-like incision in front of propodeum; promesonotal suture almost absent; metanotal groove distinct with shallow depression; petiole in profile subtriangular, in dorsal view transverse and armed with two spines; first gastral tergite in dorsal view elliptical, much longer than broad, its posterior margin with numerous small denticles. Dorsa and lateral face of head, mesosoma, petiole and first gastral tergite with dense macropunctures. Dorsum and lateral face of head,
</paragraph>
<paragraph blockId="6.[227,1127,1342,1630]" box="[227,1122,1606,1630]" pageId="6" pageNumber="7">mesosoma petiole, and first gastral tergite without long standing hairs; clypeus,</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" pageId="7" pageNumber="8">mandible and first gastral sternite with sparse long hairs. Entire body black; antenna dark brown, leg reddish brown.</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="description">
<paragraph blockId="7.[227,1128,264,1604]" pageId="7" pageNumber="8">
DESCRIPTION.
<emphasis bold="true" box="[452,536,327,351]" pageId="7" pageNumber="8">Worker</emphasis>
(
<figureCitation box="[558,657,327,351]" captionStart="Figs 46" captionStartId="6.[265,306,1258,1280]" captionTargetBox="[227,1122,256,1227]" captionTargetId="figure-2@6.[227,1122,256,1227]" captionTargetPageId="6" captionText="Figs 46. Echinopla jeenthongi, holotype worker. 4 head in full-face view; 5 body" figureDoi="http://doi.org/10.5281/zenodo.7164564" httpUri="https://zenodo.org/record/7164564/files/figure.png" pageId="7" pageNumber="8">Figs 46</figureCitation>
). Head in full-face view clearly longer than broad, elliptical, with weakly convex lateral margin and convex posterior margin.
</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" pageId="7" pageNumber="8">Eye relatively large, moderately protruding, located at mid-length of head. Frontal lobe horizontal, in full-face view covering antennal fossa only in part. Mandible short, subrectangular, anterior margin slightly convex, and its masticatory margin armed with five teeth. Clypeus broad, anterior margin almost straight, while posterior margin strongly convex. Frontal carina short, not surpassing level of anterior margin of eye. Antennal scape moderately long, steadily widened from base to apex, slightly extending beyond posterior margin of head; antennal segment II clearly narrower and longer than each of IIIVI; terminal segment (XII) almost as long as X and XI combined.</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" pageId="7" pageNumber="8">Mesosoma very elongated and subcylindrical, promesonotum clearly larger than propodeum in dorsal view. In profile dorsal outline of mesosoma weakly convex;</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" box="[227,1127,734,758]" pageId="7" pageNumber="8">promesonotal suture absent, but laterally indicated by short row of puncture;</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" box="[227,1127,765,790]" pageId="7" pageNumber="8">metanotal groove weakly impressed; propodeal outline strongly convex dorsally;</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" pageId="7" pageNumber="8">propodeal declivity weakly convex. In dorsal view mesosoma with waist-like incision in front of propodeum; promesonotum clearly longer than broad; pronotum with pair of tooth-like protrusions; maximum pronotal width slightly narrower than head excluding eyes. In dorsal view, propodeum suboval, narrowed anteriorly,</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" pageId="7" pageNumber="8">clearly longer than broad, with lateral and posterior margins convexs; proprodeal spiracle protruding laterally. Legs relatively long and slender.</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" box="[265,1127,984,1008]" pageId="7" pageNumber="8">Petiole in profile subtriangular, as long as high, ventral outline feebly concave.</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" pageId="7" pageNumber="8">In dorsal view petiole transverse, about as broad as propodeal width, its anterior face as broad as posterior face. In posterior aspect dorsal margin with two spines;</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" box="[227,660,1078,1102]" pageId="7" pageNumber="8">lateral margin without denticles or teeth.</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" pageId="7" pageNumber="8">First gastral tergite in dorsal view elliptical, much longer than broad, broader posteriorly, covering remaining tergites, posterior margin convex with numerous small denticles.</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" pageId="7" pageNumber="8">Dorsum and lateral face of head finely macropunctate, except frontal lobe punctate; mandible with dense puncture at base, smooth and shiny along masticatory margin and apex; antennal scape finely punctate. Mesosoma entirely macropunctate,</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" box="[227,1128,1297,1321]" pageId="7" pageNumber="8">except anterior-most portion punctate. Petiole and first gastral tergite macropunctate.</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" box="[227,1127,1329,1353]" pageId="7" pageNumber="8">Legs reticulate with smooth and shiny interspaces. Dorsum of head, mesosoma,</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" pageId="7" pageNumber="8">petiole, and first gastral tergite without erect hairs; clypeus, mandible, and apex of scape with sparse erect hairs; gastral sternites with dense hairs.</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" pageId="7" pageNumber="8">Entire body black; mandible reddish brown; antenna dark brown, with tip of scape reddish brown and antennal segment XII yellowish brown; coxae, trochanter, and basal half of femora yellowish brown; apical half of femora, tibiae, and tarsi reddish brown.</paragraph>
<paragraph blockId="7.[227,1128,264,1604]" lastBlockId="8.[227,1128,264,441]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis bold="true" box="[265,411,1548,1572]" pageId="7" pageNumber="8">Dealate gyne</emphasis>
(
<figureCitation box="[425,523,1548,1572]" captionStart="Figs 79" captionStartId="8.[265,306,1476,1498]" captionTargetBox="[227,1122,472,1443]" captionTargetId="figure-71@8.[227,1122,472,1443]" captionTargetPageId="8" captionText="Figs 79. Echinopla jeenthongi, paratype gyne. 7 head in full-face view; 8 body from" figureDoi="http://doi.org/10.5281/zenodo.7164566" httpUri="https://zenodo.org/record/7164566/files/figure.png" pageId="7" pageNumber="8">Figs 79</figureCitation>
). Body size slightly larger than worker. Head in full-face view longer than broad, elliptical, with weakly convex side. Compound eye relatively large, moderately protruding, positioned at mid-length of head. Ocelli present, distance between lateral ocelli longer than distance between anterior and lateral ocellus.
</paragraph>
<paragraph blockId="8.[227,1128,264,441]" pageId="8" pageNumber="9">Mandible short, subrectangular, outer margin slightly convex, its masticatory margin armed with five teeth. Clypeus broad, its anterior margin almost straight,</paragraph>
<paragraph blockId="8.[227,1128,264,441]" pageId="8" pageNumber="9">while posterior margin strongly convex. Antennal scape moderately long, steadily widened from base to apex, slightly extending beyond posterior margin of head.</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7164566" ID-Zenodo-Dep="7164566" box="[265,1121,1476,1498]" httpUri="https://zenodo.org/record/7164566/files/figure.png" pageId="8" pageNumber="9" startId="8.[265,306,1476,1498]" targetBox="[227,1122,472,1443]" targetPageId="8">
<paragraph blockId="8.[227,1121,1476,1526]" box="[265,1121,1476,1498]" pageId="8" pageNumber="9">
Figs 79.
<taxonomicName authorityName="Tanansathaporn &amp; Zettel &amp; Pinkaew &amp; Kongmee &amp; Waengsothorn &amp; Jaitrong" authorityYear="2018" box="[360,566,1476,1497]" class="Insecta" family="Formicidae" genus="Echinopla" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="jeenthongi">
<emphasis box="[360,566,1476,1497]" italics="true" pageId="8" pageNumber="9">Echinopla jeenthongi</emphasis>
</taxonomicName>
, paratype gyne. 7 head in full-face view; 8 body from
</paragraph>
</caption>
<paragraph blockId="8.[227,1121,1476,1526]" box="[227,537,1504,1526]" pageId="8" pageNumber="9">dorsal view; 9 body in profile.</paragraph>
<paragraph blockId="8.[227,1127,1563,1618]" lastBlockId="9.[227,1127,264,1392]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">Mesosoma elongated. In dorsal view, pronotum short, separated from mesoscutum by a suture, its anterolateral corner armed with a short, truncate tooth; mesoscutum large, slightly longer than broad, its anterior margin strongly convex, while posterior margin almost straight; mesoscutellum trapezoidal, slightly broader than long,</paragraph>
<paragraph blockId="9.[227,1127,264,1392]" pageId="9" pageNumber="10">anterior margin weakly convex, separated from mesoscutum and metanotum by shallow sutures; metanotum very short, separated propodeum by a deep suture;</paragraph>
<paragraph blockId="9.[227,1127,264,1392]" pageId="9" pageNumber="10">propodeum larger than mesoscutellum, anterior margin feebly concave and posterior margin convex; in profile mesopluron broad, anepisternum separated from katepi-</paragraph>
<paragraph blockId="9.[227,1127,264,1392]" pageId="9" pageNumber="10">sternum by a transverse mesoplural groove; metapleuron not demarcated from lateral face of propodeum.</paragraph>
<paragraph blockId="9.[227,1127,264,1392]" pageId="9" pageNumber="10">Petiole in profile view subtriangular, as long as high. In dorsal view petiole transverse, slightly narrower than propodeum. In posterior aspect dorsal margin with 2 spines laterally.</paragraph>
<paragraph blockId="9.[227,1127,264,1392]" pageId="9" pageNumber="10">First gastral tergite in dorsal view elliptical, much longer than broad, broader posteriorly, covering remaining gastral tergites, posterior margin convex with numerous small denticles.</paragraph>
<paragraph blockId="9.[227,1127,264,1392]" pageId="9" pageNumber="10">Sculpture, setae and colour condition similar to those of worker caste. Posterior portion of first gastral tergite with four short erect hairs.</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="discussion">
<paragraph blockId="9.[227,1127,264,1392]" box="[265,1126,754,779]" pageId="9" pageNumber="10">
REMARKS. New species belongs to the
<taxonomicName authorityName="Zettel et Laciny" authorityYear="2015" box="[710,911,754,778]" class="Insecta" family="Formicidae" genus="Echinopla" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mezgeri">
<emphasis box="[710,911,754,778]" italics="true" pageId="9" pageNumber="10">Echinopla mezgeri</emphasis>
</taxonomicName>
group (
<emphasis box="[997,1056,755,778]" italics="true" pageId="9" pageNumber="10">sensu</emphasis>
Zettel
</paragraph>
<paragraph blockId="9.[227,1127,264,1392]" box="[227,1127,785,809]" pageId="9" pageNumber="10">
&amp; Laciny, 2017). It is closely related to
<taxonomicName authority="Zettel et Laciny, 2005" authorityName="Zettel et Laciny" authorityYear="2005" box="[668,1123,785,809]" class="Insecta" family="Formicidae" genus="Echinopla" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mezgeri">
<emphasis box="[668,870,785,809]" italics="true" pageId="9" pageNumber="10">Echinopla mezgeri</emphasis>
Zettel et Laciny, 2005
</taxonomicName>
,
</paragraph>
<paragraph blockId="9.[227,1127,264,1392]" pageId="9" pageNumber="10">
the only hitherto described species of this group. Both species lack long standing hairs on the body surface. However,
<taxonomicName authority="Tanansathaporn &amp; Zettel &amp; Pinkaew &amp; Kongmee &amp; Waengsothorn &amp; Jaitrong, 2018" authorityName="Tanansathaporn &amp; Zettel &amp; Pinkaew &amp; Kongmee &amp; Waengsothorn &amp; Jaitrong" authorityYear="2018" box="[628,772,846,870]" class="Insecta" family="Formicidae" genus="Echinopla" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="jeenthongi" status="sp. n.">
<emphasis box="[628,772,846,870]" italics="true" pageId="9" pageNumber="10">E. jeenthongi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[779,838,847,870]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
can be distinguished from
</paragraph>
<paragraph blockId="9.[227,1127,264,1392]" pageId="9" pageNumber="10">
<taxonomicName authorityName="Zettel et Laciny" authorityYear="2005" box="[227,342,878,901]" class="Insecta" family="Formicidae" genus="Echinopla" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mezgeri">
<emphasis box="[227,342,878,901]" italics="true" pageId="9" pageNumber="10">E. mezgeri</emphasis>
</taxonomicName>
by the following characteristics: entire head, mesosoma, petiole and first gastral tergite finely macropunctate (finely punctate, except mesopleuron and lateral face of propodeum covered by a coarse punctuation in
<taxonomicName authorityName="Zettel et Laciny" authorityYear="2005" box="[832,947,939,962]" class="Insecta" family="Formicidae" genus="Echinopla" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mezgeri">
<emphasis box="[832,947,939,962]" italics="true" pageId="9" pageNumber="10">E. mezgeri</emphasis>
</taxonomicName>
); posterior margin of first gastral tergite with numerous small denticles (without serration in
<taxonomicName box="[1104,1127,970,993]" pageId="9" pageNumber="10">
<emphasis box="[1104,1127,970,993]" italics="true" pageId="9" pageNumber="10">E.</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="9.[227,1127,264,1392]" pageId="9" pageNumber="10">
<emphasis box="[227,311,1000,1023]" italics="true" pageId="9" pageNumber="10">mezgeri</emphasis>
); basal half of femora yellowish brown (dark brown in
<taxonomicName authorityName="Zettel et Laciny" authorityYear="2005" box="[920,1035,1000,1024]" class="Insecta" family="Formicidae" genus="Echinopla" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mezgeri">
<emphasis box="[920,1035,1000,1024]" italics="true" pageId="9" pageNumber="10">E. mezgeri</emphasis>
</taxonomicName>
); sparse short pubescent hairs on dorsum of mesosoma (very dense in
<taxonomicName authorityName="Zettel et Laciny" authorityYear="2005" box="[907,1023,1031,1054]" class="Insecta" family="Formicidae" genus="Echinopla" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mezgeri">
<emphasis box="[907,1023,1031,1054]" italics="true" pageId="9" pageNumber="10">E. mezgeri</emphasis>
</taxonomicName>
). A third species of the
<taxonomicName authorityName="Zettel et Laciny" authorityYear="2005" box="[380,578,1061,1085]" class="Insecta" family="Formicidae" genus="Echinopla" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="mezgeri">
<emphasis box="[380,578,1061,1085]" italics="true" pageId="9" pageNumber="10">Echinopla mezgeri</emphasis>
</taxonomicName>
group from West
<collectingCountry box="[775,871,1061,1085]" name="Malaysia" pageId="9" pageNumber="10">Malaysia</collectingCountry>
(Laciny
<emphasis box="[968,1015,1061,1085]" italics="true" pageId="9" pageNumber="10">et al</emphasis>
., in press)
</paragraph>
<paragraph blockId="9.[227,1127,264,1392]" pageId="9" pageNumber="10">
differs from both species by a strongly reduced puncturation of mesosoma and gaster tergite 1, and from
<taxonomicName authority="Tanansathaporn &amp; Zettel &amp; Pinkaew &amp; Kongmee &amp; Waengsothorn &amp; Jaitrong, 2018" authorityName="Tanansathaporn &amp; Zettel &amp; Pinkaew &amp; Kongmee &amp; Waengsothorn &amp; Jaitrong" authorityYear="2018" box="[511,655,1122,1146]" class="Insecta" family="Formicidae" genus="Echinopla" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="jeenthongi" status="sp. n.">
<emphasis box="[511,655,1122,1146]" italics="true" pageId="9" pageNumber="10">E. jeenthongi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[663,722,1123,1146]" pageId="9" pageNumber="10" rank="species">sp. n.</taxonomicNameLabel>
also by a non-serrate hind margin of gaster tergite 1.
</paragraph>
<paragraph blockId="9.[227,1127,264,1392]" pageId="9" pageNumber="10">
<materialsCitation country="Thailand" location="This" pageId="9" pageNumber="10" specimenCount="2" typeStatus="holotype">
HABITAT. The
<typeStatus box="[444,538,1184,1208]" pageId="9" pageNumber="10">holotype</typeStatus>
and
<typeStatus box="[590,692,1184,1208]" pageId="9" pageNumber="10">paratypes</typeStatus>
were collected from lower vegetation in evergreen forests in southern
<collectingCountry box="[545,644,1214,1238]" name="Thailand" pageId="9" pageNumber="10">Thailand</collectingCountry>
.
<location LSID="urn:lsid:plazi:treatment:A9508788742AFFBB07D5FC43FC6C1238:242660457426FFBB05B8FB1BFD6E1381" box="[654,702,1214,1238]" country="Thailand" name="This" pageId="9" pageNumber="10">This</location>
species seen to be restricted to Sunder-
</materialsCitation>
</paragraph>
<paragraph blockId="9.[227,1127,264,1392]" box="[227,280,1245,1269]" pageId="9" pageNumber="10">land.</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="distribution">
<paragraph blockId="9.[227,1127,264,1392]" pageId="9" pageNumber="10">
<materialsCitation country="Thailand" location="Surat Thani" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Surat Thani">
DISTRIBUTION.
<collectingCountry box="[466,561,1276,1300]" name="Thailand" pageId="9" pageNumber="10">Thailand</collectingCountry>
(
<collectingRegion box="[577,708,1276,1300]" country="Thailand" name="Surat Thani" pageId="9" pageNumber="10">Surat Thani</collectingRegion>
,
<collectingRegion box="[720,966,1276,1300]" country="Thailand" name="Nakhon Si Thammarat" pageId="9" pageNumber="10">Nakhon Si Thammarat</collectingRegion>
and
<collectingRegion box="[1021,1126,1276,1300]" country="Thailand" name="Phangnga" pageId="9" pageNumber="10">Phangnga</collectingRegion>
provinces)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="10" type="etymology">
<paragraph blockId="9.[227,1127,264,1392]" box="[265,1127,1337,1361]" pageId="9" pageNumber="10">ETYMOLOGY. We dedicate this species to Mr. Tadsanai Jeenthong (National</paragraph>
<paragraph blockId="9.[227,1127,264,1392]" box="[227,956,1368,1392]" pageId="9" pageNumber="10">
<materialsCitation box="[227,952,1368,1392]" collectionCode="THNHM" country="Thailand" location="Science Museum" pageId="9" pageNumber="10" specimenCount="1" typeStatus="holotype">
<location LSID="urn:lsid:plazi:treatment:A9508788742AFFBB07D5FC43FC6C1238:242660457426FFBB07D5FAFDFE4D1238" box="[227,413,1368,1392]" country="Thailand" name="Science Museum" pageId="9" pageNumber="10">Science Museum</location>
,
<collectingCountry box="[423,522,1368,1392]" name="Thailand" pageId="9" pageNumber="10">Thailand</collectingCountry>
), who donated the
<typeStatus box="[723,816,1368,1392]" pageId="9" pageNumber="10">holotype</typeStatus>
to
<collectionCode box="[851,952,1368,1391]" pageId="9" pageNumber="10">THNHM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>